(Stachys: Lamiaceae) from the - Botanical Research Institute of Texas

16 Journal of the Botanical Research Institute of Texas 5(1)
11. Leaf blades broader, 5–8 mm wide; plants frequently hispidulous or abundantly pubescent;
blade margins crenulate to serrulate _______________________________________________ S. aspera
9. Leaves ovate to elliptic or oblong, widest near the center or toward the apex; leaf margins crenate to
sharply serrate for nearly the entire length.
12. Stem sides commonly pubescent above.
13. Calyx lobes lanceolate; flowers 6 per verticil; leaf margins serrate-crenate
14. Leaves generally subsessile, the petioles thick, not longer than 6 mm; blades densely
pubescent, frequently soft-pilose or velvety; leaf margins serrate; VA and WV only ______ S. arenicola
14. Leaves generally petioled up to 15 mm long, although shorter, and relatively slender; blades
pubescent on both surfaces, but never pilose or velvety; leaf margins crenate- serrulate;
KY-TN-GA-AL-SC __________________________________________________________ S. nuttallii
13. Calyx lobes deltoid; flowers 8 or more per verticil; leaf margins crenate-crenulate.
15. Lower leaf surface densely sessile- and stipitate-glandular; blades ovate, widest below the
middle; adjacent upper verticils separate in fruit __________________________________S. eplingii
15. Lower leaf surface glandular, but not densely so; blades elliptic-ovate, widest at the middle;
adjacent upper verticils commonly crowded in fruit _________________________________ S. iltisii
12. Stem sides glabrous, the angles pubescent.
16. Fruiting calyx lobes lanceolate or deltoid-acuminate, more than half as long as the calyx tube.
17. Calyx hispidulous to strongly hispid with long, stiff hairs; stem angles moderately to
densely pubescent; leaves subsessile, with very short petioles; bracts abruptly reduced
upward from lowest flowering node ____________________________________________ S. hispida
17. Calyx sparsely to moderately pubescent with short soft hairs; stem angles moderately
pubescent to nearly glabrous; leaves with petioles 1 to 3 cm long; bracts gradually reduced
upward from lowest flowering node ________________________________________ S. subcordata
16. Fruiting calyx lobes triangular-deltoid and abruptly apiculate, less than half as long as the calyx
tube.
18. Stem nodes not bearded (sometimes obscurely fine- hairy); stem angles with scattered,
pustulate, short deflexed hairs; plants of high-elevation montane habitats _____________ S. latidens
18. Stem nodes bearded; lower stem angles copiously pubescent with 3-celled spreading hairs;
plants of low-elevation Piedmont and Coastal Plain habitats _____________________ S. matthewsii
key 2
1. Fruiting calyx lobes deltoid to broadly triangular, less than half as long as the calyx tube.
2. Stem sides pubescent; foliage glandular, often musky-scented when crushed.
3. Petioles short to absent, blades mostly rounded to truncate-cordate.
4. Lower leaf surface densely glandular with atomiferous/short- stipitate golden glands; blades
generally ovate, broadest toward the base ______________________________________________S. eplingii
4. Lower leaf surface glandular, but not densely so; blades generally elliptic, somewhat lyrate toward
base, broadest near the middle _________________________________________________________ S. iltisii
3. Petioles well developed, blades cordate at base.
5. Top of blooming stem frequently flexuous, somewhat lax; blades ovate-rounded ________________S. cordata
5. Top of blooming stem stiffish, not lax; blades oblong-elliptic _______________________________ S. nuttallii
2. Stem sides glabrous; foliage not glandular (occasionally with a few scattered glands) or musky-scented.
6. Stem nodes not bearded (sometimes obscurely fine-hairy); stem angles with scattered, pustulate, short
deflexed hairs; plants of high-elevation montane habitats ___________________________________ S. latidens
6. Stem nodes bearded; lower stem angles copiously pubescent with 3-celled spreading hairs; plants of
low-elevation Piedmont and Coastal Plain habitats ______________________________________ S. matthewsii
1. Fruiting calxy lobes lanceolate, narrowly triangular, deltoid-acuminate, or nearly subulate, more than half as
long as the calyx tube.
7. Leaf blades linear to lanceolate.
8. Leaf blade margins entire to crenate; plants generally glabrous ____________________________ S. hyssopifolia
8. Leaf blade margins serrulate with at least a few teeth; plants glabrous or pubescent.
9. Stems strict or sparingly branched; leaves sessile or barely petioled, the blades crenate to serrate
with shallow teeth _________________________________________________________________ S. aspera
9. Stems frequently lax and branched from the upper nodes; leaves obviously petioled, the blades
sharply toothed _________________________________________________________________ S. tenuifolia
7. Leaf blades wider, rounded, oblong to elliptic.