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New Cases of Intergroup Violence Among Chimpanzees in Gombe ...

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536 Wilson, Wallauer, and Pusey<br />

Fig. 4. Location <strong>of</strong> the 1998 attack on the adolescent<br />

male (square) and the path, nest<strong>in</strong>g and<br />

feed<strong>in</strong>g sites <strong>of</strong> the Kasekela focal chimpanzee<br />

that day.<br />

followed Wilkie. The focal subjects traveled together for most <strong>of</strong> the day,<br />

provid<strong>in</strong>g 2 narrative records for the day’s events.<br />

At 06:55 h, Frodo, Kris, Pax and Wilkie left the others beh<strong>in</strong>d and traveled<br />

to the southeast. At 07:40 h, those left beh<strong>in</strong>d pant-hooted, to which<br />

Frodo and others responded with grunts. The 4 males then rema<strong>in</strong>ed silent<br />

until the time <strong>of</strong> the attack, 3 h and 20 m<strong>in</strong> later. At 07:54 h, they stopped<br />

to feed on fruits <strong>of</strong> Uapaca nitida but they did not pant-hoot on arrival at<br />

the fruit tree. Just before 09:00 h they changed travel direction, return<strong>in</strong>g to<br />

the west, mov<strong>in</strong>g quietly. While they were forag<strong>in</strong>g, they appeared to be uneasy.<br />

They were all very quiet and seemed to be alert and cautious, as if they<br />

had recently encountered signs <strong>of</strong> neighbor<strong>in</strong>g chimpanzees. As the traveled<br />

south, they stopped <strong>of</strong>ten to stare <strong>in</strong>to Nyasanga Valley. They spent long periods<br />

look<strong>in</strong>g up and down the valley and listen<strong>in</strong>g for calls. More than once,<br />

one <strong>of</strong> them erected his body hair and they reached for one another as if<br />

for reassurance. For many m<strong>in</strong> at a time, they sat motionless, star<strong>in</strong>g one<br />

way then another. Pax seemed the most eager to cont<strong>in</strong>ue. He <strong>of</strong>ten moved<br />

well ahead <strong>of</strong> the others, and then waited, look<strong>in</strong>g back. The males spotted<br />

a group <strong>of</strong> red colobus but cont<strong>in</strong>ued patroll<strong>in</strong>g <strong>in</strong>stead <strong>of</strong> stopp<strong>in</strong>g to<br />

hunt.<br />

As they descended <strong>in</strong>to the valley bottom, they quickened their pace.<br />

On reach<strong>in</strong>g the stream at 10:54 h, Frodo, Wilkie, and Kris had their body hair<br />

erect and seemed tense and excited. Just before 11:00 h, Pax spotted a freshly

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