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Density and species diversity of trees in four - Makerere University

Density and species diversity of trees in four - Makerere University

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G. Eilu et al.<br />

Family<br />

No. <strong>of</strong><br />

Genera<br />

No. <strong>of</strong><br />

Species Family<br />

gradient (Ter Braak & Smilauer, 1998). The first <strong>four</strong> axes <strong>of</strong><br />

CCA <strong>and</strong> DCA, respectively, expla<strong>in</strong>ed 16.2 <strong>and</strong> 19.1% <strong>of</strong> the<br />

cumulative variance <strong>in</strong> <strong>species</strong> data (Table 4). The third <strong>and</strong><br />

<strong>four</strong>th axes with eigenvalues < 0.250 were less important <strong>in</strong> ecological<br />

terms <strong>and</strong> are not considered further. The sample scores<br />

on the first CCA axis were most strongly correlated with pH <strong>and</strong><br />

those <strong>of</strong> the second axis with altitude. Correlations between<br />

environmental variables <strong>and</strong> DCA ord<strong>in</strong>ation axes confirmed<br />

No. <strong>of</strong><br />

Genera<br />

Euphorbiaceae 16 25 Ochnaceae 2 2<br />

Rubiaceae 12 16 Olacaceae 2 2<br />

Meliaceae 8 16 Verbenaceae 2 2<br />

Fabaceae 12 13 Icac<strong>in</strong>aceae 1 2<br />

Sapotaceae 6 11 Agavaceae 1 1<br />

Sap<strong>in</strong>daceae 7 8 Alangiaceae 1 1<br />

Apocynaceae 5 8 Aquifoliaceae 1 1<br />

Moraceae 5 8 Araliaceae 1 1<br />

Ulmaceae 4 8 Arecaceae 1 1<br />

Flacourtiaceae 6 7 Asteraceae 1 1<br />

Sterculiaceae 5 7 Balanitaceae 1 1<br />

Annonaceae 4 6 Canellaceae 1 1<br />

Unknown 1 5 Chrysobalanaceae 1 1<br />

Anacardiaceae 4 4 Cyatheaceae 1 1<br />

Clusiaceae 4 4 Dichapetalaceae 1 1<br />

Simarubaceae 4 4 Ebenaceae 1 1<br />

Rutaceae 3 4 Hypericaceae 1 1<br />

Tiliaceae 3 4 Loganiaceae 1 1<br />

Rhizophoraceae 1 4 Melastomataceae 1 1<br />

Bignoniaceae 3 3 Melianthaceae 1 1<br />

Capparidaceae 3 3 Monimiaceae 1 1<br />

Oleaceae 3 3 Myristicaceae 1 1<br />

Rhamnaceae 3 3 Myrtaceae 1 1<br />

Borag<strong>in</strong>aceae 2 2 Rosaceae 1 1<br />

Cecropiaceae 2 2 Theaceae 1 1<br />

Celastraceae 2 2 Violaceae 1 1<br />

Lauraceae 2 2 — — —<br />

No. <strong>of</strong><br />

Species<br />

Table 3 Tree families, numbers <strong>of</strong> genera <strong>and</strong><br />

<strong>species</strong> recorded from Bw<strong>in</strong>di, Kasyoha, Kibale<br />

<strong>and</strong> Budongo forests <strong>of</strong> the Albert<strong>in</strong>e Rift<br />

Table 4 Eigenvalues <strong>of</strong> the first <strong>four</strong> axes <strong>of</strong> canonical correspondence analysis (CCA) <strong>and</strong> Detrended correspondence analysis (DCA) <strong>of</strong> all<br />

plots <strong>and</strong> the amount <strong>of</strong> variance expla<strong>in</strong>ed <strong>of</strong> the <strong>species</strong> data <strong>and</strong> <strong>of</strong> the <strong>species</strong>-environment relation by the CCA axes. The length <strong>of</strong> gradient<br />

is shown for DCA<br />

Axes 1 2 3 4 Total <strong>in</strong>ertia<br />

Eigenvalues (CCA) 0.480 0.259 0.228 0.128 6.757<br />

(DCA) 0.709 0.251 0.207 0.149 6.879<br />

Lengths <strong>of</strong> gradient (DCA) 5.259 2.772 2.700 2.191 —<br />

Species-environment correlations (CCA) 0.861 0.820 0.950 0.755 —<br />

Cumulative percentage variance <strong>of</strong> <strong>species</strong> data (CCA) 7.1 10.9 14.3 16.2 —<br />

(DCA) 10.3 14.0 17.0 19.1 —<br />

Cumulative percentage variance <strong>of</strong> the <strong>species</strong>-environment<br />

relation (CCA)<br />

30.1 46.4 60.7 68.7 —<br />

Sum <strong>of</strong> all unconstra<strong>in</strong>ed eigenvalues (CCA) — — — — 6.757<br />

(DCA) — — — — 6.757<br />

Sum <strong>of</strong> all canonical eigenvalues (CCA) — — — — 1.593<br />

results <strong>of</strong> CCA analysis with axis 1 most strongly correlated with<br />

pH (r = 0.565).<br />

Automatic forward selection <strong>of</strong> environmental variables by<br />

canoco (Table 6) showed that pH expla<strong>in</strong>ed the most variance<br />

(0.39) while silt expla<strong>in</strong>ed the least (0.08%). After select<strong>in</strong>g pH,<br />

altitude, ra<strong>in</strong>fall, loss <strong>of</strong> organic matter on Ignition, potassium<br />

(K), available phosphorus <strong>and</strong> slope contributed significantly (at<br />

5% level) to the model <strong>of</strong> already <strong>in</strong>cluded variables.<br />

306 Diversity <strong>and</strong> Distributions, 10, 303–312, © 2004 Blackwell Publish<strong>in</strong>g Ltd

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