The Ecology of Lake St. Clair Wetlands: A Community Profile
The Ecology of Lake St. Clair Wetlands: A Community Profile
The Ecology of Lake St. Clair Wetlands: A Community Profile
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Char1 es E. Herdendorf<br />
Center for <strong>Lake</strong> Erie Area Research<br />
Department <strong>of</strong> Zoo1 ogy<br />
<strong>The</strong> Ohio <strong>St</strong>ate University<br />
Columbus, OH 43210<br />
and<br />
C. Nicholas Raphael<br />
Eugene Jaworsk j<br />
Department <strong>of</strong> Geography and Geology<br />
Eastern Michigan University<br />
Ypsil anti, MI 48197<br />
Project Officer<br />
Walter G. Duffy<br />
Natlonal Wet1 ands Research Center<br />
U. S. Fish and Wildlife Service<br />
2010 Gause Boulevard<br />
S1 idell, LA 70458<br />
Prepared for<br />
National <strong>Wetlands</strong> Research Center<br />
Fish and Wild1 ife Service<br />
U. S, Department <strong>of</strong> the Interior<br />
Washington, DC 20240<br />
Fur salr by the Superizitt.ndrn: U! I)~c~rnenf,<br />
U.S. Guvernmrn!<br />
Printing Offrrc. U'ashrngton. D.C. 20102<br />
Biological Report 85(7.7)<br />
September 1986
tierdcndorf, Charle5 T.<br />
<strong>The</strong> ccolnqy <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Ila~r wetldnd~~.<br />
(R~oloqical rcport ; 8!7( J. i))<br />
"Sep tcarirer 1911b."<br />
Supt. <strong>of</strong> Ilocc;. no.: I 49.R0/i':ti'~(i.?)<br />
I, tictlanci t~ology--Saint (,lair, I.crke (Micii. arid Oiit.)<br />
?. Ndn--1rif luent e on ndture--Sdint ("l~flr, I d/\~ (Mi~h.<br />
drld 011t.) 3. Lic)tlar~d ccoloc~y--Great i ~ ikrs Rccjion.<br />
4, Mdrr--1trfI~ic~ntc on rratut-c--Gr-edt I crkcs Krcjl'on.<br />
1, Ral)t?~tcl , t. Nictlolds, lQ'37- . l i. ,Icrwor-ikr,<br />
Eutjc.rie, i i l. Ndtiondl Wctfdr~rls iie$c-'rdt ti C~nter (11.4.)<br />
IV. T~tlc. V. Titlc: icolarjy <strong>of</strong> 1 akr 'Liint (Idi.<br />
wctlilndi. VI. C;ilt.ics: fiiolor~it~tl t*i")or+t (W(~rlshincjl.:~r~,<br />
l1.L.) ; !3!>-7* 7*<br />
fhii report may be cited as:<br />
Herdendnrf, C. t, , C. N. Rdyhac? , and E. Jaworskl 1986. <strong>The</strong> ecology <strong>of</strong> iake<br />
<strong>St</strong>. Ciair-weliarlas: d~~~ii~lilfr~it~~i~;i:~.<br />
3.5. Fi~f;W:id!. Ser-t. Sjc?. Rep.<br />
85(7.1) 187 pa,
This report on <strong>Lake</strong> S t , <strong>Clair</strong><br />
wetlands is one <strong>of</strong> a series <strong>of</strong> community<br />
pr<strong>of</strong>iles that deal with marine and<br />
freshwater habitats <strong>of</strong> ecological impor-<br />
tance. <strong>The</strong> delta marshes, estuaries,<br />
lagoons, and channel wetlands which fri nge<br />
the Michigan and Ontario shores <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>, and the <strong>St</strong>. <strong>Clair</strong> and Detroit<br />
rivers, are among the most productive<br />
areas in the Great <strong>Lake</strong>s. Because they<br />
occur in the proximity <strong>of</strong> a densely<br />
populated, heavily industrial ized, and<br />
intense1 y farmed region, the marshes have<br />
suffered losses in both area and qua1 ity.<br />
However, the remaining marshes are vital<br />
habitats for migratory waterfowl, fur-<br />
bearers, and fish, and perform many<br />
important hydrological and ecological<br />
functions. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta is unique<br />
in the Great <strong>Lake</strong>s and fosters some <strong>of</strong> the<br />
finest coastal wetlands in the region.<br />
<strong>The</strong> biological resources <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> have attracted man since prehistoric<br />
time. Not only were fish and wlldlife<br />
consumed but the wetland plants were used<br />
for crafts, cures, and kitchen utensils by<br />
the early Indian settlers. Archaeological<br />
sites and early explorers <strong>of</strong> the region<br />
have recorded the close interactJon <strong>of</strong><br />
early man and the coastal environment.<br />
<strong>Lake</strong> <strong>St</strong>. Clafr was lldiscoveredN by<br />
Europeans on Saint Clalreqs Day, August<br />
12, 1679, and was named in honor <strong>of</strong> the<br />
saint. Early fur traders were consls-<br />
tently overwhelmed by the bountiful fish<br />
and wildlife stocks <strong>of</strong> the lake and <strong>of</strong>ten<br />
remarked in their journals about the<br />
tranquil beauty <strong>of</strong> the marshesc flora. A<br />
colorful episode <strong>of</strong> the regfon was l inked<br />
to the prohibitton era. During this time,<br />
local residents claim that the <strong>St</strong>. <strong>Clair</strong><br />
Flats, or delta, was a hub <strong>of</strong> activity for<br />
the importation <strong>of</strong> i 1 legal 8 but excel 1 ent<br />
quality* whiskey from Canada to the United<br />
<strong>St</strong>ates. Today, <strong>Lake</strong> <strong>St</strong>. Clais is one <strong>of</strong><br />
PREFACE<br />
the most intensively utilized lakes in<br />
North America with its International<br />
wetlands continuing to be significant<br />
contributors to recreational activities<br />
throughout the year. Without these wet-<br />
lands, the value <strong>of</strong> the lake would be<br />
greatly diminished.<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is noted for its<br />
storms and rapid water level changes. <strong>The</strong><br />
moderate energy produced by these storms<br />
1 imits the existence <strong>of</strong> coastal wetl ands<br />
to places where some type <strong>of</strong> natural or<br />
artificial protection is available.<br />
Corresponding1 yr the coastal marshes <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fall Into four categorf es:<br />
1) delta wetlands, 2) coastal lagoons<br />
behind protective barrier beaches, 3)<br />
estuarine tributary mouths, and 41 managed<br />
marshes protected by earthen and rip-rap<br />
dikes. According to Gowardin et al,<br />
(1979) these wetl ands would include<br />
elements <strong>of</strong> riverine, 1 acustrf ner and<br />
palustrine systems <strong>of</strong> the recently<br />
developed National Wet1 and Cl ass T f ication.<br />
<strong>The</strong> li.Jossary Geol09y (Bates and<br />
Jackson 1980) def f nes '"freshwater estu-<br />
aries" for the Great <strong>Lake</strong>s as, '%he lower<br />
reach <strong>of</strong> a tributary to the lake that has<br />
a drowned rtver moutht shows a zone <strong>of</strong><br />
transition from stream water to lake<br />
water, and is influenced by changes fn<br />
lake level as a result <strong>of</strong> sefches or wind<br />
tides." Brant and Herdendorf ( 19721 were<br />
among the f f r st investigators to descrf be<br />
the characterfstics <strong>of</strong> Great <strong>Lake</strong>s<br />
estuaries. Such estuarfes are important<br />
wetland habitats in western <strong>Lake</strong> Erie.<br />
<strong>The</strong> definitfon gfven above provides an<br />
adequate physical decrfption for the<br />
purposes sf thf s report.<br />
<strong>The</strong> information in this report Is<br />
intended to provide a baslc understandfng<br />
<strong>of</strong> the ecological relatfonships wfthln the
<strong>Lake</strong> <strong>St</strong>. CY air wetlands and the impact <strong>of</strong><br />
natural and man-i nduced disturbances on<br />
the coastal marsh community. References<br />
are provided for those seeking more<br />
detai 1 ed treatment <strong>of</strong> speci f l c aspects <strong>of</strong><br />
the coastal marsh ecology. An appendix 4s<br />
included which lists the dimensions and<br />
ownership <strong>of</strong> the major marshes, and the<br />
important biologfcal species <strong>of</strong> a1 yae,<br />
macrophytes, invertebrates, fish,<br />
amphi bians, reptiles, birds# and mammals<br />
occurring in the coastal marshes,<br />
Any questions or comments about, or<br />
requests for this publication should be<br />
addressed to:<br />
Informat-ion Transfer Speci a1 ist National Wet1 ands Research Center<br />
U. S. Fish and Wildlife Service<br />
NASA-Sl idell Computer Complex<br />
1010 Gause Boulevard<br />
Sl idell, LA 70458<br />
(50% i 546-73 10 FTS 680-73 20
FOR NlETRlC (SI) UNITS TO U.S. CUSTOMARY UNITS OF MEASUREMENT<br />
Metric (SI) units <strong>of</strong> measurement used in this report can be converted to U.S.<br />
customary units as follows:<br />
Linear measurements;<br />
mill imetres (mm)<br />
cent imetres (cm)<br />
metres (m)<br />
k i 1 ometres (km)<br />
square metres (m2)<br />
hectares (ha)<br />
square kilometres (km2)<br />
Yo1 ume measurements;<br />
cublc metres (dl<br />
cubic metres (dl<br />
cubic kilometres (kd)<br />
Mass measu r emenk<br />
grams (g)<br />
kilograms (kg)<br />
metric tons (m ton)<br />
Bate measurements;<br />
0.039 inches (in)<br />
0.394 inches (in)<br />
3.281 feet (ft)<br />
0.621 miles (mi)<br />
10.764 square feet (ft2)<br />
2.471 acres<br />
0.386 square miles (mi21<br />
35.318 cubic feet (ft3)<br />
1.308 cubic yards iyd3)<br />
0.240 cubic miles (mi31<br />
0.035 ounces (ozl<br />
2.205 pounds (lbf<br />
1.102 U.S. tons (ton)<br />
centimetres per second (cm/sec) 0.394 inches per second I1 n/sec)<br />
metres per second (m/sec) 3.281 feet per second (ft/sec)<br />
metres per second (m/sec) 1.943 nautical miles per hour (knot)<br />
cubic metres per second (m3/sec) 35.318 cubic feet per second (cfs)<br />
metres per hour (m/hr) 3.281 feet per second (ft/secf<br />
kilometres per hour (km/hr) 0.621 miles per hour (mphl<br />
degrees Celsius (oC) 9/jOC+?2 oegreesFahrenheZt (OF1
CONTENTS<br />
PREFACE ............................................................. i i i<br />
CO?4VERSX01! TABLE.. ..........................................*........ V<br />
FIGURES ............................................................. i x<br />
..............................................................<br />
TABl.ES xiii<br />
ACKNOWLEDC&lENTS ..................................................... x vi<br />
Cbdpter 1 . IN?KODUI:TION ............................................ 1<br />
1.1 Coastal Wctldnds ot the (iredt <strong>Lake</strong>s ..................... 1<br />
1.1 Compari sori <strong>of</strong> b'udstal and Inland Marshes ................ 4<br />
1.3 C~inctron drrll Valur <strong>of</strong> Coastal Wettnntlz .................. 5<br />
1.4 lake <strong>St</strong>. Cia1 r Wpt landr .................... . .. . ..... . 6<br />
Chaptcar Y . I'IIYSIL'AL l NVIRONMENT .................................... 11<br />
7.1 (;ctoIoqy ..................................... ............ 11<br />
2.2 Climat~ and Wpdthpr ..................................... 23<br />
2.3 Hydrology ............................................... 26<br />
2.4 Wdter Quality ........................................... 35<br />
Chapter 3 . BIUTIC ENVIRONMENT ...................................... 43<br />
3.1 Plankton ................................................ 43<br />
3.7 ketland V~yutatron ...................................... 47<br />
3. 3 lnvertrkrates ........................................... 54<br />
3.4 t-lsh ................................. ......*.*.*....... 58<br />
3.5 Aniphthlans and Reptiles ..................... . ....... . 66<br />
3. b Htrds ................................................... 68<br />
3 . 7 Marnrr~als .......................++.....*......,.... 73<br />
C'haytcr 4 . FC0101;ICAL PKI)CErjSI-7 .................................... 78<br />
4.1 Or igi n and f vuiution <strong>of</strong> L.akt. <strong>St</strong> . <strong>Clair</strong> Wet lands ......... 78<br />
4.2 B~olog~ cal Product ion ................... . ............. 85<br />
4.3 Cornrriuri~ ty Procpssc. s ..................................... Yl<br />
4.4 Wctlands as Habitat to Fish dnd W~ldFlfe ................ 101<br />
Chapter 5 . HUMAN iMPACIS ANO APPLIED ECOLOGY ....................... 104<br />
5.1 Wet1cfnrfl)rsturt)anc~ ..................................<br />
104<br />
5.2 Wrt 1 d nd &nrr.~hlp ar~l Managc~ni~nt ........................ 120<br />
5.3 Prospects for tho Future 126<br />
................................<br />
A U~rnens~ons <strong>of</strong> lake <strong>St</strong> . Cla~r and <strong>St</strong> . Llair<br />
1ilvt.r Wet idnd$ by tlnit~d <strong>St</strong>at~s and Canatfa<br />
Vuadrangle Maps ......................................... 141<br />
B Mean Monthly <strong>St</strong> . <strong>Clair</strong> Klvpr Flows 19OO to 1980 ......... 144<br />
C A1 yae Occurrt ng ln Coastal W~tlands dnu Ncarshore<br />
Wdters <strong>of</strong> Lzke S? <strong>Clair</strong> ................................ 146<br />
.<br />
U looplankton Occijrrt ng I n the Loastal Wfhtlands and<br />
Nc2arsnore Wdters <strong>of</strong> <strong>Lake</strong> <strong>St</strong> . Clai r ...................... 152<br />
vii
- Page<br />
Vascular Aquatic Plants Occurring in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
<strong>Wetlands</strong> ............................................ 157<br />
Benthic Macroi nvertebrates <strong>of</strong> Nearshore <strong>Lake</strong> <strong>St</strong>. Cl ai r<br />
and Connecting Waterways .............................. 164<br />
Taxonomic Listing <strong>of</strong> Freshwater Mol lusca <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
Clai r Coastal Marshes, Nearshore Waters, and Tributary<br />
Mouths ................................................ 169<br />
Habitat Preference <strong>of</strong> Freshwater Mollusca <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
Cl ai r Coastal Marshes and Nearshore Waters .............. 173<br />
Fish Known or Presumed to Utilize the Coastal <strong>Wetlands</strong><br />
<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ............... 175<br />
Amphibians and Reptiles Occurring in the Coastal<br />
<strong>Wetlands</strong> <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong> River ...... 178<br />
Birds Occurring in the Vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r and<br />
<strong>St</strong>. Clai r River Coastal <strong>Wetlands</strong> .................... . 180<br />
Mammals Occurring in the Coastal <strong>Wetlands</strong> <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong> Delta ....................... 185<br />
Ownership <strong>of</strong> Major <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Wetlands</strong> .............. 186<br />
viii
Pap <strong>of</strong> the Great <strong>Lake</strong>s drainage bastn .................... 2<br />
Map <strong>of</strong> <strong>St</strong>. Clalr Rl'vor-Lakc <strong>St</strong>, ClaJr-Detroit Rfver<br />
Ecosystem ............................................... 8<br />
Physical factors inf lutmcing tho occurrence <strong>of</strong> coastal<br />
wetlands .............................................. 11<br />
Dupositlanal fuaturcs and 1andfor.m~ <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
Oolta ................................................. 12<br />
Lfttlo Muscornoot Day, an interdlstributary bay botwoon<br />
Middle Channel and South Chdnnol <strong>of</strong> tho <strong>St</strong>. <strong>Clair</strong><br />
r?ulta .................................................. 13<br />
Woll-developed crevasse deposits adjacent to Mlddls<br />
Channel <strong>of</strong> <strong>St</strong>. Clajr Dolta ............................... 14<br />
Deltaic landforms <strong>of</strong> I)lckfnson Island, Mfchfgan .......... 15<br />
Gaoloylc cross-section <strong>of</strong> the northern portfon <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> showlng depth to bedrock. and thjcknoss <strong>of</strong><br />
glacl al, lacustrine, and deltaic deposlts 2 7<br />
................<br />
Cross-sectlon <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta showing distrfbutlon<br />
<strong>of</strong> deltaic sedlmsnts ..................................... 18<br />
Chronology <strong>of</strong> lake levsls and lake stages 3n the<br />
t-iufon and Erie baslns .................................. 1 H<br />
................<br />
Vagetatfan transoct on Uickinsen Island, <strong>St</strong>, <strong>Clair</strong><br />
Oelta, showing character <strong>of</strong> wetland soils %2<br />
Monthly wind frequency dlagrams for Wlndsor, Ontario ..... 26<br />
Monthly wfnd froquoncy for Mt. Clomens, Mlchigan ......... 26<br />
Current patterns for <strong>Lake</strong> <strong>St</strong>. Clafr undor varjous<br />
wlnd condftlons .......................................... 28<br />
iake <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Clafr Rlver drainage basfn ............ ~ ( 7<br />
Mean monthly discharyc. <strong>of</strong> the <strong>St</strong>. Claf r River<br />
:~s-:%a ................................................ 31)<br />
<strong>St</strong>. <strong>Clair</strong> Dolta distributary channels .................... 3 1
Page<br />
Hydrograph <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air and <strong>Lake</strong> Erie water levels<br />
showing the impact <strong>of</strong> a record ice jam in the <strong>St</strong>. <strong>Clair</strong><br />
Delta in March and April 1984 ..........................,e 34<br />
Distribution <strong>of</strong> specific conductance in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.. . . 36<br />
Mean water quality characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ., , . , 38<br />
Characteristics <strong>of</strong> <strong>Lake</strong> Huron water mass and Ontario<br />
nearshore water mass ..................................... 42<br />
Temperature and dissolved oxygen relationship in a <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> coastal wetland ............................... 42<br />
Concentration gradient <strong>of</strong> algal pigments across <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> .............................................. 43<br />
Mean distribution and concentrations <strong>of</strong> zooplankton<br />
groups in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ................................. 45<br />
Distribution <strong>of</strong> total rotifers and most abundant genus,<br />
j3rachlonur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ............................ 46<br />
Ontario shore1 ine <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River at Cathcart Park . . . . 47<br />
Vegetation transects on Harsens Island* <strong>St</strong>. <strong>Clair</strong> Delta . . 48<br />
Distribution <strong>of</strong> six aquatic macrophytes in the <strong>St</strong>. <strong>Clair</strong><br />
River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-Detroit River ecosystem ............. 49<br />
Emergent stand <strong>of</strong> reed grass (Phraamites australis)<br />
along Chematogan Channel, Walpole Is1 and, Ontario . . . . . . 50<br />
Emergent stands <strong>of</strong> narrow-leaved cattail (m<br />
~ t i f oia l on Little Muscamoot Bay* Harsens Island,<br />
Michigan ................................................. 51<br />
Wetland plant communities <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta .... . . , . . 53<br />
Wetland vegetation on Dickinson Island in 1964 ........... 56<br />
Wetland vegetation on Dickinson Island in 1975 .........., 57<br />
Walleye spawning and nursery areas in the Detroit River<br />
and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ....................................... 61<br />
Yellow perch spawning and nursery areas in the Detroit<br />
River and <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> ................................. 62<br />
<strong>Lake</strong> sturgeon spawning and nursery areas in the Detroit<br />
River, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, and <strong>St</strong>. <strong>Clair</strong> River ... . . . . .. . . . . . . 63<br />
Rock bass, pumpkfnseed/bluegill, largemouth bass, and<br />
smallmouth bass spawning and nursery areas in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> ................................................ 6 4
Number<br />
.........................<br />
3 8 Northern piker muskellunge, channel catfish, and carp<br />
spawning areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> 65<br />
Sport fishing areas by specjes in the vicinity <strong>of</strong> <strong>St</strong>.<br />
<strong>Clair</strong> Delta wetlands ....................L.,............ 66<br />
Least bittern (Jkobrvchu~ exllis) with young in broad-<br />
leaved cattail (Tveha latifoliq) nest .................... 73<br />
Food chain <strong>of</strong> the great blue heron (brdea herodlas)<br />
in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................................ 74<br />
Muskrat ffhousesw i n a well populated section <strong>of</strong> Magee<br />
Marsh, western <strong>Lake</strong> Erie, Ohio ........................... 75<br />
Raccoon (procvon lotar), a common predator <strong>of</strong> duck<br />
nests in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetlands ......................... 77<br />
Twelve wet1 and habitats recognized in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ..... 81<br />
Cycling <strong>of</strong> energy and material through the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
coastal wetland ecosystem ................................ 87<br />
Energy pyramid in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal marshes ..,.,.,., 88<br />
Seasonal growth <strong>of</strong> dominant submersed macrophytes in the<br />
<strong>St</strong>. Clai r River-<strong>Lake</strong> <strong>St</strong>. Cl air-Detroit River ecosystem ... 92<br />
Successional changes on Dick i nson Is1 and in response t o<br />
water level fluctuations ................................. 94<br />
Wet1 and pl ant community displacement model for Dickinson<br />
Island ................................................... 94<br />
Fall migration corridors for dabbling ducks (Tribe<br />
Anatini) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................... 98<br />
..........................<br />
Fall migration corridors for diving ducks (Tribe<br />
Aythyini) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> 99<br />
Fall migration corridors for Canada geese (w<br />
ranadensis) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................ 100<br />
Historic trends in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetland disturbance .... 105<br />
Aerial photograph <strong>of</strong> the mouth <strong>of</strong> the Clinton River*<br />
Macomb County, Michigan .................................. 107<br />
55 Aerial photograph <strong>of</strong> <strong>St</strong>. John's Marsh on Bouvier Eay, and<br />
North Channel, <strong>St</strong>. Cl a1 r Delta (April 1949) .............. 108<br />
56 Aerial photograph <strong>of</strong> North Channel, <strong>St</strong>. <strong>Clair</strong> Delta* and<br />
<strong>St</strong>. John's Marsh on Bouvier Bay (May 1980) ............... 109<br />
57 Commercial development in Marsens Island wetland ......... 11 3
Mercury concentration in sediment core from western <strong>Lake</strong><br />
Erie near mouth <strong>of</strong> Detro'it River ......................... 114<br />
Wetland losses along the Ontario shore1 ine <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> ................................................ 117<br />
Comparfson <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air coastal wetlands dfstrib-<br />
ution in 1873 and 1968 ................................... 118<br />
Location map <strong>of</strong> individual coastal wetlands <strong>of</strong> the <strong>St</strong>.<br />
Cl ai r Rfver-<strong>Lake</strong> <strong>St</strong>. Cl ai r ecosystem ..................... 120<br />
Ownership map <strong>of</strong> the Michigan portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
Delta ............................................... 121<br />
Locatlon map <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> diked coastal wetlands .... 122<br />
Aerial photograph <strong>of</strong> Dickinson Island wetlands, <strong>St</strong>.<br />
Clafr Delta (May 1980) ................................... 124<br />
xii
Number<br />
TABLES<br />
1 Comparison <strong>of</strong> emergent coastal wetlands for the<br />
Laurentian Great <strong>Lake</strong>s ................................... 3<br />
Economic value <strong>of</strong> Michigan's coastal wetlands, 1980 ...... 6<br />
Wild1 i fe use and other functions <strong>of</strong> coastal wet1 ands<br />
at low and high water levels ............................. 7<br />
Estimated dimensions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and <strong>St</strong>. Clafr<br />
River wet1 ands . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8<br />
Interpretation <strong>of</strong> the events related to the origin <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> River Delta . . . . . .. . . . . ., .. . . . . .. . . , ,. . .. .. . 19<br />
Sediment discharge rates and characteristics in the<br />
<strong>St</strong>. <strong>Clair</strong> River .......................................... 21<br />
Mean monthly temperatures for three selected stations .... 24<br />
Mean monthly precipitation for three selected stations .. , 25<br />
<strong>St</strong>. <strong>Clair</strong> River flow time and velocity from <strong>Lake</strong> Huron to<br />
<strong>Lake</strong> <strong>St</strong>. Cl air . . . . . . . . . , . . . . . . . . . . . . . . . . . . . . , . . . . . . . . . . . . 30<br />
Record storm surge levels <strong>of</strong> Canadlan stations on <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and connecting channels . . . . . . . . . . . . . . . . . . . . . . . . 32<br />
Average monthly water transfer factor parameters . . . . . . . . . 35<br />
Water level data for <strong>Lake</strong> <strong>St</strong>. Clai r . . . . . . . . . . . . . . . . . . . . . . 35<br />
<strong>Lake</strong> <strong>St</strong>. Cl air water qua1 ity measurements dur'ing the 1973<br />
growing season ........................................... 37<br />
Chemical and physical characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
and adjacent waters ...................................... 39<br />
Geochemlcal composition <strong>of</strong> sediments In Big Point Marsh,<br />
Ontario on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> , . . . . . . . . . . . . . . . . . . , . . . . . . . . 40<br />
Chemical composition <strong>of</strong> water in Big Point Marsh, Ontario<br />
on <strong>Lake</strong> <strong>St</strong>. Clafr .,.......,......... e L O a I I a I e e * r * + e e a * L e L 40<br />
Comparison <strong>of</strong> chemical characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, C1 af r<br />
and associated waters ......................ee.ss.L).ee*il*e 41<br />
xiii
3 1<br />
3 2<br />
Macrozoobenthos <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r and the lower <strong>St</strong>. <strong>Clair</strong><br />
River and Delta ..........................................<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> unionid bivalves and their glochidial<br />
host fish ..... ....*.............................."e**...e<br />
...............<br />
...............<br />
Average annual sport fishing harvest for Michigan water<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and connecting waterways<br />
Birds observed in <strong>St</strong>. <strong>Clair</strong> Delta wetlands<br />
Productivity and seasonal abundance <strong>of</strong> waterfowl in<br />
Bouvier Bay and Harsens Island wetlands ..................<br />
Species composltfon <strong>of</strong> ducks harvested in the <strong>St</strong>. <strong>Clair</strong><br />
Flats <strong>St</strong>ate Game Area (1974-75) ..........................<br />
Muskrat harvest in Michigan and Ontario bordering <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and connecting waterways .......................<br />
Factors influencing lake and marine delta morphology ..,..<br />
Morphology and re1 ative physical relationships <strong>of</strong> the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands ..................................<br />
Concentrations <strong>of</strong> nutrients and metals in wetland plants<br />
at the stage <strong>of</strong> maximum development, Big Creek Marsh*<br />
Long Point, Ontario .....................................<br />
Biomass and productivity <strong>of</strong> emergent wetl and pl ants<br />
<strong>of</strong> the Great <strong>Lake</strong>s region ................................<br />
Estimates <strong>of</strong> standing crop biomass for submergent<br />
vegetatfon in Anchor Bay, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> .................<br />
Seasonal estimates <strong>of</strong> standing crop biomass for<br />
submersed macrophytes at several locations in the <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> system .........................................<br />
Domlnant submersed aquatic macrophyte taxa <strong>of</strong> the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> system ................................... 92<br />
Vegetation changes on Dickinson Island in response to<br />
lake level fluctuatfons .................................. 95<br />
33 <strong>St</strong>atus <strong>of</strong> wildlife in the vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
and connecting waterways, 1970 ........................... 102<br />
3 4<br />
Land use change at <strong>St</strong>. Johnrs Marsh ...................... 106<br />
3 5 Adverse impacts to the various morphology types <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands ................................. 111<br />
36 Historical dredg'ing totals <strong>of</strong> Corps <strong>of</strong> Engineers<br />
projects in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, 1920-72 ...............a=.*... 112<br />
xiv
37 Metals concentrations in 1 Jvfng plant tissues from<br />
Big Point Marsh. Ontario. on <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> .............. 115<br />
38 Michigan and Ontario wetland losses on <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> .... 116<br />
39 Comparison <strong>of</strong> 1873 and 1968 <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> wetland<br />
areas .................................................... 119
In preparing a comprehensive report<br />
such as this pr<strong>of</strong>ile, it is not possible<br />
to mention a11 the individuals who pro-<br />
vided information, gufdance, and stimulus.<br />
Much <strong>of</strong> the credit goes to colleagues at<br />
<strong>The</strong> Ohio <strong>St</strong>ate University and Eastern<br />
Michigan University, including Ronald<br />
<strong>St</strong>uckey, Milton Trautman, Cl arence Taft,<br />
David Johnson, and Loren Putnam. <strong>St</strong>udents<br />
and staff <strong>of</strong> the F. T. <strong>St</strong>one Laboratory at<br />
Put-in-Bay, and the Ohfo Sea Grant Program<br />
who helped In researching various aspects<br />
<strong>of</strong> the marsh scology, are Alan Riemer,<br />
Andrea W i l son, Gary Kfrkpatrick, Mark<br />
Wageman, Laura Fay, and Suzanne Hartley.<br />
Tim Donelly <strong>of</strong> Eastern Michigan University<br />
assisted with data collection.<br />
ACKNOWLEDGMENTS<br />
Ted Ladewski <strong>of</strong> the University <strong>of</strong><br />
Michfgan, <strong>St</strong>an Bolsenga <strong>of</strong> the Great <strong>Lake</strong>s<br />
Environmental Research Laboratory (NOAA) ,<br />
Karl Bednarik <strong>of</strong> the Ohio Division <strong>of</strong><br />
xvi<br />
Wildlife, Susan Crispin <strong>of</strong> the Michigan<br />
Natural Features Inventory, Brooks<br />
Williamson <strong>of</strong> the U.S. Army Corps <strong>of</strong><br />
Engineers, Gary McCullough <strong>of</strong> the Canadian<br />
Wild1 ife Service, Joseph Leach <strong>of</strong> the<br />
Ontario Ministry <strong>of</strong> Natural Resources, and<br />
Bernard Griswol d, Thomas Edsall , Bruce<br />
Panny, and Don Schloesser <strong>of</strong> the Great<br />
<strong>Lake</strong>s Fishery Laboratory, U.S. Fish and<br />
Wildlife Service, all provided useful<br />
informat ion and encouragement.<br />
Wiley Kitchens and Walt Duffy <strong>of</strong> the<br />
National Wet1 ands Research Center provided<br />
guidance and advice throughout the<br />
project. Funding for this project was<br />
provided by the Office <strong>of</strong> the Chief and<br />
the DetroTt District, U.S. Army Corps <strong>of</strong><br />
Engineers. And we especially want to<br />
thank Sandra Herdendorf for typing and<br />
editlng the manuscript. To all those who<br />
he1 ped, we are grateful.
CHAPTER 1. INTCRODeQCTUOM<br />
1.1 COASTAL WETLANDS OF THE GREAT wetland communityr a portion <strong>of</strong> the Great<br />
LAKES <strong>Lake</strong>s - the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Clai r River<br />
system.<br />
In recent years there has been an<br />
increasing awareness <strong>of</strong> the resource value<br />
<strong>of</strong> our coastal wetlands and the urgent<br />
need to protect and conserve these<br />
ecosystems. <strong>The</strong> wetlands <strong>of</strong> the Lauren-<br />
ti an Great <strong>Lake</strong>s have been greatly altered<br />
by natural processes and cultural prac-<br />
tfces, <strong>The</strong> impacts to coastal wetlands in<br />
the Great <strong>Lake</strong>s region has become a<br />
subject <strong>of</strong> particular concern for the<br />
emerging coastal management programs in<br />
the eight states and the one Canadian<br />
province bordering the lakes (Figure 1).<br />
Traditionally, wetl and conservation<br />
efforts along the Great <strong>Lake</strong>s have been<br />
aimed at protecting waterfowl breeding, or<br />
to a lesser degree, fish spawning and<br />
nursery habitat. More recent efforts<br />
toward preservation are based on the<br />
know1 edge that wetl ands provide additional<br />
benefits? including flood control, shore<br />
erosion protection, water management*<br />
control <strong>of</strong> nutrient cycles, accumulation<br />
<strong>of</strong> sediment, and supply <strong>of</strong> detritus for<br />
the aquatic food web. Although the<br />
intrinsic value <strong>of</strong> Great <strong>Lake</strong>s wetland<br />
areas are being more fully recognized, no<br />
comprehensive studies have been undertaken<br />
to characterize the ecological relationships<br />
within them. <strong>The</strong> U.S. Fish and<br />
Wildlife Service (Herdendorf et al,<br />
1981ar bsc) inventoried the existing<br />
l fterature <strong>of</strong> physical, biological, and<br />
cultural aspects <strong>of</strong> the coastal wetl ands<br />
associated with each <strong>of</strong> the Great <strong>Lake</strong>s.<br />
<strong>The</strong>ir study pointed out many gaps in our<br />
knowledge <strong>of</strong> the resources found In Great<br />
<strong>Lake</strong>s wetl ands, partlcul arl y si te-spec1 f ic<br />
fnformation on marshes, This report is<br />
f ntended as a contrfbution toward ff 11 ing<br />
these voids by presenting a pr<strong>of</strong>ile <strong>of</strong> the<br />
For the purposes <strong>of</strong> this report,<br />
wetlands are defined as areas which are<br />
peri odi call y or permanent1 y inundated w 1 th<br />
water and which are typically character-<br />
ized by vegetation that requires saturated<br />
soil for growth and reproduction. This<br />
definition i n cl udes areas that are<br />
commonly referred to as bogs, fens,<br />
marshes, sloughsr swamps, and wet meadows.<br />
<strong>The</strong> coastal wetlands <strong>of</strong> the Great <strong>Lake</strong>s<br />
are further defined as all wetlands<br />
1 ocated within 1 km <strong>of</strong> the lake shore or8<br />
if farther from the shore, those dl rectl y<br />
influenced by water level change <strong>of</strong> the<br />
lakes or their connecting waterways.<br />
Great <strong>Lake</strong>s coastal wetl ands are<br />
highly productive, diverse communities<br />
which interface between terrestrial and<br />
aquatic envf ronments. <strong>The</strong> most obvious<br />
and unique feature <strong>of</strong> these wetlands is<br />
thelr characteristic vegetationr whlch<br />
provides a diverse community structure<br />
<strong>of</strong>fering cover and food for the animal<br />
components <strong>of</strong> the system. Because <strong>of</strong> the<br />
ability <strong>of</strong> this vegetation to siow the<br />
flow rate <strong>of</strong> water passing through?<br />
wetl ands are valuable for erosion control,<br />
trapping sediments before they reach the<br />
open lake# and attenuating the force <strong>of</strong><br />
waves to lessen the4r destructive power,<br />
<strong>The</strong> same vegetation provides a natural<br />
pol 1 ution abatement mechanism by serving<br />
as a filter for coastal trlbutaries<br />
through the reduction <strong>of</strong> the quantity <strong>of</strong><br />
nutrients and toxic pol 1 utants belng<br />
washed into the Great <strong>Lake</strong>s.<br />
Coastal wet1 ands are high1 y valued as<br />
recreational sites for activitfes such as<br />
hunting, trappfngr fishing, boatfng access<br />
to larger bodies <strong>of</strong> water, bdrdwatching?
and general aesthetic enjoyment. <strong>The</strong><br />
combination <strong>of</strong> recreatlonal desi rab ll ity,<br />
agricultural and residential potential,<br />
and the proximity <strong>of</strong> coastal wetlands to<br />
larger bodfes <strong>of</strong> water have contributed to<br />
thei r status as endangered environments.<br />
<strong>The</strong>ir unique properties are susceptible to<br />
numerous natural and human-caused factors<br />
that are now causing coastal wetlands to<br />
disappear at an alarming rate.<br />
<strong>The</strong> Laurentian Great <strong>Lake</strong>s system<br />
within the United <strong>St</strong>ates (Figure 1)<br />
extends from Duluth, Minnesota* at the<br />
western end <strong>of</strong> <strong>Lake</strong> Superior, to Massena,<br />
New York on the <strong>St</strong>. Lawrence River. It<br />
possesses a shoreline length <strong>of</strong> over 6,000<br />
km and a water surface area <strong>of</strong> 158,000<br />
km2. Herdendorf et a1 . (1981a) enumerated<br />
a total <strong>of</strong> lr370 coastal wetlands for the<br />
Great <strong>Lake</strong>s and thei r connect1 ng channel Sr<br />
for a combined wetland area <strong>of</strong> lr209 k d<br />
(Table 1). <strong>The</strong> greatest number and area<br />
<strong>of</strong> coastal wetlands ring <strong>Lake</strong> Michigan,<br />
the only Great <strong>Lake</strong> entirely wfthin the<br />
United <strong>St</strong>ates. <strong>Lake</strong> Super+or has the<br />
second highest number <strong>of</strong> wetlands, but<br />
they are relatively small in size. On the<br />
average* the 1 argest wet1 ands are found<br />
along <strong>Lake</strong> Huron and its discharge channel<br />
(the <strong>St</strong>. Clai r River); for example, the<br />
emergent wetlands <strong>of</strong> the <strong>St</strong>. Clalr Delta<br />
alone cover 36 km2. <strong>The</strong> highly<br />
industrialized <strong>Lake</strong> Erie shore has the<br />
smallest number and area <strong>of</strong> wetlands while<br />
<strong>Lake</strong> Ontario has the smallest average size<br />
<strong>of</strong> wetlands, 1 argely due to is01 ated<br />
marshes In the Thousand Islands area <strong>of</strong><br />
the <strong>St</strong>. Lawrence River. <strong>The</strong> presence or<br />
absence <strong>of</strong> coastal wetl ands is 1 argely<br />
Table 1. Cogparison <strong>of</strong> emergent coastal wetlands for the U.S. Laurentian<br />
Great <strong>Lake</strong>s .<br />
<strong>Lake</strong><br />
Total Mean Percent<br />
Shore Number <strong>of</strong> area <strong>of</strong> area <strong>of</strong> <strong>of</strong> total<br />
1 ength wet1 ands wet1 ands wet1 ands area<br />
<strong>Lake</strong> Superior and 1598 348 267 0.77 22.1<br />
<strong>St</strong>. Marys River<br />
<strong>Lake</strong> Michtgan and 2179 417 490 1.18 40.5<br />
<strong>St</strong>r. <strong>of</strong> Mackinac<br />
<strong>Lake</strong> Huron 832 177 249 1.41 20.6<br />
<strong>St</strong>. <strong>Clair</strong> River, 256 20 36 1.80 3 .O<br />
<strong>Lake</strong> <strong>St</strong>. Cl airr<br />
and Detroit River<br />
<strong>Lake</strong> Erie and 666 96 83 0.86 6.9<br />
Niagara River<br />
<strong>Lake</strong> Ontario and 598 3 12 84 0-27 6.9<br />
<strong>St</strong>. Lawrence<br />
River<br />
TOTAL/MEAN 6129 1370 1209 0.88 100.0<br />
a~ata source: Herdendorf et a?. (1981a1,<br />
3
dictated by the geomorphology <strong>of</strong> a given<br />
shoreline and the recent history <strong>of</strong> water<br />
level fluctuations. Each lake has a<br />
particular set <strong>of</strong> geomorphic features<br />
which exert control on the development <strong>of</strong><br />
coastal wet1 ands.<br />
1.2 COA4PARISON OF COASTAL AND<br />
INLAND MARSHES<br />
<strong>The</strong> coastal wetlands <strong>of</strong> the Great<br />
<strong>Lake</strong>s differ in several ways from inland<br />
wetl ands. <strong>The</strong> coastal wetl cnds are<br />
subject to temporary short-term water<br />
level changes. <strong>Lake</strong> <strong>St</strong>. Claf r wetlands<br />
are subject to inundation and dewaterfng<br />
by seiches, Long-term cycl ic water level<br />
changesr related to water budgets <strong>of</strong> the<br />
lake basins, also affect the coastal<br />
wetlands. Such fluctuationsr occurring<br />
over a period <strong>of</strong> approximately seven to 10<br />
years, may cause vegetation dieback5<br />
erosion <strong>of</strong> the wetlands* or lateral<br />
dtsp? acements <strong>of</strong> the vegetative zones <strong>of</strong><br />
wetlands. Many coastal wetlands, such as<br />
those along western <strong>Lake</strong> Erie, are exposed<br />
to relatively high wave energy.<br />
Coastal wetl ands a1 ong the Great<br />
<strong>Lake</strong>s do hot appear to exhibit the aging<br />
process associated with ini and freshwater<br />
wetlands. Because <strong>of</strong> the fluctuating<br />
water. levels <strong>of</strong> the Great <strong>Lake</strong>s, constant<br />
rejuvenation <strong>of</strong> wet1 and communi t5es<br />
occurs. As a consequencer diagrams 5n<br />
textbooks illustrating the gradual<br />
senescence <strong>of</strong> freshwater wetlands are more<br />
applicable to inland wetlands <strong>of</strong> the<br />
glaclated Midwest than to the Great <strong>Lake</strong>s<br />
coastal wetlands. Many inland freshwater<br />
wetlands undergo senescence and converslon<br />
to terrestrial environments as a result <strong>of</strong><br />
dnorganic and organic depositlon (e.g.,<br />
formation <strong>of</strong> peat deposits).<br />
Coastal wetl ands <strong>of</strong>ten display a<br />
diversity <strong>of</strong> 1 andforms not normal 1 y<br />
encountered in other wetland environments.<br />
Owing to changes in the water levels <strong>of</strong><br />
the Great <strong>Lake</strong>s since the retreat <strong>of</strong> the<br />
Pl e i skocene ice sheetsr 1 and forms such as<br />
coastal barrfersr deltas, and natural<br />
levees have been deposited. <strong>The</strong>se<br />
geamaspholagf ca? features have f n f 1 uenced<br />
the formatf on <strong>of</strong> coastal wetlands. <strong>The</strong><br />
continuing fluctuatiloc <strong>of</strong> water levels f n<br />
the Great <strong>Lake</strong>s 1s also an important<br />
varfable In determining many <strong>of</strong> the<br />
distingulshfng characteristics and the<br />
diversity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />
wetlandsr as well as those <strong>of</strong> the<br />
1 andforms the wetl ands occupy.<br />
Coastal wetlands exhi b f t topograph-<br />
ical pr<strong>of</strong>lles which are more gentle than<br />
what is generally observed in inland<br />
wet1 ands. Many l nl and wet1 ands are formed<br />
in confined glacial features such as<br />
kettle lakesp which exhibit abrupt slopes<br />
on their perimeters. Because <strong>of</strong> the low<br />
angle slope conditionss vegetation zona-<br />
tion Is more dfstlnct and better displayed<br />
In coastal wetlands. As described in this<br />
report, the <strong>St</strong>. Clafr Delta is colonized<br />
by several broad submersed and emersed<br />
wet1 and zones. In1 and wetl ands* howeverr<br />
are more conftned and do not display the<br />
extensive zonation observed in the coastal<br />
zone.<br />
Water levels in coastal wetlands are<br />
determined by climatic factors over<br />
several years. Generallyr annual higher<br />
water levels in the Great <strong>Lake</strong>s occur in<br />
late summer and lower levels occur in<br />
January. Water level changes in inland<br />
wetlands respond to immediate or short<br />
term run<strong>of</strong>f condftions. High water most<br />
<strong>of</strong>ten, occurs in the spring when run<strong>of</strong>f 1s<br />
highest; low water levels occur in early<br />
fall.<br />
Furthermore* the contribution <strong>of</strong><br />
groundwater flow is more significant to<br />
Inland wetlands, During the dry summer<br />
months with high evapo-transpiration<br />
rates, the survival <strong>of</strong> such wetlands may<br />
depend on groundwater i nf 1 ow. Great<br />
<strong>Lake</strong>s' controlled wetlands, however, shift<br />
1 andward or lakeward depending upon water<br />
level conditions over several years.<br />
Clearly, the relationship between<br />
groundwater and wetlands is intimate in<br />
inland areas, but this relationship in the<br />
coastal zone is less significant.<br />
In the Great <strong>Lake</strong>s, with few<br />
exceptions, the substrate is composed <strong>of</strong><br />
mineral sediments. Because <strong>of</strong> the<br />
f 1 ushing action <strong>of</strong> seiches and periodic<br />
water level changes, organic deposits are<br />
not common. Conversely, the hydrolsgic<br />
connect1 vfty <strong>of</strong> in7 and wetl ands is poorer.<br />
Fine mineral soils, with high organic<br />
content f n the inland marshes, contri bute
to thefr gradual senescence and account one may refer to the dollar value <strong>of</strong> a<br />
for more ac-ld-tolerant vegetation than waterfowl hunting dayl as well as to<br />
that found 5n the coastal wetl ands. Peat concepts such as margfnal lty (or scarcfty<br />
fs not typicalfy found along the coast <strong>of</strong> value) and opportunf t y costs.<br />
<strong>Lake</strong> <strong>St</strong>. Cfafr. Nevertheless, not a1 3 wetland functions<br />
(e.g,, flood water storage) are vendable<br />
In the marketplace.<br />
1.3 FU NCTlON AND VALUE OF COASTAL<br />
WETLANDS<br />
Coastal wetlands fn the Great <strong>Lake</strong>s<br />
are mu1 t f -f unct ional because these<br />
environments are part <strong>of</strong> both the up1 ands<br />
and the open-water ecosystems, It is the<br />
fnterface with the Great <strong>Lake</strong>s that<br />
mu1 t ip1 ies the wetland functtons and<br />
contr-ibutes to the "open system" dynamics.<br />
<strong>St</strong>reams and coastal waterways enhance the<br />
ecosystem connectlvf ty, whil a obstacles<br />
such as earthen berms and dikes result in<br />
coastal wetl and f ragmantation and 1 oss <strong>of</strong><br />
function. Functional loss then, can<br />
result from both bank-derlved and lake-<br />
derfved forces. An awareness <strong>of</strong> the<br />
ef f s ct <strong>of</strong> 1 ong-term lake level changes on<br />
the functlon <strong>of</strong> the coastal wetlands is<br />
begfnning to emerge, and the concept <strong>of</strong><br />
pulse stablllty fs being ascribed ta these<br />
envf ronments,<br />
Durlng the past decade, considerable<br />
research was carried out regardlng the<br />
functfon and value <strong>of</strong> wetl ands. Important<br />
general sources Include Tfner (19841,<br />
Greeson et al, (19791, and Messman et al.<br />
(19771, In reference to the Great <strong>Lake</strong>s,<br />
slgnl f icant contr7butlons were made by<br />
Herdendorf et a1 . ( 1981al b,c), Raphael and<br />
Jaworski (19791, Jaworskf t 1981) r Jaworskf<br />
and Raphael (19781, and f f 1 ton et al,<br />
(1978). According to the U, S. Fish and<br />
WfldJ f fe Service Classification (Cowardin<br />
et a1. 1979), most Great <strong>Lake</strong>s wetlands<br />
would be classified as lacustrine or<br />
palustrine,<br />
Wet1 and functions are those processes<br />
occurrf ng fn wetlands whfch are associated<br />
with the functioning <strong>of</strong> the ecosystem or<br />
with the hydrosystem. Examples <strong>of</strong> such<br />
functions are primary production and water<br />
storage, In contrast, wetland values are<br />
those wetland products or servfcss whfch<br />
satfsfy a human need. Comonly, economic<br />
values are employed to such wetland<br />
commodities and services. As a result,<br />
In response to Sectjon 404 and other<br />
permitting authority, the U.S. Army Corps<br />
<strong>of</strong> Engineers (Reppert and Sigleo 1979)<br />
developed the following list <strong>of</strong> wetland<br />
functions and ssrvlces:<br />
1. Natural biological functions<br />
a. net primary productivity<br />
b. food chain (web) support<br />
2. Habitat for aquatic and wetland<br />
speci es<br />
3. Aquatic study areas, sanctuaries<br />
and refuges<br />
4. Hydrologic support functlons;<br />
a. shoreljne protection from wave<br />
attack<br />
b. storage <strong>of</strong> storm and flood waters<br />
c. water purlflcatlon through<br />
natural filtration, sediment<br />
trapping* and nutrient cycling<br />
uptake<br />
d. groundwater recharge<br />
5. Cultural or auxll fary values<br />
fncluding consumptive and noncon-<br />
sumptfve recreation as well as<br />
aesthetic value.<br />
Except for item 5, these wetland benef fts<br />
tend to accrue to the public, and are not<br />
general 1 y vendable by a private 1 andowner.<br />
Quantlta;fve economlc value data for<br />
Great <strong>Lake</strong>s wetlands are avaflable, A<br />
study by Jaworski and Raphael t19781<br />
basicall y addressed the cultural values<br />
using a gross annual economic return<br />
method01 ogy, whereas TIlton et al, (1978)<br />
focused on ecosystem replacement values fn<br />
regard to f fsh productton, waterfowl<br />
feed fng/breedIng, nutrfent removal and<br />
control# and water supply. By combinf ng<br />
the results af these two studfes and<br />
updatfng the data to 1980# a mare complete<br />
account <strong>of</strong> the economfc value <strong>of</strong><br />
Mfchf gan's coastal wetl ands can be<br />
presented (Table 2).
