Summer roosts: Nathusius’ pipistrelle prefers tree holes and wood crevices in deciduous andpine <strong>for</strong>ests as summer roosts. Maternity colonies choose their roost sites mainly inwoodlands or near the <strong>for</strong>est edge close to open water. Sometimes crevices in the wall of ahouse close to the <strong>for</strong>est are used as a roost. Single males also occur in other habitat types(Schmidt 1991, Meschede & Heller 2000, Vierhaus 2004). Besides natural hiding places thebats accept boxes as roosts. Small narrow bat boxes 4-5 metres above the ground with freeaccess seem to be most appropriate <strong>for</strong> this species (Heise 1982, Meschede & Heller 2000).In spite of long distance migration Nathusius’ pipistrelle is very faithful to its place of origin.Females return to their home colony (Heise 1982, Schmidt 1984) and males also come backto their traditional mating areas and mating roosts each year (Meschede & Heller 2000, Boye& Meyer-Cords 2004).Associations were recorded in roosts containing Brandt’s bat, common pipistrelle, noctuleand serotine Eptesicus serotinus (Meschede & Heller 2000, L. Ohlendorf and others 2002).Winter roosts: Crevices in trees and buildings as well as in stapled wood are used as winterroosts (Vierhaus 2004). Holes and crevices in old <strong>for</strong>est and park trees are probably the mostimportant hibernacula of Nathusius’ pipistrelle groups, but there are only a couple ofaccidental observations of these roost types (Meschede & Heller 2000). In The Netherlandsmany hibernating bats were found in towns and other habitats near the coast (Grol & Lina1982).Females migrate from their maternity colony range to distant hibernation sites, thus flyingseveral hundreds of kilometres twice each year. Males do not migrate that far and stay awayfrom the region of maternity colonies in north-eastern Europe (Limpens & Schulte 2000,Vierhaus 2004). Migration obviously takes place along different routes in spring and autumn(Meschede & Helller 2000). The greatest flight distance ever recorded is 1905 kilometresfrom Latvia to southern France (Petersons 1990).Other roosts: Mating roosts are of the same types as summer roosts but most are situated inflood <strong>for</strong>ests of larger rivers, which may serve as guidelines <strong>for</strong> migration (Fiedler 1993,Boye & Meyer-Cords 2004, Arnold & Braun 2002).Foraging areas: Foraging areas are over open water, along <strong>for</strong>est edges, over reeds orpastures, and sometimes in old-growth woodlands (Austria: Bauer & Wirth 1979,Spitzenberger 2001, Denmark: Baagøe 2001a, Fennoscandia: de Jong 1993, Germany: Dense1991, Schmidt 1997, Arnold & Braun 2002, Schorcht and others 2002, Greece: Pieper 1978,von Helversen & Weid 1990, Hanak and others 2001, Hungary: Dobrosi n.y., Italy: Spagnesiand others 2000, Latvia: Petersons 1990, Luxembourg: Harbusch and others 2002, Poland:Ruprecht 1977, 1990, Jarzembowski and others 1998, Russia: Chistyakov 2001, Spain andPortugal: Benzal and others 1991, Switzerland: Gebhard 1995, The Netherlands: Limpens &Kapteyn 1991). During distant flights from a roost to a <strong>for</strong>aging area the bats followlandscape structures, e.g. <strong>for</strong>est edges, hedges, roads or <strong>for</strong>est aisles, but they also go acrossopen fields (Arnold & Braun 2002).Nathusius’ pipistrelle has a home range of 10-22 square kilometres in summer (Schorcht andothers 2002). Certain <strong>for</strong>aging areas may be 6.5 kilometres from the roost site (Boye &Meyer-Cords 2004). The size of a <strong>for</strong>aging area is 18 hectares on average in easternGermany (Eichstädt 1995). In northern Germany, four individual home ranges of females28
from a maternity colony covered a total area of 5.8 square kilometres (Schorcht and others2002). The home range of an entire colony is approximately 80 square kilometres (Meschede& Heller 2000, Arnold & Braun 2002).Abundance: In optimal mating areas 2.3-7.6 males can establish their territories per squarekilometer (Schmidt 1994a).3.11 Serotine Eptesicus serotinusForest habitats: Woodlands are of minor importance to serotines. They may <strong>for</strong>age on<strong>for</strong>est edges, but rarely hunt in a <strong>for</strong>est or roost in a tree.Summer roosts: Preferred roost types are crevices and other narrow holes in houses(Rosenau & Boye 2004). Until now maternity colonies have only been recorded in buildings(e.g. Hübner 1991, Dense 