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Rynket Klokkemorkel (Verpa bohemica) - en "ny" art for Danmark

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STlayer in th e tr ach eid wall in a characteristic manner<br />

(fig. 8). The dec ayed wood becomes grey with a<br />

soft consist<strong>en</strong>cy. Examples: Fungi Imperfecti and Pyr<strong>en</strong>omycet<br />

es.<br />

O xalic acid has be<strong>en</strong> show n to pl ay a significa nt<br />

rol e in the primar y brake down processes of wood. It<br />

is abl e to hydrolyze the hemicellulose that <strong>en</strong>velope<br />

the cellulose and thus uneover th e cellulose to fur the r<br />

brake do wn by cellulolytic <strong>en</strong>zymes. Th e <strong>en</strong>dproduets<br />

are watersoluble sugars. Oxalic acid is a low moleculal'<br />

subs tance th at can p<strong>en</strong> etrat e th e cell wall. With<br />

ca lcium it fo rms an insoluble sa lt, calcium oxal at e.<br />

T his property <strong>en</strong>a bles th e fung us to reg ulate pH, so<br />

that th e substrat e will not become to acid. Oxalic acid<br />

may also chelat e with differ <strong>en</strong>t metals, a fur the r option<br />

<strong>for</strong> pH regulation . Lar ge quantities of calciumoxa<br />

late is th er e<strong>for</strong> e found in conne ction with brown<br />

rot and in the litt er layer.<br />

Litteratur<br />

Bech-Anders<strong>en</strong> , J . 1985. Basisch e B austoffe und begr<strong>en</strong>zte<br />

Fugtigheitsverhåltnisse. Antwort<strong>en</strong> auf<br />

die Frage , waru m der Echte Haussch wamm nur<br />

in H ansem vorko mmt. - M at eri al und Organi sm<strong>en</strong><br />

Bd . 20 (4): 301-30 9.<br />

Bech-Anders<strong>en</strong> , J . 1986. The infiu<strong>en</strong>ce of the Dry<br />

Rot Fungus (Serpula lacr ymans) in vivo on Insulation<br />

Mat eri ais. - Materiel und Organi sm <strong>en</strong> Bd.<br />

22 (3) : 191-202.<br />

Bech-Anders<strong>en</strong>, J. 1987. Production, function and<br />

neutralisation of ox alic acid produced by the dr y<br />

rot fun gus and othe r brown rot fun gi, - IRG/WPI<br />

1330.<br />

1330.<br />

Chandra, T.S. & Y I. She thna 1975. Isolation and<br />

characterization of some new oxalat e-decomposing<br />

bacteria. - Antonie van Leeuw<strong>en</strong>hoek 41:<br />

101-111 .<br />

Cate, W.A. 1983. The A na to my, U ltrastructure and<br />

Che mical Composition of Wood. I: Biomass Utilization,<br />

W.A. Cate (red.), NATO ASI Series, Series<br />

A: Life Sci<strong>en</strong>ces, Vol. 67. - Pl<strong>en</strong>um Pr ess,<br />

New York & London, p. 249-269.<br />

Esau, K. 1964. A natomy of See d Plants. 4th ed. ­<br />

John Wiley & Son s, Inc. , New York, London &<br />

Sidney, 376 pp.<br />

G oring , D .A.I. 1977. A speculative pietu re of the delignification<br />

process. I: Ce llulose Ch emistry and<br />

Technology, J.e. A rthur (re d.), Washing ton D. e.,<br />

A me rican Che mical Society Symposi um Series<br />

No . 48, p. 273-277.<br />

Graustein, W.e., K. Croma ck & P. Sollins 1977. Calcium<br />

Oxalate: Occur<strong>en</strong>ce in soils and E ffect on<br />

Nutri<strong>en</strong>t and G eochemical Cycles . - Sci<strong>en</strong>ce 198:<br />

1252-1254 .<br />

Gre<strong>en</strong> , F. et al. 1991. Role of Oxalic A cid in Incipi<strong>en</strong>t<br />

Brown-Rot D ecay. Materi al und Organism <strong>en</strong><br />

(i tr ykk <strong>en</strong> ).<br />

Harms<strong>en</strong> , L. 1967. Tr æød elægg<strong>en</strong>de sva mpe og dyr.<br />

- Teknologisk In stituts Forlag, K øb<strong>en</strong>havn, 159<br />

pp .<br />

Hold<strong>en</strong>ried er, O . 1981. Kri stallbildung bei H. annosum<br />

(Fr.) Bref.<br />

Shimada, M. et al. 1991. Biochemical R elationships<br />

bet we<strong>en</strong> Biodegredation of Ce llulose and Formation<br />

of Oxalic acid in Brown-Rot Wood D ecay.­<br />

IRG/WP /1472.<br />

.l.l'..\J1 vv c i i-e t c,

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