Although methodologies are mlxed,<br />
Table 2 f s useful in establ ishing economic<br />
values within an order <strong>of</strong> magnitude and in<br />
priority. Notf ce that nutrient control J<br />
sport fishing, and fish production are<br />
clearly <strong>of</strong> more value than the traditfonal<br />
uses, i .e. , waterfowl huntingp trappfng*<br />
water supply, and commercial fishing,<br />
Conversely, these data are presented as<br />
average economic values, even though not<br />
all wetlands are a1 ike. Moreover, the<br />
methodologies did not consider surpf us<br />
value* as would be assessed by the<br />
wfllingness to pay technique.<br />
Table 2. Economic vaJue <strong>of</strong> Michigan's<br />
coastal wetlands, 1980.<br />
Economic sector<br />
Run<strong>of</strong>f Nutrient Control<br />
Sport Fishing<br />
Fish Production<br />
Waterfowl Breedbng/Feedi ng<br />
Nonconsumptl ve Recreation<br />
Waterfowl Hunting<br />
Trapping <strong>of</strong> Furbearers<br />
Water Supply<br />
Commercial FishIng<br />
a~ata source: Jaworski ( 1981).<br />
-<br />
(per (per<br />
hectare) acre)<br />
Wetland functions and values vary<br />
from wetl and t o wetland (f .e., spatially)<br />
as well as from year to year (i.e.,<br />
temporally), particularly as a result <strong>of</strong><br />
precipitation and hydrologic changes.<br />
Jaworskf et ale (19791, studied the effect<br />
<strong>of</strong> water level fluctuatfons on seven<br />
wetland complexes in the Great <strong>Lake</strong>s<br />
reg3 on, including Dickinson Island <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> Delta. As fllustrated in Table<br />
3, wild1 ife use and wetland functjons<br />
change as water levels fluctuate, As<br />
discussed in Section 2.3, law water levels<br />
occurred in 1964-65 whereas record high<br />
levels took place during the 1972 to 1974<br />
$nterlrn.<br />
very dense and the water table lies<br />
beneath the mean level <strong>of</strong> the marsh<br />
surface, It Is at such low water<br />
conditf ons that waterfowl management calls<br />
for diking and flooding <strong>of</strong> these emergent<br />
communities. Converselyr during hfgh<br />
water periods the hydrologic clrculat~on<br />
Is facilitated and lake water can enter<br />
the wetland via inlets, canals, and<br />
breaches in barrier beaches. Excessive<br />
high water is associated wlth plant<br />
community retrogression as we1 1 as erosion<br />
<strong>of</strong> the nearshore vegetation and beaches.<br />
<strong>The</strong>refore, the function and value <strong>of</strong> a<br />
coastal wetland is most dynamic, and<br />
short-term studf es <strong>of</strong> wetl and values<br />
should receive careful interpretation.<br />
In summary, a multi-functioning<br />
wetland tends to have a higher value than<br />
those with fewer functions, Converse1 y,<br />
coastal wetlands which are isolated by<br />
barriers or degraded by factors such as<br />
land drainage* exhibit fewer functions and<br />
have 1 ower- values. Moreover, wet1 ands<br />
vary in value from place to place and<br />
temporal 1 y as we1 1 r especi a1 1 y in response<br />
to lake level changes. In addition,<br />
continual wetl and acreage loss results in<br />
a marginality value <strong>of</strong> those extant<br />
wetlands. <strong>The</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />
have exceptional value due to their<br />
ecosystem relationship wfth the lake and<br />
connect1 ng channel s. <strong>The</strong> decline <strong>of</strong><br />
western <strong>Lake</strong> Erte marshes has a1 so<br />
enhanced the re1 ative value <strong>of</strong> the <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> wetlands.<br />
In recognitfon <strong>of</strong> the specfa1<br />
environmental val ue <strong>of</strong> wetl ands, Federal<br />
and <strong>St</strong>ate leglslatlon has recently been<br />
enacted to pratect these areas. In<br />
concert with the National Wetland<br />
Inventory programr each <strong>St</strong>ate in the<br />
Unlted <strong>St</strong>ates has begun the process <strong>of</strong><br />
inventorying and mapping thei r wetl and<br />
resources. Wet1 ands whose hydro1 ogic<br />
regime has been altered or destroyed may<br />
be mitigated.<br />
1.4 LAKE ST. CLAlR WETLANDS<br />
Durlng low water levels* the emergent <strong>The</strong> outlet <strong>of</strong> <strong>Lake</strong> Huron is through<br />
communftles <strong>of</strong> cattails and sedges are the <strong>St</strong>. <strong>Clair</strong> River, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, and
the Detroit River to <strong>Lake</strong> Erie (Figure 21,<br />
a distance <strong>of</strong> 139 km with a fall <strong>of</strong> 2.4 m.<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is an expansive* shallow<br />
basin having a distinctive heart shape.<br />
Although not one <strong>of</strong> the world's largest<br />
lakes, f t does rank 122 on the basis <strong>of</strong><br />
surface area (Herdendorf 1982). <strong>The</strong> lake<br />
has a surface area <strong>of</strong> 1,110 km2 and a<br />
drainage basin <strong>of</strong> 16,900 km2. A 29 km-<br />
long navigation channel, dredged to a<br />
depth <strong>of</strong> 8.2 m below low water datum<br />
( LWD) , extends f rom the mouth <strong>of</strong> the <strong>St</strong>.<br />
<strong>Clair</strong> River to the head <strong>of</strong> the Detroit<br />
River. <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has a shoreline<br />
length <strong>of</strong> approximately 272 km. <strong>The</strong> shore<br />
<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River, including main<br />
distributary channels, adds 192 km and the<br />
head <strong>of</strong> the Detroit Rfver, plus Belle<br />
Isle, add 32 km <strong>of</strong> shoreline to the study<br />
area. Of this total shoreline length <strong>of</strong><br />
496 km, approximately 188 km or 38% can be<br />
classified as coastal wet1 ands (Table 4<br />
and Appendix A).<br />
<strong>The</strong> delta is the most characteristic<br />
feature <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />
Table 3. Wildlife use and other functions <strong>of</strong> coastal wetlands at low and high<br />
water 1 eve1 s.<br />
Use/Functions <strong>of</strong> wetlands<br />
Mildlffe use<br />
Blue-winged teal (breeding)<br />
Red-w i nged b1 ackb i rd<br />
Ma1 1 ard (breeding)<br />
Sedge wren<br />
Muskrat<br />
Black tern<br />
Ye1 lowheaded b1 ackbird<br />
Great blue heron<br />
Belted kingfisher<br />
Crayfish<br />
Frogs and turtles<br />
Fish spawning (pikes)<br />
Forage fish<br />
Dabbl ing ducks (feeding)<br />
Diving ducks (feeding)<br />
tionq<br />
Peat accumul at1 on<br />
Sediment trapping<br />
Hemi-marsh<br />
Water ci rcul atf on<br />
Dominance <strong>of</strong> l and drainage<br />
Dom,inance <strong>of</strong> lake water masses<br />
Turbidity levels<br />
Export <strong>of</strong> detritus<br />
Re-suspension <strong>of</strong> In slf;ra clay<br />
a~ata source: Jaworski et a1 . (1979).<br />
Water levelb<br />
Low Water High Water<br />
b~se or function relative to water level fndi cated by 1 $nee<br />
7
<strong>Clair</strong>system. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Flats 4s a<br />
massive deposit formed by a series <strong>of</strong><br />
active and inactive distributaries at the<br />
mouth <strong>of</strong> the river. This is the largest<br />
delta in the Great <strong>Lake</strong>s system. <strong>The</strong><br />
Michigan portion <strong>of</strong> the delta has been<br />
urbanized to some extent* but the Ontario<br />
portion has been set aside as an Ind'ian<br />
reservation. <strong>The</strong> Ontario side <strong>of</strong> the<br />
delta, along with <strong>St</strong>ate wildlife areas on<br />
the Michigan side, represent the finest<br />
coastal wetlands in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
system.<br />
<strong>The</strong> shore1 lne <strong>of</strong> the <strong>St</strong>. Cl ai r River<br />
has been intensively developed for<br />
residential, commercial, industrial, and<br />
recreational use. <strong>The</strong> only extensive<br />
natural areas are on the delta wetlands<br />
(<strong>St</strong>. <strong>Clair</strong> Flats), formed where the river<br />
enters <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> vegetation <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> Delta occurs in broad<br />
arcuate zones which extend from the apex<br />
near Algonac, south into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
<strong>The</strong> zones show a progresslon from hardwood<br />
MICHIGAN<br />
Figure 2. Map <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>-Detroit River ecosystem.<br />
forests near Algonac to cattail and<br />
bulrush marshes at the lake.<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has a shallow#<br />
expansive basin with low marshy shores.<br />
Other than the delta, the best coastal<br />
marshes are found on the east shore <strong>of</strong> the<br />
lake between Mitchell Bay and the mouth <strong>of</strong><br />
the Thames River (Ontario) and in Anchor<br />
Bay between the delta and the mouth <strong>of</strong> the<br />
Cl lnton River (Michigan). <strong>The</strong> only other<br />
notable wetlands are submerged beds cf<br />
aquatjc plants in the vicinity <strong>of</strong> Peach<br />
Island and Belle Isle at the head <strong>of</strong> the<br />
Detroit Rf ver.<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, the smallest lake Jn<br />
the Laurentian Great <strong>Lake</strong>s system, has a<br />
mean depth <strong>of</strong> only 3.0 m, a maximum<br />
natural depth <strong>of</strong> 6.4 m, and a volume <strong>of</strong><br />
Table 4. Estimated dimensions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> and <strong>St</strong>. Clail; River emergent and<br />
submergent wetl ands,<br />
Coastal length Area <strong>of</strong><br />
Jurisdiction <strong>of</strong> wet1 ands wet1 ands<br />
(km) (krn2)<br />
MICHIGAN<br />
Wayne County 4.0 0.7<br />
Macomb County 6.6 11.2<br />
<strong>St</strong>. <strong>Clair</strong> County 75.2 140.7<br />
Michigan Total 85.8 152,6<br />
ONTARIO<br />
Essex County 11.5 6.8<br />
Kent County 26.5 62.8<br />
~arnbton County 64.4 161.7<br />
Ontario Total 102.4 231.3<br />
TOTAL 188.2 383.9<br />
a~ata sources: United <strong>St</strong>ates Geological<br />
Survey, Dept. Interior, 7.5-minute Quad-<br />
rangle Maps (1:24,000 scale) ; Canada<br />
Department <strong>of</strong> Energy, Mines and Resources,<br />
7.5-minute Quadrangle Maps (1:25,000<br />
scale); Direct fie1 d and aircraft obser-<br />
vations, 1983 and 1984.
4,O km3. Its deepest waters are found<br />
along a dredged navigation channel which<br />
extends from the mouth <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
River to the head <strong>of</strong> the Detroit Rfver.<br />
Approximately 98% <strong>of</strong> the water flowing<br />
into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> enters through the <strong>St</strong>.<br />
<strong>Clair</strong> Delta. <strong>The</strong> Detroit River, the on1 y<br />
outlet, drains <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> at a rate <strong>of</strong><br />
5,044 m3/sec. This yields a retention<br />
time for the lake basin <strong>of</strong> only 9.2 days.<br />
<strong>The</strong> <strong>St</strong>. Clzir River is one <strong>of</strong> the<br />
four major connecting channels on the<br />
Great <strong>Lake</strong>s, fl owl ng between <strong>Lake</strong> Huron<br />
and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> two principal<br />
cities on the river are Port Huron,<br />
Michigan, and Sarnia, Ontario, both at the<br />
head <strong>of</strong> the river at <strong>Lake</strong> Huron. <strong>The</strong> <strong>St</strong>.<br />
<strong>Clair</strong> River has two characteristic<br />
sections, the upper channel, and the<br />
lower, or delta, portion. <strong>The</strong> upper<br />
channel runs from <strong>Lake</strong> Huron to the head<br />
<strong>of</strong> Chenal Ecarte, a distance <strong>of</strong> 43 km.<br />
<strong>The</strong>re are two islands in the upper portion<br />
<strong>of</strong> the river, <strong>St</strong>ag Island and Fawn Island.<br />
At Chenal Ecarte, the river begins to<br />
branch out i n t o a number o f<br />
distributaries, forming a vast pattern <strong>of</strong><br />
islands, marshes, and waterways that are<br />
known as the <strong>St</strong>. Clalr Flats.<br />
<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River receives most <strong>of</strong><br />
Its flow from <strong>Lake</strong> Huron. Prfncipal<br />
tributaries in Michigan are the Belle,<br />
Black, and Pine Rivars. Water quality<br />
throughout the <strong>St</strong>. <strong>Clair</strong> River is<br />
genera1 1 y excel lent. Some degradation<br />
occurs i n localized areas where<br />
tributaries join the river. <strong>The</strong> width <strong>of</strong><br />
the <strong>St</strong>. Cl air River ranges from about 250<br />
to 750 m, and the maximum depth ranges<br />
between 11 and 15 m. <strong>The</strong> river's annual<br />
average discharge is about 5,070 m2/sec.<br />
Because the river's main water source is<br />
<strong>Lake</strong> Huron rather than surface run<strong>of</strong>f9<br />
seasonal f 1 ow variations are minimal .<br />
Herdendorf et al. (1981~1 1 isted four<br />
wetland complexes that are located in the<br />
lower portions <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River.<br />
<strong>The</strong> Pointe Aux Tremble Wetland (15<br />
hectares), the Pointe Aux Chenes Wetland<br />
(7 hectares), and the Russell Island<br />
Wetland (5 hectares) are all located in<br />
the North Channel <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River.<br />
<strong>The</strong> Algonac Wetland (64 hectares) is<br />
1 ocated 0.3 km f rom the shore1 f ne <strong>of</strong> the<br />
river. <strong>The</strong> Cuttle Creek Area Wetland (1,s<br />
hectares) and the Port Huron Area Wetland<br />
(1.2 hectares) are both located along the<br />
upper portion <strong>of</strong> the river.<br />
A1 though the 1 argest area <strong>of</strong> wetl ands<br />
1s found i n the <strong>St</strong>. <strong>Clair</strong> Delta, there are<br />
fragments <strong>of</strong> wetl ands around the perimeter<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Beginning at the mouth<br />
<strong>of</strong> the Detroit River, small patches <strong>of</strong><br />
rfveri ne, pal ustri ne, and 1 acustri ne open-<br />
water wetlands exist along Belle Isle,<br />
Aquatic beds <strong>of</strong> submersed vascul ar pl ants<br />
dominate as a park and other developments<br />
havs resulted in the removal <strong>of</strong> the upper<br />
communities.<br />
Along the southwestern shore <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>9 from Grosse Pointe to the<br />
Clinton River* the shoreline is inten-<br />
sively developed. Much <strong>of</strong> the shoreline<br />
is protected by seawalls, and pjers and<br />
marinas are scattered a1 ong th i s stretch.<br />
Although the shoreline is classified as<br />
1 acustrine open-water wet1 ands9 there is<br />
l i ttle wetland vegetation except for<br />
pollution-tolerant submersed aquatics such<br />
as coontail and Eurasian water-mil f<strong>of</strong>l<br />
growing in the boat channels and slips<br />
(see Appendix E for common and scientiftc<br />
names <strong>of</strong> vascular plants).<br />
Where the Clinton River empties Into<br />
<strong>Lake</strong> <strong>St</strong>. Cl airr along the western shore <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>9 there are two coastal<br />
wetland areas. South <strong>of</strong> the riverr in the<br />
Metropolitan Beach area, over 4 hectares<br />
<strong>of</strong> palustrine emergent and pal ustrine<br />
open-water wetlands exist. North <strong>of</strong> the<br />
Metropol itan Parkway highway, i .e, , along<br />
Black Creek, which is an old tributary <strong>of</strong><br />
the Cl inton River, there are sedge meadow*<br />
cattaf 1 and open-water (aquae lc bed 1<br />
communities. Because the connectyon wfth<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is poor, the water<br />
frequently stagnates resulting in lower<br />
fish and benthic invertebrate values,<br />
North <strong>of</strong> the Clinton River, located<br />
between Sand Point and Self ridge Air Force<br />
Basep is an open wetland. This wetland 7s<br />
largely a palustrine system with submersed<br />
aquatic plant communities.<br />
<strong>The</strong> shoreline <strong>of</strong> Anchor bay^ sftuated<br />
in northwest <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r is a150
largely urbanized. Residential develop-<br />
ment, marinasr and boat sljps are common<br />
in the mouths <strong>of</strong> streams as we11 as in<br />
lakeshore points. Very 1 ittle undisturbed<br />
wetland exists and much <strong>of</strong> the shorel ine<br />
is elther bulkheaded or piled with rip-rap<br />
to reduce erosion assocjated with the<br />
current high water levels. Although the<br />
entire shoreline i s classifled as<br />
lacustrine open-water wetl ands, on1 y<br />
scattered patches <strong>of</strong> emergent cattails and<br />
submersed aquatics exist along the creek<br />
mouths and behf nd bulkheads. Clumps <strong>of</strong><br />
bulrushes occur along the shorel ine9 along<br />
with strips <strong>of</strong> wild celery and other wave<br />
tolerant submersed aquatics.<br />
<strong>The</strong> first large area <strong>of</strong> deltaic<br />
wetland occurs along Bouvier Bay, and Is<br />
locallv referred to as <strong>St</strong>. Johnqs Marsh.<br />
~~~roximatel~ 1,000 hectares in sJze, this<br />
traditional 1 y p? anted in corn and other<br />
artificial waterfowl foods. Most <strong>of</strong><br />
Dickinson Island and the lower half <strong>of</strong><br />
Harsens Island together make up the <strong>St</strong>.<br />
Clafr Flats Wildlife Area, which is<br />
managed by the Michigan Department <strong>of</strong><br />
Natural Resources for waterfowl hunting.<br />
With reference to the Ontario side <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> Delta, that wetland complex<br />
i n cl udes port ions <strong>of</strong> Bassett Is1 andt<br />
Squirrel Island, Walpale Island# and <strong>St</strong>.<br />
Anne Is1 and. Bassett Is1 and consi sts<br />
mostly <strong>of</strong> cattail marsh with open-water<br />
ponds and a thin beach colonized by shrubs<br />
and herbaceous pl ants. Squi rrel Is1 and is<br />
also largely a natural cattail marsh* but<br />
the northern part, where sedge marsh would<br />
naturally occur, i s diked <strong>of</strong>f and<br />
cu1 tivated.<br />
Canada's major wetland complex in the<br />
is found both 'Ides delta focuses on Walpole Island where<br />
Of M-29? and extends from Fair<br />
Goose <strong>Lake</strong> and Johnston Bay are located.<br />
Haven to Pearl Beach. Plant communities<br />
Deciduous woodlands, shrubs, and sedge<br />
in this are by meadows form the upper part <strong>of</strong> this<br />
and sedge<br />
meadows cattaf1s8 bul and<br />
leaved and submersed aquat icr.<br />
CDnstructi on Of access roads and<br />
f'll jng Iror residential as<br />
"11 as the Of highway M-29 have<br />
fragmented this wetl and.<br />
island* but much <strong>of</strong> the natural shrub and<br />
meadow zones have been cleared and owed<br />
into row-crop agrlculture, A diked and<br />
water-level regulated cattail marsh occurs<br />
north <strong>of</strong> Goose <strong>Lake</strong>. South <strong>of</strong> Goose <strong>Lake</strong><br />
the wetlands are dominated by cattails and<br />
sedqe communities along the distributary<br />
~ i ~ 1 ~ and k1 js i the ~ 1 argest ~ ~<br />
chaknels<br />
~<br />
and shorel ine beaches. <strong>The</strong><br />
natural undeveloped, and functioning <strong>Wetlands</strong> near Johnston are open to<br />
wetland complex along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>Lake</strong><strong>St</strong>*<strong>Clair</strong>*<br />
Approximately lr200 hectares, this wetland<br />
has a continuum <strong>of</strong> environments. To the<br />
north, where elevations average 1.5 to 3 m<br />
above North Channel* swamp forest and<br />
dogwood shrub communities occur. In the<br />
mfddle <strong>of</strong> this deltaic island, several<br />
abandoned river channels bisect vast<br />
stretches <strong>of</strong> sedge meadow and cattail<br />
communit f es. Along the shore1 ine, the<br />
emergent wetlands give way to various<br />
<strong>The</strong> wetlands on <strong>St</strong>. Anne Island<br />
appear much more managed. Large corn<br />
fields on the upper and middle portions <strong>of</strong><br />
the island extend to the limit <strong>of</strong> the<br />
cattail marsh. At this point* the marshes<br />
are diked for the benefit <strong>of</strong> private<br />
waterfowl clubs. On1 y the shorel f ne<br />
fringe <strong>of</strong> the wetlands is open to <strong>Lake</strong> <strong>St</strong>.<br />
C1 air.<br />
open-water aquatic bed-type wetlands,<br />
<strong>The</strong> last major wetland area along the<br />
Ontario side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr Is in the<br />
Another major wetland complex along Mitchell Bay area. Because <strong>of</strong> the<br />
the American side <strong>of</strong> the <strong>St</strong>. Clai r Delta proximity <strong>of</strong> the managed marshes on <strong>St</strong>.<br />
is Harssns Island. Only the lower quarter Anne Island and the cultivation by<br />
<strong>of</strong> thls deltaic island remains a wetland, Canadian farmers rtght up to the lakeshore<br />
consisting largely <strong>of</strong> hybrid cattail<br />
(Typha x ~lauca) colonies along with the<br />
assocfated floating-1 eaved and submersed<br />
canal and shore1 Sne communities. <strong>The</strong><br />
center <strong>of</strong> Harsens Island is part <strong>of</strong> the<br />
<strong>St</strong>. Clafr Flats Wildlife Area and is<br />
in Kent and Lambton counties, the wetlands<br />
<strong>of</strong> Mitchell Bay consist mainly <strong>of</strong> openwater<br />
ar submerged cammunlties. Moreover,<br />
considerable diking and waterfowl nesttng<br />
improvements have taken place along<br />
Mitchell Point.
This section contains a description<br />
<strong>of</strong> the geographic location <strong>of</strong> the study<br />
areao including a discussion <strong>of</strong> topog-<br />
raphy, morphometry, and coastal geomor-<br />
phology, Although no bedrock or glacial<br />
features are directly related to the<br />
distribution <strong>of</strong> wetlands there are bio-<br />
geographical associations with more recent<br />
landforms such as natural levees, crevasse<br />
deposits, interdistributary bays, and<br />
other del talc deposits.<br />
<strong>The</strong> occurrence and distribution <strong>of</strong><br />
wetlands is determined, to a significant<br />
degree, by a combination <strong>of</strong> physical<br />
factors. In the Great <strong>Lake</strong>s, coastal wet-<br />
lands are at the interface <strong>of</strong> land, water,<br />
and atmosphere. <strong>The</strong>se three components <strong>of</strong><br />
the biosphere are delicately balanced to<br />
produce and maintain the biologi.ca1<br />
resource. In this chapter the physical<br />
parameters <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />
are exam1 ned.<br />
<strong>Wetlands</strong> occur in basins or topo-<br />
graphical depressions whlch are wet or<br />
flooded for at least several months <strong>of</strong> the<br />
year. <strong>The</strong> occurrence <strong>of</strong> the resource is<br />
therefore in part determined by water<br />
depth, which is in turn determined by the<br />
submarine topography, sed imentationr and<br />
character <strong>of</strong> 1 ake level oscillations.<br />
Another factor responsible for wetl and<br />
distribution is wave action in open water<br />
bodies and current characteristics in<br />
riverine and 1 ittoral areas. Many <strong>of</strong><br />
these factors are in turn directly related<br />
to climatic conditions. A simplified<br />
diagram illustrating these relationshjps<br />
appears in Figure 3.<br />
<strong>The</strong> more extensive wetlands <strong>of</strong> the<br />
Great <strong>Lake</strong>s are located in ancient lake<br />
bottoms or in areas <strong>of</strong> h5gh sedi mentation<br />
CHAPTER 2. PHYSICAL EIUIVOIRBWMENT<br />
I CLIMATE I<br />
1<br />
BOTTOM TOPOGRAPHY<br />
Figure 3. Physical factors influencing<br />
the occurrence <strong>of</strong> coastal wetl ands.<br />
rates. In the geological past, water<br />
levels in the Great <strong>Lake</strong>s were higher and<br />
progl aci a1 1 akes inundated the lower<br />
terrain, As the water levels <strong>of</strong> the lakes<br />
droppedr a series <strong>of</strong> clay lake plains were<br />
exposed which were colonized by vast<br />
wetl ands. Western <strong>Lake</strong> Erie, includfng<br />
the Maumee River embaymentp for examplep<br />
was an extensive wetland known as the<br />
Black Swamp. <strong>The</strong> margins <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />
Clai r, especially in Ontarf or are composed<br />
<strong>of</strong> a similar deposit.<br />
<strong>The</strong> rivers flowing f nto take <strong>St</strong>.<br />
Clai r deposited sediments a1 ong valley<br />
floors to create floodplains which were<br />
colonized by wetland communities. At the<br />
same time deposition in the lake created<br />
the <strong>St</strong>. Clafr Delta. Although most <strong>of</strong><br />
Michigan has a glacfal herttage, coastal<br />
wetlands are a product <strong>of</strong> fluvial,<br />
deltaic? and lacustrine sedimentation.<br />
<strong>The</strong> <strong>St</strong>, Clai r DeTta has been depos-<br />
ited* in past because <strong>of</strong> the shallowness<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Compared to marfne<br />
shore1 inesp ongolng coastal deposit ion<br />
which produces subaeri a1 l andforms such as<br />
the delta is not common in the Great
<strong>Lake</strong>s. In fact, no other signff icant<br />
deltas occur in the Great <strong>Lake</strong>s, <strong>The</strong><br />
combination <strong>of</strong> a shallow receivfng basfn<br />
and an abundance <strong>of</strong> sediments fs largely<br />
responsible for the deposttion <strong>of</strong> the <strong>St</strong>.<br />
Clafr Delta which fs colonized by the most<br />
extensive wetlands in the coastal Great<br />
<strong>Lake</strong>s.<br />
<strong>The</strong> progradatton or recession <strong>of</strong> the<br />
coastal zone is 1 argely influenced by wave<br />
and currents. High energy environments<br />
discourage the establ i shment <strong>of</strong> pal ustri ne<br />
and lacustrine wetlands. Also high wave<br />
energies hinder delta deposition and<br />
growth. In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wave and<br />
current energy are not excessive. Thls<br />
physical condition has allowed the<br />
development <strong>of</strong> the delta with its diverse<br />
wetland habjtats. Also the low wave<br />
energies have encouraged wetland coloniza-<br />
tion north <strong>of</strong> the Thames River along the<br />
eastern shore1 ine <strong>of</strong> the 1 ake.<br />
<strong>The</strong> climatic elements Including<br />
temperature, precfpi tation* evaporation,<br />
and wfnd velocities and dire~tions exert a<br />
control on water levels and wave<br />
parameters, <strong>The</strong>se in turn dictate wetland<br />
occurrence and types. With changes In<br />
water levels the geographfcal di stributlon<br />
<strong>of</strong> wetlands shfft or "pulse stabi71tyw<br />
occurs. Also the character <strong>of</strong> sedimentation<br />
is altered to a point where water<br />
quality (i.aer turbidity) levels are<br />
impacted.<br />
It has been determfned that wetlands<br />
in the Great <strong>Lake</strong>s colonize a diversity <strong>of</strong><br />
geomorphic settings such as backbarrfer<br />
flats, alluvial flood plains and deltas.<br />
<strong>The</strong> <strong>St</strong>. C1 air Delta (Figure 4) is rnade up<br />
<strong>of</strong> a suite <strong>of</strong> landforms which support a<br />
diversity <strong>of</strong> wet1 and types, Deltaic<br />
deposits in the Great <strong>Lake</strong>s are rare<br />
features, suggesting that fluvial sedirnen-<br />
tation is not abundant and/or wave<br />
energies are too high thus not permfttlng<br />
delta development to occur. <strong>The</strong>refore the<br />
<strong>St</strong>. Cl ai r Delta Is a unique Great <strong>Lake</strong>s'<br />
Figure 4. Depositional features and landforms <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta.<br />
12
coastal feature supporting abundant and<br />
diverse wet1 and communities not found on<br />
other Great <strong>Lake</strong>s' shore7 i nes.<br />
Since the classical study <strong>of</strong> <strong>Lake</strong><br />
Bonneville by Gilbert (18901, deltas in<br />
lakes* with only a few exceptions, have<br />
been largely ignored by geomorphologists.<br />
<strong>The</strong> major deltas <strong>of</strong> the world are located<br />
on contlnental shelves and have held the<br />
interest <strong>of</strong> geologists and geographers<br />
because <strong>of</strong> petroleum resources, agricul-<br />
tural production, and waterborne activi-<br />
ties. A1 though 1 ake deltas are smaller,<br />
they share most <strong>of</strong> the processes<br />
characteristic <strong>of</strong> marine deltas. However,<br />
a unique aspect <strong>of</strong> lake deltas is their<br />
dynamic response to re1 atively rapid<br />
changes <strong>of</strong> base level that not only<br />
produce special 1 andforms, but dl sti nctive<br />
vegetation zonation as we1 1.<br />
<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta is the largest<br />
delta in the Great <strong>Lake</strong>s basfn. Despite<br />
the delta's recreational sign1 f icance and<br />
commercl a1 importance with regard to<br />
navigation, literature on the area has not<br />
been abundant, <strong>The</strong> first significant<br />
investigation was done several decades ago<br />
by Cole (19031 who determined that the<br />
delta was beina deposited atop deep water,<br />
proglacial 1 &e clays. p ore recent1 yr<br />
Wfghtman (1961) attempted to establish a<br />
1 ate Quaternary chronology for the del tats<br />
formation. Detailed investigations by<br />
geol og 5 sts have part1 ally documented the<br />
sediment01 ogical composition <strong>of</strong> the<br />
deltaqs surface, parti cularly in Muscamoot<br />
and Goose bays (Pezzetta 1968, Mandelbaum<br />
1969). During the 1950~8 engineering<br />
studies by the U.S. Army Corps <strong>of</strong><br />
Engineers preceded construction <strong>of</strong> the 8-m<br />
deep <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel through the<br />
delta. Although not published, much <strong>of</strong><br />
these data, in the form <strong>of</strong> bore records*<br />
are on file at the Detroit District<br />
Office. Recent environmental studies<br />
associated with flooding problems on the<br />
shores <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and dredged spoil<br />
disposal s<strong>St</strong>e locations have also<br />
contributed some useful data for this<br />
outlets <strong>of</strong> the Great <strong>Lake</strong>s were changing<br />
or even blocked, causing the lake levels<br />
to oscillate several meters. <strong>Lake</strong> Erie<br />
established its present level some 4,000<br />
years ago and <strong>Lake</strong> Huron shortly<br />
thereafter. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River and its<br />
delta came to existence during these<br />
changing lake levels.<br />
As a lake delta, the <strong>St</strong>. <strong>Clair</strong><br />
exhibits several <strong>of</strong> the landform<br />
characteristics <strong>of</strong> marine deltas* such as<br />
active and inactive distributaries,<br />
interdfstrfbutary bays (Figure 5)s and<br />
crevasses or wlde breaches in the channel<br />
banks which lead into interdistributary<br />
bays (Figure 6). <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta has<br />
a classical bi rd-f oot morphology , as does<br />
the Mississippi River Delta. However,<br />
significant landform differences are also]<br />
apparent. Atypical landforms include a<br />
premodern surface at the apex <strong>of</strong> the delta<br />
and unusual ly wide distributary channels.<br />
section <strong>of</strong> the study (U.S. Army Corps <strong>of</strong><br />
Engineers 19741,<br />
Wfth tke retreat <strong>of</strong> the Late<br />
Figure 5. Little Muscamoot Bay, an interdistri<br />
butary bay between Middle Channel<br />
and South Channel <strong>of</strong> the <strong>St</strong>. Cfair 3e7ta<br />
W i sconsin ice sheet some 13,000 years ago, (August 1984). Note emergent cattails<br />
a series <strong>of</strong> moraines and till plains were (Typha an ustifolia) and solitary<br />
deposited in the Great <strong>Lake</strong>s basln. <strong>The</strong> pied-bili&~ddilymbus podiceps).
Figure 6. We1 1-developed crevasse deposits adjacent to Middle Channel <strong>of</strong> <strong>St</strong>. <strong>Clair</strong><br />
Delta. <strong>The</strong>se crevasses have been intermittently active for over 100 years. Dickinson<br />
Island is to the upper left and Harsens Island is to the lower right.<br />
<strong>The</strong> active dJstributaries--Norths<br />
Middlet and South C,hannel s--average some<br />
500 m in width and 11 m in depth.<br />
Howevert widths <strong>of</strong> 675 m and depths <strong>of</strong> 26<br />
m are not uncommon. At the mouths <strong>of</strong> the<br />
dfstrfbutarfes, channel depths decrease<br />
abruptly Indicating the presence <strong>of</strong> river<br />
mouth bars 2 to 4 m below mean lake level.<br />
As a deposftional basin, <strong>Lake</strong> <strong>St</strong>. Clafr fs<br />
relatfvely small with a maximum natural<br />
depth <strong>of</strong> 6.4 m and width <strong>of</strong> 40 km.<br />
North, Middle* and South Channels<br />
exhibit shoulder-1 i ke features or shoals<br />
along both the cutbank and point-bar sides<br />
(Ffgure 71, A sfmjlar morphology has been<br />
attrfbutsd to periodic cut and ffll<br />
associated wfth slfght base level<br />
oscillations (Butzer 1971). Borings<br />
obtained from the Corps <strong>of</strong> Engineers<br />
reveal that the distributary channels <strong>of</strong><br />
the <strong>St</strong>. ClaSr Delta are entrenched in<br />
laeustrine c1 ays which 1 ie beneath a 3.5<br />
to 5 m veneer <strong>of</strong> coarser deltaic<br />
sediments. Qn the cutbank side, where<br />
flow velocftfes may be relatlvely high<br />
during high water condltlons, an erosfonal<br />
berm usually less than 2.5 m below the<br />
water surface slopes gently from the<br />
cutbank toward the channel center, <strong>The</strong>se<br />
features may be caused by lateral erosion<br />
<strong>of</strong> the fine, sandy deltaic sedtments which<br />
over1 ie the 1 acustrfne clays. On the<br />
inside bank, especially along Middle<br />
Channel , point bar deposits characterized<br />
by ridge and swale topography are<br />
conspicuous. Here the distributary<br />
shoulders probably represent a fill<br />
deposit which is colonized by emergent<br />
vegetati on during 1 ow-water periods.<br />
Because the water level fluctuates<br />
only 45 to 60 cm seasonally, spring floods<br />
are not a normal occurrence within the<br />
delta, hence natural levees are scarcely<br />
dfscernable adjacent to modern distrib-<br />
utaries. Even though levees are poorly<br />
developeds averaging 10 to 45 cm in<br />
elevationr flooding and subsequent<br />
overbank flow does occur. Overbank flow<br />
is assocfated with breaching <strong>of</strong> levees in<br />
abnormally low areas along a levee. As a
.................<br />
...............<br />
MODERN SURFACE<br />
SUBMERGED RIVER<br />
RELICT CHANNEL<br />
Figure 7. Deltaic landforms <strong>of</strong> Dickinson Island, Michigan,
levee fs breachedr crevasse deposits are<br />
Introduced into the interdistributary bays<br />
at right angles to the channels (Figure<br />
71. With contf nued depositionp the open-<br />
water bay w i l l be filled with crevasse<br />
deposits and colon 1 zed by sedges and<br />
emergent aquatics.<br />
Crevasse channel s9 1 ocal l y known as<br />
"highwaysw (see Figure 57 in Section 5,1Ir<br />
are operative for several years.<br />
Nevertheless, deposition into the bays i5<br />
not rapid, A comparison <strong>of</strong> navigation<br />
maps reveals zhat such features may be<br />
part <strong>of</strong> the delta landscape for over a<br />
century. Pezzetta 11968) graphical1 y<br />
compared the western portton <strong>of</strong> the delta<br />
front between 1903 and 1961. Over the 58-<br />
year perjod crevasse fills were minor. In<br />
fact human-mads a1 terati ons such as<br />
dredging and bulkheadlng dominated the<br />
1 andscape. This suggests that crevasse<br />
channels are active intermittently and<br />
transport l i t tle sediment into the<br />
interdistributary bays.<br />
During the winter and early spring<br />
when <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is frozen, pack ice<br />
accumulates at the mouths <strong>of</strong> distributarjes<br />
forming ice jams. Channel flow is<br />
then dfverted lnto crevasses and some<br />
overbank flow may occur, Because the<br />
dominant grain size is sand, 1 ittle bed<br />
load sedlment is transported fr~m the deep<br />
distributaries Into the interdistributary<br />
bays. Thus, In the <strong>St</strong>, <strong>Clair</strong> Delta the<br />
filling <strong>of</strong> interdistrSbutary bays and<br />
delta growth Is a slow process.<br />
In contrast, the Mississippi Delta<br />
crevasse depos f ts rapidly convert open<br />
interdistributary bays into mud flats<br />
which are subsequently colonized with<br />
marsh grasses. During flood stage, the<br />
crevasse channels are scoured deep enough<br />
to be maintained and rapid depositlon<br />
occurs, In the past 135 years, crevasse<br />
deposf t s have transformed the open<br />
interdistributary bays <strong>of</strong> the modern<br />
Mississippi Rfver Delta into a complex <strong>of</strong><br />
marshy subdeltas (Coleman 1976).<br />
Beaches on the present delta<br />
shorel ine are poorly developed and appear<br />
transgress'ite in orf~fn, On t he Canadian<br />
side, where the beaches are somewhat<br />
better developed* the berms may reach I t o<br />
1.5 m In height and are colonfzed with<br />
sumac and small trees, Borings reveal<br />
that the principal constituents <strong>of</strong> these<br />
beaches are coarse sand or f lne gravel<br />
(0.3 m or -L.5 phi unJts) separated by<br />
layers <strong>of</strong> sand and organic materials<br />
including rafted logs, bulrush stems, and<br />
other debris. Coarse sands and fine<br />
gravels are not evident, and storm berms<br />
seldom exceed 0.7 m In elevation.<br />
Characteristic vegetation <strong>of</strong> these<br />
shorel ine features are either sedge marsh<br />
or a complex community <strong>of</strong> grasses,<br />
thistles8 and other nonwoody species.<br />
Within the interd9stributary marshes,<br />
especf a1 1 y on lower DiC~inSon and Harsens<br />
is1 ands, are arcuate-shaped features<br />
resembling beach ridges. Borings through<br />
one <strong>of</strong> these ridges revealed up to 3 m <strong>of</strong><br />
flne sand (Raphael and Jaworski 1982).<br />
<strong>The</strong> absence <strong>of</strong> washover deposits and<br />
organlc sediments indicate that these<br />
features may be regressive beaches and<br />
represent shorelines as delta accretion<br />
took place.<br />
A comparison between the eastern and<br />
western portions <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta<br />
illustrates two other distinct dif-<br />
ferences. On the CanadSan side, Chenel<br />
Ecarte and Johnson Channel are narrow,<br />
shall ow distributaries which do not carry<br />
a significant portion <strong>of</strong> the volume <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> River. Moreover9 open Inter-<br />
distributary bays are few and are<br />
colonized by marsh vegetation. Delta<br />
extension has ceased and maximum delta<br />
accretion ls now occurring to the west as<br />
evidenced by the active dlgital<br />
dtstributaries <strong>of</strong> North, Middle, and South<br />
Channels, and Chenal A Bout Rond. In the<br />
past, Chematogan and Bassett Channels were<br />
approximately 500 m wide comparable to the<br />
modern distributaries, but have been<br />
alluviated and colonized with aquatfc<br />
plants as abandonment occurred.<br />
In most deltas with large fluvial<br />
systems, lateral migration <strong>of</strong> the delta<br />
occurs because <strong>of</strong> the changes in the<br />
course <strong>of</strong> the river upvalley. <strong>The</strong><br />
premodern 1 ateral displ acements <strong>of</strong> the<br />
Mississippi River Delta originated within<br />
the alluvial valley several miles from the<br />
Gulf <strong>of</strong> Kexico% coast. Such a dlversfon<br />
process is evident in some lake deltas as<br />
well. Several older deltas <strong>of</strong> the Omo<br />
Rlverr for example, have been identi f fed
and related to the former positlon <strong>of</strong> the<br />
rfver within its alluvial valley (Butzer<br />
1971).<br />
<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta# however, has no<br />
comparable a1 luvial uall ey, and de1 ta<br />
migration has occurred from east to west<br />
at the shoreline <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>. As<br />
the Canadian distributaries degeneratedr<br />
new distributaries were created on the<br />
western side <strong>of</strong> the delta.<br />
A series <strong>of</strong> borings, cores, and<br />
exposures indicates that i n cross sect i on<br />
the <strong>St</strong>. <strong>Clair</strong> Delta is a thin and sandy<br />
deposit. An east-west cross section<br />
reveals that above the shale bedrock,<br />
1 acustrine c1 ays have been deposited over<br />
a thin deposit <strong>of</strong> glacial till (Figure 8).<br />
<strong>The</strong> coarse deltaic deposits, having a<br />
maximum thickness <strong>of</strong> 7 m rest upon blue<br />
lake clays.<br />
<strong>The</strong> north-south cross section from<br />
the apex <strong>of</strong> the delta into <strong>Lake</strong> <strong>St</strong>. Cl air<br />
S<strong>of</strong>t<br />
Clay<br />
f 11 ustrates the near-surface stratigraphy<br />
(Figure 9). Topographically however, this<br />
cross section reveals two distl nct 1 evels,<br />
a modern and the equally obvious premodern<br />
surface. <strong>The</strong> premodern surface,, standing<br />
about 1.5 rn above take <strong>St</strong>. <strong>Clair</strong>, consists<br />
<strong>of</strong> coarse* 0x1 dized sand and is confined<br />
to the apex <strong>of</strong> the delta complex. It has<br />
been dissected by long, sinuous channels<br />
which have been alluviated. Occasionally<br />
during high 1 ake levels these channels<br />
have been reoccupfed~ particularly in<br />
areas such as Dickinson Island, where<br />
human interference has been minimal.<br />
Because this hfgher surface i s<br />
topographically and sedimentologically<br />
distinct, it must be a surface which was<br />
deposited during a pre-existing higher<br />
lake level. <strong>The</strong> modern delta with its<br />
fine sand sediment (0.25 to 0.125 mn or +2<br />
to 3 phi units) is located at present mean<br />
lake level and is represented by the<br />
active crevasses and interdistributary<br />
marsh deposits.<br />
Based on the chronology <strong>of</strong> the<br />
progl acial Great <strong>Lake</strong>s*. field data, and<br />
Figure 8. Geologic cross-section <strong>of</strong> the northern portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> showing<br />
depth to bedrock, and thickness <strong>of</strong> glacial, lacustrine, and deltaic deposits,<br />
S<strong>of</strong>t<br />
Clay
61 inkc<br />
<strong>St</strong>. <strong>Clair</strong><br />
I<br />
Modern Delta Premodern Delta<br />
Bassett Harrens<br />
Island Island<br />
Walpole<br />
Island<br />
I North<br />
1 2 3 4 5 6 7 8 9 10 11 12 Miles<br />
I I I --LL--l I I I I<br />
Deltaic Sediments<br />
a ORGANIC<br />
SILTY SAND<br />
SANDY CLAY<br />
Figure 9. Cross-secti on <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta showi<br />
L .~ - A-2<br />
.- . --.-- - _I<br />
stribution <strong>of</strong> deltaic sediments.<br />
available Carbon-14 data, the relative materfals range in age from 6,100 + 80 to<br />
geomorphological events <strong>of</strong> the <strong>St</strong>. Clafr 98300 + 200 years. <strong>The</strong>se organic<br />
Delta may be determined. Sfnce the materials do not appear to behsifad and<br />
retreat <strong>of</strong> the Late Wisconsin ice sheet,<br />
the outlets <strong>of</strong> tho Great <strong>Lake</strong>s, and hence<br />
lake levels in the <strong>St</strong>. <strong>Clair</strong> basin, have<br />
oscillated sufficient1 y to construct two<br />
deltas. Compared to other proglacial<br />
Great <strong>Lake</strong>s, <strong>Lake</strong> <strong>St</strong>. Clafr was a less<br />
permanent feature as the lake bottom was<br />
exposed to subaerial modlf ication due to<br />
the f 1 uctuatf ng ice sheet which determl ned<br />
lake level conditl~n~ (Figure 10).<br />
One C-14 date retrieved from the<br />
upper portion <strong>of</strong> the lake claysr but<br />
beneath the premodern delta, f ndicates<br />
that both deltas are less than 7,300 + 80<br />
years old, Based upon older proglacial<br />
Great <strong>Lake</strong>s chronology, the premodern<br />
delta may have been deposited during the<br />
. . .580'<br />
T-- - 7<br />
12000 9000<br />
Years belore the present<br />
ALGONQUIN STANLEY NIPISSING<br />
latter stage <strong>of</strong> Algsnquf n time (Wightman Figure 10. Chronology <strong>of</strong> lake levels and<br />
19611, Additfonal C-14 dates have been lake stages in the Huron and Erie basins<br />
obtained by Mandelbaum (1969). <strong>The</strong> dated (Darr and Eschman 1970).