1992, Schmidt 1998, Rosenau 2001, Harbusch 2003). The batsroost below the ridge of a roof (e.g. Schmidt 1998), behind fascia boards (e.g. Rosenau2001), in ventilation holes of new blocks (Bauerova & Gaisler 1985), or in the extension slitsof bridges (Pérez & Ibáñez 1991). Single animals, males in most cases, sometimes use a treehole or a bat box as their roost (Rosenau & Boye 2004). The serotine changes its roost site orhanging place if the microclimate in the roost becomes uncom<strong>for</strong>table, e.g. if temperaturesrise too much (Labee & Voute 1983, Hübner 1991, Schmidt 1998, Rosenau 2001).Maternity colonies have different strategies of roost site utilization: There are records of asingle roost site used <strong>for</strong> the whole maternity period (Glas 1981, Hübner 1991), of a mainroost and a couple of near-by satellite roosts (Dense 1992), and frequent changes within aroost site network (e.g. Schmidt 1998, Dietz & Simon 1999, Rosenau 2001, Simon and others2004).In its summer roosts the serotine can be associated with the greater mouse-eared, Brandt’sbat, the common pipistrelle and the brown-long eared bat (Taake & Vierhaus 1984, Braun2003, own observation), while other small species like the whiskered bat may be pushedaway (Franke 1997).Winter roosts: Winter roosts are in cellars, mines and caves (Heidecke 1987, Haensel 1989,1992), in old buildings and crevices in walls (Hildenhagen & Taake 1982). Single specimensoccasionally hibernate in their summer roost (Lubeley 1998). No records of hibernatingserotine bats are of more than three individuals (Baagøe 2001b).Summer and winter roosts are thought to be less than 50 kilometres apart, however little isknown (Rosenau & Boye 2004).Foraging areas: In most cases the <strong>for</strong>aging areas are open fields with some woods on theedge. In agricultural landscapes the bats prefer pasture with tree rows <strong>for</strong> protection fromwinds (Verboom & Huitema 1997, Schmidt 2000, Lubeley & Bohle 2001). In addition <strong>for</strong>estedges, river banks, parks, tree rows, back yards, sports grounds, and garbage depots areappropriate <strong>for</strong>aging areas (Austria: Spitzenberger 2001, Denmark: Baagøe & Jensen 1973,Baagøe 2001a, Germany: Rosenau 2001, Braun 2003, Greece: Hanak and others 2001,Hungary: Dobrosi n.y., Italy: Spagnesi and others 2000, Luxembourg: Harbusch and others2002, Poland: Kowalski & Ruprecht 1981, Woloszyn 2001, Spain and Portugal: Benzal andothers 1991, Switzerland: Stutz & Burghard 1995, The Netherlands: Limpens & Kapteyn29
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- Page 7 and 8: ContentsPreamble1. Legal aspects...
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- Page 23 and 24: Brandt’s bat travels up to 250 ki
- Page 25 and 26: others 2004). Maternity colonies al
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- Page 33 and 34: female (Fuhrmann and others 2002).
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- Page 37 and 38: Table 2 Importance of tree holes fo
- Page 39 and 40: sites it was found that the possibi
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- Page 45 and 46: Table 6 Overview of the most freque
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- Page 49 and 50: This first approach to a woodland b
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- Page 55 and 56: oxes exposed to the sun. As an exam
- Page 57 and 58: Con: Bats use bat boxes selectively
- Page 59 and 60: 9. ReferencesABULADZE, A., BUXNIKAS
- Page 61 and 62: BEKKER, J.P. 1990. Ervaringen met v
- Page 63 and 64: BRAUN, M., & HÄUSSLER, U. 2003c. K
- Page 65 and 66: Ökologie und Verbreitung von Arten
- Page 67 and 68: (Germany). In: PRIEDE, I. G. & SWIF
- Page 69 and 70: (Nyctalus leisleri) sowie zur Besie
- Page 71 and 72: HEISE, G. 1985. Zu Vorkommen, Phän
- Page 73 and 74: JONES, G. 1990. Prey selection by t
- Page 75 and 76: (Chiroptera: Vespertilionidae). - P
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PÉREZ, J. L. & IBÁÑEZ, C. 1991.
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ROTTMANN, R., BOYE, P. & MEINIG, H.
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SCHORCHT, W., and others. 2002. Zur
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STUTZ, H.-P. B. & HAFFNER, M. 1989.
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WAGNER, M., KERTH, G. & KÖNIG, B.
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10. AuthorsDr. Peter Boye became fa