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Some Problems of Reproduction: a Comparative Study of ...

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SOME PROBLEMS OF REPRODUCTION. 37<br />

nucleus occur as follows:—The first differentiates an Apical<br />

(N a a) and a Basal nucleus (N a b) respectively. Each <strong>of</strong> these<br />

again divides to form a pair <strong>of</strong> nuclei nearly superposed, which<br />

we term and letter Apical (N s a), Subapical (N 2 sa), Subbasal<br />

(N 2 sb), and Basal (N 3 b) respectively. Each <strong>of</strong> these four<br />

again divides to form a pair <strong>of</strong> nuclei <strong>of</strong> the degree N s . The<br />

Apical pair (N 3 a + N 3 a) and Basal pair (N 3 b + N s b) are collateral;<br />

the Subapical and Subbasal pairs are superposed to<br />

form a file <strong>of</strong> four nuclei, which we name and letter Upper<br />

and Lower Subapical (N s usa + N 3 lsa), and Upper and Lower<br />

Subbasal (N 3 usb + N 3 lsb) respectively. Of the cytoplasm<br />

lining the cell-wall and surrounding the immense central<br />

vacuole, each nucleus attracts a certain portion so as to form<br />

a naked "free cell" somewhat ill-defined, and indeed anything<br />

but free. The following schema (Fig. 4) shows the for-<br />

rN>a-<br />

-N 2 b-<br />

Synergids<br />

N 3 a N 3 a Egg apparatus.<br />

(<br />

Oospbere i<br />

N 3 usa j<br />

I<br />

lN 3 lsa<br />

ilsb<br />

N 3 b N»b I<br />

Endosperm<br />

nucleus.<br />

! Antipodal<br />

! cells.<br />

FIG. 4.—Schema <strong>of</strong> divisions in embryo-sac; the vertical lines separate

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