they have not been jncluded in our<br />
interpretation, However, even if the<br />
latter dates were used they would not<br />
detract from our argument. Pezzetta<br />
(1968) also has radiogenic data that<br />
reveal that the minimum age <strong>of</strong> ,the delta<br />
sediment is 4,300 years before present<br />
(B.P. 1.<br />
With the subsequent retreat <strong>of</strong> the<br />
ice during the <strong>Lake</strong> <strong>St</strong>anley low-water<br />
stager an outlet for the <strong>Lake</strong> Huron basin<br />
to the <strong>St</strong>. Lawrence was created via North<br />
Bay, Ontario. During this stage the pre-<br />
modern sands <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta may<br />
have been exposed, oxidized, and dissected<br />
(Figure 10). Following the low-water<br />
stage, <strong>Lake</strong> Huron once again rose to the<br />
Nipissing stage and drained into the lower<br />
Great <strong>Lake</strong>s via the <strong>St</strong>. <strong>Clair</strong> River.<br />
As determined by the more recent<br />
chronology, the premodern delta may have<br />
been deposited during Nipissing time some<br />
3,500 to 5,000 B.P. and not during<br />
Algonquin time (Dorr and Eschman 1970).<br />
<strong>The</strong> premodern delta was deposited during a<br />
higher than present lake level and the<br />
subsurface sediments range In age from<br />
7,300 to 9,300 B.P. It i s suggested that<br />
fol lowf ng the <strong>Lake</strong> <strong>St</strong>anley stage (i.e.,<br />
Nipissing stage), the <strong>St</strong>. <strong>Clair</strong> River once<br />
again flowed south into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />
the premodern delta was deposited at an<br />
elevation sl ightly higher than the modern<br />
delta (Table 5).<br />
On many coasts, evidence for lower<br />
than present sea levels is determined by<br />
depositional surfaces which dip beneath<br />
younger sediments (Shelemon 1971). Had<br />
the premodern delta been deposited du r i ng<br />
the Algonquin stage and the subsequent<br />
early <strong>Lake</strong> <strong>St</strong>anley stage, evidence <strong>of</strong> that<br />
event as a depositional surface beneath<br />
the modern delta or as a paleosol should<br />
be apparent. Numerous borings suggest<br />
that the premodern delta does not plunge<br />
beneath the modern sediment. Since the<br />
stratigraphy imp1 ies that lake levels were<br />
lower than present only once, a Nipissing<br />
stage is favored for the depositton <strong>of</strong> the<br />
Table 5. Interpretation <strong>of</strong> the events related to the origin <strong>of</strong> the <strong>St</strong>.<br />
<strong>Clair</strong> Delta. a<br />
Approximate <strong>Lake</strong> stage Event<br />
Dates B.P.<br />
3,500 to Present Modern <strong>Lake</strong> <strong>St</strong>. Cl air Flow <strong>of</strong> <strong>Lake</strong> Huron continues<br />
and Algoma Phase southward; deposition <strong>of</strong><br />
modern <strong>St</strong>. Cl air River Delta<br />
and dissection <strong>of</strong> premodern<br />
surface<br />
3,500 to 5,000 <strong>Lake</strong> Nipissing Deposltion <strong>of</strong> premodern delta<br />
approximately 1.5 m (5 ftl<br />
above present mean 1 ake level<br />
5,000 to 10,500 <strong>Lake</strong> <strong>St</strong>anley <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin exposed;<br />
outlet for upper Great <strong>Lake</strong>s<br />
via North Bay* Ontario<br />
101500 to 12,500 <strong>Lake</strong> A1 gonqui n Val ders Maximum<br />
and Post-Algonquin<br />
Phases<br />
a~ata sources: Dorr and Eschman (19701, Raphael and Jaworski 619825,
premodern delta. Organic deposits, com-<br />
monly composed <strong>of</strong> large wood fragments<br />
(probably red ash) 1le beneath both<br />
deltas. Assuming that the peat accum-<br />
ulated from organic growth above the water<br />
table, it probably was deposited during a<br />
low-water period prior to the Niplssing<br />
stage ( possl bl y <strong>Lake</strong> <strong>St</strong>an1 ey 1.<br />
Following the <strong>Lake</strong> Nipissing high<br />
level, <strong>Lake</strong>s <strong>St</strong>. <strong>Clair</strong> and Huron fell to<br />
their present levels. <strong>The</strong> premodern<br />
channels were sl lghtly entrenched and the<br />
premodern surface oxidized during the last<br />
3,500 years. Towards the apex <strong>of</strong> the<br />
premodern delta, beyond the areas <strong>of</strong><br />
flood1 ng, a mix <strong>of</strong> hardwoods has colonized<br />
the oxidized soils and evidence <strong>of</strong><br />
drowning <strong>of</strong> a pre-<strong>Lake</strong> <strong>St</strong>anley delta by<br />
Niplssing high water levels is lacking.<br />
With the fa11 to the approximate present<br />
lake level, the modern delta was deposited<br />
in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Deltaic landforms occur where<br />
substantial quantities <strong>of</strong> clastic sediment<br />
are Introduced and deposited into a<br />
receiving basin. Sediment is normally<br />
eroded from a drainage basin and<br />
transported down an alluvia1 valley by a<br />
master stream and its tributaries.<br />
<strong>The</strong>refore the rate <strong>of</strong> delta development is<br />
dependent upon the flow regime and the<br />
avaf 1 abil ity <strong>of</strong> sediments.<br />
<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River maintains a<br />
relatively stable channel and i s not an<br />
alluvlal stream In the common connotatlon.<br />
Fundamentally. the river is a strait<br />
connecting two large water bodies and<br />
therefore does not have a high variability<br />
<strong>of</strong> flow. Flow measurements during the<br />
ice-free season in 1959 revealed that<br />
discharge ranged from a low <strong>of</strong> 3,900<br />
m3/sec to a high <strong>of</strong> 5,700 m3/sec. By<br />
contrast the Mississippi River flow<br />
between Baton Rouge and the Gulf <strong>of</strong> Mexico<br />
varies from a minimum <strong>of</strong> 1,400 d/sec to a<br />
maxJmum <strong>of</strong> 44,400 m3/sec (Duane 1967).<br />
<strong>The</strong>refore* for the <strong>St</strong>. <strong>Clair</strong> River, low<br />
flow was 68% <strong>of</strong> the high flow as compared<br />
to the Mississippi River where low flow<br />
was only 31% <strong>of</strong> the high flaw. <strong>The</strong><br />
discharge <strong>of</strong> the <strong>St</strong>. Clatr River is<br />
relatively constant and averages about<br />
5~097 m3/sec (International Joint<br />
Commissfon 19811, Instead <strong>of</strong> a spring<br />
flood typical <strong>of</strong> most rivers, discharge in<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fs s1 ightly Increased In<br />
late-summer when lake levels In <strong>Lake</strong> Huron<br />
are highest, Thus, overland flow and<br />
flooding Is not an annual event and -its<br />
occurrence is dependent in part upon<br />
seasonal 1 ake l eve1 condi tions.<br />
Greatest delta extension occurs In<br />
areas <strong>of</strong> highest flow and sediment yield.<br />
Flow di stribution data make it clear that<br />
the most active portlon <strong>of</strong> the delta is<br />
presently confined to the western side <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Much <strong>of</strong> the flow <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> River is carried by the large<br />
distributaries on the Amerlcan side <strong>of</strong> the<br />
delta. North, Middle, and South Channels<br />
account for approximately 95% <strong>of</strong> the flow<br />
volume whereas the principal Canadian<br />
distributary, Chenal Ecarte, accounts for<br />
5%. Although North Channel appears to<br />
have been the main channel a century agoj<br />
-. -<br />
construction and continual dredsins <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel has fncreased the<br />
f1 ow <strong>of</strong> South Channel.<br />
Because the <strong>St</strong>. <strong>Clair</strong> River is not a<br />
true river system and has few tributary<br />
streams the source area for the deltaic<br />
sediments is not solely <strong>of</strong> fluvial origin.<br />
Rather, the principal source appears to be<br />
the shorelines <strong>of</strong> southern <strong>Lake</strong> Huron.<br />
Although some organic matter is carried In<br />
suspension, the clastic fraction is<br />
dominant. In terms <strong>of</strong> composftion, quartz<br />
sand Is the most commonr but fragments <strong>of</strong><br />
feldspar, chert, carbonate minerals and<br />
igneous and metamorphic rock fragments<br />
also occur. According to Duane (19671,<br />
the mean diameter <strong>of</strong> the suspended river<br />
sediment ranges from 0.17 to 3.16 KWI while<br />
the mean diameter <strong>of</strong> the bed load varies<br />
from 1.98 to 2.64 mm. Detailed sedimen-<br />
tological studies suggest that the shallow<br />
bays <strong>of</strong> the delta are composed <strong>of</strong> fine<br />
sand (0.115 mm) deposits and the sorting<br />
Is fair (Mandelbaum 1966). It may be<br />
concluded that the <strong>St</strong>. <strong>Clair</strong> Delta is<br />
composed <strong>of</strong> sand-sized sediments. In<br />
contrast most marine deltas consist <strong>of</strong> a<br />
finer sediment fraction. Also where<br />
organic deposlts occur on the delta<br />
surface or subsurfacer they were formed in<br />
place and not transported into the de1 t a<br />
comp l ex.<br />
Estimates <strong>of</strong> the sediment discharge<br />
rates were compiled by Pezzetta (1968) and
are summarized on Table 6. <strong>The</strong> high<br />
variability is probably a reflection <strong>of</strong><br />
the time <strong>of</strong> sampling and <strong>of</strong> refinements in<br />
sampling and analytfc techniques.<br />
Furthermore, the load is not distributed<br />
uniformly from one reach to anotherr a<br />
fact which accounts for the variable<br />
transport rates,<br />
Under storm conditions there is a<br />
marked increase in the suspended load <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> River. <strong>The</strong> suspended sed-<br />
iment load is directly related to wind<br />
velocities and duration over southern <strong>Lake</strong><br />
Huron. During periods <strong>of</strong> northerly wfnds<br />
the suspend load is decreased. River<br />
velocities are competent to transport<br />
material comprised <strong>of</strong> the beach sediment<br />
from southern <strong>Lake</strong> Huron. Sediment<br />
deposits in the nearshore zone <strong>of</strong> <strong>Lake</strong><br />
Huron have a mean diameter <strong>of</strong> 0.125 mm to<br />
0.25 mm; and according to Duane (19671,<br />
most clastic sediments suspended in the<br />
<strong>St</strong>. <strong>Clair</strong> Delta have mean diameters <strong>of</strong><br />
slightly finer sizes. Particle size data<br />
suggest that the sediments <strong>of</strong> the delta<br />
are derived from reworked nearshore<br />
sediments in <strong>Lake</strong> Huron.<br />
<strong>The</strong> soil types <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
wetlands ref 1 ect the late Quaternary<br />
history <strong>of</strong> the regfon and may be divided<br />
into two broad categor'les. <strong>The</strong> soils <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> Delta proper are derived<br />
from sources adjacent to <strong>Lake</strong> Huron<br />
whereas the soil types along the eastern<br />
shore <strong>of</strong> the 1 ake are derived from lake<br />
pl a? n sediments i n southwestern Ontario.<br />
As such the delta soils general 1 y have a<br />
coarser texture than the soils <strong>of</strong> eastern<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Table $5. Sediment discharge rates and characteristics in the <strong>St</strong>. <strong>Clair</strong><br />
River.<br />
Transfer rate Location Load type Sediment<br />
grade<br />
54,700 1n3/~ear Shore <strong>of</strong> <strong>Lake</strong> Huron at Tract ion Sand<br />
head <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River<br />
61,600 d/year <strong>St</strong>. <strong>Clair</strong> River at Bay Point Suspended Sf1 t-cl ay<br />
Light (after storm)<br />
37,500 m3/& <strong>St</strong>. <strong>Clair</strong> River bottom 12 km Traction Cl ay<br />
between <strong>St</strong>. <strong>Clair</strong> 8 Marine<br />
City (after severe storm)<br />
40,100 <strong>St</strong>. <strong>Clair</strong> River Channel Tract ion ---<br />
wall between <strong>St</strong>. Cl a3 r<br />
8 Marine City<br />
19,900 d/year <strong>St</strong>. <strong>Clair</strong> River bank between Traction ---<br />
<strong>St</strong>. <strong>Clair</strong> & Marine City<br />
15,100 d/&y Surface and bottom <strong>of</strong> Chenal Total Clay<br />
A Bout Rond, 3 km below North<br />
Channel (after a storm)<br />
15,200 m3/year Entire <strong>St</strong>. <strong>Clair</strong> River Total Sand<br />
a~ata sources: Cole ( 1903 1 r Duane ( 19671, Pezzetta ( 1968).
<strong>The</strong> soi.78 <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta were<br />
derived from the shoreline and nearshore<br />
zone <strong>of</strong> <strong>Lake</strong> Huron. <strong>The</strong> sediment was<br />
transported,by the <strong>St</strong>. <strong>Clair</strong> River and now<br />
forms the delta proper. Two s<strong>of</strong>l types<br />
are evident in the delta. At the apex <strong>of</strong><br />
the landform, 1.5 to 2 m above mean lake<br />
level, Sanilac loam (Col wood fine sandy<br />
loam in Ontario) is prevalent. This soil<br />
is a very fine sandy loam and occurs on<br />
moderate (0-8) slopes. <strong>The</strong> soil formed<br />
in limey, water-laid sediment <strong>of</strong> very fine<br />
sandy loam and loamy very fine sand (U.S.<br />
Department <strong>of</strong> Agriculture 1974). A1 though<br />
the surface layer is dark-brown (0-15 cm),<br />
sand pits reveal that the deeper soil is<br />
ye1 lowish-brown (Munsell color code = 10<br />
YR 5/61, a fact suggesting slight<br />
oxidation. This soil type represents a<br />
premodern delta that was formed at higher<br />
lake levels in the geologic past.<br />
<strong>The</strong> wetlands proper are characterized<br />
by the Bach loam ("marshw soils in<br />
Ontario). <strong>The</strong> sol1 has a very fine sand<br />
Y., 1<br />
loam texture which farmed in limey<br />
lacustrine sediments (Figure 11).<br />
Occurring on land with a slope <strong>of</strong> 2% or<br />
less, the soil is <strong>of</strong>ten water l ogged, In<br />
a typlcal pr<strong>of</strong>ile the surface layer is<br />
black* calcareous, very fine sand loam, 18<br />
cm thick. <strong>The</strong> subsoil is 75 cm th9ckv and<br />
is made up <strong>of</strong> several layers <strong>of</strong> gray,<br />
calcareous, very friable* very fine sand<br />
loam. It represents the modern delta<br />
front, local depresslons~ and ancestral<br />
channels <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> dlstributaries.<br />
<strong>The</strong> Clyde loam is a poorly drained<br />
loam occupying the eastern shoreline <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. It is characterized by 0<br />
to 20 cm <strong>of</strong> very fine, very s<strong>of</strong>t, very<br />
dark gray silt-cl ay with large quantities<br />
<strong>of</strong> organic debris. From 20 to 30 cm deep.<br />
Clyde loam is dark gray, fine, firm silt<br />
and clay (Mudroch 1981). This s<strong>of</strong>l type<br />
developed on older, more f i ne-grained<br />
sediments <strong>of</strong> a lake plain and i s therefore<br />
finer and generally has a higher organic<br />
content than the delta soils.<br />
6rrvsmad Jvlm 26, 1977<br />
re.1 *.t.r,<br />
I -- 1 I 1 1 1<br />
r-"<br />
100<br />
---<br />
400 600 800<br />
--- -T -,<br />
1000 I200<br />
1<br />
1400 1600<br />
I<br />
1800 ZOO0 2200 2100 2600 2800 IOOOFI<br />
0 :a0 ZOO 100<br />
100 100 600 I00 800 900 1<br />
tread R(4p.<br />
$and and Peat alack<br />
Srrvr#ad Ocl 7, 1977<br />
' 5-7-<br />
Ebst<br />
115.5<br />
115.0<br />
171,s<br />
-----I a<br />
3000 3200 1100 1600 3804 4040 4200 1400 4600 4800 5000 5200 5400 5600 5100 6000fl<br />
-- I-- -<br />
I--<br />
(st*) 1000 i:5t IZJO lion ICOG !fn@ !do0 1300 1800 8<br />
Figure 11. Vegetation transect on Dickinson Island, <strong>St</strong>. <strong>Clair</strong> Delta, showing character<br />
<strong>of</strong> wetland soils (Jaworski et a7. 1981).<br />
22<br />
116 0<br />
115 1<br />
11s 0<br />
li4 5
<strong>The</strong> soils in the <strong>Lake</strong> <strong>St</strong>. Clafr<br />
region are basically derived from<br />
unconsolidated sediments deposited over<br />
bedrock in early Holocene time (1 .ear past<br />
20,000 years). <strong>The</strong>se s<strong>of</strong>ls contribute to<br />
the geochemical composition <strong>of</strong> bottom<br />
sediments <strong>of</strong> the wetlands. Typically, the<br />
wetland sediments investigated by Mudroch<br />
f 1981) are submerged mineral soils, with a<br />
pH ranging between 6.9 and 7.2. <strong>The</strong><br />
wet1 and sediments from the Walpole Island<br />
sector <strong>of</strong> the delta are coarser: more than<br />
half <strong>of</strong> the particles are at least sand<br />
size whereas the eastern shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> silt size particles are dominant.<br />
<strong>The</strong> absence <strong>of</strong> extensive peat or<br />
organic rich soils is noteworthy. Peat<br />
accumu1ations are poor and were probably<br />
formed in place rather than transported<br />
into the wetlands. Deltaic wetlands are<br />
<strong>of</strong>ten characterized by blanket peat<br />
deposits especially in sheltered local-<br />
ities such as bays and backbarrler flats.<br />
In the Great <strong>Lake</strong>s coastal wetlands, peats<br />
are with few exceptions (e.g.$ western<br />
Saginaw Bay) uncommon. This suggests<br />
several possibil itles: 1) wetlands during<br />
recent geologlc time were not abundant, or<br />
2) peat deposlts decomposed rapidly Or9 3)<br />
water exchange between the wetlands and<br />
lake was efficient in exporting organic<br />
material. In any caser clastic sediments<br />
are dominant in the wetland soils <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>.<br />
2.2 CLIMATE AND WEATHER<br />
<strong>The</strong> variable weather elements such as<br />
windr precipitation, air temperature,<br />
humidity* and growing season influence the<br />
physical and biological character <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. A unfque aspect <strong>of</strong> this water<br />
body Is rapid water level change due to<br />
seiches or wind tides. Also; spring ice<br />
jams can significant1 y a1 t er the water<br />
level <strong>of</strong> the lake.<br />
<strong>The</strong> climatic variables such as<br />
temperature, wind, atmospheric pressure,<br />
and precipitatfon play a stgnjf icant role<br />
in the terrestrial and lacustrfne<br />
environment <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r. <strong>The</strong>se<br />
elements are primarily responsible for the<br />
water budget <strong>of</strong> the lake as well as wave<br />
activityt the extent <strong>of</strong> the growing season<br />
in the wetlands, the distribution and<br />
length <strong>of</strong> ice cover, and the water level<br />
conditions on a daily, monthly, and<br />
seasonal bas 1 s.<br />
<strong>The</strong> climate <strong>of</strong> the lake i s<br />
characterized by cold wlnters (coldest<br />
month below 0% and warm summers (warmest<br />
month above 220C) with precipitation<br />
uniformly distributed through the year<br />
(climate type Daf based on the Koppen<br />
system). Cyclonic storms occur through<br />
the yearr but decrease in frequency durfng<br />
late June, Julys and August. Durlng this<br />
time convectional up1 ift is frequent and<br />
thunderstorms are common. Most <strong>of</strong> the<br />
moisture that falls on the Great <strong>Lake</strong>s<br />
basin as precipitation originally<br />
evaporated from the tropical At1 antic<br />
Ocean or the Gulf <strong>of</strong> Mexico (Phil1 ips and<br />
McCulloch 1972). On an average day, 10<br />
billion m3 <strong>of</strong> water in the form <strong>of</strong> water<br />
vapor are advected across the Great <strong>Lake</strong>s<br />
basin. <strong>The</strong> efficiency <strong>of</strong> precipitation<br />
mechanisms is such that only about 15% or<br />
1.4 billion m3 fa11 as rain on a given<br />
day.<br />
<strong>The</strong> frost-free season is defined as<br />
the interval in days between the last<br />
occurrence <strong>of</strong> frost in spring and the<br />
first occurrence in fall and is thus an<br />
indicator <strong>of</strong> the length <strong>of</strong> the growing<br />
season. <strong>The</strong> mean annual frost-free period<br />
for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is 160 days. With the<br />
exception <strong>of</strong> the south shore <strong>of</strong> <strong>Lake</strong> Erie,<br />
<strong>Lake</strong> Ontario* and the south shore <strong>of</strong> <strong>Lake</strong><br />
Michigan; which experience 180 f rost-f ree<br />
days annually, <strong>Lake</strong> <strong>St</strong>. Clafr has the<br />
longest frost-free period in the Great<br />
<strong>Lake</strong>s basin. <strong>The</strong> spring cooling exerted<br />
by the lake prevents premature growth <strong>of</strong><br />
the vegetation thus lessening the chances<br />
<strong>of</strong> crop loss due to late spring frosts<br />
(Eichenlaub 1979). Conversely, the slow<br />
cooling in autumn retards the occurrence<br />
<strong>of</strong> the first fall frost thus extending the<br />
growing season.<br />
C1 osel y re7 ated to the length <strong>of</strong> the<br />
frost-free season is the accumulation <strong>of</strong><br />
growing degree days as an index <strong>of</strong> the<br />
amount <strong>of</strong> heat available d~ring the<br />
growing season. <strong>The</strong> index is usually<br />
defined as the number <strong>of</strong> degrees <strong>of</strong> mean<br />
daily temperature above a base <strong>of</strong> 5,SoC.
<strong>The</strong> growing degree-day concept has been<br />
applied to delimit areas suitable for<br />
particular vegetation types and as a means<br />
<strong>of</strong> predicting the hatching date <strong>of</strong> various<br />
insects (Baker and <strong>St</strong>rub 1965). On the<br />
basis <strong>of</strong> using mean monthly temperature<br />
and the number <strong>of</strong> days in a month, the<br />
accumulated number <strong>of</strong> degrees above a base<br />
<strong>of</strong> 5.60C in a normal year is 2,340 OC in<br />
the southern portion <strong>of</strong> take <strong>St</strong>. <strong>Clair</strong> and<br />
29200 OC in the northern sector <strong>of</strong> the<br />
1 ake.<br />
Table 7 reveals that the mean annual<br />
average air temperature for three selected<br />
stations is 9.4OC. Mean monthly<br />
temperatures in January and February are<br />
well below freezing (-3.50C) but summer<br />
means exceed 220C. Such continental<br />
conditions commonly occur in regions which<br />
are remote from maritime air masses.<br />
One <strong>of</strong> the characteristics <strong>of</strong> the<br />
climate <strong>of</strong> the Great <strong>Lake</strong>s is the lack <strong>of</strong><br />
any marked seasonal i t y <strong>of</strong> prec i p i t at i on.<br />
Mean monthl y precipitation values for<br />
three stations (Table 8) average 77.83 cm.<br />
It should be noted that the Detroit and<br />
Windsor stations are located in highly<br />
urbanized areas compared to the Mount<br />
Clemens station, As noted on the table*<br />
the precipitation is 1 owest at Mount<br />
Clemens. Research in urban precipitation<br />
patterns indicates that the presence <strong>of</strong> a<br />
1 arge city such as Detroit does cause an<br />
increase in precipitation in the urban<br />
areas (Sanderson 1980). <strong>The</strong>refore9 Mt.<br />
Clemens i s more representative <strong>of</strong><br />
temperature conditions over <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>.<br />
Wind direction and frequency are<br />
significant with regard to waves, ice jams<br />
and short term (i.e., seiches) water level<br />
changes in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. According to<br />
Ayres (19641, the circulation in <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> is strongly influenced by prevail ing<br />
wind direction as are longshore currents.<br />
Figures 12 and 13 illustrate the average<br />
monthl y di rectfonal frequency <strong>of</strong> winds for<br />
Windsor and Mt. Clemens respectively. <strong>The</strong><br />
more spherical wind roses (e.g. March,<br />
April) have the greatest variabil ity in<br />
wind direction. <strong>The</strong> more skewed wind<br />
Table 7. a Mean monthly ternperatures for three selected<br />
stations.<br />
Mount Cl er,~ens, Detrolt City W i ndsor<br />
Michigan Airport Airport<br />
1940- 1969 1940-1969 1941-1970<br />
Month OF OC OF OC OF OC<br />
January<br />
February<br />
March<br />
Aprll<br />
May<br />
June<br />
July<br />
August<br />
September<br />
October<br />
November<br />
December<br />
Annual average 48.3 9.0 50.1 10.1 48.6 9.2<br />
a<br />
Data sources: Sanderson (1980); U.S. Dept. <strong>of</strong> Commerce, Weather<br />
Service.<br />
24
Table 9. a Mean monthly precipitation for three selected<br />
stations.<br />
Mount Cl emens Detroit C<strong>St</strong>y Windsor<br />
Michigan Airport Airport<br />
- -<br />
Month inches cm inches cm inches cm<br />
January<br />
February<br />
March<br />
April<br />
May<br />
June<br />
July<br />
August<br />
September<br />
October<br />
November<br />
December<br />
Annual average 28.07 71.29 30.95 78.60 32.91 83.59<br />
a<br />
Data sources: Sanderson (1980); U.S. Dept. <strong>of</strong> Commerce, Weather<br />
Service.<br />
roses (e.g., June, July, August) reveal a<br />
more prevailing wind. It is evident that<br />
predominant winds In the region are from<br />
the westerly direction# especially from<br />
the southwsst quadrant. Additional wind<br />
data for Windsor airport suggests that<br />
January, February, and March have the<br />
highest wind speeds averaging 20 km/hr<br />
while June, July, and August have the<br />
lowest at 13 km/hr. Furthermore, winds<br />
from the west-northwest have the highest<br />
speeds at 21 km/hrr and the lowesfwind<br />
speeds, 13 km/hro are from the southeast.<br />
Ice Cover<br />
A major physical feature on <strong>Lake</strong> <strong>St</strong>.<br />
Cl air is the occurrence <strong>of</strong> ice. Using 20<br />
years <strong>of</strong> datar Assel et a1 . (1983 1<br />
documented and mapped the distribution <strong>of</strong><br />
ice on the Great <strong>Lake</strong>s. <strong>The</strong> concentration<br />
<strong>of</strong> ice varies over the period <strong>of</strong> record<br />
mainly because <strong>of</strong> the shallowness <strong>of</strong> the<br />
lake. Although synoptic data on ice<br />
thickness especial 7y over the centrai<br />
portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> are poor#<br />
maximum thickness during severe winters is<br />
approximately 49 cm and maximum thickness<br />
during mild winters is about 5 cm,<br />
Normal 1y during mi d-December through<br />
February the 1 ake i s ice covered with the<br />
exception <strong>of</strong> Its exit--the Detroit River.<br />
During mild winters the only persistent<br />
ice cover in January and February occurs<br />
along the eastern shore1 ine <strong>of</strong> the 1 ake.<br />
However, during severe winters persistent<br />
ice will cover 100% <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from<br />
mid-December to mid-Apri 7.<br />
<strong>The</strong> impact <strong>of</strong> ice and ice jams on<br />
wet1 and vegetation is poorly understood,<br />
and can have negative as well as positSve<br />
effects, Ice jams migrating down<br />
distributary channels uproot aquatic and<br />
persistent vegetation from the substrate.<br />
Conversely, ice in the nearsh~re tone<br />
protects 1 acustrine wetlands by acting as<br />
a buffer and absorbing the impact from<br />
incoming waves. <strong>The</strong> coastal protectfon<br />
<strong>of</strong>fered by the ice In the nearshore zone<br />
is particularly significant during mild<br />
winters when most <strong>of</strong> the lake is not<br />
frozen and also durin~ the sprfng whe~<br />
storms frequently occur and wave energies<br />
are higher.
Jenuery February March , FEBRUARY<br />
/<br />
October I<br />
-- '$<br />
August<br />
November<br />
'i (/<br />
/ ' C 5<br />
June<br />
September<br />
December<br />
Figure 12. Monthly wi nd frequency diagrams<br />
for Windsor, Ontario, 1955-1972 (Sanderson<br />
1980).<br />
OCTOBER NOVEMBER DECEMBER<br />
Figure 13. Monthly wind frequency for Mt.<br />
Clemens, Michigan (Sel fridge Air Force<br />
Base) for period 1936-1953 (Ayers 1964).<br />
Note only velocities greater than 4 mph<br />
plotted; winds less than 4 mph shown as<br />
percent time.<br />
2.3 HYDROLOGY CirculatI~n in <strong>Lake</strong> <strong>St</strong>* Clajr<br />
n'storically water levels<br />
Of <strong>Lake</strong><br />
Investigations reveal that two major<br />
water masses occur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
C1air ranged apprOximate'y in Ayres (1964) modeled field data before and<br />
response changJng water budgets= Water after the construction <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
budgets In<br />
turn are to jnf l Ow and Cut<strong>of</strong>f Channel and found that the current<br />
Outflow as as evaporation and structure in the lake rnainta.lned semi-<br />
precipitation* <strong>The</strong> impact Of changing permanent patterns. <strong>The</strong> distributfon<br />
water levels within the basin On the pattern was altered according to changing<br />
shorelJne has been wind patterns, Leach (1980) also<br />
signi jcante HistorScal identified two discrete water masses in<br />
displacements have been documented with the 1 ake on the basis <strong>of</strong> cluster analysis<br />
the use <strong>of</strong> aerial photography over time. <strong>of</strong> physical and chemical data,<br />
Wetland vegetation transects and<br />
geographical dlstributions over time have <strong>The</strong> main flow <strong>of</strong> water entering <strong>Lake</strong><br />
a? so been determined . <strong>St</strong>. <strong>Clair</strong> is from the <strong>St</strong>. <strong>Clair</strong> River.
Thfs river contrfbutes about 98% <strong>of</strong> the<br />
total Inflow t o the lake. <strong>The</strong><br />
constructfon~ <strong>of</strong> the <strong>St</strong>. Lawrence Seaway<br />
and the opening <strong>of</strong> the South Channel<br />
Cut<strong>of</strong>f i n 1962 have increased the<br />
proportfon <strong>of</strong> <strong>St</strong>, <strong>Clair</strong> River water<br />
arriving directly into the main lake<br />
basin. <strong>The</strong> greater cross-sectional area<br />
<strong>of</strong> the Cut<strong>of</strong>f Channel for outflow which<br />
now exists and the straightening that the<br />
arti f ici a1 channel provides have decreased<br />
the frlctlonal effects and promoted a<br />
greater discharge <strong>of</strong> the st.-<strong>Clair</strong> River<br />
water directly into the main basin,<br />
In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> the extent and<br />
geometry <strong>of</strong> the ci rculatfng water masses<br />
are related to wind direction. <strong>The</strong><br />
predominant winds in the region are from<br />
the southwest quadrant (Sanderson 1980).<br />
Figure 14-D illustrates the two principal<br />
patterns in the eastern and western<br />
portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> basin derived<br />
from prevailing southwest winds. Figure<br />
14-A revsaxs a more djstinctive<br />
circulatory pattern associated with a<br />
north wind, Based upon Ayresl (1964)<br />
model, water in Anchor Bay originates from<br />
two distinct sources. Wl th southwest<br />
w'inds there are both a net diminution <strong>of</strong><br />
outflow through North Channel and the<br />
creation <strong>of</strong> a discrete gyre in Anchor Bay<br />
composed <strong>of</strong> Clinton River water. With a<br />
north wind, though, Anchor Bay becomes<br />
dominated by water from the North Channel<br />
<strong>of</strong> the <strong>St</strong>. Claf r River as the Clinton<br />
River water is reduced in the bay.<br />
<strong>The</strong> significance <strong>of</strong> the two main<br />
water masses is their relationship to the<br />
productivity <strong>of</strong> the lake. <strong>The</strong> water mass<br />
adjacent to Ontario is more influenced by<br />
nutrient load1 ngs f rom the Sydenham and<br />
Thames rivers and the smaller tritrutaries<br />
from Ontario. <strong>The</strong>se waterways drain an<br />
intensive1 y cultivated lake plain. Urban<br />
development on the south shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> also contributes to the loading.<br />
Because this water mass is more enriched<br />
and more stable than the mass associated<br />
with <strong>St</strong>. <strong>Clair</strong> River water, It has yielded<br />
greater productivity (Leach 1972, 1973 1.<br />
Due in part to the flushing time <strong>of</strong><br />
lake <strong>St</strong>. <strong>Clair</strong>, which is theoretically<br />
about 9 days, the fish communities have<br />
changed 1 fttle, with the exception <strong>of</strong> some<br />
col dwater species. According to Johnston<br />
(19771. the walleye and the yellow perch<br />
stocks over the past century have remained<br />
reasonably stable in spite <strong>of</strong> more<br />
intensive agricultural practices in the<br />
watershed, increased settlement in the<br />
coastal zoner and exploitatio~ <strong>of</strong> the<br />
resource from commercial f isheries. <strong>The</strong><br />
flushing action <strong>of</strong> relatively clean water<br />
from <strong>Lake</strong> Huron has prevented concentratlon<br />
<strong>of</strong> nutrients and eutrophication in<br />
most <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />
ater Discharae Into l ake <strong>St</strong>. <strong>Clair</strong><br />
In addltlon to the principal source<br />
<strong>of</strong> water from the <strong>St</strong>. <strong>Clair</strong> River,<br />
several other river basins contribute<br />
water to the lakes including the Blackr<br />
Belle9 Clinton* Thames, and Sydenham<br />
Rivers (Figure 15). <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River<br />
is not a true fluvial system but rather a<br />
strait connecting two large water bodies<br />
(<strong>Lake</strong> Huron and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>). It<br />
therefore does not exhibit typical river<br />
discharge characteristics. Furthermore,<br />
depositional processes associated with<br />
rivers are normally directly linked to<br />
extreme flow regjmes. But <strong>St</strong>. Clafr River<br />
flows and ultimate sediment distribution<br />
in the <strong>St</strong>. <strong>Clair</strong> Delta are more closely<br />
linked to ice conditions or high lake<br />
level condftfons.<br />
Of the total average fa11 <strong>of</strong> 2.5 m<br />
from the <strong>Lake</strong> Huron level to <strong>Lake</strong> Eriet<br />
1.5 m occur in the <strong>St</strong>. <strong>Clair</strong> River<br />
(Korkigian 1963). Its banks and bed have<br />
been relatively stable except for<br />
navigation improvementsr and sand and<br />
grave1 dredging over several decades.<br />
Figure 16 illustrates mean monthly<br />
discharge from 1900 through 1980 (Quinn<br />
and Kelley 1983). Average monthly<br />
discharge is 5,121 &/see and the range <strong>of</strong><br />
discharge is 4,254 m3/sec (February) to<br />
5,483 m3/sec (September). Detailed<br />
monthly data are included in Appendix 8,<br />
With flow velocities up to 3.2 km/hr the<br />
travel time through the system is<br />
re1 atively high, As determined by Dereckl<br />
(1983)~ the flow time <strong>of</strong> water from the<br />
international bridge at Port Huron to <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> is 21,1 hours (Table 9). <strong>The</strong><br />
flow time through the Detroit River (Belle<br />
Isle south to Geleron IsSandE fs 20.9<br />
hours, <strong>The</strong> initial travel timetable was<br />
developed for normal flow condltfons.
Based on the geomorphologic history South channels, which in turn each spllt<br />
Of the <strong>St</strong>* C1aBr Delta? the active before entering <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>e <strong>The</strong> flow<br />
portion <strong>of</strong> the landform at the present<br />
time 1s to the west. As the <strong>St</strong>. <strong>Clair</strong> is therefore proportional 1 y distributed<br />
River enters the Sk. <strong>Clair</strong> basin f t through the channels as illustrated In<br />
bifurcates near Algonac Jnto the North and Figure 17,<br />
. . . * . WATER LEFT LI PRLVIOUJ NlHb<br />
NORTH WINO<br />
-. cn~nlt EtA*Te (I JW**fOn CHI)IXEL<br />
-' ST CLSIR RIVER<br />
IU- * CLINIOW nivcn<br />
- * -WATER LLFY EY PREVIOU5 WlWU<br />
-----, - ~~lijvu~hci: OUTFLOW P ISCIPCOEHI<br />
WEST WINO C<br />
-' CUENhL fCI\RTE 4 JOHNSTON CIllNntL<br />
I--".<br />
ST. CLIlR RIVER<br />
I-- CLIITOR RIVER<br />
WATER LEFT aY PUEVIOUJ WINO<br />
NORTHWEST WIND<br />
--<br />
CIIENIL CCARTE 8 JOMNSTOW CHANNEL<br />
------L. ST. CLllR RIVER<br />
-+r - CLINTOM RIVER<br />
- ........ -L . NITER LEFT BY PREVIOUS WIND<br />
-----r . SUISURFACE CSCAPIYLnT<br />
SOUTHWEST WIND D<br />
Figure 14. Current Patterns for <strong>Lake</strong> <strong>St</strong>. Clafr under various wind conditions (Ayers<br />
1964).
Artificial modif icatlon In the past and 1925# and uncompensated navigation<br />
85 years has altered river levels and the improvements for the 7.7-m and 8.3-rn<br />
water volume i n <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. projects completed in 1933 and 1962<br />
Art.fficia1 channel changes 'In the <strong>St</strong>, respective1 y (Derecki 1982). <strong>The</strong>se<br />
<strong>Clair</strong> River s'lnce 1900 include dredging channel changes increased the discharge<br />
for commercial gravel removal between 1908 through the <strong>St</strong>. Cl air River and caused a<br />
---L. CHENAL LCARTE O JOHNBTOH CHANNEL<br />
-----* - 51. CLAIR RIVER<br />
--.CZ. CLINTON RIVER<br />
. , . , . , , . . . r WhTER LEFT BY PREVIOUS WIND<br />
SOUTH WlND<br />
--C. CHENAL CCARTe 6 JOUNSTON CHANNEL<br />
-----r- 1 ST. CLAIR RIVER ---u. CLINTON RIVER<br />
....... . W.PTER LEFT BY PREVIOUS WINO<br />
EAST WlND<br />
G<br />
E<br />
Figure 14. (Continued)<br />
29<br />
- ST.<br />
-----H.<br />
-r CRENAL ECARTE 6 JOHNLTON CHANNEL<br />
CLAlR WVER<br />
CLINTON RIVER<br />
WATER LEFT BY PREVIOU8 WlND<br />
-----+ BUSSURFACE OUTFLOW<br />
SOUTHEAST WlND<br />
--<br />
CHENAL LCARTE 6 JOHNSTOH CtiAhlNEL<br />
-- ST CLAIR RIVER<br />
--M r CLfNTOH RIVER<br />
- *=WATER LEFT BY PREVIOUS WIND<br />
NORTHEAST WIND H
Figure 15. <strong>Lake</strong> <strong>St</strong>. Cl ai r-<strong>St</strong>. Clai r River drainage basin show-<br />
ing location <strong>of</strong> precipitation and water level gages.<br />
Table 9.<br />
veloci$y<br />
<strong>Clair</strong>.<br />
<strong>St</strong>. <strong>Clair</strong> River flow time and<br />
from <strong>Lake</strong> Huron to <strong>Lake</strong> <strong>St</strong>.<br />
<strong>St</strong>atlon Tf me Distance Velocity<br />
(hr) (km) (km/hr)<br />
<strong>Lake</strong> Huron 0.0<br />
Blue Water Brldge 0.1<br />
Black River 0.9<br />
<strong>St</strong>ag Island 4.4<br />
<strong>St</strong>. <strong>Clair</strong> City 6.0<br />
Marine City 9.8<br />
Roberts Landing 12.0<br />
Algonac Lt, 12,5<br />
<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f 17.9<br />
J F M A M J J A S Q N O 1.1<br />
<strong>Lake</strong> <strong>St</strong>, Gfair 21.1 60.5<br />
Figure 16. Mean monthly discharge sf the<br />
<strong>St</strong>, <strong>Clair</strong> River 1900-1988. a~ata source: Derecki (1983).
Figure 17. Location map <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta distributary channels<br />
showing average discharge (cubic meters per second) and flow distri-<br />
bution (percent <strong>of</strong> total ) .
permanent lowering <strong>of</strong> the lake's level.<br />
<strong>The</strong> impact <strong>of</strong> these channel changes<br />
resulted in the l ~ ~ ~ r <strong>of</strong> i nthe g <strong>Lake</strong><br />
Michigan and <strong>Lake</strong> Huron water levels by<br />
0.27 m. Thfs depth superf mposed on the<br />
combined areas <strong>of</strong> <strong>Lake</strong>s Mfchlgan and Huron<br />
represents a permanent water loss <strong>of</strong> 32<br />
km3 or nine times greater than the volume<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air.<br />
<strong>Lake</strong> level changes and thef r duration<br />
play an important role in the character <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, its shoreline and its<br />
wetland connnunities. In general r high and<br />
stable water levels are most beneficial to<br />
the fish stock <strong>of</strong> the lake whereas low and<br />
stable or unstable conditions are least<br />
desf rable from an ecological standpoint.<br />
Converse1 y hf gher water levels when<br />
combined w l th storm events encourage<br />
higher flood f requencles and coastal<br />
erosion. <strong>The</strong> wetland communities <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>, Clalr respond to longer term water<br />
changes and hence adjust to such changes<br />
over time.<br />
All the Great <strong>Lake</strong>s includfng <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> experience changes in water<br />
levels due to short-term and long-term<br />
weather and cl imat ic conditions. Short-<br />
term changes are due to steep barometric<br />
gradients which may alter water levels for<br />
a period <strong>of</strong> several hours. During the<br />
winter months ice jams on the <strong>St</strong>. Claf r<br />
River cause abrupt water level changes<br />
whlch may span a period <strong>of</strong> 60 days. Long-<br />
term oscillations are related to water<br />
lnput and water output to and from <strong>Lake</strong><br />
<strong>St</strong>, <strong>Clair</strong>. Such water level changes,<br />
al though non-cycl lcr occur over a period<br />
<strong>of</strong> several years. Pre-1946 storm surges<br />
on <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and its connecting<br />
channels have been documented by Murty and<br />
Polavarapu (197.5 1 armd are summarlzed in<br />
Tab1 e 10. As noted* the water levels at<br />
Tecumsshp Ontario dropped 0,5 m and rose<br />
again over a perfod Qf approximately 48<br />
hours. Short-term depressions <strong>of</strong> water<br />
levels due to storm surges provide some<br />
flushing action and the fntroduction <strong>of</strong><br />
nutrfents fraorn the marshes fnto the bays<br />
and nearshore waters <strong>of</strong> the lake.<br />
However, the event is probably Loo short-<br />
1 ived to have signif posltfve impact,<br />
Table 118, Record storm surge levels <strong>of</strong><br />
Canadian stations o$ <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />
connecting channel s.<br />
Maximum<br />
Date <strong>of</strong> Observat l on surge<br />
storm stat i on (m) (ft)<br />
November1913 Fighting Islandb 0.48 1.56<br />
Isle Aux Pechesc 0.28 0.91<br />
October 1916 Fighting 1s1 andb 0.71 2.34<br />
Isle Aux Pechesc 0.51 1.67<br />
November 1940 ~ecumsehd 0.54 1.76<br />
a~ata source: Murty and Pol avarapu (19751,<br />
b~ower Detroit River.<br />
'upper Detroit River.<br />
d~ake <strong>St</strong>. <strong>Clair</strong>.<br />
Ice jams are a common occurrence on<br />
the <strong>St</strong>. <strong>Clair</strong> River and <strong>of</strong>ten detafn<br />
Interlake shipping. Ice blockades usually<br />
form in February and March in the <strong>St</strong>.<br />
<strong>Clair</strong> River and occasionally extend into<br />
April. On the average, ice retards the<br />
flow by 16%. Ice-jamming reduces the<br />
outflow from <strong>Lake</strong> Huron and raises its<br />
level and at the same time reduces inflow<br />
to <strong>Lake</strong> <strong>St</strong>. Clai r and lowers its level.<br />
During the winters <strong>of</strong> 1904-1905 and 1905-<br />
1906, average inflow from <strong>Lake</strong> Huron to<br />
<strong>Lake</strong> Erie decreased 1,369 m3/sec over 4<br />
months and 904 m3/sec over 5 months,<br />
respective1 y (Bajorunas 1968).<br />
During the spring <strong>of</strong> 1984, strong<br />
winds from the east and north caused a<br />
massf ve fce jam south <strong>of</strong> <strong>Lake</strong> Huron.<br />
Brash ice clogged the 62.5 km-course <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> River until the end <strong>of</strong><br />
April, and jams <strong>of</strong> 3 m or more in depth<br />
developed at the apex <strong>of</strong> the delta. This<br />
situation led to a decrease in discharge<br />
from the monthly average <strong>of</strong> 5,096 m3/sec<br />
for April, to about 2,50Om3/sec below the<br />
projected mean volume for that month.<br />
Also there was a subsequent drop fn the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> water level by more than<br />
0.76 m from March 21 to April 19 t 175.40<br />
to 174.59 m above mean sea level). In the<br />
closing days <strong>of</strong> Aprtl a convoy <strong>of</strong> Canadian<br />
and Amerfcan ice breakers, alded by<br />
unseasonably mild temperatures broke the<br />
ice jam at the upper end <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong>
Delta allowing 70 vessels anchored In the Although non-cyclicr the water levels<br />
Detrolt River and western <strong>Lake</strong> Erje to appear to have a high water period and a<br />
enter the upper Great <strong>Lake</strong>s. low water period every 7 to 10 years.<br />
Figure 18 records the impact <strong>of</strong> this<br />
ice jam in the <strong>St</strong>, Clafr River on the<br />
water level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Water<br />
level began its drop in mid-March 1984 and<br />
did not recover until the end <strong>of</strong> May 1984.<br />
<strong>The</strong> change (March 15-April 15) in the<br />
water level was 0.48 m. Ice jams<br />
temporarily a1 ter the hydrology <strong>of</strong> the <strong>St</strong>,<br />
Cl af r Delta, As the distributary channels<br />
are jammed, river water is diffused<br />
through crevasses into the bays (e.g.,<br />
Muscamoot and Goose Bays), This process<br />
is responsible for subaqueous<br />
sedimentation in the bays and the<br />
subsequent colonFzatfon by emergent<br />
wet1 ands.<br />
February 1984 was relatively mild and<br />
<strong>Lake</strong> Huron was largely ice freer but<br />
exceptionally cold weather followed and an<br />
ice cover formed over the entire lake by<br />
about March 10th (Great <strong>Lake</strong>s Commfssion<br />
1984). By the outset <strong>of</strong> April, <strong>Lake</strong> Huron<br />
again was nearly Ice free except at the<br />
head <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River. Unusually<br />
strong and persistent westerly winds<br />
reduced the movement <strong>of</strong> drifting ice down<br />
the river by blowing much <strong>of</strong> it from the<br />
south end <strong>of</strong> <strong>Lake</strong> Huron eastward.<br />
Following this event? winds from the east<br />
and north forced the jumbled ice into the<br />
<strong>St</strong>. <strong>Clair</strong> River over a 3-week period.<br />
Water level data for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
have been continuously recorded since<br />
1899. <strong>The</strong> long-term level <strong>of</strong> the 1 ake is<br />
determined by the inputs <strong>of</strong> water derived<br />
from run<strong>of</strong>f or inflow <strong>of</strong> connecting<br />
channels and tributaries to the lake, and<br />
precipitation over the lake basin. <strong>The</strong><br />
outputs are due to evaporatfon and outflow<br />
via the Detroit River. Long-term lake<br />
level oscfllations are usually thought <strong>of</strong><br />
as volumetric changes which are<br />
cl imatically controlled. Changes in 1 ake<br />
storage may be sumnarlzed as follows:<br />
Water voTume and lake level <strong>of</strong> a<br />
given 1 ake bast n are dependent upon water<br />
inflow and 0utf10wr which in turn is<br />
determined by factors such as overwater<br />
precipitation, evaporationr and run<strong>of</strong>f<br />
fnto and out <strong>of</strong> the basin. <strong>The</strong>se factors<br />
are interrelated and may be expressed as a<br />
transfer factor. <strong>The</strong> transfer factor is<br />
defined as the sum <strong>of</strong> the monthly<br />
precip itati on and run<strong>of</strong>f minus the<br />
evaporatfon and change in storage. QuInn<br />
( 1976 1 computed transfer factors for <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> (Table 11). <strong>The</strong> run<strong>of</strong>f from the<br />
land into <strong>Lake</strong> <strong>St</strong>. Clai r was determined<br />
independently for the four river basins<br />
draining fnto the lake. <strong>The</strong>y are the<br />
Be1 1 e, Cl i nton, Thames, and Sydenham<br />
Basins (Figure 15). On an annual basis<br />
and during most <strong>of</strong> the year, the transfer<br />
factor is most sensitive to run<strong>of</strong>f. <strong>The</strong><br />
evaporation however, becomes the most<br />
important factor during the high<br />
evaporation months <strong>of</strong> August to October.<br />
Figure 18 records the water level<br />
history <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r for 1900 to<br />
1984. <strong>The</strong> lake has a mean monthly<br />
elevation <strong>of</strong> 174.87 m above sea level at<br />
Father P<strong>of</strong>nt, Quebec in the Gulf <strong>of</strong> <strong>St</strong>*<br />
Lawrence. However, as the figure<br />
demonstrates, the water level in recent<br />
years has been we1 1 above this 1 eve1 .<br />
Table 12 indicates that the lake has had a<br />
record high <strong>of</strong> 175,64 m (June 1973) and a<br />
record low <strong>of</strong> 173.71 m or a range <strong>of</strong> 1.93<br />
m. Low water or chart datum is 174,25 m.<br />
This level is a fixed reference plane<br />
selected by the United <strong>St</strong>ates and Canada<br />
so that the majority <strong>of</strong> time during the<br />
navigation season the Great <strong>Lake</strong>s actual<br />
levels w i l l be above that plane. Water<br />
levels are lowest in February; based on a<br />
1900-1983 mean, the level for this month<br />
i s 174.47 m. HIghest water levels Bccur<br />
in July; the 1900-1983 average for thi s<br />
month is 174.94 m.<br />
S = (P-E) + (1-0) <strong>Lake</strong> level fluctuations occur<br />
naturally on <strong>Lake</strong> <strong>St</strong>. CSair and are the<br />
where: S = change in storage volume result <strong>of</strong> prolonged periods <strong>of</strong> precip-<br />
P = precipitation itation, drought* storm-related events, or<br />
E = evaporation ice conditions, <strong>The</strong> lake level depends on<br />
I = inflow the balance between the total water<br />
0 = outflow supplied to the lake from precipftation on
LEGEND<br />
I 1<br />
LAKE LEVELS<br />
Hydrographs are In feet above (+)<br />
or below (-) Chart Datum, the plane<br />
-'<br />
REGORDED on each lake to whrch navigation<br />
chart depths and Federal navigatton<br />
ImprOvement depths are re-<br />
PROBABLE<br />
ferred.<br />
1900-1 983<br />
Chart Datum and all other eleva-<br />
AVERAGE<br />
tions sire in feet above mean water<br />
level at Father Point, Quebec (In-<br />
MAXIMUM m,E 1973 ,973 ternattonai Great <strong>Lake</strong>s Datum - 1955). To convert feet to meters,<br />
MINIMUM 1936 1934 1926 multiply feet by o 30480.<br />
-<br />
Figure 18. Hydrograph <strong>of</strong> iake <strong>St</strong>, <strong>Clair</strong> and <strong>Lake</strong> Erie water levels, showing<br />
the impact <strong>of</strong> a record ice jam in the <strong>St</strong>. <strong>Clair</strong> Delta in March and April 1984<br />
(U,S, Army Corps <strong>of</strong> Engineers), Note the slight dip in iake Erie water levels<br />
in May in response to the <strong>St</strong>. Clai r Be1 ta ice jam.
