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472 OSBORN, THE REPTILIAN SUBCLASSES DIAPSIDA AND SYNAPSIDA. rather like a terrestrial type (Fig. io): see especially the three expanded sacral vertebrae, the dorsally expanded ilium, the broad neural spines, stout pre- and postzygapophyses, digits terminating in stout ungual phalanges, with lateral grooves for claw attachment, moderate development of the caudals. As compared with Pro/orosaurus and Kadaliosaurus, Palaohadleria was certainly clumsy, heavy-limbed, and slow moving; the powerful recurved teeth are adapted to the seizure of comparatively resistant prey. There is very considerable reason to believe that all the specimens which represent this remarkable type belong to small and immature individuals, differing in size and age, in which the ossification of the ends of the bones is still incomplete. This conjecture is confirmed by the fact that in none of the six specimens figured by Credner is the posterior cranial region ossified, and that in the development of Sphenodou this region long remains cartilaginous (Howes). The shoulder girdle, carpus, and tarsus are also imperfectly osseous. SKULL.-Length, 70 mm. shaped as in Sphenodon, that is, apparently elevated and relatively narrow, with short temporal fenestrx; with all primitive elements separate, namely, premaxillaries, maxillaries, nasals, lachrymals, prefrontals, frontals, postfrontals, postorbitals, squamosals. (The existence of separate prosquamosals is undetermined; Credner does not X mention these elements.) Supra- and laterotemporal fenestre large and of equal size. Cranium abbreviate, antorbital region more elongate than postorbital region ; nares small, terminal, separated by premaxillary septum, no preorbital opening; orbits large, with a ring of sclerotic ossicles. Premaxillaries separate, with vertical processes separating nares. Squamosal and quadrate apparently somewhat depressed. Basisphenoid perforated with two foramina as in Sphenodon, with a parasphenoid (vomer) process. tV -f DENTITION.-Upper teeth, 21-22 on each side; teeth hollow, recurved, 32,
OSBORN, THE REPTILIAN SUBCLASSES DIAPSIDA AND SYNAPSIDA. 473 ABDOMINAL RIBS.-Small, elongate, scale-like, 3 transverse series to each opposite p)air of costal ribs. GIRDLES.-Scapula large, exceptionally broad; coracoids only osseous, resembling the embryonic coracoid of Sphenodon (Fig. 6), and imperforate; no procoracoid ossification in ventral plate procoracoid cartilaginous, unless represented in Credner (Tab. xxiv, fig. i) as an osseous element overlapping the coracoid. Interclavicle a rhombic anterior plate (as in Stegocephala) with elongate, narrow posterior bar. Large curved clavicles. Pelvic girdle composed of a continuous ventral plate, more rounded pubis with incisio-obturatoria, elongated ischia and dorsally expanded ilium, with a prominent anterior or pubic, and a broad but somewhat less promlinent ischiadic peduncle. LIMBS.-Relatively heavy and compact, the ends of all the limb bones cartilaginous. Hind limbs a little longer than fore limbs. Propodials of nearly equal size (that is, radius and ulna equal tibia and fibula). Humerus expanded distally more than proximally, with supposed radiocondylar (ectepicondylar) foramen. Femur straight, flattened; tibia and fibula equal in size. Manus longer than pes, composed of more slender elements. Carpus composed of 8 separate elements. Phalangeal formula in manus and pes 2, 3, 4, 5, 3. Phalanges stout, cylindrical; terminal phalanges laterally compressed and grooved for claw. Tarsus (Fig. ii) composed of at least seven elements, namely, a large astragalus (tibiale + ? intermedium), calcaneum (fibulare) and five distal tarsalia. RELATIONSHIPs.-Credner enumnerates (op. cit., page 548) fourteen characters in which this animal resembles Sphenodon puncta/us. These are without exception primitive characters and speak for the remarkably primitive, persistent nature of the New Zealand species. Paleeohatteria, however, is still more primitive, in: (i) its solid pelvic plates, (2) its five distal tarsalia, (3) its simple caudal vertebrae without breaking points, (4) its rhomboidal interclavicle. It resembles Prolorosaurus, and is inore specialized than Spizenodon, in: (i) the presence of three sacrals, (2) the dorsal expansion of the ilium, (3) the reduction of the proximal row of tarsals to two elements. All the progressive characters of Palcohatteria point toward the Dinosauria rather than toward the Rhynchocephalia. Genus Kadaliosaurus Credner.' MATERIALS.-A single specimen from the Mittel Rothliegenden (Lower Permian) associated with remains of Stegocephala (Branchiosaurus, Archegosaurus, etc.) Size.-Portion preserved, 225 mm.-about equal to the distance between the humerus and the femur. VERTEBRA:.-Amphiccelous perforated by large continuous notochord, hypocentra if present not exposed to view. Rudimentary diapophyses on the dorsals. Dorsal vertebra elongate; 20 or more dorsals (15-I6 only preserved); the back is certainly long; 2 sacrals. The 8 anterior caudals preserved apparently indicate that the tail was not very long. RIBs.-The ribs indicate a deep, lightly constructed chest (longest rib 33 mm.). Ribs apparently hollow, capitulum and tuberculum not distinctly separated, capitula attached apparently to anteroinferior portion of the centrum or between centra, without union with a diapophysis. A large separate sacral rib and probably a second one concealed. Four rather strong anterior caudal ribs united with vertebre anteriorly. I Die Stegocephalen und Saurier, etc., VIII. Theil. Zeitschr. d. Deutsch. geol. Ges., Bd. XLI, I889.
Page 1 and 2: MEMOI RS OF THE Se t r H isto-ry. V
Page 3: VOLUME I, PART VIII. THE REPTILIAN
Page 6 and 7: 452 OSBORN, THE REPTILIAN SUBCLASSE
Page 34 and 35: Po.Qi 480 OSBORN, THE REPTILIAN SUB
Page 36: 482 OSBORN, THE REPTILIAN SUBCLASSE
Page 40 and 41: Fig. 17 a. Stereosternum tumidum Co
Page 42 and 43: | J j 486 OSBORN, THE REPTILIAN SUB
Page 62 and 63: ol 506 OSBORN, THE REPTILIAN SUBCLA
Page 65 and 66: INDEX TO VOLUME I. [New names of ge
Page 67 and 68: Aphelops profectus, 358, 373 superc
Page 69 and 70: Hoplophoneus pr1mevus, 394 robustus
Page 71 and 72: Pleurosaurus, 463, 468, 494, 495, 4
Page 73 and 74: Sesia albicornis, 226, 229, 230, 27
Page 75: Trochilium tipuliformis, 286 tricin
Page 78:
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