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the british - Corning Museum of Glass

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xii INTRODUCTION.<br />

1. Tegumentary System. The skin is sufficiently distinct.<br />

After a few hours' maceration in fresh water (Sag. bellis),<br />

<strong>the</strong> epi<strong>the</strong>lial and pigmental cells are easily removed with<br />

a hair-pencil, leaving <strong>the</strong> outer layer <strong>of</strong> muscular fibre bare.<br />

If <strong>the</strong> specimen be immersed in spirit for a day or two<br />

(A. dianthus), <strong>the</strong> integument may be separated in flakes,<br />

which, under <strong>the</strong> microscope, are seen to be composed <strong>of</strong> a<br />

multitude <strong>of</strong> short corrugated fibrillæ, set in no definite<br />

direction, interspersed with clear granules, pigment grains,<br />

and cnidæ.<br />

An examination <strong>of</strong> <strong>the</strong> living animal (dianthus, bellis,<br />

crassicornis, Halc, chrysan<strong>the</strong>llum, Cor. viridis, &c.) shows<br />

that <strong>the</strong> skin is composed <strong>of</strong> three elements, though <strong>the</strong>se<br />

cannot always be separated. A layer <strong>of</strong> epi<strong>the</strong>lial ciliated<br />

cells forms <strong>the</strong> first tunic: <strong>the</strong>se are constantly in process<br />

<strong>of</strong> being thrown <strong>of</strong>f from <strong>the</strong> true skin, in <strong>the</strong> form <strong>of</strong><br />

mucus; but in some cases (Phellia,Edwardsia) <strong>the</strong>y<br />

entangle foreign matters, and retain <strong>the</strong>ir cohesion as an<br />

investment more or less dense, and more or less firmly<br />

adherent to <strong>the</strong> skin. Below this is <strong>the</strong> true skin, <strong>of</strong> a<br />

more granular character, and carrying, imbedded in its<br />

thickness, a multitude <strong>of</strong> cnidæ, whose discharging points<br />

are directed outwards. Intimately connected with this<br />

layer, but still lying sufficiently beneath it to be regarded<br />

as a distinct stratum, are <strong>the</strong> pigment-cells, which impart<br />

<strong>the</strong> colours to <strong>the</strong> animal.<br />

The tentacles <strong>of</strong> Aiptasia and An<strong>the</strong>a (less conspicuously<br />

also <strong>of</strong> S. bellis) are lined with a dense layer <strong>of</strong> cells, forming<br />

to <strong>the</strong> naked eye a dark brown lining. Some peculiarities<br />

<strong>of</strong> <strong>the</strong>se cells I have detailed (at page 167, infra): it is<br />

probable that this layer may have some special function yet<br />

unrecognised.<br />

2. Muscular System. In most species <strong>the</strong> muscular framework<br />

<strong>of</strong> <strong>the</strong> body is beautifully distinct, and <strong>the</strong> tissue is<br />

readily isolable. The column is a cylinder <strong>of</strong> muscular<br />

tissue, consisting <strong>of</strong> two layers, <strong>the</strong> outer composed <strong>of</strong><br />

transverse, <strong>the</strong> inner <strong>of</strong> longitudinal, fibres. The transverse<br />

fibres are <strong>the</strong> more strongly marked: <strong>the</strong>y average<br />

about 0001 inch in diameter, and are never striate.<br />

The cylinder which forms <strong>the</strong> column, is closed in most<br />

species by two extremities, which are flat, like <strong>the</strong> top and<br />

bottom <strong>of</strong> a tin canister: <strong>the</strong> former is <strong>the</strong> disk, <strong>the</strong> latter<br />

<strong>the</strong> base. Each <strong>of</strong> <strong>the</strong>se is but a continuation <strong>of</strong> <strong>the</strong> same

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