Table 11. take <strong>St</strong>, <strong>Clair</strong> average monthly water transfer factor parameters. a<br />
Cubic &rs oar ntglrth<br />
Parameter J F M A M J J A S O N D<br />
Precipitatfon 23 20 25 34 25 34 31 31 25 23 28 28<br />
Tributary run<strong>of</strong>f 156 190 342 277 136 62 37 28 25 42 79 153<br />
<strong>Lake</strong> evaporation 25 25 11 8 48 37 34 45 59 51 42 20<br />
Change in storage -76 42 79 42 17 17 -6 -17 -37 -48 -25 14<br />
a Data source: Quinn (1976).<br />
Table 12. Water level data for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. a<br />
Parameter Meters Feet Remarks<br />
Low water datum 174.25 571.70<br />
Month1 y water levels<br />
Average 174.87 573.72 1900-1983 data base<br />
Record high 175.64 576.25 June 1973<br />
Record low 173.71 569.90 January 1936<br />
a~ata source: U.S. Army Corps <strong>of</strong> Engineers.<br />
the lake and run<strong>of</strong>f from the surrounding<br />
drainage area and the amount <strong>of</strong> water<br />
discharged from the Detroit River or by<br />
evaporation. <strong>The</strong> relationship between<br />
long-term changes in lake level and the<br />
climatic events which cause the changes is<br />
usually quite evident, even though there<br />
is generally a time lag in the response <strong>of</strong><br />
1 ake levels to imbalances in the system.<br />
<strong>The</strong> high levels <strong>of</strong> 1973 on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
as well as the other Great <strong>Lake</strong>s were<br />
attributed to a 16% increase In precip-<br />
itation and a 24% decrease in evaporation<br />
during 1972 compared to the long-term<br />
average for both precipitation and<br />
evaporat I on.<br />
2.4 WATER QUALITY<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> waters are usually<br />
we11 mSxed by both horizontal and vert-ical<br />
currents. <strong>The</strong> two major factors affectfng<br />
current patterns and water quality are the<br />
flow-through current <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
River (Figure 19) and the seasonal water<br />
temperature changes, Variations in water<br />
density, largely caused by changes in the<br />
water temperature and amount <strong>of</strong> suspended<br />
sol ids, and winds affect the 1 ateral<br />
movement at the lake surface, while tur-<br />
bulence created by storms, and rec-<br />
reational and commercfal boatfng actlv-<br />
ities affect the vertical movement (U.S.<br />
Army Corps <strong>of</strong> Engineers 1974).<br />
Water quality informatfon fs presented<br />
here f n terms <strong>of</strong> nutrlents and<br />
contaminants which either enhance or 1 imit<br />
wetland productivlty. Of particular concern<br />
are phosphorusr nltrogenr df ssol ved<br />
oxygen, alkal lnftyt pH, major fans, trace<br />
elementst and toxic organic compounds.<br />
Tharmaf structure, ! fght penetration" and<br />
sediment load are also addressed f n terms<br />
<strong>of</strong> Impacts to wetland stab4llty and<br />
habftats,
Figure 19. Distribut.ion <strong>of</strong> specific conductance in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
(Bricker et al. 1976). Note low values from the <strong>St</strong>. <strong>Clair</strong> River and<br />
high values at the mouths <strong>of</strong> the Thames River and Clinton River.<br />
In March 1970 the Canadlan government<br />
announced that 5,500 kg <strong>of</strong> commercially<br />
caught walleye I L !LjAB@)<br />
from <strong>Lake</strong> <strong>St</strong>. Cl afr were destroyed because<br />
<strong>of</strong> mercury contamination. In the early<br />
19705s mercury concentrations in fish in<br />
<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> ranged from 0.3 ppm to over<br />
5 ppm, the higher values were usually<br />
found in larger fish and in predacious<br />
species. <strong>The</strong> United <strong>St</strong>ates and Canada<br />
adopted a concentration <strong>of</strong> 0.5 ppm in fish<br />
muscle as a maximum for commercial fish,<br />
Both governments closed commerci a1 fishing<br />
in the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>,<br />
as well as walleye fishing in western <strong>Lake</strong><br />
Erie,<br />
<strong>The</strong> mercury pollution was largely<br />
attributed to waste discharges from chlor-<br />
alkali processing plants on the <strong>St</strong>. <strong>Clair</strong><br />
and Detroit Rivers which began operation<br />
in the 1950s. Several <strong>of</strong> these plants<br />
ceased operation in the early 1970s and as<br />
a result, mercury contamination has<br />
diminished in the environment. Levels <strong>of</strong><br />
total mercury In walleye collected in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> have decl jned from a mean <strong>of</strong><br />
over 2 ppm in 1970 to 0.5 ppm in 1980. In<br />
western <strong>Lake</strong> Erie, 1968 levels <strong>of</strong> mercury<br />
were 0.8 ppm as compared to only 0.3 pprn<br />
in 1976. <strong>The</strong> rapid environmental response<br />
subsequent to the cessation <strong>of</strong> major point<br />
source d fscharges at Sarnia, Ontario, and
at Wyandotte, Mfchlgan* can be attributed<br />
to rapid flushing by the <strong>St</strong>. <strong>Clair</strong>-Detrolt<br />
Rivers system and rapid burial <strong>of</strong><br />
contaminated sediments by the high load <strong>of</strong><br />
suspended sediment delivered to western<br />
<strong>Lake</strong> Erie.<br />
Few comprehensive water quality<br />
investigations have been conducted on <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and measurements in the coastal<br />
wetlands are rare. <strong>The</strong> Michigan Water<br />
Resources Commission (1975) conducted a<br />
thorough study <strong>of</strong> the open lake and bays<br />
along the American shore in 1973 (Table<br />
13 1 and Leach (1972, 1973, 1980) reported<br />
on conditions in CanadIan waters (Figure<br />
20). <strong>The</strong> U.S. Envl ronmental Protection<br />
Agency operates a national water qua1 i ty<br />
data storage and retrieval system<br />
(STORET). Pub1 ished and unpubl i shed <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> data from several Federal,<br />
state, provincial, and local agencies have<br />
been entered into this STORET for the<br />
period 1967-1982. Table 14 represents a<br />
retrieved STORET summary (mean values) for<br />
22 parameters measured in the <strong>St</strong>. Cl air<br />
Rfver, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Detroit River, and<br />
western <strong>Lake</strong> Erie during this period.<br />
Mudroch and Capobianco (1978) determfned<br />
the geochemical composition <strong>of</strong> the<br />
surface sediments (0 to 20 cm) and the<br />
overlying rnarshwater in Big Point Marsh on<br />
the Ontario shore <strong>of</strong> <strong>Lake</strong>-<strong>St</strong>. ClaOr<br />
(Tables 15 and 16). Mineralogically the<br />
sed irnents are composed <strong>of</strong> ill ite, quartz,<br />
calclte, dolomite, K-fe1 dspars,<br />
pl agi ocl ose, and mi nor amounts <strong>of</strong> chl ori te<br />
and kaol inite. <strong>The</strong> water can be<br />
characterized as hard with high levels <strong>of</strong><br />
nutrients. <strong>The</strong> pH <strong>of</strong> the marsh sediment<br />
is about 7.0, while the overlying water<br />
ranges between 8.1 and 8.4.<br />
Two important features which<br />
determine the water temperature <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> are the lake's shallowness and<br />
the high flow rate from <strong>Lake</strong> Huron.<br />
Because <strong>of</strong> the shallow depth <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>, it warms quickly in the spring and<br />
cools rapidly In the fall. <strong>The</strong> lake is<br />
too shallow to stratify thermally;<br />
therefore the water is almost always<br />
isothermal from surface to bottom, <strong>The</strong><br />
high inflow from the <strong>St</strong>. Clalr River<br />
Table 13. <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> water quality measurements for 1973 growing season, a<br />
Parameter<br />
-lake--<br />
July Aug Sept July Aug Sept July Aug Sept July Aug Sept<br />
Secchi depth (rn) 1.9 1.6 2.2 2.0 1.3 1.7 2.0 1.4 2.3 0.6 0.9 1.2<br />
Conductlvlty(umohs/cm) 184 203 202 181 213 205 183 214 210 340 273 274<br />
Temperature OC 19.1 22.3 21.2 19.4 23.5 22.0 20.2 22.9 21.9 23.3 24.6 21.1<br />
pH (units) 8.6 8.6 8.8 8.5 8.7 8.8 8.6 8.6 8.7 8.5 8.8 8.5<br />
DIssolvedoxygen(mg/l) 8.1 8.2 7.2 8.3 8.7 7.7 6.9 8.0 7.6 8.5 7.9 7.1<br />
Suspendedsollds (mg/l) 7.3 11.6 1.8 4.0 12.0 15.0 4.3 6.9 12.0 35.7 11.7 26.3<br />
Dlssolved solids (mg/lf 127 129 -- 127 120 -- 127 124 - 218 153 --<br />
Alkalfnlty (mg/l) 80 81 90 78 80 88 80 80 90 114 90 108<br />
Hardness (mg/l) -- -- 95 -- -- 95 -- - 96 -- -- 119<br />
Sodium (t~~/l) 5.1 6.1 5.2 5.1 5.5 5.3 5.3 5.6 5.1 15.8 9.7 12.7<br />
Magnesium (mg/l) 7.8 -- 8.0 8.0 - 7.9 8.0 - 8.0 12.1 -- 9.8<br />
Calclurn (rng/l) -- 26.1 24.9 -- 26.0 25.0 -- 26.9 25.3 -- 31.0 31.7<br />
Potassfurn (mg/l) -- -- 0.8 -- -- 0.9 -- -- 0.8 -- -". 1.6<br />
Sulfate (mg/l) -- 18.8 17.7 -- 17.5 17.5 -- 18.7 17.9 -- 19.1 22.9<br />
Phosphorus, total (ug/l) 29 60 15
Figure 20. Mean water quality characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from April<br />
through November 1971 (Leach 1972). Note increased concentrations near the<br />
southeast shore in the vicinity <strong>of</strong> the Thdmes River mouth.
Table 14. Chemical and physical characgeristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />
adjacent waters (mean record 1967-1982).<br />
Parameter<br />
Temperature OC<br />
Di ssolv%fed oxygen mg/l<br />
D,O, saturation %<br />
Conductivity (250C) umohs/cm<br />
Df ssol ved so1 f ds mg/ 1<br />
Suspended sol ids mg/l<br />
Secchi depth m<br />
Alkalinity mg/l<br />
pH<br />
Calcium, total<br />
units<br />
mg/l<br />
Magnes i um, total mg/ 1<br />
Potassi um, total mg/l<br />
Sod 1 urn, total mg/l<br />
Chloride, total mg/l<br />
Sulfate, total mg/l<br />
Fl uori de, total mg/l<br />
Silica, dissolved ug/1<br />
Ammonia, dissolved<br />
Nitrate + Nitrite, diss.<br />
ug/l<br />
ug/l<br />
Phosphorus, total ug/l<br />
Phosphorusr dissolved ~g/l<br />
Chlorophyll p ug/l<br />
<strong>St</strong>. ~lairb <strong>Lake</strong> Detrolt Western<br />
Uni t s River <strong>St</strong>. <strong>Clair</strong> River <strong>Lake</strong> Erie<br />
a ~ ta a source: U.S. Environmental Protection Agency, Large <strong>Lake</strong>s<br />
<strong>St</strong>ation, Grosse Ile, Michigan, STORET Data System.<br />
b~eadings do not reflect mean concentration at the <strong>St</strong>. <strong>Clair</strong> River due<br />
to location <strong>of</strong> sampling stations near major outfall s.<br />
(approxlmatel y 5,000 m3/sec) quick1 y<br />
rep1 aces the water in the lake with water<br />
from <strong>Lake</strong> Huron (in about 9 days). Thus,<br />
the temperature <strong>of</strong> Laks Huron has a strong<br />
influence on the temperature <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>. This is especially true in Anchor<br />
Bay and along the Michigan shore because<br />
most <strong>of</strong> the water from the <strong>St</strong>. <strong>Clair</strong> River<br />
enters the lake via the North Channel<br />
(33%) a t the east side <strong>of</strong> the bay.<br />
Ljke <strong>Lake</strong> Erie, highest temperatures<br />
are reached in August. On the Ontarto<br />
shore, water temperatures average about<br />
240C in July and August. On the western<br />
side <strong>of</strong> the lake, which is influenced more<br />
by the cool water from <strong>Lake</strong> Hurcn,<br />
temperatures are usually 2 to 40C lower.<br />
Temperatures are highest in the shallow<br />
coastal wetlands, reaching over 28OC, <strong>The</strong><br />
typfcal summer thermocl ine <strong>of</strong> deeper 1 akes<br />
in the Great <strong>Lake</strong>s system is non-exlstent<br />
in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />
In the fa1 1, the 1 ake Is warmest on<br />
the western sfde* a reversal <strong>of</strong> the summer<br />
pattern. Water is coolest in the shallows<br />
near shore and warmest near the mouth <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> River. Normally <strong>Lake</strong> <strong>St</strong>,<br />
<strong>Clair</strong> Ss almost completely ice covered by<br />
mid-January. Ice break-up usually occurs<br />
in early March.<br />
Mid-summer water transparency (Secchf<br />
dfsc) In <strong>Lake</strong> <strong>St</strong>. @lair normally ranges<br />
from 0,7 m along the Michigan shore to 2,6
Table 15. Geochemical composition <strong>of</strong><br />
sediments in B+g Point Marsh, Ontario, on<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Parameter Concentration ( ppm)<br />
Sf02<br />
A1 203<br />
Fe2~3<br />
CaO (%I<br />
Na2o<br />
K20<br />
Ti02<br />
MnO<br />
'205<br />
C r<br />
Ni<br />
Cd<br />
Co<br />
Cu<br />
Organic C (%I<br />
63.7<br />
93.7<br />
%ata source: Mudroch and Capobianco<br />
(19781,<br />
km near the center <strong>of</strong> the lake to 0.6 m<br />
along the Ontario shore (Herdendorf 1970).<br />
In the nearshare reglons the water color<br />
is brownish-green. Water in the vfcinfty<br />
<strong>of</strong> the navigation channel Is a d$stincttva<br />
cloudy-green but near the center <strong>of</strong> She<br />
lake away from the channel is a cfear-<br />
green.<br />
Table 16. Chemical composition <strong>of</strong> waterd<br />
Big Point Marsh, Ontario, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Parameter<br />
Ammonia-Nitrogen<br />
Nitrate-Nitrogen<br />
Nitrogen, Kj<br />
Phosphorus, total<br />
Potass 1 urn<br />
Cal cl um<br />
Magnes f urn<br />
Sod i um<br />
Alkalinity, total<br />
Lead<br />
Zinc<br />
Chromi um<br />
Copper<br />
Cadmi um<br />
Min. Max,<br />
(mg/l) (mg/l)<br />
a Data source: Mudroch and Capobianco<br />
(1978).<br />
<strong>The</strong> minimum intensity <strong>of</strong> subsurface<br />
light that permits photosynthesls has been<br />
set at 1.0% <strong>of</strong> incident surface light<br />
(Cole 1983). <strong>The</strong> zone <strong>of</strong> a lake from the<br />
surface to a depth at which 99% o f the<br />
surface light has been absorbed is known<br />
as the euphotic zone. Below this depth,<br />
primary productivity is considered n i 1.<br />
<strong>The</strong> relationship between Secchi disc<br />
transparency and the thickness <strong>of</strong> the<br />
euphotic zone provides a simple way to<br />
estimate the photosynthetically active<br />
region <strong>of</strong> a lake. A ratio <strong>of</strong> 3.0 to 3.5<br />
has been empirically determined for this<br />
relationship, yielding a thickness <strong>of</strong> the<br />
euphotfc zone rangfng from about 2 m along<br />
the shores to over 9 rn near the center <strong>of</strong><br />
the lake,<br />
Nitrogen* phosphorus, and sil ica are<br />
the primary lfmfting nutrients in <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>. Bicarbonate alkalintty is in such
great abundance (>80 mg/l) that a source<br />
carbon for primary productivity is never<br />
lackfng. Relatively high nitrate (>400<br />
vg/1It phosphorus (>ZOO ug/l) and silica<br />
(>2 mg/7) in the nearshore waters <strong>of</strong> the<br />
wetland bays (Table 13) reflect high<br />
watershed inputs, Leach (1972) also found<br />
high nutrient loading from the Thames<br />
River (Table 17). <strong>The</strong> mid-lake lower<br />
concentrations reflect low nutrient<br />
loading from <strong>Lake</strong> Huron water. <strong>The</strong> mean<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> concentrations <strong>of</strong> nutrients<br />
lie between values for tha upper Great<br />
<strong>Lake</strong>s and those <strong>of</strong> western <strong>Lake</strong> Erie<br />
(Schelske and Roth 197318 yielding a<br />
mesotrophic classification for this lake.<br />
On the basis <strong>of</strong> water chemistry,<br />
Leach (1980) identified two distinct water<br />
masses i n <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>: 1) a<br />
northwestern mass consisting primari 1 y <strong>of</strong><br />
<strong>Lake</strong> Huron water flowing from the main<br />
channels <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River and 2) a<br />
southeastern mass <strong>of</strong> more stable water<br />
enriched by nutrient loadings from Ontarlo<br />
tributaries and shoreline urban devel-<br />
opment. <strong>The</strong> margins <strong>of</strong> the water masses<br />
shift according to the wind di rectf on and<br />
characteristics <strong>of</strong> the two water masses<br />
for the period 1971-1975 is shown in<br />
Figure 21.<br />
Because thermal stratification does<br />
not occur* due to the lake's shallowness<br />
and rapid flow <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River,<br />
dissolved oxygen concentrations are<br />
usually at saturation. In Bradley Marsh,<br />
north <strong>of</strong> the Thames River, Mudroch and<br />
Capobianco (19781 found that the<br />
temperature and dissolved oxygen<br />
saturation in marshwater during the<br />
growlng season are inversely related<br />
(Figure 22), but the 0, saturatian did not<br />
fa1 l below 75%. <strong>The</strong> Lcoastal marshes are<br />
shallow and well mixed by wind, and<br />
consequent1 y the water temperature Is<br />
affected di rectly by air temperature. <strong>The</strong><br />
diurnal temperature changes are 1 arge8 for<br />
example in July 1976, they found<br />
marshwater temperature at 0600 hours to be<br />
20°C and at 1500 hours* 27%. <strong>The</strong>se<br />
changes coupled with wind maintain high 0<br />
levels In the marshwater during thg<br />
summer.<br />
speed, but the-overall discreetness <strong>of</strong> the<br />
distribution 1s maintained during the ice-<br />
free periods <strong>of</strong> the year ( April-December) . <strong>The</strong> Michigan Water Resources<br />
A compression <strong>of</strong> the water quality Commission (1975) reported that<br />
Table 17. Comparison <strong>of</strong> cJemica1 characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> and associated waters.<br />
De1 t a <strong>Lake</strong> Thames<br />
Parameter (units) Channels <strong>St</strong>. <strong>Clair</strong> River<br />
Chlorophyll 3 (vg/l) 1.2<br />
Pheopigments (ug/ll 0.4<br />
Part. org. carbon (mg/1) 0.7<br />
Sol. react. phosphate (pg/1) 0.6<br />
Total phosphate (vg/l) 2.5<br />
Nitrate (pg/l) 17 8<br />
Nitrite (vg/l) 1.6<br />
Sol. react. silicate (mg/l) 0.3<br />
Total a1 kal tnity (mg/l) 87<br />
Sul fate (mg/l) 15.3<br />
Chloride (mg/l) 8.1<br />
'Flat3 source: Leach ( 19721,
sooi<br />
1 ". . .." . . ---- - - .<br />
' CHLOROPHYLL 5? (ug/L) 1<br />
Figure 21. Characteristics <strong>of</strong> <strong>Lake</strong> Huron<br />
water mass (A) and Ontario nearshore water<br />
mass (B) in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from 1971 to<br />
1975 (Leach 1980). Solid line is mean<br />
water qua1 i ty for each water mass and<br />
dashed line is prediction based on a<br />
central station in each water mass.<br />
chlortnated hydrocarbon concentrations in<br />
sedfments are low throughout the lake.<br />
Lindane, aldrin, endrin, heptachlor<br />
(detectfon limits 0.001 mg/kg)r and<br />
chlorodane (detectton 1 imi t 0.005 mg/kg)<br />
concentrations were below detection, and<br />
dfeldrfn (detection limlt 0.001 mg/kg) was<br />
only detected at a few stations, then at<br />
maximum levels <strong>of</strong> only 0.006 rng/kg, DOE*<br />
TDE, and DDT (detection 1 Smit 0.002 mg/kgl<br />
were below detection at most stations, but<br />
high concentrations (0.039 mg/kg) were<br />
found In L'Anse Creuse Bay near the<br />
Clinton Rfver splllway. PCB (detection<br />
limits 0.05 mg/kg) levels were below<br />
detectfon in most nearshore areas, but<br />
concentrations ranged from 0.50 to 1.10<br />
mg/kg near the splllway, indicating that<br />
the Cl inton River is an important source<br />
<strong>of</strong> PCBfs.<br />
#<br />
o Temperature<br />
Apr ~ 6 y , I .lul Aug Sep 1976<br />
Figure 22. Temperature and dissolved<br />
oxygen relationship in a <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
coastal wetland, near the mouth <strong>of</strong> the<br />
Thames River, Ontario (Mudroch and<br />
Capobianco 1978).
3.1 PLANKTON<br />
Phvto~l Sessf 1 e A-<br />
Planktonic and sessfle algae,<br />
although not as important as macrophytes<br />
in the coastal marshes9 are signif icant<br />
primary producers <strong>of</strong> organfc matter in<br />
wetlands, converting the sun's energy into<br />
chemical compounds that in turn are used<br />
as food by animals and nonphotosynthetic<br />
mtcro-organisms. In the nearshore and<br />
open waters <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, however9<br />
alga productivity dominates. Phyto-<br />
plankton production and distribution are<br />
fnfluenced by sun1 ight, temperature,<br />
nutrients, morphometry, water movementsr<br />
grazing by zooplankton, and other factors.<br />
Many inorganic elements are required for<br />
algal cell growthp incl uding nitrogens<br />
phosphorus, potassium, calcium, and iron.<br />
Algae reproduce rapidly when phosphorus is<br />
added to the water9 and continue to<br />
reproduce as more phosphorus is added.<br />
However, nitrogen and other nutrients must<br />
also be present if algal production is to<br />
contlnue, Recycling <strong>of</strong> nutrients within a<br />
marsh may be sufficient to promote algal<br />
blooms for several years after the initial<br />
loadfng (Sawyer 1954).<br />
Pieters (1893) was the first botanist<br />
to describe the algae <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r<br />
largely from the marshes <strong>of</strong> Anchor Bay.<br />
He noted that over 60 species <strong>of</strong> green<br />
algae, largely desmids, were associated<br />
with the vascular aquatlc plants <strong>of</strong> the<br />
coastal wetlands. W i nner et ale (19701<br />
and the Michigan Water Resources<br />
Commission (1975) have made more recent<br />
studies <strong>of</strong> phytoplankton populations in<br />
Ontario and Michigan nearshore waters<br />
respectively. <strong>The</strong> results <strong>of</strong> these in-<br />
vestigations are summarized in Appendix C.<br />
Diatoms are the most abundant forms <strong>of</strong><br />
pl anktonlc algae in the surface waters <strong>of</strong><br />
the lake. r mr and<br />
are important genera throughout<br />
the water column. Filamentous forms such<br />
as -r are common where sol id substrate<br />
is available, whjle Sp$roqaa is<br />
found in open waters <strong>of</strong> the marshes.<br />
<strong>The</strong> distribution <strong>of</strong> the algal<br />
pigment, chlorophyll gr in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
can be used as an indicator <strong>of</strong> the<br />
re1 ati ve abundance <strong>of</strong> phytoplankton.<br />
Leach (1972) found that starting at the<br />
delta channels, the concentrations <strong>of</strong><br />
chlorophyll g and pheopi gments increase<br />
toward the southeast (Figure 231, reaching<br />
the highest levels (20 pg/l) fn the<br />
marshes at the mouth <strong>of</strong> the Thames River.<br />
Figure 23. Concentration gradient <strong>of</strong><br />
algal pfgments across <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> from<br />
the <strong>St</strong>. <strong>Clair</strong> Delta to the mouth <strong>of</strong> the<br />
Thames River (Leach 1372).
<strong>The</strong> strongest positfve correlat~on was<br />
Found during the spring plankton blooms.<br />
SIl ica also has a strong relationship<br />
to the phytoplankton composftion. De-<br />
pletion <strong>of</strong> sil lca toward the end <strong>of</strong> the<br />
summer greatly reduces the percentage <strong>of</strong><br />
df atoms in the phytoplankton community.<br />
In the southern portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>,<br />
diatom concentration falls from 95% <strong>of</strong> the<br />
total phytoplankton population in May to<br />
10% <strong>of</strong> the population in July. Con-<br />
verse1 y, cyanophytas (b1 ue-greens) i n-<br />
crease from 0% <strong>of</strong> the total population in<br />
May up to 25% <strong>of</strong> the population in July<br />
(Leach 1972).<br />
<strong>The</strong> term periphyton generally refers<br />
to micr<strong>of</strong>loral growth, particuTar1 y<br />
sessile algae, on submerged substrate.<br />
Modifiers are normally used to indicate<br />
type <strong>of</strong> substrate: epipelic - growfng on<br />
sediment; epilithic - growing on rock;<br />
epiphytic - growing on macrophytes; and<br />
epfzooic - growing on animals (Wetzel<br />
1975). <strong>The</strong> periphyton <strong>of</strong> western <strong>Lake</strong><br />
Erie consists <strong>of</strong> predominant1 y 1 lttoral<br />
communitfe~r most commonly associated with<br />
wet1 and vegetation, A greater number <strong>of</strong><br />
species occurs in the littoral zone than<br />
in the llmnetlc zone <strong>of</strong> the lake due to<br />
the greater diversity <strong>of</strong> habitats<br />
available in the nearshore region,<br />
Although no detalled studies have been<br />
done in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlandsr<br />
Millie (1979) examined the epiphytlc<br />
diatom flora <strong>of</strong> aquatic macrophytes fn<br />
marshes along the south shore <strong>of</strong> western<br />
<strong>Lake</strong> Erie. Three common species <strong>of</strong><br />
wetland macrophytes were studied as hosts,<br />
narrow-leaved cattail (w -usti -<br />
white water lily (<br />
and swamp smartweed (<br />
a), 157 were new<br />
ds. Centrlc forms,<br />
were the most common taxa, but each marsh<br />
possessed a dlstfnct flora and<br />
suecessJona1 pattern. Millie attributed<br />
this heterogeneity to the diversity <strong>of</strong><br />
littoral habitats in the marshes*<br />
particularly differences in chemical and<br />
physical factors, Because <strong>of</strong> the nearness<br />
and similarity <strong>of</strong> the <strong>Lake</strong> Erie marshes to<br />
the wetlands along the Ontario shore <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. Claj r, comparable peri phyton<br />
communlties would be expected for both<br />
1 akes,<br />
In Lawrence <strong>Lake</strong>, Michi ganr A1 1 en<br />
( 1971) measured the annual net production<br />
<strong>of</strong> aquatic macrophytes and their attached<br />
algal forms. He found that the epiphytic<br />
algae were responsible for approxf matel y<br />
31% o f the total 1 ittoral production in<br />
the lake. Brock (19701, working in the<br />
Everglades, observed that epiphytf c algae,<br />
rather than macrophytes were responsible<br />
for the majority <strong>of</strong> the primary production<br />
<strong>of</strong> bl adderwort ( sp. 1 com-<br />
munities. Diatoms were found to be the<br />
most abundant form <strong>of</strong> 1 ittoral periphyton,<br />
both in number and biomass, In several<br />
Ontario 1 akes (<strong>St</strong>ockner and Armstrong<br />
1971). Likewiser it is anticipated that<br />
epiphyttc diatoms are an important<br />
component <strong>of</strong> the energy base in <strong>Lake</strong> <strong>St</strong>.<br />
Cl a1 r marshes.<br />
By definition, plankton are floating<br />
organisms whose movements are more or less<br />
dependent on currents. However# some<br />
zoopl ankters exhibit active swimming<br />
movements that aid in maintaining vertical<br />
posltion. Zoopl ankters are diverse in<br />
their feeding habits. Herbivores graze on<br />
phytoplankton, periphyton, and macrophytes<br />
within the coastal marshes. Carnivores<br />
prey on attached protozoans and other<br />
zooplankters, while omnivores feed at all<br />
trophlc levels. In turn, zooplankton are<br />
important fish and waterfowl food. Every<br />
fish species and many duck species<br />
urilizing the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands eat<br />
zooplankters during some portion <strong>of</strong> the1 r<br />
lffe cycle.<br />
<strong>The</strong> anlmal components <strong>of</strong> the wetland<br />
plankton (Figure 24) consist <strong>of</strong> three<br />
groups: protozoans, rotifers (Figure 251,<br />
and microcrustaceans (primarily<br />
cl adocerans and copepods) . Common zoo-<br />
pl ankters, including sessile or periphytic<br />
protozoans# occurring fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
wetlands are listed in Appendix D.<br />
<strong>The</strong> first study <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
zooplankton, and one <strong>of</strong> the earl lest<br />
1 imnol ogical investigations in the Great<br />
<strong>Lake</strong>s, was organized by Refghard (1894)<br />
and included work by many <strong>of</strong> the leading
Figure 24. Mean distribution and concentrations <strong>of</strong> zoo-<br />
plankton groups in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> for April through Novem-<br />
ber 1972 (Leach 1973). Note correlation between total<br />
zooplankton population and a1 gal pigments.
CARFA ENLARGED /<br />
Figure 25. Distribution <strong>of</strong> total roti fers and most abundant genus,<br />
Brachionus, in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> during July-September 1973 (Bricker et al.<br />
" m N b t e high concentration in the vicinity <strong>of</strong> the Clinton River Cut<strong>of</strong>f<br />
Channel.<br />
specfalists <strong>of</strong> that perlod. Species lists<br />
<strong>of</strong> Protozoa (Smfth 18941, planktonic<br />
Rstlfera (Jennings 18941s Gladocera (Bi rge<br />
1894)r and Copepoda [Marsh 1895) were<br />
compiled from samples collected among<br />
aquatic plants fn Anchor Bay and at other<br />
nearshore stations along the west shore <strong>of</strong><br />
take <strong>St</strong>, Clai r, Planktonic organ1 sms<br />
remained negl scted untll Winner et a1 .<br />
(2970) obtained zooplankton data from two<br />
estuary mouths on the Canadian shore.<br />
Leach ( 1973 ) col lected zooplankton samples<br />
from the South Channel <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
Daita* Mitchell Bay* the estuary mouth <strong>of</strong><br />
the Thames Rivsrr and other nearshore and<br />
<strong>of</strong>fshore stations on <strong>Lake</strong> <strong>St</strong>. Clalr.<br />
Brfcker et a7, (1976) documented the zoo-<br />
plankton populatfons <strong>of</strong> Anchor Bay, the<br />
nearshore region in the vicinity <strong>of</strong> the<br />
Clinton River estuary* and <strong>of</strong>fshore re-<br />
gions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> results <strong>of</strong><br />
these collections are included in Appendix<br />
0.<br />
<strong>The</strong> distribution <strong>of</strong> zooplankton in<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is largely dependent on the<br />
prevailing currents <strong>of</strong> the lake. Other<br />
inf l oences, such as temperature, algal<br />
abundance, and chemical characteristics<br />
a1 so determine the concentratlons <strong>of</strong><br />
zooplankton. Because these factors vary<br />
greatly throughout the lake, the<br />
distributions <strong>of</strong> zooplankton also vary<br />
consf derabl y in different portlons <strong>of</strong> the<br />
lake (Bricker et al. 1976). A persistent<br />
eddy structure In the southeastern portion
<strong>of</strong> the lake (Figure 141 promotes<br />
entrajnrnent <strong>of</strong> the waters and allows<br />
nutrient and algal levels to increase<br />
above other portions <strong>of</strong> the lake (Leach<br />
1972). Because <strong>of</strong> the high phytoplankton<br />
levels in this area, the zooplankton<br />
biomass is greater here than in any other<br />
regton, <strong>The</strong> rapid flow conditions <strong>of</strong><br />
water through the delta and the Michigan<br />
portion <strong>of</strong> the lake inhibits any great<br />
concentration <strong>of</strong> zooplankton; therefore,<br />
the biomass is greater on the Ontario side<br />
(Figure 24). Despite rapid flow rates,<br />
zooplankton biomass is elevated near the<br />
Cl lnton Rfver Cut<strong>of</strong>f Canal (Figure 25) due<br />
to nutrient loading from the watershed<br />
(Bricker et al, 1976).<br />
Because <strong>of</strong> the rapid flushing rate <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (9 days) and the dense<br />
subrnergent macrophyte populations competing<br />
for available nutrients, the total<br />
biomass <strong>of</strong> plankton in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is<br />
low. Zooplankton that succeed the best in<br />
this environment are those that have a<br />
short 1 ife span and a fast turnover rate.<br />
For that reason* rotlfers are the most<br />
abundant forms <strong>of</strong> zooplankton found. Mean<br />
numbers <strong>of</strong> rotifers ranged from<br />
approximately 62/1 in July and 69/1 In<br />
August down to 10/1 in September (Leach<br />
1973 1 . <strong>The</strong>se organisms are thirty times<br />
more abundant at the Clinton River Cut<strong>of</strong>f<br />
Canal (Bricker et a1. 1976). Crustaceans<br />
have a consistently low abundance in the<br />
lake ranging from a high <strong>of</strong> around 6/1 to<br />
a low <strong>of</strong> 0.3/1. As with rotifers, the<br />
peak abundance occurs in July and the low<br />
occurs in September. Cyclopoid and<br />
ca1 anoi d copepods, and cl adocerans (water<br />
fleas) are the predominant microcrustaceans<br />
found in the lake (Figure 24).<br />
Protozoans vary fn abundance but have the<br />
second highest occurrence following the<br />
rot1 fers. <strong>The</strong>y compose an average <strong>of</strong> 37%<br />
<strong>of</strong> the total zooplankton communlty.<br />
3.2 WETLAND VEGETATltON<br />
Although wetland communities exist in<br />
the <strong>St</strong>. <strong>Clair</strong> River (Figure 261, Clinton<br />
RJver, Marsac P<strong>of</strong>ntt Swan Creakp and <strong>St</strong>,<br />
<strong>Clair</strong> Delta complexes, only the latter<br />
area has been w'ell studied. <strong>The</strong>refore?<br />
the description <strong>of</strong> the pl ant communities<br />
Figure 26. Ontario shore1 ine; <strong>of</strong> <strong>St</strong>. <strong>Clair</strong><br />
River at Cathcart Park (August 1984),<br />
herein is based on the <strong>St</strong>, <strong>Clair</strong> Delta<br />
wet1 andsr particular7 y those on DJck f nson<br />
Is1 and. Important 1 iterat~re sources, by<br />
wetland area, are: <strong>St</strong>. John" Marsh<br />
(Roller 19761, Dfckinson Island (Jaworski<br />
et al. 1979, 19813, Canadian sfde <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> Delta (Eayly 1975)? and the<br />
enti re del ta complex ( Jaworski and Raphael<br />
1978; Raphael and Jaworskf 1982). Lyon<br />
(1979) mapped the wetlands communfties on<br />
the Michigan side <strong>of</strong> the <strong>St</strong>. C1 alr Delta<br />
using the wetlands classfficatfon <strong>of</strong> the<br />
U, S, Fish and Wildlife Service and a map<br />
scale <strong>of</strong> 1:24b000, Hayes (1964) surveyed<br />
the plant communities <strong>of</strong> a wet prairle on<br />
Harsens Is1 and,
Several distinct environmental types*<br />
or pl afit zones, have been identified<br />
wfthin the <strong>St</strong>, <strong>Clair</strong> Delta wetland complex<br />
located in northeastern <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
<strong>The</strong>se zones are part <strong>of</strong> a vegetation<br />
continuum (Figure 27). For example, on<br />
the 11.3 km2 Dickinson Island* the plant<br />
comrnuniPles grade from cattafl marsh along<br />
the southwest shore to upland hardwoods on<br />
the northeast point. <strong>Lake</strong> level<br />
fluctuations control the development and<br />
areal extent <strong>of</strong> the various zones<br />
(Jaworski and Raphael 1978). <strong>The</strong><br />
important aquatic macrophytes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> are 1 isted in Appendix Er and<br />
distribution maps <strong>of</strong> several species are<br />
presented In Figure 28,<br />
1.5 me Certain species, such as the<br />
cattails and the bulrushes can withstand<br />
moderate wave energy. Common vascular<br />
pl ants Include the following specfes:<br />
(eel grass,<br />
(pickerel weed)<br />
(yellow water Illy)<br />
water smartweed)<br />
(Eurasian<br />
(hybrid cattail)<br />
(hard-stem bul rush)<br />
(three-square<br />
(sago<br />
pondweed)<br />
Char9 sp. (muskgrass or stonewort).<br />
This type includes plant communities<br />
such as those present in Goose Bay,<br />
Gooseneck Pond <strong>of</strong> Fisher Bay, and Little<br />
Muscamoot Bay. Species are largely sub- LOW* narrow beaches <strong>of</strong> fine sand and<br />
merged and emergent aquatic macrophytes. stands <strong>of</strong> emergent vegetation (Ff gu r e 29)<br />
Habitats consist <strong>of</strong> low-wave energy lake are found on the Canadian sfde <strong>of</strong> the<br />
bottoms and nearshare environments <strong>of</strong> bays delta. <strong>The</strong> sand extends down less than 1<br />
where the water depths range from 0.3 to m and exhibits alternatqng layers <strong>of</strong> clean<br />
.3<br />
SW.<br />
July 1972 dlarcs~ooi Bay<br />
DOGWQOR<br />
MCAROW SE THREE<br />
E SQUARE BULRUSH<br />
-IY 0<br />
0<br />
5<br />
-5 +-- T-. r-- ?--I<br />
0<br />
ASPEN<br />
2 00<br />
V € r 20 V+(l.letbw notfc rtal*<br />
--r---7. - - - ,-<br />
8<br />
400 LOO 800 1W0 3600 3800<br />
"71-__7__7,-- T---- --<br />
FEET<br />
October 1974<br />
FEET<br />
I<br />
40W<br />
BULRUSH<br />
- --- 0<br />
---..<br />
---- 5<br />
1000 3600<br />
Figure 27. Vegetation transects on Harsens Island, <strong>St</strong>. <strong>Clair</strong> Delta, showing impact <strong>of</strong><br />
rising water levels (Jaworski and Raphael 1976).<br />
4%
MICHIGAN<br />
ANCHOR BAY<br />
PORT<br />
HUROiL<br />
MICHIGAN<br />
ANCHOR BAY<br />
ONTARIO<br />
/ ST. CLAlR S<br />
MICHIGAN<br />
PORT<br />
HURON<br />
Potarnogeton crispus Potanlogeton ri chardsoni i Butornus umbel 1 atus<br />
(curly pondweed) (Richardson's pondweed) (flowering rush)<br />
PORT HURsEU<br />
MICHIGAN hllCHlGAN MICHIGAN<br />
ANCHOR BAY<br />
Val 1 i sneria americana<br />
(wil d celery)<br />
ANCMOR BAY<br />
Heteranthera dubia<br />
(water stargrass)<br />
Myriophyl lum spicatum<br />
(water-mil foil )<br />
Figure 28. Distribution <strong>of</strong> six aquatic macrophytes in the <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>-Detroi t River ecosystem (Schloesser and Manny 1982, unpubl ished data supplied by<br />
