134 ANDRZEJ BIEÑKOWSKI 27-30. Aedeagus: 27 - Ch. taygetana, 28 - Ch. cilissa (holotype), 29 - Ch. villiersi ruficornis (paratype), 30 - Ch. kinabaluensis
A STUDY ON THE GENUS CHRYSOLINA 135 truncate and appendiculate at apex, twice curved in lateral view, and resembles that of Ch. (Chalcoidea) tesari or Ch. (Pezocrosita) pusa, but never occurs in <strong>the</strong> members of <strong>the</strong> sub<strong>genus</strong> Allohypericia. The last abdominal sternum in male of Ch. seriepunctata bears a semicircular apical impressi<strong>on</strong>, like in some species of <strong>the</strong> sub<strong>genus</strong> Pleurosticha, but never occurs in <strong>the</strong> species of <strong>the</strong> sub<strong>genus</strong> Allohypericia. Therefore, I can not assign Ch. seriepunctata to any sub<strong>genus</strong> described before. MATERIAL Ch. seriepunctata: Amur reg.: between Malaja Pera and Bolshoj Ergel rivers, 6.6.1958: 1 male; Primorski Krai: envir<strong>on</strong>s of Kamenushka, 2.8.1989: 1 female, S. KHVYLIA leg.; Shkotovo, Maikhe, 3.6.1927: 1 female, REZVOJ leg. 67. KASAP (1988) studied <strong>the</strong> type specimen of Ch. bruneli, redescribed this species and figured <strong>the</strong> aedeagus. This species has colorati<strong>on</strong> which is rare <strong>with</strong>in <strong>Chrysolina</strong> and resembles that of Ch. koktumensis and Ch. nigrovittata: “Elytra yellowish red <strong>with</strong> black l<strong>on</strong>gitudinal bands <strong>on</strong>e al<strong>on</strong>g <strong>the</strong> suture, o<strong>the</strong>r al<strong>on</strong>g <strong>the</strong> middle of each elytr<strong>on</strong>, rest of <strong>the</strong> body shining black”(KASAP 1988). Unfortunately, KASAP (1988) did not describe <strong>the</strong> shape of maxillary palpus, positi<strong>on</strong> of antennal inserti<strong>on</strong>, shape of prothoracic hypomer<strong>on</strong>, positi<strong>on</strong> of elytral epipleur<strong>on</strong>, structure of tarsi, pygidium, and <strong>the</strong> last abdominal sternite, presence or absence of humeral calli and hind wings. Therefore, it is impossible to solve <strong>the</strong> questi<strong>on</strong> of <strong>the</strong> systematic positi<strong>on</strong> of this species. 68. Ch. indica has metasternum not extending forward, <strong>with</strong> anterior border unmarginate at middle. Such a structure of metasternum is present in allied Asian genera, namely, Humba CHEN, 1934, Sphaerolina BALY, 1871, and Ambrostoma <strong>MOTSCHULSKY</strong>, <strong>1860</strong>, and in an African <strong>genus</strong> Sphaeratrix GISTEL, 1848. On <strong>the</strong> o<strong>the</strong>r hand, wholly margined metasternum is typical of <strong>Chrysolina</strong>. Within this large <strong>genus</strong>, <strong>on</strong>ly three African species, namely, Ch. superba, Ch. ambrostomoides (both are close to Mediterranean Ch. americana), and Ch. beatricis (unknown to me) have unmargined metasternum. The structure of prosternum in Ch. indica (broadened backwards, <strong>with</strong> deep emarginati<strong>on</strong> at apex) looks like that of members of an Asian <strong>genus</strong> Ambrostoma and Australian <strong>genus</strong> Promechus BOISDUVAL, 1835 and str<strong>on</strong>gly differs from that of o<strong>the</strong>r <strong>Chrysolina</strong> members. BECHYNÉ (1950) designated Ch. indica as <strong>the</strong> type species of <strong>the</strong> sub<strong>genus</strong> Timarchomima BECHYNÉ, 1950: 65. I c<strong>on</strong>sider Timarchomima to be a m<strong>on</strong>otypic <strong>genus</strong> <strong>with</strong>in <strong>the</strong> subtribe Chrysolinina. Besides, I was able to examine some o<strong>the</strong>r former members of Timarchomima, namely, Ch. clavareaui and Ch. templet<strong>on</strong>i. The shape of prosternum and metasternum in <strong>the</strong>se species is typical of <strong>Chrysolina</strong>. Ch. templet<strong>on</strong>i was designated by BECHYNÉ, 1950 as <strong>the</strong> type species of <strong>the</strong> sub<strong>genus</strong> Timarcholina, so <strong>the</strong> latter name should be treated as a valid subgeneric name.
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