A study on the genus Chrysolina MOTSCHULSKY, 1860, with a ...

106 ANDRZEJ BIEÑKOWSKI
the subgenus Ch. (Chrysocrosita), and Ch. subcostata poretzkyi JACOBSON - in the
subgenus Ch. (Pleurosticha). The genus Cecchiniola JACOBSON is included in the
genus Chrysolina as a subgenus. Timarchomima (with a single species T. indica) is
regarded as a separate genus within the subtribe Chrysolinina, but not a subgenus of
Chrysolina. Therefore, Timarcholina is a valid subgeneric name for the other former
members of Timarchomima. Lectotypes of Taeniosticha poricollis MOTSCHULSKY,
Chrysomela pterosticha FISCHER DE WALDHEIM, Chrysolina nikinoja nikinoja BECHYNÉ,
Ch. nikinoja exgeminata BECHYNÉ, Crosita alaschanica JACOBSON, and C. przewalskyi
JACOBSON are designated. Ch. blanchei FAIRMAIRE is recorded from Turkey, Ch. aveyronensis
BECHYNÉ - from Spain, Ch. numida REICHE and Ch. pardoi CODINA PADILLA -
from Tunisia, Ch. koenigi JACOBSON - from Kyrghyzstan, Ch. brunnicornis bermani
L.MEDVEDEV - from Chita reg., Ch. lopatini MOHR - from Altai, Ch. grata grata - from
Sinai, Ch. analis - from Algeria, Ch. marginata borealis - from Taimyr, and Ch. eurina
- from European Russia (Moscow reg.) for the first time.
Key words: Entomology, Coleoptera, Chrysomelidae, Chrysolina, checklist, taxonomy.
INTODUCTION
Chrysolina is a very large and diverse genus of leaf-beetles. Most species are
distributed in Europe, Asia and Africa. A small number of species inhabits
N. America (including introduced European ones). Some species were introduced
into Australia. The last complete catalogue of Chrysolina was published by
WEISE (1916). In this catalogue the species were arranged alphabetically, not
corresponding to the subgenera. A taxonomical review of the genus by BECHYNÉ
(1950, 1952) did not include all species known at that time, and some new
subgenera and species were described later.
The present checklist is the first step of my taxonomical revision of the genus
Chrysolina at the subgeneric level.
All available names of generic and specific levels belonging to Chrysolina
are included. The names, which were first proposed as infrasubspecific ones and
not arised to subspecific or specific rank, are unavailable (International Code of
Zoological Nomenclature 1999, Art. 10.2). Therefore, I did not include such
names in the checklist. The original rank of some old synonyms, which were cited
after WEISE (1916), is still unknown to me. Probably, some of them are
infrasubspecific and thus unavailable.
Papers devoted to systematics and distribution of certain taxa are numbered,
and their respective numbers (in square brackets) are given in the checklist. In the
checklist, subgenera, species within each subgenus, and subspecies within each
species are arranged alphabetically.
I studied original descriptions of all the subgenera and most of the species,
many other taxonomical papers (see References), „Coleopterorum Catalogus”
(WEISE, 1916), and 45-135 volumes of „Zoological Record”. Specimens of 277
species of Chrysolina were examined (marked with asterisk in the checklist).