R. L. <strong>St</strong>uckey).
Fiaura 29. Emeraent stand <strong>of</strong> reed qrass<br />
(fhragmi tes austbl is) a1 ong he ma tog an<br />
Channel, Wal pole Island, Ontario (August<br />
sand and organic-rich sand or peat.<br />
<strong>St</strong>randed or transgressive, beach ridges<br />
are low, sandy features on the delta<br />
Islands that can be up to 100 m wide.<br />
Normally these ridges are only slightly<br />
above the water level. Characterfstfc<br />
vegetation <strong>of</strong> these sandy habltats<br />
Includes the following vascular species:<br />
bindweed).<br />
( staghorn sumac)<br />
(touch-me-not,<br />
(swamp thistle)<br />
This marsh type is prevalent within<br />
abandoned channels and fn cattail marshes<br />
where open water and sandy sediments<br />
occur. Bulrushes are also common In<br />
shallow water along the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />
shore and in bays, as in Anchor Bay.<br />
Water depths range from 0.6 to 1.2 m.<br />
Representative plants include the<br />
followfng species:<br />
Scir~us acut~ (hard-stem bul rush)<br />
Ce~ha1 anthyg occidental I 5<br />
(buttonbush)<br />
Char3 sp. (muskgrass or stonewort)<br />
Various emergent and submergent<br />
species.<br />
Cattai 1 Marshes<br />
<strong>The</strong>se marshes are broad zones located<br />
on the lower portions <strong>of</strong> the delta islands<br />
where water depths exceed 15 cm. <strong>The</strong>y are<br />
also found on inundated shoulders <strong>of</strong> river<br />
channels and bottoms <strong>of</strong> shallow ponds and<br />
bays. Pure colonies <strong>of</strong> cattails (lyebq x<br />
are associated with peaty or<br />
clayey sediments (FJgure 30). In small,<br />
shall ow open i ngs in the cattall marshes,<br />
duckweeds (hmna minor and SDI rode1 q<br />
~glyrhlzg), water-mil foils (Nvrio~t).ylJm<br />
al_tecniflorum and d % ~icatum), and<br />
bl adderwort (Ib;triculariq. vulgari~) are<br />
abundant.<br />
Sedae Marshes<br />
Thfs mdian type comprises a narrow<br />
zone between the cattail marsh and the<br />
grassy meadow or the dogwood shrub zones.<br />
Sedge mavshes are also found along river<br />
channels* at the base <strong>of</strong> old shore1 ines<br />
now stranoed in cattail marshes, and along<br />
the edge <strong>of</strong> eroding lake and bay<br />
shorelines. <strong>The</strong>se marshes form on<br />
occasional 1 y flooded wet sites where<br />
permanent water depths do not exceed 15<br />
cm. Common species present include the<br />
fol lonring:<br />
ana ad ens is (bluejoint<br />
mex strictq (tussock sedge)<br />
!2E.S% (tussock sedge)<br />
!a% (tussock sedge)<br />
1 anuainosq (tussock sedge)<br />
Carex sartwell i I (tussock sedge)
Figure 30. Emergent stands <strong>of</strong> narrow-<br />
leaved cattail (Typha angustifolia) on<br />
Little Muscamoot Bay, Harsens Island,<br />
Michigan (August 1984).<br />
So1 a w<br />
gfffcinale$ (common comfrey)<br />
(nightshade).<br />
iN&&u@m<br />
<strong>The</strong>se habitats include artificial<br />
canals and openings in the marshes<br />
(ponds), Water depths are general 1 y<br />
shallow, 0.3 to 0.6 m, and sediments are<br />
usually clayey or organic. Communities<br />
can be quite variable. Typical species<br />
fnclude the following:<br />
ellow water lily)<br />
(white water lily)<br />
Lemna minor (duckweed)<br />
Wederi g 5;ordata (pickerel weed)<br />
,- ~anadens-ls (waterweed)<br />
mjgm (water smartweed)<br />
Potamoael;on 5:ris~uq (curly pondweed)<br />
Potamq&&Qn sp~. (oondweeds)<br />
Ce~hal anthus o~cid&ntal is<br />
(buttonbush)<br />
W sp. (muskgrass or stonewort)<br />
C1 ildo~hora sp. ( f il amentous green<br />
algae).<br />
Abandoned distributary channels in<br />
the delta are generally shallow, less than<br />
1 m deep, and have silty or peaty<br />
sediments. <strong>The</strong> variable plant communities<br />
include the following species:<br />
Nuohar advena (yellow water Illy)<br />
$uberosa (white water lily)<br />
M~rioptydJm<br />
--<br />
tr!lterniflor~ (little<br />
water-mll foil<br />
Wttaria latifolfa (common<br />
arrowhead)<br />
,cutus (hard-stem bulrush)<br />
Sci rpus americanus (three-square<br />
bulrush)<br />
(buttonbush).<br />
This type 4s a zone <strong>of</strong> transition<br />
vegetation or ecotone, between the sedge<br />
marshes and the hardwood community. This<br />
zone 1 fes sl ightly above the water table<br />
and is infrequently flooded. <strong>The</strong><br />
communities consist <strong>of</strong> a mixture <strong>of</strong><br />
grasses, herbs and shrubs, and water-<br />
tolerant trees, including the fallowing<br />
species:<br />
po~ul uz Qemuloide~ (quaking aspen)<br />
(red ash)<br />
d osier<br />
~alustrSs (swamp rose)<br />
Sol i &gg spp. (goldenrods)<br />
sp. [panic grass)<br />
(rattlesnake
Carey strictp (tussock sedge)<br />
wamp mi 1 kweed)<br />
ush)<br />
(marsh fern)<br />
sf 1 verweed) ,<br />
This type also represents a tran-<br />
sition into up1 and hardwood vegetatfon.<br />
<strong>The</strong> depth to the water table is normally<br />
0.5 to 1.0 m in this zone. Communities<br />
are mlxed shrubsr water-tolerant treesr<br />
and some understory plants typical <strong>of</strong> the<br />
meadows. Prominent plants include the<br />
following species:<br />
1- (eastern<br />
cottonwood<br />
PODU~Y+ tremu181das. (quaking aspen)<br />
Fraxiw Pennsvlrnnca (red ash)<br />
Gornus ~tolonifera (red osier<br />
dogwood<br />
Qxnuj racemosa (gray dogwood)<br />
oalmata (wlld grape)<br />
sp. (hawthorn).<br />
During low-water periods, these shrubs and<br />
swamp trees invade the sedge marshes.<br />
However, during high water condi tions, as<br />
fn the 1970~~ flooding and dle back <strong>of</strong><br />
woody plants in the shrub zone are common.<br />
<strong>St</strong>ands <strong>of</strong> oak-ash hardwoods occur in<br />
the upper portion <strong>of</strong> Dickinson, Harsens,<br />
<strong>St</strong>. Anne, Squirrel, and Walpole is1 ands at<br />
elevations ranging from 1 t o 3 m above<br />
mean lake level. <strong>The</strong> major species <strong>of</strong><br />
these stands are Eraxinus eann+~lvanica<br />
(red ash) and (swamp white<br />
oak), Assocfated hardwoods occurring In<br />
this type include the following species:<br />
American elm)<br />
<strong>The</strong> wetland plant communities <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> Delta occur in broad, arcuate<br />
zones which extend from the vlcinity <strong>of</strong><br />
Algonac* Mlchfgan, southward into <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> (Figure 311, In spfte <strong>of</strong> extensive<br />
cultural mod1 fications, sufficient natural<br />
wetland vegetation remains to reconstruct<br />
the communlty zonation, On the various<br />
islands <strong>of</strong> the delta, five major<br />
macrop hyte communities can be mapped.<br />
<strong>The</strong>se communitfest from the driest to the<br />
wettest habitats, include: oak-ash<br />
hardwoods, dogwood meadow9 sedge marsh,<br />
cattail marsh, and bulrush marsh. Other<br />
fdentlfiable communities which are 1 imited<br />
In extent include: canal-pond-abandoned<br />
channel, lakeshore and bay bottom8 and<br />
reed grass communities.<br />
<strong>Lake</strong>ward <strong>of</strong> the hardwoods i s a<br />
complex <strong>of</strong> communfties referred to as the<br />
dogwood meadow. This vegetation zone<br />
occurs as a transftion zone between the<br />
hardwood forest and the wetter sedge<br />
marsh. It appears that periodic high lake<br />
level and human disturbance (such as<br />
burning and mowing) prevent woody species<br />
from dominating this zone. Major plant<br />
species herein are bluejoint grass<br />
(ww -1, s<strong>of</strong>t rush<br />
(m gffusus) , gray dogwood, red os i er dogwood, and quakfng aspen.<br />
<strong>The</strong> sedge marsh forms a narrow zone<br />
between the dogwood meadow and the cattail<br />
marsh, as well as along eroding lake and<br />
bay shorel ines, particularly on the<br />
Michigan side <strong>of</strong> the delta. This<br />
community also occurs along lower dlstributary<br />
channels and on old shorel i nes.<br />
<strong>The</strong> sedge marsh is associated with the<br />
alkaline, inorganic soils which are either<br />
saturated or occasional ly flooded.<br />
However, permanent Inundation <strong>of</strong> the sedge<br />
tussocks results in dieback <strong>of</strong> the sedges.<br />
Major plant species are sedges <strong>of</strong> the<br />
genus Carex and bluejoint grass. Carex<br />
stricta is particularly abundant on<br />
Dickfnson Island, whereas L lacustri~ is<br />
common on Walpole Island,<br />
Located on the lakeward margin <strong>of</strong> the<br />
delta, where water depths exceed 15 to 30<br />
cm, is the broad cattail marsh zone.<br />
Zones <strong>of</strong> cattail also occur on inundated<br />
shoulders <strong>of</strong> the river channels, on lower<br />
ends <strong>of</strong> distributary channels, and on<br />
crevasses. Pure stands <strong>of</strong> cattail are<br />
assocfated with peaty or clayey sediments.<br />
<strong>The</strong> hybrid cattail (TyDha x glauc(a)r which<br />
attafns 2 to 3 m in height and grows in
ANCHOR BAY<br />
a OAK-ASH<br />
HARDWOOD<br />
DOGWOOD MEADOW<br />
0 DREDGED DISPOSAL<br />
0 CULTIVATED LAND<br />
Figure 31. Wetland plant communities <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta (Raphael and Jaworski<br />
1982).<br />
53
circular colonlesP is the domlnant<br />
species. Some narrow-leaved (L,<br />
and broad-1 eaved cattall s<br />
1 can also be identified.<br />
<strong>The</strong> bulrush marsh i s found in<br />
abandoned river channels, in 1 arge open-<br />
water areas <strong>of</strong> the cattafl marsh* and on<br />
sand bars in low-wave energy areas <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and Anchor Bay. Colonies <strong>of</strong><br />
bul rush are associated with sandy<br />
substrates and water depth <strong>of</strong> 0.5 to 1,25<br />
m. <strong>The</strong> prfncipal species <strong>of</strong> the marsh is<br />
hard-stem bulrush. Buttonbush i s<br />
associated with the bulrush community in<br />
abandoned distributary channels where some<br />
water flow occurs.<br />
In canals, abandoned river channel sr<br />
marsh open1 ngs, and other stagnant water<br />
environments, a variety <strong>of</strong> floating and<br />
submersed aquatic communities occur.<br />
Water depths range from 0.24 to 1.0 m and<br />
water circulation is minimal except for<br />
wind-driven currents. Organic substrates<br />
are common along with turbidity due to<br />
humlc acids and suspended particulate<br />
organic matter. Whereas species diversity<br />
fs the ruler abundant plant species<br />
include duckweedsp pickerel weed,<br />
waterweed, water-mllfoll, and water<br />
smartweed. Other associated species are<br />
whfte water lily<br />
<strong>Lake</strong> and bay-bottom communities occur<br />
along low wave energy bottoms and<br />
nearshore anvi ronments such as Little<br />
Muscamoot Bay. Water depths generally<br />
range between 0.3 and 2.0 m, Major<br />
species in this zone include wild celery,<br />
muskgrass or stonewort (w sp,), and<br />
flexed naiad. Brown (1983) reported the<br />
dominance <strong>of</strong> along with pondweeds,<br />
Eurasian wat foil, waterweed* and<br />
varfous emergenks growfng about Sand<br />
Island In Anchor Bay. Because <strong>of</strong> higher<br />
wave energy, lake and bay-bottom<br />
communities are less widespread on the<br />
Canadian slde <strong>of</strong> the delta. In <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>, Schloesser and Manny (1982) found<br />
few macrophy%esp which excludes<br />
, beyond 1,25 km from shore.<br />
Although conspicuous where pressnt,<br />
the feed grass community I s <strong>of</strong><br />
insufficient dJstrlbutfon to map in the<br />
<strong>St</strong>. C1air Delta. Dense colonies <strong>of</strong> reed<br />
grass ( )I usually less<br />
than 0.5 hectares in extent* are located<br />
In scattered patches on the drowned<br />
shoulders and natural levees <strong>of</strong> North,<br />
Middle, and South channels.<br />
Wetland vegetation is not static from<br />
year to year (Section 4.3) . Dynamic<br />
change occurred In the plant cornrnunitles<br />
during the interlm 1964 and 1975 as water<br />
levels In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fluctuated from<br />
very low levels to extreme high water<br />
conditions, Dicklnson Island, which has a<br />
full complement <strong>of</strong> wetland zonest and<br />
except for the dredged material disposed<br />
in the early 1970~~ had 1 i ttle development<br />
(Figures 32 and 33). In particular, large<br />
shifts occurred among the bulrushsubmersed<br />
commun ! ties and the emergent<br />
sedges and cattai 1 s.<br />
A literature search by Herdendorf et<br />
a1. (1981~) revealed no plant species in<br />
the <strong>St</strong>. Clatr Delta complex which appeared<br />
on Federal or <strong>St</strong>ate lists <strong>of</strong> endangered<br />
species. However, there Is a very small<br />
patch <strong>of</strong> the rare plant, wild rice<br />
) located on lower Hareover,<br />
Wagner (1977)<br />
reported the fo1 low1 ng threatened vascul ar<br />
plant species (Michigan) for Harsens<br />
Island: marsh sedge ( ~ i s t v l i s<br />
auberul.a) frtnged orchid<br />
(Habenarln 1, Hill's thi stle<br />
(CIrs1~ hi11 ii) , panrc grass (Panicurn<br />
JelbergjJ,), and sand grass (m<br />
ourouraa) *<br />
In 1976 and 1977, the U. S. Fish and<br />
Wildlife Servfce (Hfltunen 1980; Hfltunen<br />
and Manny 1982) conducted surveys <strong>of</strong> the<br />
macrozoobenthos <strong>of</strong> the lower <strong>St</strong>. Clafr<br />
River and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>se surveys<br />
Jndicatad that the <strong>St</strong>. <strong>Clair</strong> Rfver and<br />
Delta support a dfverse and abundant<br />
macrozoobenthos. <strong>The</strong> number <strong>of</strong> taxa and<br />
densfty found fn the river system was
signJficantly greater than that found in<br />
Anchor Bay or the open portions <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>, Cl afr (Table 181,<br />
In the <strong>St</strong>. <strong>Clair</strong> River and Delta,<br />
oligochaete worms are the most abundant<br />
macrozoobenthos (55%). Sixteen species<br />
are commons with the tubif icid genera<br />
and<br />
d the naidld genera and<br />
dominating. Dfpteran 1 arvae<br />
are the second most abundant group (26%).<br />
By number, the chironomids compose 95% <strong>of</strong><br />
the dipterans. Crustaceans are among the<br />
less abundant forms and are represented by<br />
the amphipods <strong>of</strong> the four genera,<br />
Gamm.ar_us* my !Zmgaux, and<br />
Ponto~oreia. <strong>The</strong> first three are<br />
permanent residents <strong>of</strong> the river wetlands;<br />
the fourth populates the deep waters <strong>of</strong><br />
the Great <strong>Lake</strong>s and is probably transient<br />
from <strong>Lake</strong> Huron. <strong>The</strong> insect orders,<br />
Trichoptera and Ephemeroptera together<br />
compose on1 y 1% to 2% <strong>of</strong> the total number<br />
<strong>of</strong> macrobenthos. Caddisfl ies are not as<br />
numerous as the mayflies, but are<br />
represented by a greater number <strong>of</strong> genera.<br />
<strong>The</strong> mayfl fes are 1 argel y Nexaaen l a (90% 9.<br />
Although the density <strong>of</strong> mayflies and<br />
caddisfl ies Is low, these organ isms are a<br />
significant part <strong>of</strong> total fauna because<br />
the biomass <strong>of</strong> their populations is<br />
greater than that <strong>of</strong> other insect groups<br />
(Hi 1 tunen 1980).<br />
<strong>The</strong> richness <strong>of</strong> the benthic fauna <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is evident in the high<br />
diversity <strong>of</strong> taxa present in the wetlands<br />
<strong>of</strong> Anchor Bay and elsewhere in the lake<br />
(Table 181. <strong>The</strong> moderate abundance <strong>of</strong><br />
01 igochaetes as compared to other groups<br />
indicates the quality <strong>of</strong> the benthic<br />
environment is high. Hi ltunen and Manny<br />
(1982) found the mean percent composition<br />
<strong>of</strong> the pol lution-tolerant 01 igochaetes<br />
ranged from 25% in Anchor Bay to 49%<br />
elsewhere in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> values<br />
are far below the standard <strong>of</strong> 60%<br />
01 igochaetes in pol 1 uted environments<br />
proposed by Goodnfght and Whit1 ey (1960).<br />
<strong>The</strong> presence <strong>of</strong> pollution-intolerant<br />
Ephemeroptera and Trichoptera in<br />
substantial numbers (300/m2) at a<br />
nearshore station in Anchor Bay 3 supports<br />
the conclusion that the benthic<br />
environment <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr is not<br />
severe1 y impai red by pollutjon (Hiltunen<br />
and Manny 1982).<br />
Macrozoobenthos constitutes an<br />
important source <strong>of</strong> food for fish and<br />
waterfowl in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Claf r<br />
River wetlands. Price (19631, working In<br />
western <strong>Lake</strong> Erier showed that rainbow<br />
smel tr alewives~ glzzard shad, spottail<br />
shiners* trout-perch, yellow perch,<br />
channel catf ishr white bass, and walleye<br />
feed heavily at different life stages on<br />
01 igochaetesr leeches* cladocera, ostra-<br />
cods, amphlpodsr mayfl less caddisfl iesr<br />
rnf dges, snails, and bivalves. Although<br />
1 fttle published work exists on the food<br />
habits <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River fishes, the<br />
aforementioned fish species and the<br />
invertebrates that they feed upon are all<br />
abundant In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>.<br />
<strong>Clair</strong> River (Hiltunen 1980). Dawson<br />
(1975) has observed that many <strong>of</strong> the<br />
macrozoobenthos reported for Anchor Bay<br />
are also important sources <strong>of</strong> food for<br />
waterfowl inhabiting that bay. A list <strong>of</strong><br />
the important benthic invertebrate species<br />
<strong>of</strong> the <strong>St</strong>. Clai r River-<strong>Lake</strong> <strong>St</strong>. Cleir<br />
ecosystem is presented in Appendix F.<br />
Duri ng four different stages <strong>of</strong> the<br />
Pleistocene Epoch, which itself lasted<br />
from about one million to about ten<br />
thousand years agor the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />
basin was covered with glacial ice.<br />
Within the glaciated region all benthic<br />
organ1 sms were destroyed, but each time<br />
the ice receded they reinvaded the<br />
previously ice-covered region. <strong>The</strong><br />
Pleistocene mollusks <strong>of</strong> the intsrgl acial<br />
lakes and their paleoecology are well<br />
documented by La Rocque (1966). Many<br />
species, particularly the unioni d bivalves<br />
and the 1 arge prosobranch snails8 require<br />
a continuous waterway for mfgration.<br />
Glochidia may be carried over long<br />
distances while attached to their fish<br />
host. <strong>The</strong> rich molluscan fauna <strong>of</strong> the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> region was largely<br />
repopulated from the Ohfo-Mississippi<br />
system when glacf a1 meltwaters in this<br />
region flowed south. Additf onal ly, Clarke<br />
(1981) speculated that most small snails<br />
(pulmanates) and small clams (sphaeri fds)<br />
are carried about imbedded fn the feathers<br />
<strong>of</strong> water birds, in mud attacked to their<br />
feet or clamped to the appendages <strong>of</strong><br />
1 argep flying aquatlc insects. Appendix G<br />
provides a lysting <strong>of</strong> the mollusks
0 5000 Feet<br />
t=TFL-r-e 7--rlf=74<br />
0 1500 Meters<br />
Figure 32. Wetland vegetation on Dickinson Island in 1964.<br />
L-3 Developed<br />
. .<br />
assoclated with the coas.ta1 wetlands and descended from land snails, must come to<br />
nearshore waters. Append fx H contains the surface periodf cally for air.<br />
Information on the hab 1 t at preference <strong>of</strong><br />
these spectes.<br />
Most aquat tc snalls are vegetarians.<br />
<strong>The</strong> veneer <strong>of</strong> livfng algae which covers<br />
most submerged surfaces is their chief<br />
We1 ?-vegetated portions source <strong>of</strong> food, but dead plant and animal<br />
<strong>of</strong> unpolluted embayments* marshes, beach materjal is frequently ingested.<br />
ponds, and slugglsh tributary mouths are Dissolved oxygen is an Important Ifmiting<br />
the most productJve localities for factor; most gilled species require high<br />
freshwater snails Jn <strong>Lake</strong> <strong>St</strong>. Clafr. <strong>The</strong>y concentratfons. w<strong>St</strong>h limpets such as<br />
Ilve an submerged vegetation* on rocks and mrissb being found only where the water<br />
on the bottom at the water's edge and out remains near saturation (Pennak 1978).<br />
to a depth <strong>of</strong> several meters. Two However, -1omp dec.ls_umr and Amnlico'ia<br />
subclasses, pro sob ranch^ and Pulmonata, l.imasa have been col lected in water with<br />
are we1 1 represented In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> less than 2 ppm oxygen (Harman 1974). <strong>The</strong><br />
coastal marshes. <strong>The</strong> former group 1s concentration <strong>of</strong> dissolved sol Ids in <strong>Lake</strong><br />
characterized by internal re y <strong>St</strong>. <strong>Clair</strong>, particular1 y bicarbonate<br />
gJlls (~tenidia)~ or as in , alkalinity at over 80 mg/l, provfdes<br />
external gflls, and an oP@rcuIum used to adequate essential materials for shell<br />
seal the shell aperaturee <strong>The</strong> latter construct;Son. Appendix W shows the<br />
group does not have gfllst but obtajns habitat preferences for the 47 species <strong>of</strong><br />
oxygen through a riIung-like" pulmonary gastropods which have been reported for<br />
cavjty, Pul monate snaf 1 s p whlch have the coastal marshes and Rearshore waters.<br />
56
Shore1 ine<br />
- 0 5000 Feet<br />
D<br />
0 1500 Meters<br />
Bul rush-Submersed (sparse)<br />
Cattails (some die back)<br />
a Dogwood Meadow<br />
Woodlands<br />
0 Developed<br />
Disposal Site<br />
Figure 33. Wet1 and vegetation on Dicki nson Island in 1975. Previous shore1 ine refers<br />
to 1964 shore shown in Figure 32.<br />
Dl ec- <strong>The</strong> bivalved molluscan<br />
fauna <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> consists <strong>of</strong> three<br />
families. <strong>The</strong> majority <strong>of</strong> the species<br />
belong to the Unionidae (freshwater<br />
mussels or naiades) and Sphaerlidae<br />
(fingernail clams). <strong>The</strong> third family,<br />
Corbicul fdae (1 fttle basket clams), fs<br />
represented by an f ntroduced Asiatic<br />
species, Bivalves are most abundant<br />
nearshore* especially in water less than 2<br />
m deep, <strong>St</strong>able gravel and sand substrates<br />
with a good current support the largest<br />
populations. Common1 y mussels inhabit<br />
substrates free <strong>of</strong> rooted vegetat i on, but<br />
there are numerous exceptionsv i nc1 udi ng<br />
Anodonta arandis and<br />
<strong>St</strong>omach contents <strong>of</strong> unionid mussels<br />
are commonly mud, desmidsv dfatoms, and<br />
other unicellular a1 gae, protozoans*<br />
rotifers, flagellates, and detritus. <strong>The</strong><br />
1 argest populations <strong>of</strong> mussels develop<br />
below areas where disintegration <strong>of</strong> rich<br />
vegetation is occurring (Churchill and<br />
Lewis 1924).<br />
<strong>The</strong> female pocket-book mussel<br />
(Wsflb ysntrfcc&a) is capable <strong>of</strong><br />
extending and pulsatl ng the posterior edge<br />
<strong>of</strong> the mantle fn such a way that it<br />
resembles an injured mf nnow (Clarke 1981).<br />
This activity attracts fish such as<br />
bluegill, white crappje, smallmouth bass,<br />
and yellow perch. and Increases the<br />
opportunitf es for juvenile mussels<br />
(glochidia) to attach themelves to a fOsh<br />
after they have been ejected from the<br />
parent. <strong>The</strong> larvae are released by the<br />
parent when its light rsnsftlve spots are<br />
stimulated* for example, by the shadow <strong>of</strong><br />
a passing fish. %veral unionid blvalves<br />
possess special mantle structures adapted<br />
to lure fDsh their vicinity. <strong>The</strong><br />
glochidia <strong>of</strong> each species <strong>of</strong> freshwater<br />
mussel (correspo~ds Oa vel dgsr larvae af<br />
marine bivalves) musk attach to the gills
Table 18. Macroziobenthos <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the lower <strong>St</strong>. <strong>Clair</strong> River<br />
and Delta (1977).<br />
Major<br />
taxa Compos<strong>St</strong>ion Density Composition Density Composition Density<br />
(% (no./m2) (8) ( no./mZ) (%I (no./m21<br />
Nemert i nea<br />
Nematoda<br />
Hi rund i nea<br />
01 igochaeta<br />
Polychaeta<br />
Amphipoda<br />
Isopoda<br />
Diptera<br />
Ephemeroptera<br />
Col eoptra<br />
Lepldoptera<br />
Trlchoptera<br />
Hydracarlna<br />
Gastropoda<br />
Pel ecypoda<br />
Totals<br />
a Data sources: Hiltunen (19801, Hiltunen and Manny (1982).<br />
b- Absent<br />
+ Present but not enumerated.<br />
and fins <strong>of</strong> a particular fish species or<br />
small group <strong>of</strong> species (Table 19) before<br />
further development can take place, Most<br />
glochidia never accomplish this, but those<br />
that do succeed remain attached for a few<br />
weeks as they metamorphose into tiny<br />
mussels. <strong>The</strong> young mussels then drop from<br />
the fish to take up an independent lSfe on<br />
the lake bottom, moving about and<br />
siphoning water for respiration and to<br />
obtain plankton as a source <strong>of</strong><br />
nourishment, Appendix H indicates the<br />
habitat preferences <strong>of</strong> the 64 species <strong>of</strong><br />
pelecypods which have been reported for<br />
coastal marshes and nearshore waters.<br />
3.4 FlSH<br />
<strong>Lake</strong> <strong>St</strong>. ClaJr, along with the <strong>St</strong>.<br />
<strong>Clair</strong> Rfver and the Detroit River* forms<br />
the connecting waterway between <strong>Lake</strong> Huron<br />
and <strong>Lake</strong> Erie, and is therefore an<br />
important fish conduit. <strong>The</strong> bays and wet-<br />
lands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, especially in the<br />
<strong>St</strong>. <strong>Clair</strong> Delta area at the mouth <strong>of</strong> the<br />
<strong>St</strong>, Cl air River, are important spawning<br />
and nursery areas for many species that<br />
support major fisheries in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
as well as <strong>Lake</strong> Huron and <strong>Lake</strong> Erie<br />
(Johnston 1977; Goodyear et al. 1982arb,c;<br />
Hatcher and Nester 1983). <strong>St</strong>udies<br />
conducted at a number <strong>of</strong> widely separated<br />
sltes in the Great <strong>Lake</strong>s indicate that the<br />
nearshore waters are spawning and nursery<br />
grounds for most fishes (Boreman 1976).<br />
More than 70 species <strong>of</strong> fish have been re-<br />
corded as residentsr or migrants, in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>; <strong>of</strong> these, 48 species are depen-<br />
dent on or known to use the marshes and<br />
shallow areas <strong>of</strong> the lake (Appendix I).<br />
Although many species <strong>of</strong> fish utilize<br />
coastal wet1 ands, comparative1 y few
Table 19. <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> unionid bivalves and their glochidial host fishea<br />
Fish hostsb AM5 FUS QUA ELL ALA LAS SIM AN0 STR TRU PRO CAR LEP ACT LIG LAM<br />
Northern pfke X<br />
Carp X X<br />
Golden shiner X<br />
Creek chub X X<br />
White sucker X<br />
N hog sucker X<br />
N Sh redhorse X<br />
01 bull head X<br />
Ye bull head X<br />
Br bull head X<br />
Ch catfish X X<br />
Bk stickleback X<br />
White bass X X<br />
Rock bass X X X<br />
Gr sunfish X X X X<br />
Pumpkinseed X<br />
Or-sp sunfish X<br />
Bluegill X X X X<br />
Sm bass<br />
Lm bass X X X X<br />
Wh crappie X X X X X X X<br />
B1 crappie X X X X<br />
Iowa darter . X<br />
Johnny darter X X<br />
Ye perch X X<br />
Wa1 1 eye<br />
Freshwater drum X X X X<br />
Mottled sculpin X<br />
Mudpuppy X<br />
(amphibian)<br />
a~ata sources: Full er (1974) r Clarke (1981)<br />
b ~ -<br />
northern CAMB - ~1ic-<br />
Sh- shorthead FUS - Fusc;onafa flpyg<br />
B1-bl ack QUA - guduh and P,<br />
Ye-ye1 1 ow ELL - Elliat_io dllatata<br />
Br-brook ALA -<br />
Gr-green LAS -<br />
Or-sp-orange-spotted SIM -<br />
Sm-smal lmouth AN0 -<br />
Lm-1 argemouth STR -<br />
Wh-white TRU -<br />
PRO -<br />
CAR -<br />
LEP -<br />
ACT -<br />
LIG -<br />
LAM -<br />
C<br />
X X X<br />
X X<br />
X X X<br />
X X X<br />
X X
specles are strongly dependent an aquatic<br />
vegetation for spawntng, feeding8 or<br />
cover. <strong>The</strong> more common wetland-dependent<br />
species found fn <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and<br />
occurring frequently in the coastal<br />
wetl ands I nc1 ude 1 ongnos<br />
-1, bowfin (m<br />
pike (&&,& luciu@, gras<br />
-) , central mudmlnnow (Y-a<br />
m), golden shiner (<br />
blacknose sh<br />
, blackchln shiner (&&roois<br />
mic shfner f<br />
1 sand shiner<br />
1 P lake chubsucker f<br />
, rock bass (<br />
1, green sunf<br />
smallmouth bass<br />
generally associated with lotic waters and<br />
clean sand or gravel bottoms8 but they are<br />
also common in coastal lake waters, where<br />
they may util ize deep lacustrine coastal<br />
wetlands with sand or gravel bottoms. <strong>The</strong><br />
bet.erod~ll)~ tadpole madtom (lig&dus<br />
-9, banded killifish (Egn&ulus<br />
green sunfish, rock bass, and smallmouth<br />
bass, are signif lcant game species. Among<br />
-haw) r pumpkinseed (l&pomi~ commercial and game species the walleye<br />
-1, Iowa darter ( F a - u ) r ($tizostedioo y. -1 and native or<br />
and brook stickleback (w inconslam). introduced salmonids apparently have<br />
Several other species are largely little direct association with coastal<br />
restricted to vegetated waters but are wetl ands, Other specles <strong>of</strong> recreational<br />
either uncommon or rare In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. or commercial Importance, including ye1 low<br />
<strong>The</strong>se are the spotted gar (-<br />
perch (parca -1, whlte bass<br />
$xuh$u), muskellunge (&&x -1 , (w -1, and freshwater drum<br />
pugnose shiner (- -1 and ( ~ i n o t u -1, r<br />
as we1 1 as<br />
pugnose minnow (m ~miliu). important forage species, including spot-<br />
However, a signif icant native, selfsustaining<br />
populatfon <strong>of</strong> muskellunge<br />
exists in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Haas 1978) and<br />
tail shiner (Notroo hudson_ius) and<br />
emerald shiner (- j&hgchm,)r<br />
are ubiquitous in the coastal zone <strong>of</strong> <strong>Lake</strong><br />
spawns in the delta wetlands. Here, it is<br />
harvested by the Chippewa Indians on the<br />
Walpole Island Indian Reserve, Ontario.<br />
<strong>St</strong>. <strong>Clair</strong> and may be locally commmon in<br />
certain coastal wetl ands,<br />
Known ffsh spawning areas fn the <strong>St</strong>,<br />
Several fish species found in the <strong>Clair</strong> system have been documented by the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin are common to U.S. Fish and Wildlife Service (Goodyear<br />
abundant in wetlands and adjacent waters, et a1. 1982arbrc1, including site specific<br />
although they are <strong>of</strong>ten associated with information on ffsh spawning and nursery<br />
non-vegetated habitats as well.<br />
-1,<br />
<strong>The</strong> gl z-<br />
goldfish<br />
areas for sturgeon (- ful\~asce&),<br />
alewife (w a s e u d o m ) , rainbow<br />
trout (SalmP -1, smelt (Osmru.s<br />
-1, white sucker (&&ostom~<br />
-1, smallmouth bass, rock bass,<br />
yellaw bull head (m na$al f S) r<br />
walleye, and ye1 low perch. Goodyear and<br />
her colleagues summarized information on<br />
a17 past and presently known spawning<br />
areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong><br />
River. <strong>The</strong>se areas include the marshy and<br />
occur in quiet* low-gradient waters with<br />
bottoms <strong>of</strong> mud or clay. <strong>The</strong>se species are<br />
general 1 y cover-oriented and may be common<br />
in sheltered riverine and l acustrine<br />
coastal wet1 ands along <strong>Lake</strong> <strong>St</strong>. Cl air.<br />
All except the gizzard shad, goldfish,<br />
bl untnose rnf nnon, and fathead minnow are<br />
significant game spectes,<br />
shallow bay areas along the shore1 ine and<br />
the wetlands <strong>of</strong> the delta. While the<br />
relationship <strong>of</strong> submersed aquatfc<br />
macrophytes in 1 ittoral waters to f i shery<br />
productf vi ty is not fully understood,<br />
prel lminary results <strong>of</strong> recent studies<br />
indicate that the submersed littoral<br />
macarophyte habitat is important to fish<br />
production in these waters (Schl oesser and<br />
Manny 1982).<br />
Walleye migrate through <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> and use the channels <strong>of</strong> the <strong>St</strong>.
<strong>Clair</strong> Delta for spawning areas (Figure<br />
34). Tagging studles (Ferguson and Derksen<br />
1971; Wolfert st a1. 1975) indicated<br />
considerable movement <strong>of</strong> wall eye between<br />
<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and adj<strong>of</strong>ning lakes Erie<br />
and Huron. Each lake has many known and<br />
some suspected spawni ng areas. Sf nce<br />
walleyes prefer to spawn close to their<br />
p 1 ace <strong>of</strong> or i g i n ( Eschmeyer 1950; Ferguson<br />
and Derksen 1971) there appears to be a<br />
number <strong>of</strong> semi-di screte stocks coexisting<br />
in these lakes. Tagging studies have not<br />
been sufficiently intense, however# to<br />
delimit these stocks within time and<br />
space.<br />
Yellow perch (w flayescm,sI is<br />
one <strong>of</strong> the most abundant fish in <strong>Lake</strong> <strong>St</strong>.<br />
Clalr and is known to spawn over aquatic<br />
vegetation in most inshore areas <strong>of</strong> the<br />
lake (Figure 351, but it is not known<br />
whether discrete stocks exist. Commercial<br />
fishermen belleve that <strong>Lake</strong> Huron fish can<br />
be identified by shape and color in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. Similarlys Ohio fishermen<br />
postulate that a postspawning movement <strong>of</strong><br />
large yellow perch comes into the western<br />
basin <strong>of</strong> <strong>Lake</strong> Eries presumably from <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>re is no evidence to<br />
support or reject these views, <strong>The</strong> scant<br />
tagging that has been carried out with<br />
yellow perch suggests that they are<br />
capable <strong>of</strong> moving between the lake systems<br />
(Johnston 1977). However, the maintenance<br />
<strong>of</strong> a number <strong>of</strong> age-groups in <strong>Lake</strong> <strong>St</strong>,<br />
<strong>Clair</strong> is in direct contrast to the<br />
situation in <strong>Lake</strong> Erie.<br />
<strong>St</strong>urgeons a threatened species in<br />
Mfchigan and an endangered species in<br />
Ohlor are common in the deeper portions <strong>of</strong><br />
the delta and channels. <strong>The</strong> major known<br />
spawning area for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> sturgeon<br />
is the North Channel <strong>of</strong> the <strong>St</strong>, Clalr<br />
River (Figure 36). <strong>The</strong>se sturgeon migrate<br />
up to the North# South, and Middle<br />
channels to spawn at a site in the North<br />
Channel ; young-<strong>of</strong>-the-year sturgeon<br />
migrate from this spawning site to the<br />
marsh between Bouvier and Goose Baysv<br />
where they are found among the rushes<br />
(Goodyear at al. 1982a~b). <strong>The</strong> channels<br />
are also productive fishing areas for<br />
muskellunger smallmouth bass# 1 argemouth<br />
bass* freshwater drum* and northern pike,<br />
Flgures 37 and 38 depict spawning and<br />
nursery areas for several <strong>of</strong> these<br />
species.<br />
[ /fi\ WALLEYE<br />
Spawning area<br />
2.:. Nursery area<br />
Figure 34. Walleye spawning and nursery<br />
areas in the Detr<strong>of</strong>t River and <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> (Goodyear et al. 1982b,c),<br />
<strong>Lake</strong> <strong>St</strong>. CSair has been acclaimad one<br />
<strong>of</strong> the best fishing areas in Elarth America<br />
(Figure 39). This fishery is fostered by<br />
the marsh habitats which sustain species<br />
number and diversity, Sport fishing in<br />
wetlands exhfbits high sconomlc values,<br />
Jaworski and Raphael ( 1878) calcul atad the<br />
value <strong>of</strong> Mfchigan's coastal wetlands with
Detroit River<br />
~atrolt<br />
YELLOW PERCH<br />
Spawnlnp area<br />
AY. Nuraery area<br />
Figure 35. Yellow perch spawning and<br />
nursery areas in the Detroit River and<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Goodyear et at, 1982b,c).<br />
respect to sport f l shfng at $ll6/wetland<br />
hectare/year, <strong>Lake</strong> <strong>St</strong>. Cl air exhibits the<br />
highest sport flshery value for Great<br />
lakes coastal wet1 ands because <strong>of</strong> its<br />
proximity to Detroit and the high qua1 i ty<br />
<strong>of</strong> angl i ng for wa1 l eyer smallmouth bass,<br />
and yellow perch (Table 20). Current<br />
estimates indicate that the <strong>St</strong>, Clafr<br />
Delta wetlands average 8 to 16 angler-<br />
day s/wetl and hectare/year (Werdendorf et<br />
a%. 1981~1.<br />
Major game fish species whfch furnish<br />
a recreational fishery in Bouvier Bay<br />
Wetland, known also as <strong>St</strong>. Johnts Marsh,<br />
include northern pike, 1 argemouth bass9<br />
ye1 low perch, bull heads* and various<br />
panflsh (Posplchal 1977). Recreational<br />
fishing is popular from shore sites as<br />
well as from boats In the shallow bays and<br />
channels <strong>of</strong> <strong>St</strong>. Johnrs Marsh, Within the<br />
<strong>St</strong>. <strong>Clair</strong> Flats <strong>St</strong>ate Wildlife Area on<br />
Dickinson Island and on Harsens Island,<br />
recreational fishing is popular for<br />
muskellunge, northern pike, wall eye,<br />
yellow perch, smallmouth bass, channel<br />
catfish (fctalurs punctatus), bull heads,<br />
bluegill other sunfishes, crappie, rock<br />
bass, white bass, lake sturgeonr coho<br />
salmon (Qncorhv- steel head<br />
trout and other species. <strong>The</strong> area is also<br />
a prolific producer <strong>of</strong> bait-minnows. Bow<br />
and arrow fishing for carp is a growing<br />
springtime activity, which generates 3,000<br />
to 5,000 angler-days <strong>of</strong> recreation per<br />
year (Pospichal 1977). Ice fishing<br />
(commonly by hook and line) Is also an<br />
active sport in January and February.<br />
Common species pursued include ye1 1 ow<br />
perch (85%) s sunfish (10%)r walleyer and<br />
northern pike. More than 95% <strong>of</strong><br />
recreation ice fishing is for wetland<br />
associated species.<br />
Other than species composition and<br />
recreational use <strong>of</strong> the flsh fauna <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>r knowledge <strong>of</strong> the wetland<br />
fishery <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> Is lacking in<br />
several respects. Other important con-<br />
siderations include: 1) actual relation-<br />
ships <strong>of</strong> the species to the wetlands in<br />
terms <strong>of</strong> their utilization for spawning,<br />
nursery, and feeding areas, 2) fish<br />
community structure, 3 1 niche occupation,<br />
and 4) interspecific relationships within<br />
wetlands. <strong>The</strong> coastal wetlands <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. ClaJr appear to support a mlxed fish<br />
fauna <strong>of</strong> warmwater and coolwater species.<br />
In addl ti on to wetland dependent speciesr<br />
many species common in other coastal<br />
habltats are also common in coastal<br />
wetlandsE dependlng on condition <strong>of</strong><br />
shelter, bottom type, water clarity, water<br />
depth, and density <strong>of</strong> vegetation. Because<br />
<strong>of</strong> the obvious voids in fisheries<br />
information wfthin coastal wet1 ands,<br />
research in this area ranks high priority.
MlCHlGAN<br />
LAKE STURGEON<br />
Spawning area<br />
.* .+ Nursery area<br />
Middle Channel<br />
outh Channel<br />
Figure 36. <strong>Lake</strong> sturgeon spawning and nursery areas in the Detroit River, <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>, and <strong>St</strong>. <strong>Clair</strong> River (Goodyear et a1 . 1982a,b,c).<br />
63
Figure 37. Rock bass, pumpkinseed/biuegill, largemouth bass, and smallmouth bass<br />
spawning and nursery areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Goodyear et al. 1982b).
FISHING AREAS<br />
f_l_j YELLOW PERCH<br />
SMALLMOUTH BASS<br />
MICHIGAG ONTARIO<br />
Figure 39. Sport fishing areas by species In the vicinity <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta wetlands<br />
(<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Advisary Committee 1975).<br />
33 AMPHlBlANS AND REPTILES<br />
<strong>The</strong> assemblage <strong>of</strong> amphlbfans and<br />
reptlles reported For the <strong>Lake</strong> <strong>St</strong>. ClaSr<br />
regfon can be divided into four major<br />
groups: 11 those specfes almost always<br />
found In wetlands and somehow dependent on<br />
wetland habitats to the degree that they<br />
are uncommon elsawherer 2) aquatic and<br />
semiaquatic species found in a wide range<br />
<strong>of</strong> law1 and wet habf tats, including coastal<br />
wetlands. 3) terrestrial species which<br />
enter water or wetlands only fnc5dentally<br />
or to breed* and 4) aquatfc and<br />
semiaquatic specles found prSmar4l y in<br />
upland or lotic waters and not common to<br />
coastal wetlands (Herdendorf st a7.<br />
1981~). <strong>The</strong> latter are not disussed in<br />
thls report. Four species recorded In the<br />
literature as occurring fn the coastal<br />
zone <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr are largely wetland<br />
dependent: mudpuppy (i&&uxi<br />
JMGW<br />
red-spotted newt ( Y&k<br />
, eastern fox snake (w<br />
eastern massasauga<br />
1. <strong>The</strong> mudpuppy is<br />
largely an open lake species which is most<br />
sonmon around large 1 acustrine wetlands<br />
with submersed vegetation, whereas the<br />
red-spotted newt is found primarily in<br />
small ponds and sheltered palustrine or<br />
riverine wetlands. <strong>The</strong> eastern fox snake<br />
is common in large coastal emergent-type
Table 20. Average annual number <strong>of</strong> sport fish takenafrom Michigan waters <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. Clai r and connecting waterways (1975-1979).<br />
Species <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Detroi t Total<br />
<strong>St</strong>. Clafr River River individuals<br />
<strong>Lake</strong> trout<br />
Rai nbow/steel head<br />
Brown trout<br />
Coho salmon<br />
Chi nook salmon<br />
Wal leye/sauger<br />
Bass<br />
Northern pike/musky<br />
Yellow perch<br />
Sunf ish/bluegill<br />
Crappie/white bass<br />
Bull head/catf ish<br />
Rock bass<br />
Sucker<br />
Whitefish/cisco<br />
Other<br />
Angler days 1,079,031 320,701 4529376 lr 852,108<br />
Fishermen 100,622 32,523 41,674 174s 819<br />
a Data source: Michigan Department <strong>of</strong> Natural Resources (unpublished data).<br />
wetlands, but the rarer eastern massasauga<br />
is primarily restricted to swamp forest<br />
and bogs.<br />
Amphibians found in a wide range <strong>of</strong><br />
lowland wet or aquatic habitats in the<br />
<strong>Lake</strong> <strong>St</strong>. Clafr coastal zone include the<br />
blue-spotted salamander (&nbvstoma 1 atec a), spotted salamander ( A m<br />
n@&ulatu!q), four-toed salamander (Hemi-<br />
Utv1 i -1, eastern tiger sa1 a-<br />
mander t,4mbvsta =), gray treefrog<br />
(fjYLB yersIcQ&x), northern spring peeper<br />
(liyljl ~ ruci fer) western chorus frog<br />
(pseudacris triseriata), Blanchardqs<br />
crfcket frog C-5 -1 r<br />
bullfrog (m -1, green frog<br />
(m m1 ano-I, pickerel frog<br />
(Baas pa1 ustris) , and northern leopard<br />
frog (m ploiens). <strong>The</strong> salamanders may<br />
<strong>of</strong>ten occur in moist terrestrial habitats,<br />
but the1 r larvae are always aquatjc.<br />
Aquatic and seml-aquatic reptiles which<br />
probably occur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />
wetlands or their border lands include the<br />
northern ringneck snake (Digdoahfs ~unctatlls<br />
edwar_dsl), eastern milk snake<br />
(mm -1 northern<br />
water snake (&&& SL northern<br />
brown snake (<strong>St</strong>orerb L 1, northern<br />
red-bellied snake (sore* p, ~sg@ito -<br />
1par:ulatg), Butlerts garter snake (m<br />
eastern gar<br />
Several amphfblans <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>.<br />
Clafr coastal zone are largely terres-<br />
trial, although they are found in lowland<br />
areas and are dependent on water for<br />
breeding. Consequent1 y the adults ar<br />
l arvae may occur in coastal wetlands or
their margfns at least seasonal 1 y. <strong>The</strong>se<br />
specles 1 nclude the red-backed salamander<br />
)C the Amerfcan<br />
d the wood frog<br />
trial reptf le5<br />
may occur in the margins <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
coastal wetlands or incidentally in the<br />
wetlands themselves<br />
smooth green snak<br />
m ) r blue racer<br />
1, and black rat snake (<br />
I.<br />
Although few species <strong>of</strong> reptiles and<br />
amphibians are absolutely dependent on the<br />
coastal wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> for<br />
habitat, the continued abundance <strong>of</strong> most<br />
species in the coastal tone is probably<br />
dependent on the presence <strong>of</strong> extensive<br />
coastal wet? ands. <strong>The</strong> relatively un-<br />
disturbed coastal hab 'f t at a1 ong most <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> Delta probably supports a<br />
present herpet<strong>of</strong>auna similar to that <strong>of</strong><br />
pre-settl ement ti mesr both in composition<br />
and relatf ve abundance <strong>of</strong> specles.<br />
Appendix J Ilsts the specles known or<br />
presumed to occur in or adjacent to the<br />
coastal wetlands; because <strong>of</strong> the scarcity<br />
<strong>of</strong> published records, this list is<br />
somewhat specul atlve.<br />
species 1s wetland-dependent and was once<br />
common and restricted to coastal emergent<br />
wetlands and wetland margins <strong>of</strong> western<br />
<strong>Lake</strong> Erie and the coast <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air.<br />
<strong>The</strong> black rat snake is also listed as<br />
threatened In Michfgan and may occur fn<br />
drier margins af <strong>Lake</strong> <strong>St</strong>, Clafr coastal<br />
wetlands. <strong>The</strong> eastern massasauga is a<br />
candidate for Federal listing on the<br />
threat.ened/endangered speci es 1 ist. <strong>The</strong><br />
following species, considered rare in<br />
Michigan, may also occur in these coastal<br />
wetlands: eastern spiny S<strong>of</strong>tshellr spotted<br />
turtle ( 18 and fourtoed<br />
salamander,<br />
<strong>The</strong> extensive wetlands <strong>of</strong> the <strong>St</strong>.<br />
<strong>Clair</strong> Delta are utilfzed for nesting and<br />
during migration by waterfowl wading<br />
birds, gulls, terns, and shorebirds.<br />
Appendix K lists the Important bird<br />
species which occur in the vicinity <strong>of</strong> the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands. Waterfowl<br />
commonly observed in the wetlands include<br />
mallard (BagS whvn-), black duck<br />
(81195 -1, blue-wlnged teal (m<br />
$i%cocs). wood duck (&1X -1, American<br />
wigeon (m -18 and northern<br />
shoveler (u ~lvpeata) . In addi tion,<br />
canvasback (m val f +=i-1 scaup<br />
(m spp.1, ruddy duck (Ilxylrra<br />
&gnat and hooded merganser<br />
<strong>The</strong> presence <strong>of</strong> water, food, and<br />
cover, and the relative isolation from the<br />
cultural development which occurs around<br />
most other bodies <strong>of</strong> water are among the<br />
factors that contribute to the importance<br />
<strong>of</strong> wetlands in maintaining the coastal<br />
zone herpet<strong>of</strong> auna. <strong>The</strong> recreational or<br />
(w ~ 1 1 a t u s utilize ) the<br />
habitat for nesting . Several endangered<br />
commerci a1 importance <strong>of</strong> the coastal and threatened species (Michigan) <strong>of</strong> birds<br />
herpet<strong>of</strong>auna is unknown8 but the bullfrog, have been reported In the coastal zone.<br />
green frog* eastern spiny s<strong>of</strong>tshell, and<br />
snapping turtle are edible species which<br />
may be harvested. In addition, reptiles<br />
Osprey (Pandi_nn ha1 iaet~~5.1 and the Caspian<br />
tern (<strong>St</strong>erna -1 have been reported in<br />
scattered localities along the shore1 I ne.<br />
and amphi bians such as the northern water Nest1 ng colonies <strong>of</strong> ye1 law-headed<br />
snake and the snapping turtle, serve as<br />
food sources for many species <strong>of</strong> fish,<br />
birds, and mammals, or may themselves be<br />
blackbird ( 18<br />
black tern (m -1, and common<br />
tern (Sf;arna hi run&? have been observed<br />
predators <strong>of</strong> economf tally important, fish in several wetlands. <strong>The</strong> common tern is<br />
and wild1 ife. Beyond ~m3ra1 background presently on the Michigan endangered<br />
on the occurrence and relative abundance<br />
<strong>of</strong> reptiles and amphibians in the coastal<br />
specles list.<br />
wetlands* little publfshed information<br />
exists pertaining to papu1atlon8 community<br />
structurer and dynamics <strong>of</strong> the<br />
Waterfowl are a most Important<br />
component <strong>of</strong> the bIotic envf ronment. <strong>The</strong><br />
importance <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />
herpet<strong>of</strong> auna,<br />
to ducks, geese, and other waterfowl is<br />
<strong>The</strong> eastern Cox snake is listed as<br />
threatened by +Re <strong>St</strong>ate <strong>of</strong> Michtgan, This<br />
based primarily on the location <strong>of</strong> these<br />
wetlands in relation to waterfowl flyways,<br />
the productivity (in terms <strong>of</strong> waterfowl
foods) <strong>of</strong> the wetlandso and the declfne <strong>of</strong><br />
other coastal wetland systems. Except for<br />
that <strong>of</strong> mallard, black duck and blue-<br />
winged teal, nesting appears to be a<br />
secondary use <strong>of</strong> these wetlands with<br />
feeding and resting during migratlon bejng<br />
<strong>of</strong> primary importance.<br />
Based on waterfowl feeding and<br />
resting use durfng migratfon as well as on<br />
duck harvest and production <strong>of</strong> young, the<br />
wetlands and associated open waters <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> may be the most important<br />
wetland system in the Great <strong>Lake</strong>s region<br />
with the possible exception <strong>of</strong> Long Point<br />
in <strong>Lake</strong> Erie. With reference to the Great<br />
<strong>Lake</strong>s-Chesapeake Bay migration torrider*<br />
the principal flight paths <strong>of</strong> the tundra<br />
swan, formerly whist1 ing swan, (CYQILUS<br />
-1. canvasback, bufflehead<br />
( -1, and ruddy duck in-<br />
clude a resting stopover fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
(Bell rose 1976). In reference to water-<br />
fowl harvesting in Michigan, <strong>St</strong>. Clai r<br />
County, which includes most <strong>of</strong> the wetland<br />
on the American side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr, is<br />
the leading harvest county (Carney et a1 .<br />
1975). Production data for the Bouvier<br />
Bay (including <strong>St</strong>. John" Marsh) and<br />
Harsens Is1 and wetlands reveal an average<br />
<strong>of</strong> 240 young produced per km2 EHerdendorf<br />
et al. 1981~).<br />
Important 1 lterature sources re-<br />
garding the waterfowl use <strong>of</strong> the wetlands<br />
are Jaworski and Raphael (1978) as we11 as<br />
Herdendorf et al. (1981~). <strong>The</strong> latter<br />
source is especially important in regard<br />
to the Bouvier Bay, Dickinson Island, and<br />
Harsens Island wetlands. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong><br />
Flats <strong>St</strong>ate Game Area comprises the lower<br />
portions <strong>of</strong> Harsens Island and Dickinson<br />
Island, plus the two former areas <strong>of</strong> the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> National Wfldllfe Refuge,<br />
one at the mouth <strong>of</strong> North Channel in<br />
Anchor Bay, and another in between the end<br />
<strong>of</strong> Middle Channel and Channel A Bout Rond.<br />
In Ontario, the Canadian Wildl ife<br />
Service (Dennis and Chandler 1974, Dennis<br />
et a1. 1981) have documented the util ity<br />
<strong>of</strong> wetlands with regard to waterfowl. In<br />
terms <strong>of</strong> total use durlng migration, the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands rank second to<br />
Long Point (<strong>Lake</strong> Erie), Ontario. <strong>The</strong><br />
coastal marshes are presently the most<br />
important staging areas in southern<br />
Ontario for mallard, black duck, Canada<br />
1, and tundra<br />
ons <strong>of</strong> .the North<br />
asback* redhead<br />
1, and tundra swan also<br />
util ize the <strong>Lake</strong> <strong>St</strong>. Claf r marshes during<br />
migratlon staging periods.<br />
Walpole Island possesses extensive<br />
marshes, particularly at its southern end<br />
where the shore fronts on <strong>Lake</strong> <strong>St</strong>. Cl ai r.<br />
<strong>The</strong>re are flve major islands that make up<br />
the 24,000-hectare Wal pole Is1 and Indi an<br />
Reserve. Here many nesting species <strong>of</strong><br />
birds rare in both the region and Ontario<br />
as a whole have been observed, including<br />
canvasback, redhead, ruddy duck, northern<br />
harrier (Circus little gull<br />
(m -1 , Forsterts tern (w<br />
-1, and king rail (&Uui 3<br />
(Goodwin 19821, Wooded areas e<br />
island include sections <strong>of</strong> oak savannah<br />
which accommodate a rookery <strong>of</strong> blackcrowned<br />
night-herons )<br />
and great egrets (- allus).<br />
As indicated In Table 21, a diversity<br />
<strong>of</strong> waterfowl rely on the <strong>St</strong>. <strong>Clair</strong> Delta<br />
wetlands. Waterfonl use during migration<br />
is more Important than use <strong>of</strong> these<br />
wet1 ands for breeding purposes. Moreover,<br />
ducks are much more numerous than are<br />
swans and geese. With regard t o dabbling<br />
ducks, the most abundant migrants include:<br />
mallard, black duck, American wigeon as<br />
we1 1 as pintail (&as<br />
teal, and green-winged<br />
Among the more abundant migrant diving<br />
ducks are canvasback, great6<br />
(&j&ygj -1, lesser scaup<br />
ilfflnfs) * 11 a<br />
goldeneye ( and buffle-<br />
head.<br />
<strong>The</strong> use <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta<br />
wetlands for feeding by both dabbling and<br />
diving ducks fs <strong>of</strong> prime importance (Table<br />
22). Plant foods from shallon marshes and<br />
waste gralns from nearby agricultural<br />
fields are especially important in the<br />
diet <strong>of</strong> dabbl ing ducks, parti cul arly<br />
mallards and black ducks as well as Canada<br />
geese, whereas submersed aquatic plants<br />
and thei r associated i nrertebrate fauna<br />
constitute important fwd items far diving<br />
ducks such as canvasback, redhead* and<br />
scaup (Jaworski and Raphael 1978). Corn
Table p.<br />
1977).<br />
Birds observed in <strong>St</strong>. <strong>Clair</strong> Delta wetlands (1975-<br />
Species Bouvi er Dickinson Harsens<br />
~ayb Is1 and Is1 and<br />
Tundra swan<br />
Canada goose<br />
Snow goose<br />
Ma1 1 ard<br />
Black duck<br />
Gadwall<br />
Pintail<br />
Amerlcan wigeon<br />
Green-wi nged teal<br />
Bl us-w i nged teal<br />
Northern shoveler<br />
Wood duck<br />
Red head<br />
Ring-necked duck<br />
Canvasback<br />
Greater and lesser scaups<br />
Common go1 deneye<br />
Buff 1 ehead<br />
Ruddy duck<br />
Hooded and comon mergansers<br />
Horned grebe<br />
Piad-bi 1 led grebe<br />
Be1 ted kingf lsher<br />
Sedge and marsh wrens<br />
Great blue heron<br />
Green-backed heron<br />
Great and cattle egrets<br />
Black-crowned night-heron<br />
American and least bitterns<br />
K4ng and Virginia rails<br />
Sora<br />
Common moorhen<br />
Amerfcan coot<br />
Herring gull<br />
Ring-btlled gull<br />
ForsPerss and black terns<br />
Comon tern<br />
Eastern meadow1 ark<br />
Ye1 1 ow-headed blackbird<br />
Red-w i nged bl ackbi rd<br />
Common snipe<br />
American woodcock<br />
cooperfs hawk<br />
Red-tat led hawk<br />
Rough-1 egged hawk<br />
Northern harri er<br />
'~ata source: Herdendorf at a1. ( 1981~1.<br />
b~ote: Bouvier Bay survey included <strong>St</strong>. John's Marsh; other<br />
surveys less i ntense,
- - -<br />
Summer Fa1 1 Spring<br />
Adult Young Total Duration Total Duration<br />
'$able 22. Productivity and seasonal abundance <strong>of</strong> watesfowl in Bouvier<br />
Bay marshes and Harsens Island interior wetlands (1974).<br />
Area and pai rs<br />
waterfowl ( no/ km2 pop. (wks) pop. (wks)<br />
Dabbl lng ducks 35 222 41800 4-10 4r900 3-4<br />
Divers 2 14 200 - 5~000 3-4<br />
Geese 1 4 0 0 0 0<br />
Swans 0 0 0 0 0 0<br />
--<br />
Total 38 240 5 r 000<br />
Dabblfng ducks 33 210 1,400 4-10 1~500 4-5<br />
Divers 4 22 100 10 2,400 4-5<br />
Geese 1 5 - - - -<br />
Swans 38 237 lr500 - 3 * 900 -<br />
--<br />
Total 76 474 3 r 000<br />
a~ata sources: Herdendorf et al. (1981c)r Michigan Department <strong>of</strong> Natural<br />
Resources (unpubl lshed data),<br />
and other grains grown on the Harsens<br />
Island <strong>St</strong>ate Game Area and on the upland<br />
portions <strong>of</strong> the Canadian side <strong>of</strong> the delta<br />
attract dabbl ing ducks and Canada geese.<br />
In contrast* submersed aquatic plants i n<br />
the diet <strong>of</strong> diving ducks include wild<br />
celery, pondneeds, and waterweed.<br />
<strong>The</strong> significance <strong>of</strong> the plant and<br />
invertebrate food base <strong>of</strong> the <strong>St</strong>, <strong>Clair</strong><br />
Delta wetl ands in relation to waterfowl<br />
can not be overstated. An important study<br />
by Dawson (1975) <strong>of</strong> Anchor Bay in northern<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, indfcates that only a<br />
small percentage <strong>of</strong> the preferred aquatic<br />
pl ants and invertebrates avail able to<br />
migrating waterfowl fn <strong>Lake</strong> <strong>St</strong>. Claf r<br />
wetlands are currently being utilized.<br />
Waterfowl food supplies are especially<br />
important during sprf ng migration because<br />
preferred foods are general 1 y less<br />
abundant and breeding females must arrive<br />
in good condition on the breedfng grounds.<br />
Thus the spring migration is less<br />
restricted to specific feeding areas and<br />
waterfowl can be observed in lakes and<br />
wetlands whSch they did not frequent in<br />
the fall. For exampf er note in Table 22<br />
that divfng ducks are more numerous Jn the<br />
relatively shallow Bouvfer Bay and Harsens<br />
Island wetlands in sprfng than in the fall<br />
migration season. Waterfowl migratjon is<br />
discussed in more detail in Sectton 4.3.<br />
With regard to production <strong>of</strong><br />
waterfowl In the wetlands along take <strong>St</strong>.<br />
<strong>Clair</strong>, these wetlands appear to be <strong>of</strong><br />
moderate importance on1 y. Some pre-<br />
lfmlnary data collected by JaworskS and<br />
Raphael (1978) suggested that Michigan's<br />
coastal wetl ands have a duck nesting<br />
density <strong>of</strong> 32 breedjng pafrs per km2 and<br />
that goose and swan nestlng was<br />
Insfqniffcant, Data reveal that the pairs
<strong>of</strong> dabbling ducks slfghtly exceed this<br />
prel iminary average, but that paf rs <strong>of</strong><br />
diving ducks were much less than the man<br />
(Table 22) . Eterdendorf et al. (1981~1<br />
reported that ma1 1 ard, blue-winged teal#<br />
and cinnamon teal are the most common<br />
nesters in the delta wetlands along wlth<br />
smaller numbers <strong>of</strong> black duck, wood duck,<br />
pintall, redhead, and ruddy duck. Small<br />
but signi f icant numbers <strong>of</strong> the re1 at1 vel y<br />
rare redhead duck nest in <strong>St</strong>. Johnrs<br />
Marsh, on lower Harsens Island* and along<br />
coastal Walpole Island. Both mallard and<br />
blue-winged teal are cl assif fed as meadow<br />
nesters, but the mal 1 ard exhibits great<br />
versatility In regard to selection <strong>of</strong><br />
potential nesting sites.<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has long been renowned<br />
as an excel? ent duck huntfng area.<br />
Hunting pressure from both the land and<br />
water is common on the American and<br />
Canadlan sides <strong>of</strong> the delta. <strong>The</strong> main<br />
species harvested in these coastal<br />
wetlands are the mallard, along with<br />
lesser numbers <strong>of</strong> black duck, pintail, and<br />
teal (Table 23) . However, In open-water<br />
areas <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, where hunting<br />
from boats and floating blinds prevails*<br />
the majorfty <strong>of</strong> the ducks harvested are<br />
divers# i.e., scaup and the now restricted<br />
canvasback and redhead, rather than<br />
dabbling ducks.<br />
<strong>The</strong> type <strong>of</strong> shores range from sandy<br />
beaches to mudflats. <strong>The</strong> beaches attract<br />
small flocks <strong>of</strong> shorebirds In migration.<br />
Those species commonly seen feeding are<br />
bfrds, such as herons and egrets, gather<br />
to feed in the mud flats near larger<br />
streams.<br />
Nesting species in the <strong>St</strong>. <strong>Clair</strong><br />
Flats (Figure 409 include such specIes as<br />
Table 23. Species composition <strong>of</strong> ducks<br />
harvested In the <strong>St</strong>., <strong>Clair</strong> Flats <strong>St</strong>ate<br />
Game Area (1974-1975).<br />
Annual<br />
Spechs harvest Percent<br />
(avge <strong>of</strong> total<br />
individ.)<br />
Ma1 1 ard, 4,362<br />
including domestic<br />
Black duck and hybrids 498<br />
Plntail 242<br />
Green-w 1 nged teal 173<br />
American w l geon 13 8<br />
Ri ng-necked duck 78<br />
Bl ue-w i nged teal 7 1<br />
Scaup 34<br />
Gadwall 30<br />
Wood duck 28<br />
Merganser 19<br />
Buf fl ehead 18<br />
Northern shoveler 10<br />
Common go1 deneye 9<br />
Redhead 4<br />
Ruddy duck 3<br />
Oldsquaw 0<br />
Canvasback 0<br />
Scoter 0<br />
- -<br />
TOTAL 5,717 100.00<br />
a~ata source: Jaworski and Raphael (1978).<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands support the<br />
following nesting waterbirds In wooded<br />
habitats: great blue heron (<br />
-1, black-crowned night-h<br />
green-backed heron (- .-US),<br />
great egret, American woodcock (3~01<br />
OD=<br />
-1, be1 sher (Cerv1 Q D<br />
bald eagle Jeuc-1, and<br />
osprey.<br />
Investigations in 1954 <strong>of</strong> a rookery<br />
on Dfcki nson Is1 and found approximately<br />
350 pairs <strong>of</strong> great blue herons and 20<br />
pairs <strong>of</strong> black-crowned night-herons a1 ong<br />
with seven pafrs <strong>of</strong> great egrets. <strong>The</strong>Ir<br />
numbers have declined over the past three<br />
decades. Nearby Harsens Island supports a<br />
rookery <strong>of</strong> bl ack-crowned night-herons<br />
(U.S. Army Corps <strong>of</strong> Engineers 1974).<br />
S<br />
Herons are a solitary bl ).d except<br />
when breeding. <strong>The</strong>y are colonfd nesters
Figure 40. Least bittern (Ixobr chus<br />
exilis) with young in broad-leaved __Y-r catta 1<br />
(Typha latifolia) nest (Michigan Department<br />
<strong>of</strong> Natural Resources).<br />
who requlre a habitat with an abundance <strong>of</strong><br />
twigs and sticks for nest-building. <strong>The</strong>se<br />
long-legged waders prefer a diet <strong>of</strong> f ishp<br />
but w i l l also feed on insectsr<br />
crustaceans, amphibians, reptiles,<br />
mo1 lusca, and rodents (Figure 41). <strong>The</strong>y<br />
are strong fliers and are known to fly<br />
great distances in search <strong>of</strong> food, For<br />
example in <strong>Lake</strong> Erie, herons fly 15 km<br />
from their rookery on West Sister Island<br />
to feed fn the marshes at Locust Point on<br />
the Ohio mainland (Meeks and H<strong>of</strong>fman<br />
1980).<br />
Wading birds usually forage in the<br />
shore1 i nes <strong>of</strong> the tributary streams and<br />
rfthin the coastal marshes <strong>of</strong> the <strong>St</strong>.<br />
Clatr Delta. <strong>The</strong>ir diet consists <strong>of</strong> fish<br />
for the most part, but crayfish and<br />
fnsects are also consumed in significant<br />
quantities.<br />
Short-eared owls (m<br />
red-tai 1 ed,<br />
1 have been<br />
reported fn <strong>St</strong>, John's Marsh in Bouvier<br />
Bay (Pospichal 1977). Bald eagle and<br />
osprey nest in wooded habitats assxiated<br />
with delta wetlands (Kelley et a1. 19639.<br />
3.7 MAMMALS<br />
<strong>The</strong> wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> are<br />
important habitat for several species <strong>of</strong><br />
mammals (Appendix L). <strong>Wetlands</strong> are<br />
essential for muskrat (I)nda.tra gjhethi-1<br />
and mink (w -1. <strong>The</strong> Virginia<br />
opossum (U~&J&IS 1, striped<br />
skunk (w whiti,~)~ and red fox<br />
(m u)<br />
are commonly observed in<br />
the wetlands. Furbearers are a valuable<br />
resource fn many coastal wetlands*<br />
particularly those associated with the<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Delta, including <strong>St</strong>. John's<br />
Marsh (a portion <strong>of</strong> Bouvier Bay wetlands),<br />
Dickinson Island* Harsens Island* Hal pole<br />
Is1 and, and Mitchell Bay wetlands.<br />
Fifteen species <strong>of</strong> mammals have been<br />
reported from the delta fslands (Hayes<br />
1964, U.S. Army Corps <strong>of</strong> Engineers 19749~<br />
ossum, eastern<br />
f10ridanus)~ Eur<br />
Limited deer and upland game hunting is<br />
permltted on the delta i slands.<br />
<strong>The</strong> pri nci pal mmal inhabit+ ng <strong>Lake</strong><br />
<strong>St</strong>. Claf r wetlands in terms <strong>of</strong> occurrence<br />
and economf c value is the muskrat. Based<br />
on fts feeding and lodging habitsf the<br />
density <strong>of</strong> muskrats per unit <strong>of</strong> coastal<br />
wetland can be estimated, Jaworski and<br />
Raphael ( 19781 determfnsd that Macomb
DEA'D PLANTS AND<br />
MINERAL NUTRIENTS<br />
Figure 41. Food chain <strong>of</strong> the great blue heron (Ardea herodias) in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> (U.S. Amy Corps <strong>of</strong> Engineers 1974).
