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A Review of the Genus Eunice - Smithsonian Institution Libraries

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22 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY<br />

3. bases scoop-shaped<br />

The scoop-shaped bases are folded around <strong>the</strong> lower edge <strong>of</strong> <strong>the</strong><br />

neuropodium and usually enclose <strong>the</strong> emerging bases <strong>of</strong> <strong>the</strong><br />

subacicular hooks; <strong>the</strong> scoop-shaped region is usually angled,<br />

and so it reaches far<strong>the</strong>r dorsally on <strong>the</strong> posterior face <strong>of</strong> <strong>the</strong><br />

parapodium than on <strong>the</strong> anterior face.<br />

55. median inflated ventral cirri<br />

1. narrow tips short and button-shaped<br />

2. narrow tips tapering<br />

3. narrow tips digitiform<br />

4. narrow tips absent<br />

In nearly all species, median ventral cirri have a distinctly<br />

marked tip lacking <strong>the</strong> distinct glands present in <strong>the</strong> bases; <strong>the</strong><br />

presence <strong>of</strong> <strong>the</strong>se glands makes it possible to recognize <strong>the</strong><br />

presence <strong>of</strong> <strong>the</strong> base, without <strong>the</strong> narrow tip.<br />

56. median ventral cirri<br />

1. tapering<br />

2. digitiform<br />

This character is scored only for species in which <strong>the</strong> bases are<br />

not inflated.<br />

57. posterior ventral cirri<br />

1. basally inflated<br />

2. without basal inflation<br />

58. bases <strong>of</strong> posterior inflated ventral cirri<br />

1. ovate or spherical<br />

2. thick, transverse welts<br />

3. triangular welts<br />

4. scoop-shaped<br />

59. posterior inflated ventral cirri with narrow tips<br />

1. short and button-shaped<br />

2. tapering<br />

3. digitiform<br />

4. absent<br />

These characters parallel <strong>the</strong> ones already characterized for<br />

median setigers. In most species, <strong>the</strong> posterior ventral cirri lack<br />

basal inflations; however, when inflated, <strong>the</strong> shape <strong>of</strong> <strong>the</strong> bases<br />

is <strong>of</strong>ten different from that present in median setigers.<br />

60. posterior ventral cirri<br />

1. tapering<br />

2. digitiform<br />

3. short, nearly tubercular<br />

4. broadly triangular, nearly scoop-shaped<br />

61. anterior notopodial cirri<br />

1. basally inflated<br />

2. medially inflated<br />

3. tapering<br />

4. digitiform<br />

5. clavate<br />

62. median notopodial cirri<br />

1. basally inflated<br />

2. medially inflated<br />

3. tapering<br />

4. digitiform<br />

5. clavate<br />

63. posterior notopodial cirri<br />

1. basally inflated<br />

2. medially inflated<br />

3. tapering<br />

4. digitiform<br />

5. clavate<br />

These character states are listed in <strong>the</strong> same order for all three<br />

regions <strong>of</strong> <strong>the</strong> body; both outgroup species have character state<br />

1 throughout <strong>the</strong> body. In most species, <strong>the</strong> notopodial cirri<br />

differ in shape from one end <strong>of</strong> <strong>the</strong> body to <strong>the</strong> o<strong>the</strong>r.<br />

64. notopodial cirri<br />

1. articulated throughout body<br />

2. articulated in anterior setigers<br />

3. without articulations<br />

Both outgroup species have character state 3. Notopodial cirri<br />

are rarely articulated in species without articulated ccratostylcs;<br />

most species with articulated notopodial cirri have articulations<br />

limited to <strong>the</strong> anterior end, usually about <strong>the</strong> first 20 setigers in<br />

a specimen with 100-120 setigers.<br />

Pectinate Setae<br />

65. anterior pectinate setae<br />

1. furled<br />

2. flat<br />

66. anterior pectinate setae<br />

1. distally tapering<br />

2. distally flaring<br />

67. median and posterior pectinate setae<br />

1. furled<br />

2. flat<br />

68. median and posterior pectinate setae<br />

1. distally tapering<br />

2. distally flaring<br />

Onuphids <strong>of</strong>ten have narrow, furled, tapering pectinate setae<br />

with relatively few teeth in anterior setigers and wider, flaring<br />

pectinate setae in median and posterior setigers. A similar<br />

pattern can be found in some eunicids, but most <strong>of</strong> <strong>the</strong>m appear<br />

to have similar pectinate setae in anterior and posterior setigers.<br />

The median and posterior pectinate setae appear to differ in a<br />

few cases, but documentation is difficult because <strong>the</strong>se setae<br />

are small and are <strong>of</strong>ten broken in posterior setigers where <strong>the</strong><br />

limbate setae, which protect pectinate setae in anterior setigers,<br />

are few in number or absent.<br />

Falcigers and Spinigers<br />

Compound falcigers differ a great deal in shape from anterior<br />

to posterior end and from superior to inferior positions in a<br />

single fascicle. The differences are mainly in <strong>the</strong> length and<br />

shape <strong>of</strong> <strong>the</strong> appendages, but <strong>the</strong> shafts may also change along<br />

<strong>the</strong> length <strong>of</strong> <strong>the</strong> body. In addition to <strong>the</strong> characters scored here,<br />

<strong>the</strong>se shape differences should be consulted for additional<br />

characterizations <strong>of</strong> <strong>the</strong> different species.

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