County* Michigan wetlands have sustained<br />
yield <strong>of</strong> 16 muskratsfhectare. This is a<br />
relatively high yfeld for coastal wetlands<br />
which usually range from 5 to 15/hectare.<br />
<strong>The</strong> food supply <strong>of</strong> these omnivorous<br />
herbivores may be diverse, especi a1 1 y<br />
durf ng periods <strong>of</strong> stress. Baumgartner<br />
(1942) has identified 34 food items for<br />
muskrats in Michigan. <strong>The</strong> preferred food<br />
is cattail, bulrush, and bluejoint grass.<br />
<strong>The</strong>se emergent plants are common in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> wet1 ands,<br />
Little site specific research has<br />
been conducted on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> muskrats,<br />
but work done on this species in western<br />
<strong>Lake</strong> Erie can provide some insight.<br />
Several structures observed In <strong>Lake</strong> Erie<br />
coastal marshes are associated with the<br />
activity <strong>of</strong> muskrats (Bednarik 1956). <strong>The</strong><br />
most noticeable structure 1s the muskrat<br />
"house", a dome-shaped pile <strong>of</strong> emergent<br />
vegetation, primarily cattail (Figure 42).<br />
<strong>The</strong> average house varies in slze from 1 to 2.5 rn In diameter at the base and from 0.7<br />
to 3 m in height. <strong>The</strong>y are located in<br />
stands <strong>of</strong> emergent vegetation or along the<br />
periphery <strong>of</strong> such stands. <strong>The</strong>y are <strong>of</strong>ten<br />
constructed on protuberances in the marsh<br />
bottom, util izing plants in the Immedi ate<br />
area. <strong>The</strong> majority <strong>of</strong> houses are<br />
constructed during the fall, chiefly from<br />
October to November. Building activity<br />
occurs mainly during perlods <strong>of</strong> darkness.<br />
Typical densities <strong>of</strong> active muskrat houses<br />
in Sandusky Bay marshes average 4/hectare.<br />
Small houses have one living chamber and<br />
larger houses have two or three lfvfng<br />
chambers above the water 1 ine. <strong>The</strong> 1 iving<br />
Figure 42. Muskrat "houses" in a well-<br />
populated section <strong>of</strong> Magee Marsh, western<br />
<strong>Lake</strong> Erie, Ohio (Bednarik 1956).<br />
chamber is about 35 to 50 cm in height and<br />
is formed by the muskrat chewing out the<br />
vegetation. Most houses have two<br />
underwater exits or plunge holes.<br />
Other structures made by muskrats are<br />
associated with feeding activity. Rafts<br />
are constructed from stems <strong>of</strong> plants piled<br />
in a circular fashion to serve as a<br />
feed1 ng platform. "Feeding bogsw are<br />
covered, floating platforms which are<br />
smaller in dimensions than the muskrat<br />
house and are no higher than 40 cm,<br />
averaging 60 cm in diameter. <strong>The</strong>se<br />
structures are usual 1y located some<br />
distance from the larger muskrat house and<br />
serve as protected feeding sites. Itpush-<br />
upsn are built only in the winter and are<br />
small, hollowr dome-shaped shells <strong>of</strong><br />
submergent vegetatton over a plunge hole<br />
in the ice. <strong>The</strong>se protected plunge holes<br />
allow the muskrat to extend the area over<br />
which it can forage since It can travel<br />
greater dfstances under the ice.<br />
<strong>The</strong> reproductive cycle <strong>of</strong> the muskrat<br />
on Sandusky Bay <strong>of</strong> <strong>Lake</strong> Erie has been<br />
documented by Bednari k (1956) and Donohoe<br />
(1966). Reproductive activity begins in<br />
January and ends In September, with the<br />
greatest acti vl ty occurring I n February<br />
and March. <strong>The</strong> time <strong>of</strong> ff rst mating is<br />
determined in part by the time <strong>of</strong> ice<br />
breakup. <strong>The</strong> gestation period varies f ram<br />
20 to 28 days. Females usually have two<br />
1 itters <strong>of</strong> young per season although some<br />
may have three litters. Placental scar<br />
counts indicate that the mean number <strong>of</strong><br />
young per 1 itter Is l l r but Bednarik<br />
concluded that at the time <strong>of</strong> birth, the<br />
average Iltter size is eight. Sandusky<br />
Bay has habitats similar to those <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> and Is located on the south<br />
shore <strong>of</strong> <strong>Lake</strong> Erie approximately 100 km<br />
southeast <strong>of</strong> Detroit.<br />
Predation by mink occurs but is not<br />
an important influence on the muskrat<br />
population because <strong>of</strong> the low numbers <strong>of</strong><br />
mink in the marshes. Predation on juv-<br />
enlle muskrats by raccoons and Norway rats<br />
occurs In early summer and is not an<br />
Important mortal f t y factor. Some mortal -<br />
fty is attributed to hemorrhagic or<br />
"Erringtonts" disease, This disease can<br />
cause signif icant fluctuatjons in the<br />
muskrat populatfsns 3n western <strong>Lake</strong> Erie<br />
marshes (Bednarik 19561 and presumably<br />
also in lake <strong>St</strong>. CPaSr marshes,
<strong>The</strong> annual harvest <strong>of</strong> muskrat in the<br />
Michigan countfes and Ontario areas<br />
borderfng <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> is glven in Table<br />
24. <strong>The</strong> harvest from Mlchigan and Ontario<br />
marshes can exceed 100,000 individuals per<br />
year. With a pelt plus carcass price <strong>of</strong><br />
approximately $10.00 per animals muskrat<br />
trapping Is a million dollar a year<br />
Industry in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> region,<br />
Jaworskl and Raphael (1978) observed that<br />
furs from the coastal wetlands are <strong>of</strong> high<br />
quality8 hence the fur prices in these<br />
areas are usually high. <strong>The</strong> recent<br />
decline In Ontario production is<br />
attributed to roughly a 30% decline in<br />
-<br />
Table 24. Muskrat harvest in Michigan and<br />
Ontario bordering Ldke <strong>St</strong>. <strong>Clair</strong> and<br />
connect1 ng waterways.<br />
Year <strong>St</strong>. <strong>Clair</strong> Macomb Wayne Total<br />
1965 2,675 1,272 112 4,059<br />
1966 15,515 4,084 2,700 22.299<br />
1967 24,335 2,658 392 27,385<br />
1968 13,395 5,469 1,006 19.870<br />
1969<br />
-<br />
16,063 6,012 1,484 23,559<br />
1970 30,350 3,479 1,149 34,978<br />
Annual<br />
mean 17,055 3,829 1,141 22,025<br />
Year Dover Twnshp. Walpole Is. Total<br />
1974 23,512 67 880 91,392<br />
1975 22,591 73 r 796 96,387<br />
1976 21,174 31,212 52,386<br />
1977 13 r 174 14,790 31,964<br />
1978 11,658 15,385 27 1 043<br />
1979 12,715 37,340 50,055<br />
1980 15,138 26,891 42 029<br />
1981<br />
1982<br />
8,886<br />
9,215<br />
26,363<br />
29,780<br />
35t249<br />
38,995<br />
Annual<br />
~ a r 15,785 ~<br />
35,937 51,722<br />
a Data sources: Jaworski and Raphael (19781<br />
and George Ducknorth, Ontar30 Ministry <strong>of</strong><br />
Natural Resources (pers, cm. ) .<br />
wetland area during the same period IG.<br />
Duckworthv Ontarlo Ministry <strong>of</strong> Natural<br />
Resources; pers. corn.<br />
Again* little direct information is<br />
available on the ecology <strong>of</strong> raccoons in<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands# but western <strong>Lake</strong><br />
Erie studies can be helpful in<br />
understanding this ant ma1 . Several as-<br />
pects <strong>of</strong> the life history <strong>of</strong> the raccoon<br />
in Sandusky Bay marshes were investigated<br />
by a trapping and telemetry study (Urban<br />
1968, 19701, <strong>The</strong> density <strong>of</strong> the raccoon<br />
population was estimated to be 17/km2.<br />
<strong>The</strong> juvenile to adult female ratio was<br />
1.2xl.0, indfcating a moderate<br />
productivity for the raccoon population.<br />
<strong>The</strong> mean weights <strong>of</strong> both adults and<br />
juveni 1es increased from sprlng untll<br />
early winter and then decreased over the<br />
winter.<br />
<strong>The</strong> telemetry portion <strong>of</strong> the study<br />
provided information on raccoon movements,<br />
home ranger and denning. General lyr raccoons<br />
spend the daytime period in or near<br />
dens. <strong>The</strong> amount <strong>of</strong> nocturnal movement is<br />
related to the size <strong>of</strong> the home range,<br />
Raccoons w i th 1 arger home ranges move<br />
greater distances. Raccoons move at a<br />
mean rate <strong>of</strong> 162 m/hr. Marshland f s the<br />
major habitat type encompassed in an<br />
average night <strong>of</strong> travel and the habitat<br />
type in which raccoons spend the most<br />
time. Raccoons spend approximately 73% <strong>of</strong><br />
the time in the vicinity <strong>of</strong> shallow water<br />
(Figure 43). Male juvenile raccoons<br />
disperse from the marsh in the fall.<br />
Raccoons do not appear to search out<br />
waterfowl nests, since little change<br />
occurs in their movement patterns when<br />
waterfowl nesting is initiated. Dikes<br />
receive high usage in proportion to the<br />
amount <strong>of</strong> area they represent in the<br />
marsh. Movements <strong>of</strong> female raccoons<br />
encompass more wooded area per night than<br />
the male raccoons. Home range for the<br />
average raccoon is 51 hectares in size.<br />
Food items <strong>of</strong> raccoons in Sandusky<br />
Bay marshes (<strong>Lake</strong> Erie) include muskrat,<br />
crayf ishr fish, duck eggs, p1 ant materf a?<br />
seeds, and birds (Andrews 1952~ Bednarik<br />
1956, Urban 1968). Fish, crayfish, and<br />
plant material were the chlef food items
In all seasons. Muskrat fur was found in<br />
only 8% <strong>of</strong> raccoon scats collected year<br />
round, but In 47% <strong>of</strong> scats collected in<br />
the sprfng. Andrews (1952) noted that<br />
chln~ney crayfish 1 Is a<br />
favorite food <strong>of</strong> raccoons in the summer,<br />
Raccoons were responsible for the<br />
termlnat-lon <strong>of</strong> 39% <strong>of</strong> the waterfowl nests<br />
built on the Sandusky Bay dikes in 1967<br />
and 1968 (Urban 1970).<br />
Raccoons are the most signlf icant<br />
furbearer In the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetlands<br />
Sn terms <strong>of</strong> pelt value <strong>of</strong> the total<br />
harvest. <strong>The</strong> density <strong>of</strong> raccoons in <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> wetlands has been estimated at<br />
about 0.3jhectare by Jaworskl and Raphael<br />
Figure 43. Raccoon (Procyon lotor), a (1978) <strong>The</strong>y fndlcate a value per pelt<br />
common predator <strong>of</strong> duck nests in <strong>Lake</strong> <strong>St</strong>. PIUS carcass <strong>of</strong> about $40.00 for the early<br />
<strong>Clair</strong> wetlands. to middle 1970s. <strong>The</strong>refore, the 38,000<br />
hectares <strong>of</strong> wetlands wfthfn the study area<br />
has a total raccoon value <strong>of</strong> about<br />
$456 r 000.
4.1 ORIGIN AND EVOLUTlOM OF LAKE ST.<br />
CLAlR WETLANDS<br />
<strong>The</strong> relationship <strong>of</strong> water levels,<br />
topography and the geomorphic devel opment<br />
<strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal zone is<br />
significant to b fogeographlcal development<br />
and maintenance <strong>of</strong> the wetlands and<br />
their areal development. This is particularly<br />
true for the <strong>St</strong>. <strong>Clair</strong> Deltar which<br />
is constantly changing its morphology.<br />
Because <strong>of</strong> shifting channels, variable<br />
flow regimes, and sedimentatfon, the<br />
geological nature, (and hence the<br />
b~ologlcal characteristic <strong>of</strong> the 1 f ttoral<br />
zone and wetlands) has not been constant.<br />
Sfnce not a11 rivers which debouch<br />
into a basin deposft deltas, the marine<br />
processes are slgniffcant. A delta is not<br />
solely a praduct <strong>of</strong> river deposition.<br />
Basicall yp del taf c 1 andforms are a product<br />
<strong>of</strong> both river and marine processes. <strong>The</strong><br />
rfver contributes the sediments to the<br />
receiving bas1 n. Whether the sediments<br />
accumulate in the mouth is dependent upon<br />
the marine processes such as wave action<br />
and long-shore currents? and the geo-<br />
morphlc character <strong>of</strong> the submarf ne topo-<br />
graphy. If there is a dominance <strong>of</strong> the<br />
marine process the sediments may be trans-<br />
ported down coast and a delta may not be<br />
extensive, Conversely? a rlver domlnated<br />
system contributing an excess <strong>of</strong> sediment<br />
coupled wjth a low wave climate w i l l<br />
prsmote deltaic development.<br />
Several <strong>of</strong> our concepts <strong>of</strong> delta<br />
development have been obtained from the<br />
M$ssiss$ppi deltafc plain, Because <strong>of</strong> <strong>of</strong>f<br />
and gas exploratfan, rfver and marine<br />
CQmm63rCer and f he extensive wet1 ands in<br />
the arear the Missfssippi River Delta has<br />
been extensive1 y studied geologicall yr<br />
blologica~lyr and geographically. It<br />
CHAPTER 4, ECOLOGllCAL PROCESSES<br />
serves as a model by which other marine<br />
and 1 acustrjne deltas are compared.<br />
Traditlonally deltas have been<br />
classified on the basis <strong>of</strong> their<br />
morphology and descriptive terms such as<br />
arcuate, b i rd-foot, and cuspate have been<br />
applied to the various delta shapes.<br />
Using this classification the <strong>St</strong>. <strong>Clair</strong>?<br />
like the Misslssippi Delta may be<br />
described as bird-foot, It is generally<br />
assumed that a given set <strong>of</strong> processes is<br />
responsible for each deltaic morphology.<br />
Although* morphological sf mil aritles are<br />
evident, significant differences also<br />
occur (Table 25).<br />
A unfque aspect <strong>of</strong> a delta's<br />
geomorphology is related to its changfng<br />
base levels. <strong>The</strong> base level <strong>of</strong> all marine<br />
deltas appears to have been eustatf call y<br />
stable over the past 4*000 years. In the<br />
Misslssippi Delta? because <strong>of</strong> the stable<br />
sea level? the ancestral streams deposited<br />
a serles <strong>of</strong> delta lobes along the<br />
Louisiana coast. A t <strong>Lake</strong> <strong>St</strong>. Clalr,<br />
changing base levels have produced two<br />
distinct deltas at different levels, the<br />
premodern and the modern. Examples <strong>of</strong><br />
this base level change are apparently rare<br />
with regard to deltaic deposition.<br />
Marine deltas are <strong>of</strong>ten characterized<br />
by subsidence due to sediment loading.<br />
Geosyncl i nal downwarping and sediment<br />
compaction in response to sediment ac-<br />
cumulation in the Mississippi Delta has<br />
allowed up to 122 m <strong>of</strong> deltaic deposits to<br />
accumulate since the termination <strong>of</strong> the<br />
Late Wisconsin glacial ice (Morgan 1970).<br />
With continued sedimentation and<br />
subsidence, the shtftjng delta lobes<br />
overlapped# burying older deltaic<br />
envf ronments. <strong>The</strong>reforer the traditional<br />
sedimentologlcal units, top-setr fore-set,
Table 25. Factors influencing lake and marine delta morphology. a<br />
Factors Characteristics <strong>St</strong>. <strong>Clair</strong> Delta Mississippi Delta<br />
Geomorphic: Base 1 eve1 Long- and short-term <strong>St</strong>able<br />
variations<br />
Flow regime Seasonally constant Seasonally variable<br />
Channel d iversl on Downstream (spring Wfthfn alluvial<br />
ice jams) valley (sprfng floods)<br />
<strong>St</strong>ructural behavior <strong>St</strong>able<br />
<strong>of</strong> basin <strong>of</strong> deposition<br />
Sediment compaction<br />
and significant<br />
subsidence<br />
Dominant sedlment size Fine to coarse sand Clay to silt<br />
Relative rate <strong>of</strong> S1 ow<br />
sedimentat ion<br />
Marine: Relative wave energy Low Low<br />
Rapid<br />
Offshore pr<strong>of</strong>ile Flat to gently Flat to gently<br />
slopfng sloping<br />
Vegetative: Vegetatlon Deciduous trees to Predominantly fresh<br />
zonation freshwater marsh to brackish to salt<br />
marsh<br />
Control <strong>of</strong> vegetation Short-term 1 ake level Salinity, substrate<br />
distribution changes composition, and tidal<br />
range<br />
a Data source: Jaworski and Raphael (1973).<br />
and bottom-set beds, identified on the<br />
margins <strong>of</strong> <strong>Lake</strong> Bonneville by Gilbert<br />
(1890) do not occur in the Mississfppi<br />
Delta complex.<br />
Subsurface data reveal that the <strong>St</strong>.<br />
<strong>Clair</strong> Delta is stable. Sediments ac-<br />
cumulating in the delta are not <strong>of</strong><br />
sufficient thickness to allow any<br />
significant downwarping to occur, Borfngs<br />
on several beaches suggest that the base<br />
<strong>of</strong> these depositional features Is at<br />
apgroxlmately low water datum. Apparently<br />
the beaches have not subsided as the<br />
cheniers have done on the flanks <strong>of</strong> the<br />
Mississippi Delta,<br />
Isostatic rebound i n the <strong>St</strong>. Clafr<br />
basin, due to the waning <strong>of</strong> Wisconsin ice<br />
fo1 lowing the Valders maximum some 10~500<br />
to 1Zt500 years ago, has been minlmaf ,<br />
Pertinent zero isobases since the Valders<br />
maximum ice advance are the Algonquin and<br />
Nipissing hinge 1 friest which are oriented<br />
east-west and located in the central<br />
portion <strong>of</strong> <strong>Lake</strong> Huron. <strong>The</strong>refore, crustal<br />
displacement due to regional glacial<br />
rebound or 1 ocal ired sed imen%atlcn does<br />
not appear to have direct1 y influenced the<br />
vertical and horizontal morphology <strong>of</strong> the<br />
<strong>St</strong>. Claf r Dslta, <strong>The</strong> <strong>St</strong>, <strong>Clair</strong> Delta has*<br />
in fact, extended itself across an es-<br />
sentiall y rlgid platform.
Although structural stabil tty and<br />
mfnimum subsfdence characterize the <strong>St</strong>.<br />
<strong>Clair</strong> Delta, significant changes in the<br />
level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> in the past few<br />
millenla have contributed to the mor-<br />
phology <strong>of</strong> the delta. Based upon a<br />
structural framework* deltas may be de-<br />
posited in one <strong>of</strong> three geologic<br />
envlronments: a subsiding area, a stable<br />
area, or an elevating area. <strong>The</strong><br />
Mtssissippf Delta with its thick ac-<br />
cumulation <strong>of</strong> recent sediments* is a<br />
classic example <strong>of</strong> a subsiding delta.<br />
Although the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin is<br />
geologically stable, the effect <strong>of</strong> a "tower<br />
lake level on the delta complex is similar<br />
to elevatlng the 1 and, Since base level<br />
in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has dropped in the 1 ast<br />
4,000 years, the focus <strong>of</strong> depostion has<br />
shifted lakeward creating the modern <strong>St</strong>,<br />
Cl ai r Delta and wetl and habitat.<br />
<strong>The</strong> configuration <strong>of</strong> deltas is<br />
related to <strong>of</strong>fshore slope" relattve wave<br />
power in the nearshore zone, and riverine<br />
Influence (Wrlght and Coleman 1972). A<br />
delta such as the Mississippi with its<br />
digltal distributaries and marshy<br />
embayments 1s dominated by Its rlver since<br />
Its <strong>of</strong>fshore slope is flat to convex and<br />
the nearshore wave power is low. Although<br />
long-term wave data have not been recorded<br />
for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, wave energy must be<br />
considered to be low since the maximum<br />
fetch for wave generation for the delta is<br />
only 40 km. <strong>The</strong> subaqueous delta front is<br />
shallow and concave, causing waves<br />
generated In the shallow lake to attenuate<br />
lakeward <strong>of</strong> the delta, Also, marshes have<br />
become establ ished on exposed <strong>of</strong>f shore<br />
areas between Johnson Bay and the -2 m<br />
contour suggesting that law wave energy<br />
conditfons prevafl, Although there may be<br />
an occasional dominance <strong>of</strong> martne<br />
pracessss as evidencad by the presence <strong>of</strong><br />
transgressive beaches, f t must be<br />
concluded that the geometry <strong>of</strong> the <strong>St</strong>.<br />
<strong>Clair</strong> Delta, like that <strong>of</strong> the Mississippi,<br />
is prfncfpally dTctated by Its rfver,<br />
<strong>The</strong> pkevtous discussion suggests that<br />
although the <strong>St</strong>, Clafr and Mississippi<br />
river deltas have similar appearances<br />
several dissfmf l ar features and p rocssses<br />
prevafl . For example, the domfnant<br />
sediment size in the Misslssfppi Delta 1s<br />
silt to clayo whereasr the <strong>St</strong>, <strong>Clair</strong> Delta<br />
Is fine t o coarse sand. Alsos the<br />
relative rate <strong>of</strong> sedfmentation in the <strong>St</strong>.<br />
Clafr Delta is slow compared to the rapid<br />
rate <strong>of</strong> the Misslssfppi Delta. <strong>The</strong><br />
wetl ands dlscussed in this community<br />
pr<strong>of</strong>ile exhibit a dfversity <strong>of</strong> form.<br />
However, thef r distributfon can be re1 ated<br />
to fundamental geomorphic features some <strong>of</strong><br />
which are unique. Jaworski et a1. (1979)<br />
examined and developed geomorp hic model s<br />
for the coastal wetlands <strong>of</strong> the Great<br />
<strong>Lake</strong>s as a step to understanding their<br />
origin, stability, and distribution.<br />
<strong>The</strong> occurrence, distrfbution, and<br />
diversity <strong>of</strong> coastal wetlands are, in<br />
part, determined by the morphology <strong>of</strong><br />
coastal <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Perhaps in no<br />
other b f ogeograp hic envi ronment i s the<br />
relationship between 1 and forms and<br />
vegetation so evident. All wetlands occur<br />
in topographical depressions created by<br />
nature or more recently by humans. <strong>The</strong>se<br />
topographical lows may be erosional fn<br />
origin such as river channels or<br />
depositional fn origin such as lagoons or<br />
floodbasins.<br />
Although elevations <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
Delta are low, a diversity <strong>of</strong> landforms<br />
exists (Sectlon 2.1) . When the entire<br />
shorel ine <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air is considered,<br />
the habitat diversity is even greater.<br />
Twelve wetl and sett 1 ngs have been<br />
identified in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Figure 44).<br />
A l l but one s e t t i n g (f.e.,<br />
diked/disturbed) are <strong>of</strong> a natural origin.<br />
<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta exhibits a greater<br />
diversity <strong>of</strong> wetland settings because it<br />
is a product <strong>of</strong> river as we71 as marine<br />
processes. <strong>The</strong> habitats beyond the<br />
deltaic plain are fewer in number but<br />
unfque to the lake shoreline. A third<br />
suite <strong>of</strong> wet1a:jd habltats occurs both In<br />
the delta complex and on the shorel jne.<br />
<strong>The</strong> followfng section discusses the<br />
12 wetland habitats recognized in <strong>Lake</strong> <strong>St</strong>.<br />
Claf r. <strong>The</strong> deltaic habitats are descrfbed<br />
first followed by the coastal envi ronmsnts<br />
<strong>of</strong> wetland occurrence. Finally, wetl ands<br />
occurrfng in both the <strong>St</strong>. Clafr deltaic<br />
and coastal plains are described:<br />
1, are shall ow submerged<br />
features <strong>of</strong> the df stributary channels
1. River shoulders<br />
<strong>Lake</strong><br />
4. Crevasse deposits<br />
5. Shallow interdistributary bays<br />
6. Deep water interdistributary bays<br />
7. Transgressive beaches<br />
10. Transitional areas<br />
Premodern<br />
2. Abandoned channels<br />
8. Shallow shelves<br />
9. Barrierllagoons<br />
<strong>Lake</strong><br />
Plain<br />
12. Diked areas<br />
Figure 44, Twelve wetland habitats recognized in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> De1 ta. Paralleling<br />
the shore1 ine <strong>of</strong> the channels, the<br />
submerged river shoals extend 35 m into<br />
the rlver and have maximum depths <strong>of</strong> 2<br />
m. <strong>The</strong> sediment consists <strong>of</strong> fine sand.<br />
<strong>The</strong> feature is reasonably continuous<br />
along the length <strong>of</strong> the channels but fs<br />
normally absent on the outsfde or<br />
cutbank side <strong>of</strong> the channel. <strong>The</strong><br />
feature attains a maximum width at<br />
submerged point bars. At the channel<br />
shoreline, reeds such as the robust<br />
are dense (Figure<br />
29); however, submergent aquatics are<br />
dominant. An aggressive fnvader,<br />
Euraslan water-mll fol 1 has become more<br />
abundant in the last decade in the<br />
distributary channels (Schloesser and<br />
1 1. Eustuarinelflood<br />
basin<br />
Manny 19823, It appears to be a common<br />
submersed macrophyte and a minor<br />
nuisance to recreational boating In the<br />
delta region as well as in Anchor Bay,<br />
Furthermore, the plant denslty appears<br />
to be greater in the non-commercial<br />
navigation channels (i.e., North and<br />
Middl e channels).<br />
2. dissect the<br />
up1 and surface <strong>of</strong> the delta and merge<br />
into the modern delta when traced<br />
lakeward. <strong>The</strong> channels are long and<br />
linear and <strong>of</strong> varfable width. <strong>The</strong><br />
1 argest channel on Dickfnson Island is<br />
some 500 m wjde, but usual channel<br />
wfdths are about 100 m. <strong>The</strong> sediments<br />
have a fine texture and a high organfc
content suggesting gradual decay over<br />
time, <strong>The</strong> boundary between the<br />
premodern delta upland and the<br />
abandoned channel is usual1 y distinct<br />
especial1 y at the apex <strong>of</strong> the delta.<br />
During years <strong>of</strong> low lake level several<br />
<strong>of</strong> the ancient channels are above the<br />
water table and plant stress may occur.<br />
Emergents such as hard-stem bul rush<br />
(Scfrow+ -1, arrowhead (<br />
lati f ol la), and three-square<br />
colonize the deeper water<br />
nels (1 m) as do water<br />
ccasionally buttonbush<br />
3. are generally<br />
similar to the abandoned<br />
channels discussed except that they are<br />
st111 in the process <strong>of</strong> decay.<br />
Chematogen Channel 3s a dlstrlbutary<br />
channel which contfnues to conduct <strong>St</strong>.<br />
Clalr Rlver water to <strong>Lake</strong> <strong>St</strong>. Clalr.<br />
<strong>The</strong>se channels are malnl~ on the<br />
Ontario sido <strong>of</strong> the delta ind were<br />
responslble for the depositfon <strong>of</strong> the<br />
Canadian portion <strong>of</strong> the delta. Since<br />
that time this area was abandoned and<br />
the more recent depositlonal act iv'i ty<br />
began in Michfgan. As abandonment<br />
occurred, the channels were gradual 1 y<br />
s l l t e d so that only narrow<br />
distributaries remain today. Such<br />
channels not only dlssect the premodern<br />
delta but the modern surface as well.<br />
<strong>The</strong> sedlment <strong>of</strong> the channel fflls .Is<br />
fine sand, but some silts have been<br />
deposited from the adjacent 1 ake pl afns<br />
as well. Since these channels occur on<br />
the modern delta, slopes are more<br />
gentle. In contrast abandoned river<br />
channels are on the premodern surface<br />
and slopes to the channels more<br />
distinct. <strong>The</strong> channel fills <strong>of</strong> the<br />
recent1 y abandoned channels are <strong>of</strong>ten<br />
submerged beneath 15 cm <strong>of</strong> water and<br />
colonized by sedges E- spp.1 and<br />
more sporad?call y by blue-joint grass<br />
spp. and pondweeds (e.g., curly<br />
pondwsed ~ r i s ~ colon9 u ) ze<br />
the narrowfng active channels.<br />
occur on the f7anks<br />
<strong>of</strong> dfstrlbutary bays and have a lobate<br />
form, Since they are flood or hlgher<br />
energy deposits they are composed <strong>of</strong><br />
coarser sand. Channel depths are<br />
approximatel Y 4 m and are not<br />
COI onjzed, but the sand deposits<br />
adjacent to the "highwaysn support<br />
commun itf es <strong>of</strong> bul rush ~PP. 1.<br />
Most crevasse deposits are at the lower<br />
end <strong>of</strong> the active dfstributary channels<br />
and thus exposed to wave act Ion.<br />
Because <strong>of</strong> higher wave energies and<br />
perhaps due t o Ice damage and<br />
recreational boating the emergent<br />
aquatics are not particularly dense,<br />
<strong>The</strong> landward side <strong>of</strong> the crevasse<br />
deposfts appear to be better protected<br />
from wave action and thus support a<br />
denser stand <strong>of</strong> emergent vegetation<br />
including not only bulrush fn deeper<br />
habitats but cattails (Iy&h spp.) in<br />
shallower areas <strong>of</strong> the deposit. Also<br />
crevasse deposits closer to <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> (farther down the distributary<br />
channels) are exposed to greater wave<br />
actfon and are sparsely colonized with<br />
bulrush.<br />
aallow interdlstributgILy_ bavg occupy<br />
extensive areas between the open lake<br />
and the equally obvious higher<br />
premodern deltas. On the flanks these<br />
shallow wetland hab i tats are bordered<br />
by natural levees. Occasionally,<br />
particularly in Michigan, narrow<br />
discontinuous beaches occur. <strong>The</strong><br />
sediment In the bays was derived from<br />
crevasses during high water periods and<br />
are thus silty-sand size particles and<br />
mostly mineral in character. Organic<br />
sedfmsnts are sparse and discontinuous.<br />
A t the surface the organic<br />
accumulatlonr where present, ranges in<br />
thickness from 5 to 15 cm. Water<br />
depths average 5 to 30 cm. <strong>The</strong> shallow<br />
Interdlstrfbutary bays are almost<br />
excl us I vel y col on i zed by dense stands<br />
<strong>of</strong> cattails <strong>of</strong> which the hybrid TvDha x<br />
standlng 2 to 3 m high<br />
dominates. A t slightly higher<br />
elevations sedges are interspersed with<br />
cattafls. <strong>The</strong> high cattail produc-<br />
tivity and the low organic accumulation<br />
suggests that these habitats are<br />
flushed durJng high water periods. <strong>The</strong><br />
sedimentation process is relatively<br />
complete throughout the deltaic plain,<br />
though a few small lakes (e.g., Goose<br />
lake) stfll persjst.
interdfstrf butary bays. <strong>The</strong> landward<br />
slde is bordered by low, poorly<br />
developed transgressive beaches. <strong>The</strong>ir<br />
flanks are characterized by slightly<br />
higher, poor1 y developed natural levees<br />
which are permanently breached by<br />
crevasse channels. <strong>Lake</strong>ward these bays<br />
are exposed to wave action <strong>of</strong> the open<br />
lake. As clastic sediments are<br />
introduced into these bays, the basins<br />
w i l l evolve Into shallow interdistributary<br />
bays. Water depths range<br />
from 0.5 to 1.75 m and the bay bottom<br />
consists <strong>of</strong> fine sand. Occasionally<br />
in sheltered areas si 1 t deposits w i 11<br />
accumulate, but organic deposits are<br />
sparse through the habitat. Wave<br />
action prohibits the colonization <strong>of</strong><br />
dense emergents; however, n! spp.<br />
are prevalent. <strong>The</strong> deep water bays are<br />
confined to the Michigan portion <strong>of</strong> the<br />
de1 ta where the distributary channels<br />
are most active and where natural<br />
levees extend into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Common submersed aquatics include<br />
Characeae and f f s<br />
(Schloesser and Manny 1982). <strong>The</strong><br />
d istributjon <strong>of</strong> submersed macrophytes<br />
may be due to a lack <strong>of</strong> suitable<br />
substrate for attachment or excessive<br />
wave actlon.<br />
are best<br />
developed wlthin the <strong>St</strong>. <strong>Clair</strong> Delta<br />
but do sporadically occur along the<br />
Ontario shore1 ins. <strong>The</strong> beaches are<br />
wave deposited features standing<br />
approximately 30 cm above the lake. As<br />
noted in Figure 4, older beach deposits<br />
occur landward <strong>of</strong> the active<br />
transgressive beach in the shallow<br />
i nterdistributary bays. <strong>The</strong>y are<br />
composed <strong>of</strong> stratified coarse sand,<br />
gravel and organic debris. Although<br />
they vary in width, most beaches are<br />
about 5 m wfde. Borings through the<br />
beach deposits reveal alternating<br />
layers <strong>of</strong> coarse clastics and thin (1<br />
cm) wave deposited organic 1 enses.<br />
This clearly suggests that the features<br />
were deposited during periods <strong>of</strong><br />
erosion. During higher wave energy<br />
conditions the beaches are not<br />
breached; but rather overwash occurs,<br />
causing rafted debris to be deposited<br />
atop the shallow interdistributary<br />
marsh. Woody vegetation includfng<br />
eastern cottonwood (P-, 1,<br />
7. *<br />
willows (531 i x spp.) and staghorn sumac<br />
(m colonize the beaches.<br />
During higher water years sedges (Carex<br />
strictq) r reed-canary grass (<br />
udinace_a), and jewelweed (-<br />
spp.) commonly colonize the beaches.<br />
<strong>The</strong> latter three species are more<br />
common on the overwash deposits<br />
landward <strong>of</strong> the beaches.<br />
8. Shallow shelf enviroqg&i& border the<br />
nearshore zone <strong>of</strong> most <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>. <strong>The</strong> shelves are composed <strong>of</strong><br />
silt-sized clastic fragments. <strong>The</strong><br />
shelves are approximately 200 m wide,<br />
1.5 m fn depth and are turbid<br />
especially following storms. <strong>Wetlands</strong><br />
colonizing this habitat exist as a low<br />
dense fringe parallel to the shore and<br />
remaln exposed to lake effects such as<br />
wave action. This habitat is best<br />
displayed along the eastern shore <strong>of</strong><br />
the lake and is colonized wfth<br />
emergents and submergents. <strong>The</strong><br />
dominant vegetation is emergent<br />
macrophytes such as cattail. Other<br />
wetland species include lake sedge<br />
(Carex -1, large stands <strong>of</strong><br />
pickerel weed (Wderia EPCP-) ,<br />
water-mil foil ( k l v r l m<br />
hetero-<br />
-1, and white water lily<br />
(Nvmr>haea odorata) (Mudroch 19811,<br />
9, Barrier/l a-lexs~ occur south <strong>of</strong><br />
Mftchell Bay along the eastern<br />
shoreline <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong><br />
barriers are poorly developed and stand<br />
approximately 50 cm above the lake.<br />
However, they do provide some<br />
protection from wave action and<br />
seiches, especially during 1 ower water<br />
years. <strong>The</strong> lagoon is approximately<br />
1,000 m in width and i s characterized<br />
by a si 1ty bottom. Morphologically<br />
this habitat is similar to the shallow<br />
she1 f environment except for the<br />
occurrence <strong>of</strong> a low barrier, <strong>The</strong><br />
bottom sediments not only Include silts<br />
but 1 oose, partial 1 y decomposed organic<br />
fragments which may be j.~ or may<br />
have been transported into the lagoon<br />
by streams from the adjacent lake<br />
plain. Emergent macrophytes dominated<br />
by cattalls ( 1 occur in<br />
the area. <strong>The</strong> landward side <strong>of</strong> these<br />
wetlands has a gentle natural slope<br />
which in most fnstances along <strong>Lake</strong> <strong>St</strong>,<br />
<strong>Clair</strong> is abrupt1 y termfnated by
artificial dikes. It is important to<br />
note that the Ontario shoreline north<br />
<strong>of</strong> the Thames River has not experlenced<br />
signlf lcant coastal recession in recent<br />
years (Boulden 1975) whereas the south<br />
shore <strong>of</strong> the 1 ake has. This suggests<br />
that the barrier and adjacent wetlands<br />
retard wave action on this coastal<br />
reach.<br />
10. are broad zones<br />
i a1 wetl ands from<br />
well-drained upland surfaces. In some<br />
instances, such as on Harsens Island<br />
and along the eastern shoreline <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>r the transitional areas have<br />
been incorporated in diked farmland or<br />
water level-control led wetl ands and<br />
thus do not exist. However, good but<br />
rather iselated examples occur on<br />
Dickinson Island and east <strong>of</strong> <strong>St</strong>. John's<br />
Marsh in -bIichigan. <strong>The</strong> sediments<br />
consist <strong>of</strong> fibrous peat macr<strong>of</strong> ragments<br />
which are underlain by fine# mottled<br />
sand. <strong>The</strong> transition zone is normal 1 y<br />
not permanent1 y flooded and water<br />
tables are about 15 to 30 cm below the<br />
surface. <strong>The</strong> lakeward slde <strong>of</strong> this<br />
environment Is characterized by sedge<br />
sediments are derived from flood events<br />
and are usually fine sand and stlt wlth<br />
a sign1 f icant organic content. Natural<br />
levees adjacent to the channel are<br />
poorly developed and vegetation<br />
zonation difficult to detect. <strong>The</strong><br />
water level -in this wetland habitat is<br />
vartable and depends In part on the<br />
rdver dfscharge. Durlng dry years the<br />
wetlands may be stressed because <strong>of</strong> a<br />
low water table. <strong>The</strong>refore the wetland<br />
may be seasonal in nature. A t the<br />
confluence <strong>of</strong> river and lake, water<br />
tables are most <strong>of</strong>ten at or very near<br />
the surface and the floodbasin w i l l<br />
sustain dense emergents in most years.<br />
At the mouth <strong>of</strong> the Clinton Rlver dense<br />
stands <strong>of</strong> cattails have been observed<br />
for many years. Upstream the marshes<br />
grade into bottom1 and swamps.<br />
-.<br />
12. are a cultural modification<br />
<strong>of</strong> a wetland rather than a<br />
physical entity. Included in this<br />
category are transitional zones which<br />
have been dfked and drained for farming<br />
or recreational purposes. Since these<br />
tussocks (u spp.) interspersed with<br />
sparse cattails. Where water tables<br />
wetlands transcend physical boundaries<br />
such as soil/sediment types and slopest<br />
their morphologfcal framework is less<br />
are deeper, the transition zone is meaningful. On Harsens Island the<br />
colonized by a meadow which includes Michigan Department <strong>of</strong> Natural<br />
but is not limited to red osier dogwood<br />
(Carnus &lonifera)r gray dogwood (C.<br />
Resources has diked and drained much <strong>of</strong><br />
the shallow distributary bay area and<br />
1, panic grass (j2gmiam spp.1. the transition zone. <strong>The</strong> land cover <strong>of</strong><br />
and bluejoint grass. On the landward the central portion <strong>of</strong> the island is<br />
embl ing aspen (m in corn and grain crops which are utilis<br />
abundant. Because <strong>of</strong> ized as part <strong>of</strong> a waterfowl hunting<br />
the variable elevation and depth to<br />
groundwater thJs zone exhibits a<br />
management scheme. Wal pole Is1 and is<br />
the largest island in the delta. A<br />
diversity <strong>of</strong> wetl and plants. Similar large part <strong>of</strong> the island, which is now<br />
wetland species have been observed on<br />
the premodern surface <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
cultivated mainly for corn and soybean,<br />
was a1 so a shallow interdisciplinary<br />
Delta. Here local depressfons are bay. <strong>The</strong> diked marshes along the east<br />
underlain by clay pan which results in<br />
a perched water table which commonly<br />
shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> have long been<br />
used for private duck hunting. Wasupports<br />
dogwood/meadow vegetation,<br />
ter level in these marshes is regulated<br />
by pumps, dikes, and canals. <strong>The</strong><br />
fl, habltats are<br />
characterized by marsh vegetation<br />
marshes are landlocked and water depths<br />
range from 10 to 50 cm (Mudroch 1981).<br />
bordering a rfver which debouches into<br />
the lake, <strong>The</strong> Clinton River and<br />
Emergents colonize these marshes. Dominant<br />
macrophytes are the cattail and<br />
several small streams tributary to <strong>Lake</strong> sedge. Also large stands <strong>of</strong> pickerel<br />
<strong>St</strong>. Clafr support such wetlands, <strong>The</strong><br />
extent <strong>of</strong> the vegetated wetland is<br />
weed, water mil foil (M rio hyllum<br />
heteropbyll um) , and white*<br />
<strong>of</strong>ten restricted by a steep backslope occur in the channels within these<br />
paralleling the flood plain. <strong>The</strong> diked wetl ands.
Table 26 summarizes the identified<br />
morphological wetland settings Sn <strong>Lake</strong> <strong>St</strong>.<br />
Clalr in terms <strong>of</strong> selected physical<br />
characteristics, For thf s investigation<br />
the coastal and deltaic wetlands are<br />
classified into four major vegetative<br />
zones:<br />
1. Submergent (e.g., pondweeds,<br />
coontai 1 1<br />
2, Emergent (e.g.$ cattail, bulrush)<br />
3. Sedge/meadow (e.g. sedge* canary<br />
grass)<br />
4. Shrub (e.g., dogwood, willow, sweet<br />
gal el.<br />
Wetland diversity exhibited by submergent,<br />
sedgep and shrub vegetation is most<br />
evident in the abandoned channels (2) and<br />
the transitional (10) zones. Here,<br />
topography is variable, wave or river<br />
current action is low or not applicable<br />
and turbidity is low. Conversely the<br />
wetl and community is less diverse along<br />
river shoulders (l), crevasse (4), and<br />
estuarine/flood basfn (11) environments.<br />
<strong>The</strong>se rather monotypfc wetlands appear to<br />
be assocfated with hiqher wave/current<br />
action and higher turbidity levels. <strong>The</strong><br />
exception is the shallow interdistributary<br />
bay (5) environment which is composed <strong>of</strong><br />
dense and robust stands <strong>of</strong> cattails.<br />
Perhaps the combined perfodic exposure and<br />
a hlgher organic content <strong>of</strong> the substrate<br />
account for the dominance <strong>of</strong> the<br />
emergents.<br />
4.2 BlOLOGlCAL PRODUCTION<br />
Examination <strong>of</strong> energy flow in coastal<br />
wetlands shows that the entire<br />
heterotrophic component <strong>of</strong> the wetland Is<br />
dependent on organic matter produced<br />
during photosynthesis. In many cases,<br />
utilization <strong>of</strong> the material is through<br />
grazing <strong>of</strong> lfving tissues. Many species<br />
<strong>of</strong> waterfowl feed on various parts <strong>of</strong><br />
aquatfc hydrophytes. <strong>The</strong> seeds <strong>of</strong><br />
muskrat, mice, and deer browse heavily on<br />
several species <strong>of</strong> aquatic hydrophytes.<br />
Phytophagous fnsects are the food supply<br />
for several species <strong>of</strong> birds, fishes,<br />
reptiles, and mammals (Tilton et ale<br />
1978).<br />
A more significant form <strong>of</strong><br />
uttlization is the direct consumption <strong>of</strong><br />
detritus or alternatively, the microbial<br />
populations associ ated with organic<br />
particulate matter. Numerous inverte-<br />
brates re1 y completely on these food<br />
sources. <strong>The</strong>se organisms in turn form the<br />
food supply for fish, amphibians,<br />
reptiles, birds, and mammals.<br />
In addition to plants being a direct<br />
or indirect food source for many species<br />
<strong>of</strong> animals, aquatic plants provSde cover<br />
and nesting areas for waterfowl. Several<br />
species <strong>of</strong> fish9 such as the northern pike<br />
(b 3 uc i spawn in vegetated wetl and<br />
areas and muskrats (w ðiu)<br />
prefer emergent vegetation for construction<br />
<strong>of</strong> feeding platforms and houses.<br />
Nutrient cycl ing is important to the<br />
continuation <strong>of</strong> primary and secondary<br />
production in wetlands (Figure 45).<br />
Coastal wetland vegetation immobilizes<br />
certain amounts <strong>of</strong> nutrients, a portlon <strong>of</strong><br />
which are released upon senescence and<br />
decay <strong>of</strong> the plants. Depending on the<br />
sedimentation characteristlcs <strong>of</strong> the<br />
wetlandsr nutrients are stored in the<br />
wetland as organic sediment or peat. In<br />
the marsh sol 1 sr microb 1 a1 processes<br />
transform some <strong>of</strong> the nutrients from<br />
organfc to inorganic forms. <strong>The</strong> net<br />
effect <strong>of</strong> these processes is generally a<br />
reduction In the concentration <strong>of</strong><br />
nutrients in water flowing through the<br />
wetl and. <strong>The</strong>refore, the coastal marshes<br />
are important in control 1 i ng nutrient<br />
loading to nearshore waters <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
Clai r (Tilton et a?. 1978).<br />
<strong>St</strong>udies <strong>of</strong> metals uptake by aquatic<br />
plants provide some information on<br />
nutrient cycling. Mudroch and Capobianco<br />
(1978) studied the uptake <strong>of</strong> metals from<br />
the water and sedtrnent by emergent,<br />
submersedl and floatfng-leaved plants fn<br />
marshes along the Ontario shore <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. Cl a i r. <strong>The</strong>y found that submersed
Table 26. Morphology and relative physical relationships <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />
wetlands.<br />
Wet1 and Water Dominant Current/<br />
envl ronment Substrate deptha vegetatf on wave actionb Turbidfty<br />
(m)<br />
1. River shoulders sand -6.0 submergent high/l ow moderate<br />
2. Abandoned river silt/ -1.0 emergent/ 1 ow/NA 1 ow<br />
channel s organics subme rgent/<br />
sedge/scrub<br />
3. Recent1 y aban- silt/ -0.5 emergent/sedge med/NA high<br />
dwned channel s organics<br />
4. Crevasse deposlts sand -1.5 emergent high/med high<br />
5. Shallow intar- fine sand/ -0.3 emergent 1 ow/low 1 ow<br />
distributary bays arganics<br />
6, Deep water inter- f Ins sand -1.5 emergent/ high/hfgh high<br />
distributary bays 5u bme rgent<br />
7. Transgrasslve coarse i-0.5 sedge/scrub low/high N A<br />
beach deposlts sand/<br />
organics<br />
8. Shal low shelves silt/cl ay -1.0 emergent/ low/high htgh<br />
submergent<br />
9. Barrler/lagoon si lt/cl ay -1.0 emergent/ low/Jow hlgh<br />
compl ex submergent<br />
10. Transi tlonal fine rand +0.3 emergent/ NA/NA N A<br />
areas submergent<br />
11, Estuarfne<br />
floodbasin<br />
sand/s i 7 t +O. 2 emergent high/NA high<br />
12. Diked areas variable -0.1 emergent/ low/l ow moderate<br />
submergent<br />
" Below level <strong>of</strong> lake<br />
+ Feature usually above lakt3 level.<br />
NA Not appl fcable.
plants accumulated the highest concen-<br />
trations, considerably higher than the<br />
source water and sediment, and that these<br />
elements were retained in the plant<br />
tfssues until decay processes incorporated<br />
them into surface sediments.<br />
1. ENERQV 3. WATER 6. SEDIMENTS<br />
2. GASES 4. NUTRIENT$ 8. ANIMALS<br />
INVERTEBRATE<br />
Figure 45. Cycling <strong>of</strong> energy and material<br />
through the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal wetland<br />
ecosystem (Ti1 ton et a1 . 1978).<br />
Mudroch (1980) also studied the<br />
geochemical composition <strong>of</strong> sediment,<br />
uptake <strong>of</strong> nutrients and metals by<br />
macrophytes, and nutrient and metal<br />
composition in the water at Big Creek<br />
Marsh on the northeast shore <strong>of</strong> <strong>Lake</strong> Erie<br />
in Long Point Bay. <strong>The</strong> maximum<br />
concentrations <strong>of</strong> most metals (Pb, Nip Car<br />
Crr and Zn) in marsh sediments were loner<br />
than concentrations found in <strong>Lake</strong> Erje<br />
surf ici a1 muds, presumably due to uptake<br />
by the aquatic plants In the mud,<br />
Submerged plants (Ghim Spet Mvr1clgMlhm<br />
sp., and sp.) accumulated larger<br />
quantities <strong>of</strong> Cat Pbr Cur Nf, Cr, and Cd<br />
than emergent plants (m spp.1, but<br />
nutrfent concentrations were relatively<br />
uniform for all species (Table 27). She<br />
found that the biomass production <strong>of</strong> the<br />
macrophytes In the marsh was related to<br />
the subhydric soil fertility as we11 as<br />
the nutrient content in the marshwater.<br />
She concluded that short-term supplies <strong>of</strong><br />
nutrient-rich sewage to the surface<br />
subhydrlc soil layer can have a prolonged<br />
effect on macrophyte production by<br />
enrichment <strong>of</strong> the nutrient pool maintained<br />
In the perennial plant system.<br />
<strong>The</strong> flow <strong>of</strong> energy through a food<br />
chain is <strong>of</strong>ten represented by a pyramid<br />
which illustrates the quantitative<br />
relatfonshfps among the varfous trophic<br />
levels (Figure 46). Juday (1943) was one<br />
<strong>of</strong> the earliest investigators to Introduce<br />
trophic levels concepts, developed the<br />
Table 27. Concentrations <strong>of</strong> nutrients and metals in wetland plarlts at the<br />
stage <strong>of</strong> maximum development, Big Creek Marsh, Long Point, Ontario.<br />
Species<br />
a Data source: Mudroch (19801,<br />
-<br />
K N P C a<br />
-- - - .-. - -----
'-\<br />
--._--_------SOLAR ENERGY<br />
Figure 46. Energy pyramid in <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> coastal marshes (Upper Great <strong>Lake</strong>s<br />
Regional Commission).<br />
concept from studies <strong>of</strong> freshwater<br />
wetlands. He determined the vardous<br />
components <strong>of</strong> the aquatic population in<br />
Weber <strong>Lake</strong>, Wisconsinr as they existed in<br />
midsummer. <strong>The</strong> dissolved organic matter<br />
composed about 60% o f the total pyramid,<br />
the fish9 only 0.5Xr and the other animals<br />
slightly less than 5% <strong>of</strong> the total.<br />
In coastal marshes there are <strong>of</strong>ten<br />
four major sources <strong>of</strong> energy for aquatic<br />
consumers: 1) marsh detritus, 2) phyto-<br />
plankton production, 3) detritus from<br />
terrestrial sources brought in by drainage<br />
and, 4) planktonic material carried into<br />
the marsh from the open lake, Although<br />
much research remains to be done on food<br />
chain production and ecosystem energy<br />
re1 ationshi ps9 particular1 y <strong>of</strong> freshwater<br />
wetlandsr there are a few general<br />
principles which appear to have validity:<br />
PI food cycles rarely have more than five<br />
trophic levels, 2) the greater the<br />
separation <strong>of</strong> an organism from the basic<br />
source <strong>of</strong> energy (solar radiation), the<br />
less the chance that it w i l l depend solely<br />
upon the precedjng trophic 1 eve1 for<br />
energy* 3) at successively higher levels<br />
in the food cycle, consumers seem to be<br />
progressive1 y more efficient 4n the<br />
utilization <strong>of</strong> food supply, and 4) In<br />
wetl and successl on productivity and<br />
photosynthetic efficiency increase Prom<br />
01 igotrophy to eutrophy and then decl ins<br />
as the marsh undergoes senescence<br />
(Jaworski et ale 19779.<br />
A1 though no specf fjc works have been<br />
published on the fntricats structure <strong>of</strong><br />
energy flow in Great <strong>Lake</strong>s coastal<br />
wet1 ands* Ti %ton 6% a1 , (1978) have<br />
general ized the important processes based<br />
on studjes in other wetlands'. <strong>The</strong><br />
conversion <strong>of</strong> solar energy -into bjomass by<br />
autotsophs is perhaps the most important<br />
process. Conversion into a form avail able<br />
to heterotrophic organisms serves as the<br />
foundation for several complex and dynamic<br />
food webs.<br />
<strong>The</strong> coastal wetland community<br />
possesses two basic complexes <strong>of</strong><br />
interrelationships: 1) invertebrates ( primarily<br />
insects)o fishes, bf rdsr and<br />
mammals which util ize 1 ivi ng p7 ant tissues<br />
and 2) organisms which utilize detrftus or<br />
dead plant tissues, Living plant tissue<br />
(7 .e., algae, 'leaves, and<br />
rhizomes) serve as food for phytophagous<br />
animals such as stem boring and leaf<br />
mining insects as well as certain aphids<br />
and beetles. Many specfes <strong>of</strong> waterfowl<br />
graze extensively on plant material r and<br />
muskrats are important pl ant consumers.<br />
<strong>The</strong> next higher trophic level in the ftrst<br />
complex consists <strong>of</strong> animals whlch prey<br />
upon the phytophagous organisms. Spiders,<br />
predatory beetles, dragonfllesr certsln<br />
fishes, frogs, birdsr and small<br />
insectivorous mammals are important<br />
organisms <strong>of</strong> this upper trophic level.<br />
<strong>The</strong> second complex consists <strong>of</strong> a vast<br />
number <strong>of</strong> insect larvae which rely on<br />
organic detritus as a dlrect energy source<br />
or by stripping microbial populations from<br />
the surface <strong>of</strong> organic particles,<br />
Gastropods and annelids are also important<br />
organ isms in the detritophagous complex.<br />
Whatever residual that is not util fzed by<br />
these animals is subjected to further<br />
decomposition by bacterf a1 and fungal<br />
populations, As with the phytophagous<br />
complexr there exists in the detritophagous<br />
complex a wide spectrum <strong>of</strong> animals<br />
which prey on the detritus-feeding<br />
organisms. Several species <strong>of</strong> insects,<br />
amphibians# waterfowlr and mammals compose<br />
this level, and many <strong>of</strong> these species are<br />
not selective in thelr prey* utilizing<br />
organ f sms from both compl exes.<br />
Teal (19629 found that a smaller<br />
percentage <strong>of</strong> the total energy represented<br />
f n the primary product 1 on <strong>of</strong> wetl ands
passes through the phytophagous complex<br />
than the detritophagous complex.<br />
Submergent vegetation tends to be<br />
inhabited and grazed more heavily than<br />
emergent forms (Krecker 1939) because the<br />
submergent aquatic macrophytes lack the<br />
more impenetrable structural tissues<br />
prevalent in the emergent type. Estimates<br />
<strong>of</strong> the proportlon <strong>of</strong> material processed<br />
through each complex (Ti lton et a]. 19781,<br />
favors the detritus web (80% t o 95%) over<br />
the grazing web (5% to 20% 1. Ti 1 ton et<br />
a1 . (1978) a1 so point out that the<br />
Importance <strong>of</strong> an organism to an ecosystem<br />
may exceed its role in energy flow,<br />
Muskrats utilize only a fraction <strong>of</strong> the<br />
available energy stored in the 1 lve plants<br />
they cut and harvest In the wetlands* but<br />
they may be <strong>of</strong> signiffcant value to the<br />
detritophagous complex. Similarlyo the<br />
teeming populations <strong>of</strong> phytophagous<br />
insects may consume on1 y a small fraction<br />
<strong>of</strong> plant tissuet but through this activity<br />
may reduce the growth <strong>of</strong> host plants and<br />
the primary production <strong>of</strong> the wetland<br />
ecosystem.<br />
<strong>The</strong> energy budget for cattai 1 (Iyehg<br />
sp. marshes in Minnesota has been<br />
investigated by Bray (1962) and may<br />
provide insight for <strong>Lake</strong> <strong>St</strong>. Clafr<br />
wetlands. During the growing season the<br />
distribution <strong>of</strong> the various components <strong>of</strong><br />
solar radiation energy was as follows:<br />
a1 bedo (reflection1 22.0<br />
evapotranspiration 38.4<br />
conduction-convectlon 38.5<br />
primary production 1.1<br />
Thf s apparently 1 ow ut 11 f zat i on <strong>of</strong> sol ar<br />
radiation by sp. is cons'lstent wlth<br />
other wetland studfes and supports the<br />
general view that most plant communltfes<br />
utilize only 1% to 2% <strong>of</strong> the total solar<br />
energy for primary production (Ttlton el<br />
al, 1978). sp. In Austrian<br />
wetlands ranges from 1-24 t o 2.0% for Ma<br />
through July (Sieghardt 1973) and<br />
sp. in Georgfan estuarfes utilizes 1,4% <strong>of</strong><br />
total solar radiation (Tea? 19621, <strong>The</strong><br />
uni formity <strong>of</strong> these results suggests that<br />
emergent wetlands in the take <strong>St</strong>. Claf r<br />
system utfl ize approximately 1.2% <strong>of</strong> solar<br />
radiation for prdmary production.<br />
Emergent wet1 and communities are<br />
among the most productive areas on earth.<br />
West1 ake (1963) estimated that freshwater<br />
emergent macrophytes have a net primary<br />
productivity <strong>of</strong> 3,000 to 8,500 g/rnZ/yr<br />
comparable in productivity to salt marshes<br />
and tropical rajn forests. Productivity<br />
<strong>of</strong> freshwater submerged macrophytes ranges<br />
from 400 to 2,000 g/m2/yrr less than ha1 f<br />
<strong>of</strong> thefr marine counterpart. Site<br />
specif ic primary productivfty studies <strong>of</strong><br />
coastal emergent wetlands in <strong>Lake</strong> <strong>St</strong>,<br />
<strong>Clair</strong> do not exist. Table 28 lists the<br />
maximum blomass and productivity <strong>of</strong><br />
several emergent species (common in <strong>Lake</strong><br />
<strong>St</strong>. Clalr) for other areas <strong>of</strong> the Great<br />
<strong>Lake</strong>s region. Extrapolation <strong>of</strong> these data<br />
to the cattail marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r<br />
can provide an approximate and reasonable<br />
estimate <strong>of</strong> product lvi ty, Jv~ha marshes<br />
appear to be one <strong>of</strong> the most productive<br />
communities, yielding vaf ues (approxfmately<br />
2,500 g/mz/yr) near the lower end<br />
<strong>of</strong> the range expected for freshwater<br />
emergent macrophytes.<br />
<strong>Lake</strong> <strong>St</strong>, Cl ai r, particularly Anchor<br />
Bay and Mitchell Bay, are known for thetr<br />
we1 1 developed submerged aquatic<br />
macrophyte beds. Dawson (19751 found that<br />
considerable spatial varfation in standing<br />
crop exists in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> bays (Table<br />
29). He est'imated mean dry weights <strong>of</strong><br />
varlous submerged communities to range<br />
from 4 g/m2 in sha71ow areas characterized<br />
by sparse cover <strong>of</strong> stonewo<br />
to 316 g/m2 in areas domfna<br />
<strong>of</strong> water-milfoll (<br />
1. Schloesser ( 1982)<br />
monthly abundance <strong>of</strong> submersed macrophytes<br />
in the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>. elair<br />
during the 1978 growing season (Table 30).<br />
He found that the amount <strong>of</strong> submersed<br />
vegetation was low in early sprfng, No<br />
vegetatfon was Sound at the mouth <strong>of</strong> the<br />
Clinton Rtver or the head <strong>of</strong> the Detrolt<br />
River (Belle Isle) in April or May, and<br />
the only vegetation collected at <strong>St</strong>ag<br />
Islands Algonac and Anchor Bay during<br />
these months consfsted <strong>of</strong> decabSng or<br />
dormant material f ram the previous growf ng
Table 28. Biomass ,arid productivity <strong>of</strong> emergent wetland plants <strong>of</strong> the<br />
Great <strong>Lake</strong>s region.<br />
Species<br />
'~ata source: Tilton et a1. (1978).<br />
Maximum Net<br />
Common name b i omass product 1 on<br />
broad-1 eaved cattai l<br />
hybrfd cattail<br />
giant bur reed<br />
common arrowhead<br />
manna grass<br />
reed grass<br />
wild rice<br />
sp i ke-rush<br />
lake sedge<br />
beaked sedge<br />
river bulrush<br />
great bul rush<br />
s<strong>of</strong>t rush<br />
Mean<br />
(g/m2) (g/m2/yr)<br />
Tabla 29. Estimates <strong>of</strong> standing crop biomass for submergeni vegeta-<br />
tion in Anchor Bay, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (September, October 1974).<br />
Dominant Weedbed Range in Percent <strong>of</strong><br />
taxa in community area (km2) dry welght (g/m2) submergents<br />
TOTAL 108.8 4-3 16 100.0<br />
"~ata source: Dawson ( 1975).
Table 30. Seasonal estimates <strong>of</strong> standing crop biomass for submersed<br />
rnacrophytes at several locations in the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> sys~em (1978).<br />
Dominant<br />
Locatl on June July Rug Sept kt taxa<br />
<strong>St</strong>. <strong>Clair</strong> Rfver (<strong>St</strong>ag Is.) 72 267 103 38 15 SPP<br />
<strong>St</strong>. Clafr Delta (Algonacl 54 97 158 174 284<br />
Anchor Bay (Sand Is,) 85 113 135 133 146<br />
Clfnton River (mouth) 24 63 118 33 16 V a l l i w<br />
Detroit River (Belle Isle) 0 38 52 38 17 Yalli-<br />
iaowkaB<br />
a~ata source: Schloesser (1982).<br />
season. By mid-June dominant plants<br />
sprouted new growth at stem and leaf<br />
nodes, and by mid-July new growth which<br />
did not orfginate from overwintering<br />
dormant material was found at a17<br />
locations (Table 30). Higher maximum<br />
bfomass values were generally observed at<br />
locations where dormant overwinterfng<br />
vegetatjon was observed in early spring.<br />
Flgure 47 shows that the maximum biomass<br />
was reached In July in the <strong>St</strong>. <strong>Clair</strong> River<br />
(267g/m21; in August In the Gl inton River<br />
mouth (118 g/m2f and Detroit River head<br />
(53 g/m2); and in October in the <strong>St</strong>. <strong>Clair</strong><br />
Delta (284 g/m21 and Anchor Bay f 146<br />
g/m2). <strong>The</strong> submersed macrophytes became<br />
senescent and the biomass values decreased<br />
by late fall at all locations. <strong>The</strong><br />
results <strong>of</strong> thfs study indfcated that<br />
little or no submersed macrophyte growth<br />
takes place before June or after October<br />
in the rfver or lake. <strong>The</strong> dominant taxa<br />
found at each location are given in Table<br />
31,<br />
4.3 CQMMUNiTY PROCESSES<br />
Given the transittonal or ecotone<br />
character <strong>of</strong> wetlandsr the <strong>Lake</strong> <strong>St</strong>. Clafr<br />
wetlands are even more dynamic due to the<br />
pulsing nature <strong>of</strong> the lake level changes<br />
and the connectivlty due to the numerous<br />
distributary channels and embaymentst<br />
particularly wlthin the <strong>St</strong>. <strong>Clair</strong> Delta.<br />
By outlfning the community processes in<br />
these dynamlc and interrelated envi ron-<br />
ments, the high prlmary and secondary<br />
producti vi ty can be more full y understood.<br />
<strong>The</strong> <strong>Lake</strong> <strong>St</strong>. Clalr wetlands are<br />
actually composed <strong>of</strong> a variety <strong>of</strong> habitats<br />
f ncl ud 1 ng open ponds, cattai?/reed<br />
marshes, earthen dikes, barrler beaches,<br />
delta flats, and wooded swamps.<br />
Col7ectfvely these habitats are known as<br />
the coastal marsh community, Each habitat<br />
attracts its own species <strong>of</strong> plants* blrdsr<br />
mamnalsp reptiles, amphibianst and fn some<br />
cases, fish. <strong>The</strong> result Is more varfety<br />
in plant and animal life than in any other<br />
area <strong>of</strong> equal size fn the intarior <strong>of</strong> the<br />
bordering states and province. <strong>The</strong><br />
overall conditions <strong>of</strong> the uncontrolled<br />
coastal marshes are stf 17 very primitive,<br />
some having been visited by no more than a<br />
handful <strong>of</strong> people in the last several<br />
decades. W i thln those marshes where<br />
natural processss are allowed to take<br />
place, zonatfon and succession in response<br />
to changing envf ronmsntal condftions are
ANCHOR BAY<br />
MICHIGAN<br />
Sand Island<br />
\ LAKE<br />
Figure 47. Seasonal growth (dry weight) <strong>of</strong> dominant submersed macrophytes in the <strong>St</strong>.<br />
<strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-Detroi t River ecosystem (Schloesser 1982).<br />
Table 35 Dominant submersed aquatic macrophyte taxa <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
sys tern.<br />
Taxa<br />
-<br />
<strong>St</strong>. <strong>Clair</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Lake</strong> <strong>St</strong>. Clinton Detroit<br />
R'iver Delta Cl ai r River River<br />
(<strong>St</strong>ag Island) (Algonac) (Anchor Bay) (mouth) (Belle Isle)<br />
%ata sources: Dawson ( 19751 r Schloesser (1982).
among the important commun l t y processes.<br />
Water level fluctuationr and the resultant<br />
plant and anfmal responser is <strong>of</strong>ten the<br />
most sfgnificant drivjng force.<br />
Because the water level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> is not stable, but has oscillated<br />
over 2 m during the period 1900 to 1977<br />
(Jaworski et a1. 19793, the wetland<br />
vegetation a1 ong the 1 ake margi ns exhibits<br />
pulse stability. <strong>The</strong> concept <strong>of</strong> pulse<br />
stability imp1 ies that it is these water<br />
level fluctuations which result tn the<br />
maintenance <strong>of</strong> the wetl and communities.<br />
Thus* even though wetland plant<br />
communfties retrogress during hlgh lake<br />
level periods and shift lakeward during<br />
low water level conditionsr these<br />
successional changes prevent the wet1 ands<br />
from being dominated by either aquatic or<br />
up1 and communities. Moreovers coastal<br />
wetlands do not become senescent or<br />
exhibit conversion t o terrestrial<br />
environments as compared to inlands<br />
pal ustrine wetl ands.<br />
It is important to distinguish<br />
between annual (or seasonal 1 water level<br />
fluctuations and longer term oscillations,<br />
With regard to annual water level changes<br />
fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, the average magnitude<br />
is approximate1 y 0.5 m, with highest<br />
levels occurrtng in June-July and lowest<br />
levels during the winter months. This<br />
seasonal water level pattern is opposite<br />
that in shallowr freshwater swamps which<br />
may dry up during the middle <strong>of</strong> the<br />
growing season. Cattails appear to be<br />
favored by low water levels in winter<br />
because oxygen diffuses through exposed<br />
stalks down into the roots and rhizomes<br />
(Jaworski et al. 1979).<br />
However, it is the longer term lake<br />
level osclllatlons which have the greatest<br />
effect on vegetation succession. In the<br />
Great <strong>Lake</strong>s the hydroperiod for these<br />
longer term oscillations is 8 to 20 years<br />
(U.S. Dept. <strong>of</strong> Commerce 1976). Near<br />
record low water levels - at about 174 m<br />
above datum (IGLD) - occurred in 1964-<br />
1965, whereas record highwater levels - at approximately 176 m above datum (IGLD) -<br />
took place during the 1972 to 1974<br />
<strong>The</strong> latepal shifts <strong>of</strong> the plant<br />
communities in response to water level<br />
fluctuations are illustrated by Figures 32<br />
and 33 <strong>of</strong> Section 3-2. DickYlnson Island<br />
was utiljzed for this dtscussion because<br />
it has an intact envi ronmental gradient<br />
over which vegetation shifts can occur.<br />
El sewhere on Warsens Island as we1 1 as on<br />
the Canadian side <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta9<br />
the cattail marsh ends abruptly against<br />
dikes or canals, and much <strong>of</strong> the sedge end<br />
other drier habftat communities have been<br />
devel oped.<br />
<strong>The</strong>se successional changes can be<br />
documented by a series <strong>of</strong> transects taken<br />
over time across the same survey 1 ine. As<br />
indicated by Figure 48, water level<br />
changes in the coastal wetlands <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. Clai r caused dramatic vegetation<br />
changes. Beginning in July 1964s one can<br />
observe extensive comunjties <strong>of</strong> cattail r<br />
sedge, and dogwood meadow, Howevers<br />
during the high water period <strong>of</strong> the 1970~~<br />
widespread ddeback occurred as far inland<br />
as the shrub/swamp frfnge which borders on<br />
the oak-ash hardwoods. During such<br />
condltlonsr much <strong>of</strong> the cattall marsh<br />
consisted <strong>of</strong> open water as well as<br />
communities <strong>of</strong> bulrushes and submersed<br />
aquatics. Notice that by the summer <strong>of</strong><br />
1977 r considerable reestablishment began<br />
to take place as water levels dropped<br />
somew hat.<br />
A plant commundty displacement model<br />
can be used t o predict community<br />
succession associated with water level<br />
fluctuati ons. As f 11 ustrated in Figure<br />
49s each wetland plant community has been<br />
placed at its proper elevation in relatlon<br />
to the long-term mean water level. <strong>The</strong><br />
length <strong>of</strong> the l ine under the plant<br />
community's name lndjcates the amount sf<br />
vertical shifting which that community<br />
exhibits. To use PhSs model* note that<br />
the communities shOft lakeward (to the<br />
left) durlng low water levels and displace<br />
communities along thef r lakeward margfn.<br />
To simulate S U C C ~ S S during ~ ~ ~ rfsing lake<br />
levels, shift the comunfttes to the right<br />
a1 ong the1 r respective dlspl acement 1 f nes,<br />
As one moves upward along the<br />
envi ronmenisl gradtent, the extent <strong>of</strong><br />
cornunity displacement is less and core<br />
areas persfst durfng both low and hlghinterim.<br />
Because <strong>of</strong> thess CYC~IC<br />
water water pert ads, f herefore* the shrub/swamp<br />
level oscillations, succession is a1 ways fringe zone shifts least and Os displaced<br />
taking place in the plant communities. less by the adjacent $1 ant comunf iy.
WEST<br />
Feet Drowned<br />
Tree-Shrub<br />
678<br />
670<br />
<strong>Lake</strong> Dead<br />
Cottonwood<br />
Cottonwood<br />
June 1975 EAST<br />
Meters<br />
A<br />
July - Qct. 1977<br />
July 1964<br />
Mixed<br />
- - . - - - - - - - - - - - - - - - -<br />
T-"I ---1-7- - T-*---T--- 7-----r--- --T- -7- -- --r<br />
+ 1x , _1T_11_1_T1_1r11_7-~T_I_ - -<br />
0 500 1000 1500 2000 2500 3000 3500 4000 4500 5000 5500 6000 6500 Ft.<br />
r-- --7""- ,<br />
0 200 400 600 800 1000 1200 1400 1800 1800 Meters<br />
Figure 48. Succes~iorial changes on Dickinson Island in response to water level fluctu-<br />
ations,<br />
flll UCTCHS<br />
hi." 1<br />
Shri~b/Swa~lip I<br />
I - - -<br />
Meadow -- ,,<br />
Sedge Mdrstl i -<br />
"" . . - . - - - - .- .- Llf6.0<br />
. . - . - - - - .- .- Llf6.0<br />
Mcan iakc Level 1<br />
Figtars 43. Wetland plant community dis-<br />
placement rnode'l for Dickinson island<br />
showing predicted response to water level<br />
changes,<br />
176<br />
175<br />
174<br />
As shown in Table 32, the wetland<br />
pl ant communities not on1 y change location<br />
but alter their areal extent in response<br />
t o these long-term water level<br />
oscillations. <strong>The</strong> dominance <strong>of</strong> the<br />
emergent cattail and sedge marsh during<br />
low-water levels is clear. In contrast,<br />
during high water periodsr the wetlands<br />
exhibit extensive floating leaved and<br />
submersed communities along with the<br />
cattails. <strong>The</strong> cattail marsh actual 1 y<br />
appears more widespread during high water<br />
conditions than it is because the dead<br />
cattail stalks <strong>of</strong>ten remain in place for<br />
several years. Notice, toor that the area<br />
<strong>of</strong> the shrub/swamp changes very 1 ittle as<br />
the water levels change. This situation<br />
prevails because the shrub/swamp community
Table 32. Vegetation cianges on Dickinson Island in response to<br />
lake level fluctuations.<br />
Percent <strong>of</strong> total areab<br />
Vegetation class Low water Mean level Hiah water<br />
1964 1975<br />
Open water<br />
Floating leaved/<br />
Submersed<br />
Emergents (cattails)<br />
Sedge marsh<br />
Dogwood meadow<br />
Shrub/Swamp<br />
Developed areas<br />
a Data source: Jaworski et a1 . (1979).<br />
b~ote: Total area lr130 hectares (2,800 acres).<br />
is slightly higher in elevation than the<br />
high water mark and it is difficult to<br />
separate dieback shrub from live brushy<br />
vegetation. Also? cattail is an ag-<br />
gressive invader compared to sedges and<br />
dogwood.<br />
<strong>The</strong> zonation exhibited by the wet1 and<br />
plant communities has previously been<br />
illustrated by Figures 26 and 33 and<br />
described in Section 3.2. <strong>The</strong>se plant<br />
zones, in order from upland to aquatic<br />
sites, are: shrub/swamp? dogwood meadow?<br />
sedge marsh? cattail marshr and open water<br />
floati ng/submersed commun f t i es . In many<br />
wetlands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> urban<br />
development and other land uses have<br />
progressed from the adjacent upland areas<br />
lakeward to the emergent marsh. On1 y on<br />
Dickinson Island, and to some extent in<br />
Bouvier Bay (i.e., <strong>St</strong>. John's Marsh)? does<br />
a complete environmental gradient? or full<br />
zonation <strong>of</strong> wetlands* exist today.<br />
Species diversity <strong>of</strong> invertebrates<br />
tends to be hlgher fn <strong>St</strong>. <strong>Clair</strong> Delta<br />
wetland ecosystems than f n nearshore<br />
benthic and pr<strong>of</strong>undal benthic zones <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Invertebrate productivtty<br />
also tends to be higher in wetlands<br />
compared to open water areas and ft<br />
appears that invertebrate productivity<br />
tends to. be higher in emergent wetlands<br />
compared to submergent wet1 ands (Ti 1 ton et<br />
al, 1978). Invertebrate populations in<br />
coastal wetlands are a food source for<br />
higher trophic levels in the food webt<br />
comprising a major fraction <strong>of</strong> the diet <strong>of</strong><br />
numerous species <strong>of</strong> fish* reptiles?<br />
amphibians, blrdsr and mammals.<br />
Several species <strong>of</strong> forage fish<br />
(spottail? black nose^ and emerald shiners9<br />
feed or spawn In wetland ecoysystems,<br />
<strong>The</strong>se forage fish are consumed by<br />
piscivorous fish as well as birds and<br />
mammals. toss <strong>of</strong> wetlands whjch produce<br />
forage fish can cause a reduction in the<br />
numbers <strong>of</strong> piscivorous fish as we11 as<br />
birds such as the kingfisher and common<br />
mergansers (Ti 1 ton et a1 . 1978).<br />
Zonation has also been demonstrated<br />
for larval fish populations in western<br />
<strong>Lake</strong> Erie (Cooper et al. 1981ar 1981b.<br />
1984; Mizera et ale 3.9811. A series <strong>of</strong><br />
studies in the estuaries and open waters<br />
<strong>of</strong> western <strong>Lake</strong> Erie has shown that the<br />
estuaries <strong>of</strong> the Maumee and Sandusky<br />
Rivers contained the hfghest densities <strong>of</strong>
larval fish when compared with other<br />
nearshore and <strong>of</strong>fshore areas. Glzzard<br />
shad, white bass, and freshwater drum<br />
dominated the estuarfne populatfons. <strong>The</strong><br />
highest densjty <strong>of</strong> yellow perch was found<br />
in nearshore areas assocfated wfth sandy<br />
bottoms, particularly north <strong>of</strong> Woodtick<br />
Peninsuia and in the vicinity <strong>of</strong> Locust<br />
P<strong>of</strong>nt. <strong>The</strong> followtng depth/denslty re-<br />
lationship was observed for thls species:<br />
<strong>The</strong> same general relationship was found<br />
for most species, fndfcatlng a greater<br />
preference for spawning and nursery<br />
grounds fn the coastal areas. A sfmilar<br />
pattern is suspected for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Surveys conducted by Hatcher and Nester<br />
E1983)~ although not in the lake proper,<br />
show high densities Jn the <strong>St</strong>. <strong>Clair</strong> Delta<br />
and at the head <strong>of</strong> the Detroit River.<br />
Johnson (1984) belleved that the most<br />
important factor influencing production <strong>of</strong><br />
flsh larvae in #%stern <strong>Lake</strong> Erie marshes<br />
is predatfon. Although no information Is<br />
avaflable on predator-prey tnteractlons<br />
involving larval fJsh In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
wetlands* related research has Indicated<br />
habftat structure can be Smportant in<br />
medtatlng the outcome <strong>of</strong> aquatic predator-<br />
prey interactions. Glass (1971) found<br />
argemouth bass ( a4.L<br />
predat f on succes s the<br />
Ity <strong>of</strong> habitat structure increased.<br />
Crowder and Cooper (19791 concluded that<br />
areas <strong>of</strong> hSgh structural complexity should<br />
create an effective refuge for prey and<br />
onhance their survfval, However,<br />
(19791 found prey-sized bluegill (<br />
1 were strongly attracted to<br />
shade produced by I m diameter floats in<br />
wh~ch no submerged structure was present,<br />
He ~oncluded prey fish in shade can detect<br />
a predator (in sun) long before the<br />
predator can detect the prey.<br />
<strong>The</strong>refore prey specles and prey-<br />
sized fSsh may act1 vely escape predators<br />
by dfsappearfng Snto areas <strong>of</strong> high<br />
struct~ral complexity or by seeklng shade<br />
so as to gajn the visual advantage.<br />
Consequentlys fish larvae in wetlands may<br />
use floating habitat types to gain the<br />
vjsual advantage while other larvae may<br />
use submergent or emergent habitat types<br />
to hide from potential predators, Open<br />
water areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />
marshes may be expected to have low larval<br />
use during the day because <strong>of</strong> the<br />
increased success <strong>of</strong> visual predators in<br />
this habitat type, but these areass<br />
particular1 y the uncontrolled wetlands,<br />
may be Important to fish larvae at night,<br />
<strong>The</strong> coastal marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
attract large numbers <strong>of</strong> migratory<br />
waterfowl. Located at a crossing point on<br />
two major flyways, these marshes attract<br />
ducks from eastern Canada headlng for<br />
wintering grounds on the Mississippi River<br />
bottoms and ducks from the prairie<br />
provinces <strong>of</strong> Canada which winter along the<br />
Atlantfc coast. Beds <strong>of</strong> wfld celery are<br />
partjcul arl y attractive to great numbers<br />
<strong>of</strong> canvasbacks (m 1, red-<br />
heads ( ), and scaups<br />
(w<br />
spp. migrating southeastward to<br />
thef r wintering grounds on Chesapeake Bay<br />
(Andrews 1952).<br />
Lincoln (1935) introduced the concept<br />
that all populatlons <strong>of</strong> migratory birds<br />
adhere to their respectfve flyways as they<br />
make thef r semt annual fl lghts between<br />
breeding grounds and wintering grounds.<br />
Four distinctive flyways have been<br />
Identified for North America, two <strong>of</strong> whqch<br />
cross western <strong>Lake</strong> Erie and <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong>: the Atlantic Flyway and the<br />
Mississippi Flyway, <strong>The</strong>se routes are used<br />
by all <strong>of</strong> the migratory waterfowl and<br />
other waterbirds which frequent the<br />
coastal wetlands. Each flyway has its own<br />
individual population <strong>of</strong> birdsp even for<br />
those specfes which have a broad<br />
contJnenta1 distribution. <strong>The</strong> breeding<br />
grounds <strong>of</strong> two or more flyways may, and<br />
<strong>of</strong>ten dor overlap broadly so that during<br />
the nesting season extensive areas may be<br />
occupfed by bjrds <strong>of</strong> the same species, but<br />
which belong to dffferent flyways.<br />
In the fall, the Atlantlc Flyway<br />
recsivas accretion <strong>of</strong> waterfowl from<br />
several fnterlor migration paths starting
at the breeding grounds on the Arctic<br />
tundra. Canada goose )V<br />
and df vfng ducks8 incl SI<br />
redheads and scaup come from their<br />
breeding grounds on the great northern<br />
plains <strong>of</strong> central Canaaa, follow the<br />
general southeasterly trend <strong>of</strong> the Great<br />
<strong>Lake</strong>s? rest in the <strong>St</strong>. <strong>Clair</strong> Delta, cross<br />
<strong>Lake</strong> Erje in the islands region, and<br />
continue over the mountains <strong>of</strong><br />
Pennsylvania to w i nter along the Atlantic<br />
coast in Chesapeake and Delaware bays.<br />
Concurrent1 y, dabbl ing ducks such as<br />
1, black ducks<br />
1, and blue-winged teals<br />
1 that have gathered in<br />
staging areas <strong>of</strong> southern Ontario<br />
(fncludfng the <strong>St</strong>. <strong>Clair</strong> Delta) also leave<br />
these feeding grounds, cross western <strong>Lake</strong><br />
Erie and proceed southwest over a course<br />
that leads down the Ohio and Mississfppi<br />
valleys (Mississippi Flyway), However,<br />
part <strong>of</strong> this duck population, upon<br />
reaching the <strong>St</strong>. <strong>Clair</strong> Flats, swings<br />
abruptly to the southeast, crosses the<br />
Appalachian mountains and winters along<br />
the Atlantic coast (Lincoln 1950).<br />
<strong>The</strong> Mississippi Flyway is easily the<br />
longest migration route <strong>of</strong> any in the<br />
Western Hemisphere. Its northern terminus<br />
is the Arctic coast <strong>of</strong> Alaska, whlle its<br />
southern end 1 ies In the Patagonia region<br />
<strong>of</strong> Argentina. Although the main path <strong>of</strong><br />
the flyway lies to the west <strong>of</strong> the Great<br />
<strong>Lake</strong>s major branches follow the southern<br />
trend <strong>of</strong> <strong>Lake</strong> Michigan and the<br />
southwestern trend <strong>of</strong> <strong>Lake</strong> Erle and the<br />
Maumee Rf ver valley. Some <strong>of</strong> the black<br />
ducks ma1 1 ardsr and teals that cross the<br />
Great <strong>Lake</strong>s in the viclnfty <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> do not turn abruptly to the<br />
southeast, but continue on to the<br />
southwest as members <strong>of</strong> the Mississippi<br />
Flyway bound for the Gulf <strong>of</strong> Mexico coast<br />
rather than the Atlantic seaboard. Fall<br />
migration is at Its peak in September and<br />
October, but the main shorebird passage is<br />
underway in August.<br />
Sprlng migration begins in late<br />
February with the appearance <strong>of</strong> rlng-<br />
billed gulls ( 1 In <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. Ma 11 bring<br />
h waterfowl mov-ntr ducks and loons<br />
( sp.1 appearing 5n open leads as<br />
blackbf rds (<br />
large numbers starting in late March. <strong>The</strong><br />
huge Canada goose movement normally takes<br />
place In early April. With April and May<br />
comes the major push <strong>of</strong> spring migration,<br />
especizlly the landbirds. Migrants are<br />
less selective than breeding birds In<br />
their choice <strong>of</strong> habitat, nevertheless,<br />
waterb i rds prefer shore1 lnes wf th pockets<br />
<strong>of</strong> vegetation. Coastal marshes and stream<br />
mouths comnly attract migrating dabbl f ng<br />
ducks. <strong>The</strong> open water shorelines<br />
concentrate the diving duck migrants and<br />
other waterbirds including loons, grebes,<br />
cormorants (<br />
For over three decades after Lincoln<br />
defined and mapped his four waterfowl<br />
flyways <strong>of</strong> North America, little new work<br />
was pub1 ished on the subject. <strong>The</strong>n,<br />
Bell rose (1968) pub1 ished his outstanding<br />
treatise on waterfowl migratlon corrldors<br />
east <strong>of</strong> the Rocky Mountains. In addition<br />
to these data and ground observatlons~<br />
Be1 1 rose used such advanced techn I ques as<br />
visual sightings from aircraft and radar<br />
su rvei l 1 ance. One impetus for undertaking<br />
this revision was the Increasing hazard to<br />
aircraft from migrating birds.<br />
Be1 l rose found that Llncol n s f 1 yway<br />
concept, although still valid on a grand<br />
scale, tended to oversfmpllfy the<br />
migratjon process. He observed that<br />
flyways were In reality e complex <strong>of</strong><br />
corridors (Figures 50-52). Each corridor,<br />
in turn? Is a web <strong>of</strong> routes as opposed to<br />
a single, narrow band rigidly followed by<br />
the waterfowl. <strong>The</strong> migration corridors<br />
represent passageways sac h connecting a<br />
serles <strong>of</strong> waterfowl habitats. Migration<br />
corridors differ from flyways in being<br />
smaller and more precisely deflnsd as to<br />
specles. Bell rose cons lders the f l yways<br />
proposed by Lincoln to be prfmarily<br />
geographical and secondarily biological,<br />
while his corridors are primarily<br />
biological and secondarif y geographical.<br />
As waterfowl migrate between breeding<br />
grounds and wfntering areas, they stop to<br />
rest and feed in wet1 ands, <strong>The</strong>se wetl ands<br />
2re referred to as nconcentratlon areas."<br />
<strong>The</strong> coastal wetl ands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Claf rr<br />
the lower Detroit Riverr and western <strong>Lake</strong>
DABBLING DUCKS<br />
ESTIMATED NUMBER<br />
USING A CORRIDOR<br />
PENNSYLVANIA<br />
Figure 50. Fa1 1 migration corridors for dabbling ducks (Tribe Anatini ) across<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Be1 l rose 1968).<br />
Erie provide some <strong>of</strong> the finest<br />
concentration areas <strong>of</strong> this type along the<br />
flyways. <strong>The</strong>se areas are characterized by<br />
an abundance <strong>of</strong> waterfowl foods, as well<br />
as by low wave energy and low human<br />
disturbance. Canvasbacks, redheads*<br />
animal foods, Canada geese and mallards<br />
also feed heavf ly on waste grains fn<br />
agricultural fields. Food availability<br />
may be more important than food<br />
preference, especially during the sprf ng<br />
migration when food supplies are less<br />
abundant. Food availabil ity f n wetlands<br />
is reduced by extreme high and low water<br />
levels, heavy siltationr turbidity, heavy<br />
hunting pressure* and other disturbances.<br />
Jones (1982) examined the food<br />
preference <strong>of</strong> wintering go1 deneye, greater<br />
scaup, and lesser scaup at the mouth <strong>of</strong><br />
the Detroit River. He determined that 30<br />
food taxa were utilized by the waterfowl<br />
including 10 plant and 20 animal<br />
references. Wild csler<br />
1, pondweeds (<br />
and other plant speci es made up over 60%<br />
<strong>of</strong> the food budget <strong>of</strong> most <strong>of</strong> the sampled
- -<br />
DIVING DUCKS<br />
ESTIMATED NUMBER<br />
USING A CORRIDOR<br />
Figure 51. Fa1 1 migration corridors for diving ducks (Tribe Aythyini) across<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Be1 lrose 1968).<br />
waterfowl. Miller (1943) and Bell rose<br />
(1976) have found that other diving ducks*<br />
particularly canvasbacks and redheads*<br />
feed on submersed aquatics such as those<br />
noted above as well as waterweed Elc&&<br />
s2amku&)<br />
In western <strong>Lake</strong> Erie marshes,<br />
Bednar f k (19751 found that preferred<br />
natural foods <strong>of</strong> diving ducks* such as<br />
wild celery* appeared to be more adversely<br />
affected by turbidity and siltation than<br />
foods <strong>of</strong> dabbling ducks and geese. In<br />
general, Great <strong>Lake</strong>s conservation agencies<br />
do not encourage the creation <strong>of</strong> resident<br />
wintering flocks# particularly in the <strong>Lake</strong><br />
<strong>St</strong>. Clafr region* because <strong>of</strong> the problem<br />
<strong>of</strong> waterfowl starvation during severe<br />
winters. Waterfowl that reach the spring<br />
breeding grounds in good condition tend to<br />
exhibit greater nesting success than those<br />
which are undernourished (Bell rose 1976).<br />
Q&l&lng duck <strong>The</strong>se ducks are so<br />
ca1 led because they normal 1 y do not dive<br />
below the water for food, but merely<br />
dabble on the bottom In shallow water.<br />
Annually about 17r500r000 ma1 1 ards and<br />
pintail s migrate down fl ight corrldors<br />
from Canada to the United <strong>St</strong>ates east <strong>of</strong><br />
the Rocky Mountains (Bellrose 19681, <strong>The</strong><br />
l argest portion, about 12r275r000s enter<br />
the geographfcal conffnes <strong>of</strong> the<br />
Mississippi Flyway from the northern Great<br />
Plains, About 20% <strong>of</strong> these birds continue<br />
across the Mississfppf Flyway and move<br />
down the Atlantic Flyway. In addition,<br />
anotRer 650,000 black ducks move south<br />
from Ontario and Quebec.<br />
_
CANADA GEESE<br />
ESTIMATED NUMBER<br />
USING A CORRIDOR<br />
75,100 - 150,000<br />
5,100 - 25,000<br />
Figure 52. Fall migration corridors for Canada geese (Branta canadensis) across<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Bellrose 1968).<br />
Several corridors carrying dabbllng As the name suggests,<br />
ducks cross the Great <strong>Lake</strong>s region (Flgure these ducks normally dive below the water<br />
50). An estimated 65,000 mallards, 35,000 for food. About 40200r000 diving ducks<br />
wigeons, and 25r000 pintalls move eastward annually migrate south into the United<br />
along the Chesapeake Bay Corridor. <strong>The</strong> <strong>St</strong>ates east <strong>of</strong> the Rockies (Bell rose<br />
corrldor starts In the upper Mfss'lssfppi 113681, Slightly over 609% <strong>of</strong> these are<br />
Rf ver Val ley and Progresses eastward scaups. Redheads are second fn abundance<br />
through Wlsconsin, Mlchiganr and Ohlo. It at ZO%5 while canvasbacks and ring-necked<br />
encompasses the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr ducks each form about 7% <strong>of</strong> the<br />
and <strong>Lake</strong> Erie f ram Algonacr Mlchfganr to populaticn. As with the dabbling ducksr<br />
Sandusky Bay. From these marshes it is a nume;.ous diving duck mfgration corridors<br />
645 km* non-stoP flight to Chesapeake Bay cross the Great <strong>Lake</strong>s region (Figure 51).<br />
where most <strong>of</strong> these ducks winter, <strong>The</strong><br />
Bl ack Duck Corridor extends southwest~ard <strong>The</strong> Southern Michigan Corrfdor takes<br />
from eastern Ontario5 across <strong>Lake</strong> <strong>St</strong>. the maln flow <strong>of</strong> diving duck passage from<br />
CIair to the confluence <strong>of</strong> the Wabash and eastern WJSCO~SSR~ across southern Mich-<br />
Qhfe, riversr and on south to Arkansas. lgan to Saginaw Bay and the <strong>Lake</strong> <strong>St</strong>.<br />
Approximately 35#QOO black ducks use this Cl a i r-Detroi t River-<strong>Lake</strong> Erte wet1 ands<br />
path, areas. Diving ducks congregate on SagJnaw<br />
100
Bay to the extent that peak numbers<br />
include 22,000 lesser scaupsr 22r000<br />
redheads, and 7,000 canvasbacks.<br />
Approximately 160 km to the south, peak<br />
populations <strong>of</strong> 380,000 lesser scaupsr<br />
260r 000 canvasbacks, and 42r000 redheads<br />
have been observed fram <strong>Lake</strong> <strong>St</strong>. Cl ai r to<br />
western <strong>Lake</strong> Erie. Although as many as<br />
15,000 diving ducks may winter on the<br />
Detroit River, at least 700,000 fly on<br />
Prom <strong>Lake</strong> <strong>St</strong>. Clalr to wintering grounds<br />
in the Atlantfc Flyway. This route is<br />
known as the Chesapeake Bay Corridor and<br />
is a similar route to the one taken by<br />
mallards and pintails.<br />
More than any other species<br />
<strong>of</strong> waterfowl, Canada geese have radical1 y<br />
altered their migration routes In the past<br />
few decades. Bell rose (1968) attrf buted<br />
this great change tn their migration<br />
habits to their rapid adoption <strong>of</strong> newly<br />
created waterfowl refuges, <strong>The</strong>y are still<br />
in the process <strong>of</strong> evolvtng new migration<br />
corridors, Currently, about lr300s000<br />
Canada geese leave Canada in the fall for<br />
wintering grounds in the United <strong>St</strong>ates.<br />
<strong>The</strong> largest number, 600~000 use the<br />
Atlantic Flyway, while another 475,000<br />
take the Mississippi Flyway. Most <strong>of</strong> the<br />
Atlantic Flyway crossings <strong>of</strong> the Great<br />
<strong>Lake</strong>s take place over <strong>Lake</strong> Ontarfor but<br />
one corridor uses the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> (Figure 52). <strong>The</strong> main migration<br />
corridor for Canada geese i n the<br />
Mississippi Flyway extends down the west<br />
shore <strong>of</strong> <strong>Lake</strong> Michigan,<br />
Mississippi River valley.<br />
then down the<br />
Snow geese (m 1 use<br />
the Misstssippi Flyway. Each October<br />
about 450r000 birds 1 eave Canada for<br />
wfntering grounds on the coastal marshes<br />
<strong>of</strong> Louisiana. <strong>The</strong> main corridors follow<br />
the east and west shores <strong>of</strong> <strong>Lake</strong> Michiganr<br />
converging fn the Mississippi River valley<br />
north <strong>of</strong> Loulsi ana. <strong>The</strong> easternmost<br />
flfght corridor, used by about 15,000<br />
geese, runs from the south end <strong>of</strong> James<br />
Bay to the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />
western <strong>Lake</strong> Erie, then turns south-<br />
westward across Indiana, A somewhat<br />
Jarger number <strong>of</strong> birds uses a corridor<br />
that extends from James Bay through<br />
Saginaw Bayr and then merges with the<br />
flight path from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>,<br />
4.4 WETaNDS AS HABITAT TO FISH AND<br />
WlLDLlFE<br />
Because the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />
are composed <strong>of</strong> a variety <strong>of</strong> habitats<br />
including open ponds, cattail/reed<br />
marshesr earthen dikes, barrier beachesr<br />
delta flats, and wooded swamps, many<br />
species <strong>of</strong> plants and animals are<br />
attracted to the coastal marsh community.<br />
Of particular importance to fish and<br />
nil dl i fe, both resident and transient<br />
species# are the following considerat~ons:<br />
1) food chain production and energy flow,<br />
2) ffsh productionr spawningr and nursery,<br />
31 waterfowl migration, wintering and<br />
nesting, 4) mammal forager and 5)<br />
invertebrate habftat. Dole (1975) er-<br />
timated the density <strong>of</strong> wildlife in the<br />
vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Table 33).<br />
Most animals showed a stable trendr but a<br />
few important species are decreasing in<br />
abundance.<br />
Although there are scattered patches<br />
<strong>of</strong> wetlands around the perimeter <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>, the wetlands in the <strong>St</strong>. <strong>Clair</strong><br />
Delta provide the most valuable habitat<br />
for fish and wlldllfe, includtng water-<br />
fowl. Mot only ara these deltaic wetlands<br />
productlver they are diverse and exhibit<br />
high connectivity to the <strong>St</strong>. <strong>Clair</strong> River<br />
channels as well as to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
With regard to fish spawning in these<br />
wetlands, important species include the<br />
following: lake sturgeon* northern piker<br />
rnuskell unget carp, golden shiner, emerald<br />
shiner, common shinerr spottai l shlner,<br />
spotffn shiner, channel catf ishr bull-<br />
heads# rock basst pumpkf nseed, bl ueglll ,<br />
sunfish, and walleye (Figure 39). It has<br />
been suggested that ice fishing$ as f n<br />
Little Muscamoot Bay <strong>of</strong> Harsens Islandp is<br />
particul arl y good because wet1 ands are an<br />
important source <strong>of</strong> fish foods in winter<br />
(W. C. Bryant, MDNRI Ffsheries Division,<br />
pers. corn.). Howevers except for gtzzard<br />
shad, most f I sh mlgrate to deeper waters<br />
in fall and return to the shallows<br />
nearshore envi ronrnents fs1 lowing ice<br />
breakup (Werner and Manny 19791, Because<br />
wetlands warm up more quickly than <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong> in the sprfnge these coastal<br />
environments may be a crftical source <strong>of</strong><br />
food for ftsh attracted to the sharelines,
Gable 33, <strong>St</strong>atus <strong>of</strong> wildlife in ttge vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> and connecting waterways (1970).<br />
Wild1 lfe group<br />
and species Density Trend<br />
Big game<br />
White-tailed deer<br />
Waterfowl<br />
Ducks<br />
Geese<br />
Small game<br />
Cottontail rabbit<br />
Ring-necked pheasant<br />
Ruffed grouse<br />
Gray squ i rrel<br />
Fox squirrel<br />
Woodcock<br />
Mourning dove<br />
Bobwhite qua11<br />
Furbearers<br />
Muskrat<br />
M'lnk<br />
Beaver<br />
Weasel<br />
Raccoon<br />
Skunk<br />
Opossum<br />
Badger<br />
Non-game<br />
Woodchuck<br />
Red fox<br />
Gray fox<br />
Crow<br />
Rod squirrel<br />
Coyote<br />
Raptars<br />
Rare and endangered<br />
Bald eagle<br />
American osprey<br />
Unusual or unique<br />
Sandhi1 1 cranes<br />
Golden sag1 e<br />
a Data source: Dole (1975).<br />
1 ow i ncreasi ng<br />
high<br />
rned f um<br />
rned i urn<br />
high<br />
1 ow<br />
1 ow<br />
medi um<br />
1 ow<br />
high<br />
1 ow<br />
high<br />
rned i um<br />
1 ow<br />
rned i urn<br />
rned i um<br />
high<br />
high<br />
1 ow<br />
rned i urn<br />
med l um<br />
rned i um<br />
high<br />
1 ow<br />
1 ow<br />
rned f urn<br />
rned i urn<br />
rare<br />
stabl e<br />
increasing<br />
stable<br />
stab1 e<br />
stabl e<br />
decreasing<br />
stable<br />
stabl e<br />
stable<br />
stab1 e<br />
stabl e<br />
stabl e<br />
decreasl ng<br />
stable<br />
'lncreas'lng<br />
increasing<br />
stable<br />
stab1 e<br />
stable<br />
stabl e<br />
stabl e<br />
stable<br />
stabl e<br />
stabl e<br />
stabl e<br />
decreasing<br />
decreasing<br />
stabl e<br />
translent
With regard to wildlife habitats the<br />
<strong>St</strong>. <strong>Clair</strong> Delta wetlands are also<br />
exceptional, <strong>The</strong> main reasons for this<br />
outstandfng habitat qua1 l ty are: 1) large<br />
areas <strong>of</strong> relatively pristine wetlandsJ 2)<br />
habitat diversity (several wetlands<br />
exhibit cornpl ete envl ronmental con-<br />
tlnuums), and 3) relatively few barriers<br />
to movements. Dickinson Island and the<br />
Bassett Is1 and-Goose <strong>Lake</strong> areas represent<br />
relatively pristine wetl ands with direct<br />
hydrologic connection to <strong>Lake</strong> <strong>St</strong>. Clafr<br />
and few barriers to movements or<br />
migration. In Figure 31, one can readily<br />
see the habitat diversity, juxtaposition<br />
<strong>of</strong> environments, and mix <strong>of</strong> open water and<br />
vegetation which contributes to high<br />
wild1 ife values (We1 l er and Spatcher<br />
1965).<br />
<strong>The</strong> importance <strong>of</strong> the coastal<br />
wetlands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> with<br />
reference to waterfowl has a1 ready been<br />
discussed in Section 3.6. A1 though some<br />
waterfowl breeding does occur in the delta<br />
wetlands, the primary importance <strong>of</strong> the<br />
<strong>St</strong>, <strong>Clair</strong> River wetlands is in regard to<br />
resting and feeding habitat during<br />
migration. Because <strong>of</strong> availability <strong>of</strong><br />
preferred plant and invertebrate foods<br />
(Dawson 19751, as well as fts location<br />
along the Atlantic and Missfssippi Flyways<br />
(Be1 1 rose 1968, 19761, these wetlands<br />
serve as concentration areas and stopover<br />
points for both dfving and dabbling ducks.<br />
Habitat degradation along western <strong>Lake</strong><br />
Erie and Saginaw Bay, due to excessive<br />
turbidity, siltation and water quality<br />
decline (Martz and Ostyn 1977; U.S.<br />
Department <strong>of</strong> Interior 19771, i s<br />
associated with greater re1 iance on the<br />
<strong>St</strong>. <strong>Clair</strong> Delta wetlands. In this regards<br />
the waterfowl habltat deficien~les<br />
predicted for southeast Michigan as <strong>of</strong> the<br />
year 2000 are most significant (U.S.<br />
Department <strong>of</strong> Interior 1971).<br />
Large numbers <strong>of</strong> furbearers as we11<br />
as birds, reptiles, and amphibians have<br />
been identified in the <strong>St</strong>. <strong>Clair</strong> Delta<br />
wetlands (Herdendorf et al, 1981~).<br />
Muskrats abound in the cattafl marshes and<br />
raccoons are particul arly numerous where<br />
large trees occur. In comparison to the<br />
Canadian wetl ands, whfch consist largely<br />
<strong>of</strong> cattafl marshes and open-water areass<br />
the Dickinson Island wetlands on the<br />
American side <strong>of</strong> the delta exhibit much<br />
greater habitat diversity. Given access<br />
by boat via the abandoned channel which<br />
winds across Dickinson Island, both<br />
consumptive and nonconsumptive users can<br />
easfly enjoy this rich wildlife resource.<br />
To evaluate fish, wildlife and other<br />
values <strong>of</strong> wetlands, the Michigan Depart-<br />
ment <strong>of</strong> Natural Resources has developed a<br />
field evaluation manual (Wolverton 1981).<br />
However, the <strong>St</strong>. <strong>Clair</strong> Delta wetlands have<br />
not been rated by thfs agency.<br />
Neverthel essJ because <strong>of</strong> its exceptional<br />
fish, waterfowl, and other wildlife<br />
values, along with its accessibility to<br />
users and re1 at i vel y undeveloped nature,<br />
particularly Dickinson Is1 and# these<br />
wetl ands may we11 be among the mast<br />
valuable in the Great <strong>Lake</strong>s region.
5.4 WETLAND DISTURBANCE<br />
CHAPTER 5. HUMAM 81MPA67S AND APPLiED ECOLOGY<br />
A new awareness has arisen in the<br />
last decade over the alarming loss <strong>of</strong><br />
wetlands. In Michigan, 71% <strong>of</strong> the coastal<br />
wetlands have been lost (Jaworski and<br />
Raphael 1978) and much <strong>of</strong> the remaining<br />
42r420 ha have been altered because <strong>of</strong><br />
degraded water quality, changes in<br />
hydrology, and other factors, However,<br />
losses and degradation are not solely due<br />
to human intervention. Western <strong>Lake</strong> Erie<br />
has suffered a significant loss <strong>of</strong> aquatic<br />
vegatatlon, possfbl y because <strong>of</strong> changing<br />
1 ake 1 eve7 s, water movements, and carp<br />
feeding habits (Core 1948; <strong>St</strong>uckey 1971).<br />
Saiches and flooding may temporarily<br />
reduce emergent wetlands; however,<br />
regeneration usually occurs. Converse1 y,<br />
human impacts such as dredging, filling,<br />
diking* and draining have a permanent<br />
adverse impact on the resource base.<br />
Adverse impacts in the <strong>St</strong>. <strong>Clair</strong><br />
wetlands began wlth the exploitation <strong>of</strong><br />
fish and wildlife prior to European<br />
contact and continued Into the 1850s.<br />
Howeverr these impacts were modest and the<br />
losses somewhat replaceable through<br />
natural reproducllon. Impacts which fol-<br />
lowed were more permanent. We w i l l<br />
discuss the adverse impacts within a<br />
historical framework spanning about 125<br />
years (Figure 53). This is how our<br />
present problems emerged and this is the<br />
way they have been addressed.<br />
Historicallyr modlficatfon <strong>of</strong> the<br />
wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> began shortly<br />
after European contact. Historical<br />
accounts suggest that the wetlands <strong>of</strong> Lak?<br />
<strong>St</strong>, <strong>Clair</strong> were not excessively exploited<br />
by the fur traders as were other wetlands<br />
<strong>of</strong> the Great <strong>Lake</strong>s such as those along<br />
western <strong>Lake</strong> Erie. An abundance <strong>of</strong><br />
wildlife and fish attracted farmers from<br />
the settlement at Detroit. <strong>The</strong> desirable<br />
quallty and quantity <strong>of</strong> fish and wildlife<br />
led to the establlshment <strong>of</strong> fishing and<br />
huntlng clubs. In the mid-19th century*<br />
one such club, the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Fishing<br />
and Shooting Club, spent $80r000 to<br />
improve the property it occupied in the<br />
delta (Jenks 1912).<br />
Historic maps reveal that the<br />
wet1 ands were not impacted sfgnif icantly<br />
In the 1850s (Meade 18571, Although<br />
maritime commerce was carried on through<br />
the lake and its distributary channels,<br />
emergent aquatics were abundant and<br />
diverse throughout the <strong>St</strong>. <strong>Clair</strong> Delta and<br />
the mouth <strong>of</strong> the Clinton River. Jenks<br />
(1912) noted that wild rice (Zlrsnls<br />
aauatlca) was abundant as were at least<br />
116 specfes and 25 varieties <strong>of</strong> wetland<br />
macrophytes, including sedges. Fish and<br />
wildlife were exploited in a non-<br />
commercial capacity in the area. In 1857<br />
New Baltimore was the only community in<br />
the northern part <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Significant wetland disturbances are<br />
associated with two interrelated cultural<br />
processes: agricultural development, which<br />
was followed by urban growth. <strong>The</strong><br />
surveyor-general <strong>of</strong> the United <strong>St</strong>ates<br />
reported in 1815 that a large part <strong>of</strong> the<br />
southeastern regfon <strong>of</strong> the Michigan<br />
territory was swamp and practically<br />
worthless. As early as 1826 attempts were<br />
made for reclamation; but It was not until<br />
September 1850 that a general swampland<br />
law was enacted. Its purpose was to allow<br />
for draining and dikfng <strong>of</strong> nworthless<br />
public lands, lying as marshes or subject<br />
to periodic overflow by adjacent water-<br />
coursesw (Donaldson 19701,<br />
<strong>The</strong> 1850 swamp act stimulated<br />
significant wetland alteration. By 1873
Drec?ging, Filling, Rulkheadir~g<br />
- URBANIZATION -<br />
Figure 53. Historic trends in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetland disturbance,<br />
the land cover between the Detroit and establishment <strong>of</strong> the <strong>St</strong>, <strong>Clair</strong> Ship Canal,<br />
Clinton Rivers was in agriculture. <strong>The</strong> a 6 m channel dredged through South<br />
north shore <strong>of</strong> the lake a1 so supported Channel for commerclal navigation.<br />
agriculture. Establ ished villages such as<br />
New Baltimore grew, and new urban centers When the Grand Trunk Railway, the<br />
such as Anchorville were founded. thirdmode, wasconstructed, it dissected<br />
<strong>Wetlands</strong> at the head <strong>of</strong> the Detr<strong>of</strong> t Rlver the remaining wetlands in several places.<br />
were also being dredged. On the <strong>St</strong>, <strong>Clair</strong> However, on this shore1 ine most <strong>of</strong> the<br />
Delta, hardwoad forests <strong>of</strong> the premodern wetlands had a1 ready been drained so the<br />
deltawere cleared for farmland. About effect <strong>of</strong> this railroad was not<br />
half <strong>of</strong> the modern delta on Harsens Island particularly adverse.<br />
was diked.<br />
Several sportsmen's clubs were<br />
Three <strong>of</strong> trans~ortat'on which between 1850 and 1873 in<br />
were by the mid-1870s Ontarlo on portions <strong>of</strong> the delta which are<br />
provided improved access to the shoreline<br />
by boat, For example, in<br />
and An ra'lway was Ontario the <strong>St</strong>e, Anne Club and the Canada<br />
constructed the perimeter <strong>of</strong> the Cl ub were founded, while In Mlchigan the<br />
lake Detroit to A1gonacv <strong>The</strong> Grande Paint- and the Old Clubs were<br />
was On the h4gher ground' establ j shed. Hotels Lo accommodate summer<br />
generally av<strong>of</strong>dfng the remaining<br />
tourjsts from Detroit were constructed on<br />
between New BaltimOre and<br />
the South Channel levee. Since the levees<br />
*lgonacr the bed was constructed were law and subject to periodic flood'ing,<br />
through <strong>St</strong>. John's Marsh# a shall0w jnter- bul k-headf ng began to appear. Iso1 ated<br />
distributary bay col onired with emergent fill jng and bulkheadjng was most fntense<br />
aquatics, a1 ong South Channel f the ~aSn navigatfon<br />
A second mode <strong>of</strong> transport with channel) and In ii few Isolated localities.<br />
adverse impacts on the delta was the <strong>The</strong> first road on the <strong>St</strong>, ClaJr Delta was<br />
105
constructed on the west side <strong>of</strong> South<br />
Channel to allow access and settlement. In<br />
contrast, the shore1 ine from the ihames<br />
River northward remained unaltered except<br />
for 1 imited settlement in the northern<br />
port1 on <strong>of</strong> Wal pol e Is1 and.<br />
Channel ization for commercial nav-<br />
igatfon began in earnest In 1886. <strong>The</strong><br />
U. S. Congress authorized the deepening <strong>of</strong><br />
the ClSnton River (Figure 54) to 2.4 m,<br />
and <strong>Lake</strong> <strong>St</strong>. Claf r and the <strong>St</strong>. <strong>Clair</strong> Flats<br />
Channel to a depth <strong>of</strong> 8.4 m. By 1892 the<br />
<strong>St</strong>. <strong>Clair</strong> Rlver was dredged as we1 1. On<br />
the lower portlon <strong>of</strong> South Channel,<br />
channels were dredged through the natural<br />
levees into Big Muscamoot Bay and<br />
bulkheading and backfilling for summer<br />
houslng contlnued. By 1903 the modern<br />
delta wetlands (the shal low inter-<br />
distributary bay) <strong>of</strong> Harsens Island<br />
continued to be drained, diked, and used<br />
for agriculture.<br />
<strong>The</strong> period 1900-1930 was the<br />
beginning <strong>of</strong> urbanlzation. Channels and<br />
roads were constructed a1 ong Middle and<br />
South Channels. <strong>The</strong> electric rail road was<br />
removed and replaced by highway M-29,<br />
linking Algonac with communities to the<br />
west. Recreational boating facflfties<br />
began to appear. Urban areas and<br />
waterways expanded at the expense <strong>of</strong><br />
agriculture. Urbantzation more than<br />
doubled and agricultural land use declined<br />
to one-half in the <strong>St</strong>, John's Marsh area<br />
(Table 34 and Figures 55 and 56). Direct<br />
lmpacts on the wetlands contlnued.<br />
In 1945 Intense commercIa1 and urban<br />
growth began. Urban strip development<br />
extended from Detroit t o Algonac.<br />
Dredging, f i 11 f ng, and channel ization<br />
consumed and drained a sjgnificant portion<br />
af <strong>St</strong>, John's Marsh. Urban development<br />
also extended along the perimeter <strong>of</strong><br />
Harsens Is1 and. To improve waterfowl<br />
habitat, most <strong>of</strong> the remalnlng portion <strong>of</strong><br />
Marsens Island was diked and planted i n<br />
corn and other crops. Although most <strong>of</strong><br />
the Ontarto portfon <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Flats<br />
stfll remadns as the Walpole Island Indian<br />
Reserve, south <strong>of</strong> Wftchell Bay wetlands<br />
have been drained and channelized for<br />
agriculture. Dfkes were constructed to<br />
a1 1 e vi ate f 1 ood i ng and control water<br />
levels fn formerly unrestricted wet1 ands.<br />
<strong>The</strong> diking projects on Harsens Island and<br />
fn Ontario extended into the cattail<br />
marsh, Sedge and dogwood/meadow wet1 ands<br />
which are located at or just above the<br />
water table? were the areas initially<br />
drained for agrfculture. An analysis <strong>of</strong><br />
coastal land use, based on 1973<br />
topographfc maps during record high lake<br />
levels reveals that the only significant<br />
areas <strong>of</strong> intact wetlands in Michigan were<br />
within the interdistributary basins <strong>of</strong> the<br />
<strong>St</strong>. Cl ai r Flats and a small area south <strong>of</strong><br />
the Clinton River. In 1985 the Michigan<br />
wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> represented<br />
only some 29000 hectares <strong>of</strong> the land<br />
cover.<br />
Commerci a1 and recreational traffic<br />
has also had a considerable impact on <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. Bulk cargo and recreational<br />
boating have requ 1 red channel deepening.<br />
New channel and marine construction on the<br />
<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> shoreline continues.<br />
Public maintenance dredging to accommodate<br />
1 ake carriers and private recreational<br />
craft is estimated to be 122,000 m3<br />
annual 1 y ( Rap hael et a1 . 1974) exclusive<br />
<strong>of</strong> the <strong>St</strong>. Clalr River. <strong>The</strong> construction<br />
<strong>of</strong> highway 1-94 from Detroit northward to<br />
Port Huron has improved access to the <strong>Lake</strong><br />
<strong>St</strong>, <strong>Clair</strong> shoreline. Reduced driving time<br />
from Anchor Bay and Algonac to Detrolt has<br />
encouraged suburban devel opment,<br />
partlcularly along the western shoreline<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. All <strong>of</strong> these factors<br />
have stimul ated development <strong>of</strong> the<br />
shoreline and have had adverse impacts on<br />
many <strong>of</strong> the wetlands.<br />
In part, because <strong>of</strong> the exceptional<br />
fish and wildlife productivity and<br />
Table a34. Land use change at <strong>St</strong>. John's<br />
Marsh.<br />
Land<br />
use -.LcuL- ..1974<br />
ha acres ha acres<br />
Urban 126 313 320 793<br />
Agriculture 822 2,035 431 1,066<br />
Waterways 21 52 109 269<br />
a Data source: Roller (1977 1.
Figure 54. Aerial photograph <strong>of</strong> the mouth <strong>of</strong> the Clinton River, Macomb County,<br />
Michigan, showing a coastal wetland (Verchev's Marsh) isolated from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> by<br />
channelization and residential development (June 1975).
Figure 551, Aerial photograph <strong>of</strong> <strong>St</strong>. John's Marsh on Bouvier Bay, and North Channel,<br />
<strong>St</strong>, <strong>Clair</strong> Delta (April 1949), Note wetlands established during low lake levels. Com-<br />
pare with Figure 56,
Figure 56. Aerial photograph <strong>of</strong> North Channel, <strong>St</strong>, <strong>Clair</strong> De'l ta, and <strong>St</strong>. John's Marsh<br />
on Bouvier Bay (May 1980). Compare with Figure 55, Note wetlands lost during high<br />
lake levels, residential development, dredge spoil sites, and highway (M-29) bi fur-<br />
eating marsh.
ecreational value <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalrr<br />
visitors from Mfchigan and Ontarfo<br />
continue to be attracted to the lake. <strong>The</strong><br />
lake is readfl y accessible to the urban<br />
areas <strong>of</strong> metropolitan Detroit and Windsor<br />
as well as numerous small urban areas such<br />
as Flint, Saginaw, Port Huron, and Sarnia.<br />
<strong>Lake</strong> <strong>St</strong>. Cl air is the most valuable Great<br />
<strong>Lake</strong>s area for non-salmonid sport fish in<br />
Mlchlgan. For example, in 1975* nearly<br />
half <strong>of</strong> the total Great <strong>Lake</strong>s fishing<br />
effort was expended on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
(Jaworski and Raphael 1978). Furthermorer<br />
fur (rwskrat and raccoon) production <strong>of</strong><br />
the area is among the highest in the<br />
<strong>St</strong>ate.<br />
Wlth increased commerce 4n the Great<br />
<strong>Lake</strong>s, a cut<strong>of</strong>f channel was dredged at<br />
South Channel where it debouches into <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. To accomodate Seaway traffic,<br />
a 8-m channel was dredged in 1952 through<br />
the interdistributary marsh for a dfstance<br />
<strong>of</strong> 10 km. Seaway Island, a large disposal<br />
site now occupies a portion <strong>of</strong> the<br />
Jntardistributary marsh. To improve<br />
recreational boating facillties and<br />
mitigate against further erosion,<br />
bulkheading and filling along all<br />
urbanized shore1 ines have occurred over<br />
the decades. Seawalls extendfng we1 1<br />
above hfyh water levels have replaced the<br />
natural levees which extended south from<br />
Harsens Island into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, <strong>The</strong><br />
number <strong>of</strong> crevasse channels which<br />
transported water, sediment, and nutrients<br />
into Muscamoot Bay has decreased.<br />
<strong>The</strong> degeneratfon <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
wetlands follows a historical pattern<br />
similar to that <strong>of</strong> other large wetland<br />
complexes in the United <strong>St</strong>ates, Initially<br />
stlrnulated by the 1850 swamp acts,<br />
agricultural activity occurred. A second<br />
signfffcant stlmulant was access5billty<br />
dus t o ssveral transportatton<br />
devslapments. Inter-urban electric<br />
raf 1 roads, excursfon vessels, and<br />
eventually, roads and interstate highways<br />
made the wetlands and the shoreline<br />
readily accessible. Ef f l cient transport<br />
routes have encouraged urban and suburban<br />
expanston, changfng summer residences to<br />
year-round housing ,<br />
By contrast, access to the delta and<br />
the shoreline <strong>of</strong> Ontario is more dffficult<br />
because <strong>of</strong> the lack o f roads.<br />
Furthermore, southern Ontario is less<br />
populous and much <strong>of</strong> the eastern portion<br />
<strong>of</strong> the delta is an Indtan reserve, <strong>The</strong>se<br />
physical and social factors have not<br />
encouraged as much wetland loss as seen in<br />
Michigan* except at the northern end <strong>of</strong><br />
Walpole Island which has experienced<br />
wetl and loss.<br />
With the exception <strong>of</strong> the electric<br />
railroad through <strong>St</strong>. John's Marsh9 wetland<br />
losses occurred on the periphery <strong>of</strong> the<br />
resource; that is, modification began at<br />
the boundary between the marsh and the<br />
adjacent upland. In the <strong>St</strong>. <strong>Clair</strong> Delta<br />
agriculture was initiated on the premodern<br />
surface and gradual 1 y expanded southward<br />
to the transition (dogwood/meadow) zone<br />
and eventual f y into the shall ow inter-<br />
distributary bays. <strong>The</strong> natural levees<br />
were modifled by bulkheading in the 1870s.<br />
Filling, aided by bulkheads, extended into<br />
the deep interdistributary bays and onto<br />
the river shoulders. This pattern <strong>of</strong><br />
development has continued to the present<br />
because wetland boundaries are difficult<br />
to ascertain, especial 1 y i n the Great<br />
<strong>Lake</strong>s where temporary lower lake levels<br />
provide a false perception <strong>of</strong> the position<br />
<strong>of</strong> the shore1 ine.<br />
Adverse impacts to the artificial<br />
(diked wetrands1 and 11 natural<br />
envi ronments identif led in section 4.1 as<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands are shown in Table<br />
35. With the exception <strong>of</strong> the recently<br />
abandoned channels, which are in Ontar i or<br />
and the transgressive beaches which are<br />
generally Is01 ated, al? the wet1 ands have<br />
had adverse impacts. Bulkheads and fills<br />
have been constructed on the river<br />
shoul ders, crevasses, and margins <strong>of</strong> the<br />
deep-water Interdistributary bays for over<br />
a century. To construct roads, spoil was<br />
dredged from the landward side <strong>of</strong> the main<br />
channels which inadvertently led to the<br />
channel ization <strong>of</strong> the shallow bays.<br />
Bulkheads are a common structural feature<br />
along many shorelines <strong>of</strong> the lake and<br />
delta and appear to be detrimental to<br />
submersed aquatics ( Schl oesser and Manny<br />
1982). Bulkheads constructed on river<br />
shoulders occupy the shallowest portion <strong>of</strong><br />
the rlvsr. Consequently, as dredge and<br />
f i 11 occursr submersed wetl ands are<br />
df spf aced i akeward where bottom attachment<br />
fs more difffcult because <strong>of</strong> stronger<br />
currents and greater depths.
Table 35. Adverse impacts to the various morphology types <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> wetlands.<br />
Wet1 and<br />
morphology type<br />
Impact Exampl e<br />
1. River shoulders bulkheaded, South Channel<br />
dredged dl sposal<br />
2. Abandoned river drainedr agriculture Harsens Island<br />
channels<br />
3. Recently abandoned no change<br />
channels<br />
4. Crevasse deposits channel 1 zed,<br />
bulkheaded<br />
Bassett Channel<br />
South Channel<br />
5. Shallow inter- di ked/drained, Harsens Is1 and<br />
dlstrlbutary bays agrtculture~<br />
channel i zed<br />
6. Deep lnterdis- bulkheadedr<br />
tributary bays channel ized<br />
Harsens Is1 and<br />
7. Transgress1 ve minor urban Big Muscamoot Bay<br />
beaches settlement,<br />
bul kheaded<br />
* ,<br />
8. Shallow shelves dikedr filled, Mitchell Bay<br />
channel i zed<br />
9. Barrier/l agoon channel fzed<br />
*<br />
~orth <strong>of</strong> Thames River<br />
10. Transi tional areas diked, agriculturer<br />
filled for residenti<br />
a1 devel opment<br />
Harsens Is1 and<br />
11. Estuarl ne/<br />
floodbasf n<br />
12. Diked<br />
diked/drainedr <strong>St</strong>. John's Marsh<br />
agriculture/urban<br />
d i ked/drai ned<br />
East shore1 i ne<br />
(orfgf nally shelves) (O~tarlo)
To accommodate rec reattonal boating<br />
and commerci a1 shippl ng, channels to<br />
depths <strong>of</strong> 8 m have been dredged on the<br />
perimeter and In the basin <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />
<strong>Clair</strong>. Also there is considerable<br />
residential pressure on the shore1 fne to<br />
include the wetlands <strong>of</strong> the basin,<br />
Bu?kheadtngr ftlling, dtking9 and djsposal<br />
<strong>of</strong> dredged materials have occurred since<br />
the early 1900s.<br />
Because <strong>of</strong> the need for commercial<br />
navigation and recreational boating fn<br />
take <strong>St</strong>. <strong>Clair</strong>, a significant amount <strong>of</strong><br />
channel dredging has occurred. <strong>The</strong> U.S.<br />
Army Corps <strong>of</strong> Engineers, which Is<br />
responrj-i ble for harbor maintenance, has<br />
conducted dredging and disposal activities<br />
at the Cl inton and <strong>St</strong>. Claf r Rfvers and<br />
the 8 m navigation channel bisecting the<br />
lake, In Ontario the Department <strong>of</strong> Public<br />
Works mafntains the <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f<br />
Channel and the harbor at Wallaceburg,<br />
east <strong>of</strong> the <strong>St</strong>, Clalr Rlver Delta. To<br />
maintafn channels at the proscribed<br />
depths, the aceumul ated sedlment was<br />
normal 1 y hydraul Jcall y dredged and<br />
dfsposud <strong>of</strong> in convenient areas such as in<br />
designated portions <strong>of</strong> the open lake.<br />
<strong>The</strong> volume <strong>of</strong> materfals dredged for<br />
take <strong>St</strong>. ClaJr and the <strong>St</strong>, <strong>Clair</strong> River<br />
navjgatlon projects Is shown on Table 36.<br />
Over the 52-year perJodp about 782,800 m3<br />
were dredged from U.S, project sites in<br />
the two areas. More recent data indicate<br />
that from 1971 through 1984 dredging<br />
activity in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> totaled 382r300<br />
m3, <strong>The</strong> average for the 14-year period Is<br />
27t300 m3 annually which is considerably<br />
less than the historlcal average <strong>of</strong><br />
313,200 m3 for <strong>Lake</strong> <strong>St</strong>. Clalr. On the<br />
Cl inton RIver the 1971-1984 total was<br />
45,000 d, which approximates the historic<br />
average.<br />
<strong>The</strong> Ontario maintenance dredging and<br />
new work dredging averaged about 195,800<br />
m3 annually. Most <strong>of</strong> the dredging in<br />
Ontario is confinad to the <strong>St</strong>, <strong>Clair</strong><br />
Cut<strong>of</strong>f Channel (182,400 d/year).<br />
In 1970 the U.S. Congress enacted<br />
Pub1 dc Law 91-611 (River and Harbor Flood<br />
Control Act), which authorized the Corps<br />
to construct, operate, and malntain<br />
confined disposal sites for containment <strong>of</strong><br />
polluted dredged spoil in the Great <strong>Lake</strong>s<br />
for a period not to exceed 10 years. At<br />
that timer the <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel<br />
and <strong>Lake</strong> <strong>St</strong>, Clajr sediments were 100%<br />
polluted9 and the Clinton River was 85%<br />
polluted (Raphael et al. 19741. <strong>The</strong><br />
pollutant was primarily mercury, although<br />
the conventional pollutants were also<br />
beginning to reach limits set by the U.S.<br />
Table 36. Hiituricaa dredging total 5 <strong>of</strong> Corps <strong>of</strong> Engineers projects in <strong>Lake</strong><br />
<strong>St</strong>. Cldir 1320-1972.<br />
---<br />
Mafntenance New work Annual<br />
d redgi ng dredgl ng Total average<br />
Channel oF Laire <strong>St</strong>. <strong>Clair</strong> 2~894r400 13~634.600 1Sr529,000 317,900<br />
Cl inton Rlver 237 r 400 96,900 334,300 6,400<br />
<strong>St</strong>. Cl air River 3 ,125,300 20~879~200 24r004r500 461,600<br />
a Data source: Raphael et aS, (19741.<br />
i 12
Envl ronmental Frotactfon Agency. To<br />
comply wlth PL 91-631, two conflned<br />
disposal sites with a combtned cdpaclty <strong>of</strong><br />
1.5 mllfion m3 were constructed at the<br />
north end <strong>of</strong> Dfckinson Island. Both sltes<br />
were placed on tho premodern surface <strong>of</strong><br />
the <strong>St</strong>. <strong>Clair</strong> Delta. <strong>The</strong> west site<br />
enclosed an area <strong>of</strong> about 22 hectares and<br />
the east site about 48 hectares (U.S. Army<br />
Corps <strong>of</strong> Engineers 1974). In Ontarfo the<br />
<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel was constructed<br />
in the early 1950s to accommodate Seaway<br />
traffic along South Channel, <strong>The</strong> channel<br />
was dredged through the fnterdistrlbutary<br />
bay and was 1 inked to the main navigation<br />
channel in <strong>Lake</strong> <strong>St</strong>. Clalr. Dredge spoll<br />
was put Sn a disposal sltu (Soaway Island)<br />
along the east bank <strong>of</strong> South Channel. <strong>The</strong><br />
facflity covers an area <strong>of</strong> about I1 km2,<br />
which was prlmarlly colonized wfth<br />
emergent macrophytes typical <strong>of</strong> an Inter-<br />
dl stributary bay snvl ronment.<br />
Private or contract dredglng (under<br />
Sec.10 permit <strong>of</strong> the U. S. River and<br />
Harbor Act <strong>of</strong> 1899) is done by local<br />
cftizens or homeowners to fmprove access<br />
to <strong>Lake</strong> <strong>St</strong>. Clalr or to construct<br />
boatslfps along the lake or the<br />
dfstributaries (Figure 57). <strong>The</strong> data are<br />
not normally tabu1 ated; however, based on<br />
our estimates, private dredging volumes<br />
probably average about 7,600 to 11,400 m3<br />
annually. SSgnff tcant areas <strong>of</strong> dradglng<br />
for restdential development have occurred<br />
Figure 57. Commercial development tin Har-<br />
sens Is1 and wetland (August 1984).<br />
on the shoreline <strong>of</strong> Anchor Bay, <strong>St</strong>. John's<br />
Marsh, at the mouth <strong>of</strong> the Clinton RIver<br />
and along the distrfbutary channels <strong>of</strong> the<br />
dsl ta, parttcul arl y North and South<br />
channels. Prfvate dredging in btarfo is<br />
lim~ted to approach channels for<br />
recreational craft Jn Mitchell Bay and a<br />
marina complex near Belle River on the<br />
south shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr.<br />
Dfking fragments coastal wetlands,<br />
separating the managed parcels from the<br />
adjacent littoral and upland environments.<br />
General1 y, as the slre <strong>of</strong> the wetland is<br />
diminfshed, so Is the vegetative and<br />
wildlife diversity (Larsen 1973).<br />
Furthermore, dik 1 ng <strong>of</strong> wet1 ands reduces<br />
vegetation diversity because water level<br />
controls are established to maintain a<br />
particular vegstatfon type or envlron-<br />
mental condftlon. <strong>St</strong>abll $zing water<br />
levels tends to eliminate those plant<br />
species whfch raqul re low-water periods<br />
for regenerat 1 on, and promotes domfnance<br />
<strong>of</strong> a few tolerant spectes such as cattafl<br />
( spp. 1 and woody plants (Keddy and<br />
Reznfcek 1984).<br />
<strong>The</strong> hydrologic isolation <strong>of</strong> dlked<br />
systems 1s reflected In their greatly<br />
reduced flsh use and simpllfled<br />
1 nvertebrate commun5t~esr particularly<br />
where wlnter ice and summer temperatures<br />
reduce df ssol ved oxygen and otherwise<br />
reduce the water qua1 ity. <strong>The</strong> effect <strong>of</strong><br />
earthen dtkes on waterfowl mfaratlon,<br />
other wild1 ffe movementsr and prtrdatfon Is<br />
not well known. Howevert DennTs and North<br />
(1981) have shown a dramatlc decline in<br />
the use <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> marshes by<br />
true marsh-dwellfng specles such as the<br />
Amerlcan wigeon (<br />
), green-wfnged tea2 (<br />
ood duck (&.&<br />
1968-69 to 1976-77 the use by these<br />
dabbl lng ducks decl fned by 73% durf ng the<br />
spring pertod. Autumn marsh use also<br />
declfned by 40%. Howeverr the<br />
investigators noted that sprlng use<br />
generally provides a batter f ndicatlon <strong>of</strong><br />
the value <strong>of</strong> the wetlands to mfgrant<br />
waterfowl si nee the managemnt techn l quas<br />
whfch were used during the fall huntfng<br />
seasons are not Implemented during the<br />
sprf ng,
Discovery <strong>of</strong> mercury contaml natfon In<br />
tha <strong>St</strong>. Clafr Rfver-<strong>Lake</strong> <strong>St</strong>. Clafr-Detrolt<br />
RJver-<strong>Lake</strong> Erie system in 1970 generated<br />
great pub1 l c Interest In the mercury<br />
levels <strong>of</strong> water and flsh, Subsequent<br />
banntng <strong>of</strong> sport and commercial ffshlng<br />
led to testing for trace mercury by<br />
varjous <strong>St</strong>ate and Federal agencies to<br />
def f ne the extent <strong>of</strong> the pollution and to<br />
determf ne its major sources. Two chemfcal<br />
plants, Dow Chemical Corporatlon, Sarnia*<br />
Ontarfo, and BASF Wyandotte Chemical<br />
Corporatlon, Wyandotte, Mfchfgan, were<br />
fdentff fed as major contributors to the<br />
mercury pollution <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />
<strong>Lake</strong> Erie (Waiters et al. 19741, <strong>The</strong>se<br />
companfes manufactured chlorf ne gas and<br />
caustlc soda by the alactrolytlc process,<br />
whlch lnvolves the slectrolysis <strong>of</strong> brfne<br />
to produce chlorlne gas and a mercury-<br />
sodium metal ama'lgarn. <strong>The</strong> amalgam 1s<br />
contacted wlth water to produce sodium<br />
hydroxf de, the mercury metal belng<br />
recycled ta tho afectrotytfc coll, <strong>The</strong><br />
rate <strong>of</strong> accfdental mercury release from<br />
these two plants has been sstfmated to be<br />
25 kg per day from 1950 to 1970 for the<br />
Dow Chsmlcal plant and S to 10 kg per day<br />
from 5939 to 1970 for the OASF Wyandotte<br />
plant (Federal Water Oualfty<br />
Admfnfstratfsn l970).<br />
Waltars @t at, (19741 calculated that<br />
228 metrlc tons <strong>of</strong> mercury pol Jution had<br />
been laadad to the top 20 cm <strong>of</strong> western<br />
<strong>Lake</strong> Erie sediment durlng tho p~rlod 1939<br />
to 1971, Sodlment cores takan at the<br />
mouth <strong>of</strong> the Detr<strong>of</strong>t Rfver and wastern<br />
<strong>Lake</strong> Erica y folded surfaca mercury valutls<br />
up to 3.8 ppm whleh generally decreased in<br />
concentratton exyonentfally wfth depth<br />
f FIgurs $81. Several yoars after the<br />
chlor-alkalf plants diminished aperation<br />
ths area was agafn carad (Wflson and<br />
Wdlturs 19789 vfth analyses shovfng that<br />
recent deposits covered the high1 y<br />
contrrnlnated sedfnrcsnt wfQh a thrn layer <strong>of</strong><br />
new material vhlch had mercury<br />
concentrat l ons approaching backgraund<br />
lsvsls 6B,b ppm). As a result af these<br />
discharges, marcury fn flsh <strong>of</strong> take <strong>St</strong>.<br />
Clnlr and western <strong>Lake</strong> Erfe was a major<br />
contaminant problem In the early 1970s.<br />
Levels <strong>of</strong> total mercury in walleye<br />
C L 1 col lectsd from<br />
Figure 58. Mercury concentration in sedi-<br />
rrient core from western <strong>Lake</strong> Erie near<br />
mouth <strong>of</strong> Detroit River (Halters et al.<br />
1974). Note mercury-enriched surface zone<br />
over1 yi ng sediment havi ny natural back-<br />
ground mercury level s (dashed 1 ine).<br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> have decl ined from over 2<br />
ug/g In 1970 to 0.5 ug/g In 1980. In<br />
western <strong>Lake</strong> Erie, 1968 levels <strong>of</strong> mercury<br />
were 0.BA3ug/g as compared to only 0.31<br />
ug/g in 1976. <strong>The</strong> rapid environmental<br />
response subsequent to the cessation <strong>of</strong><br />
the point source discharges at Sarnia,<br />
Ontarlo, and Wyandotte* Michigan, can be<br />
attributed to rapid flushing <strong>of</strong> the <strong>St</strong>.<br />
Cl a1 r-Detroit Rf ver system, the high load<br />
<strong>of</strong> suspended sediment delivered to western<br />
<strong>Lake</strong> Erier and t h e high rate <strong>of</strong><br />
productivity (International Jolnt Com-<br />
missjon 1981).<br />
Mudroch and Capob-ianco (1978) studied<br />
the relatlonshlp between the concentration<br />
<strong>of</strong> seven metals In sediment and<br />
marshwater, and uptake <strong>of</strong> these metals by<br />
emergent, submersed, and fl oat1 ng-l eaved<br />
plants growing In wetlands located on the<br />
east shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>y found<br />
a hf gh correl atton between metals and<br />
organfc carbon contents In the sediments.<br />
<strong>The</strong> accumulation <strong>of</strong> metals In plants<br />
varied fram specfes to species (Table 37)<br />
and showed a complex relatfanshlp with<br />
metal concentratfons i n the sed i ment .<br />
Water-mf l foil ( spp.) and<br />
pondweed ( lated more<br />
metal than the ather aquatic plants.
Table 37. Metals concentratiorls in livirly plant tissues from Big Point<br />
Marsh, Ontario on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. a<br />
Metals (ggm)<br />
Specles Pb Zn C r N! Cd Go Cu<br />
a Data source: Mudroch and Capobianco (1978).<br />
Metal concentrations In roots <strong>of</strong> broad-<br />
leaved cattat purple<br />
loosestrjfe ( , and<br />
pickerel weed were<br />
found to be hlgher than in the aboveground<br />
biomass. <strong>The</strong> metals taken up by marsh<br />
flora are retafned In the tissues and<br />
after decay they become a part <strong>of</strong> the<br />
marsh surface sediment.<br />
Jaworskf and Raphael (1976) compiled<br />
comprehensfve data on wetland loss for the<br />
Mlchfgan sjde <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. In 1873<br />
the U.S. portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
supported 7,274 hectares <strong>of</strong> wetland<br />
vegetation (TabTe 381. By 1973 this<br />
habitat had dwlndled to 2,020 hectares.<br />
Significant losses not on1 y occurred on<br />
the <strong>St</strong>. <strong>Clair</strong> Delta and <strong>St</strong>. John" Marsh<br />
but on the entire margin <strong>of</strong> the lake as<br />
well. Gaukler Polnt at the head <strong>of</strong> the<br />
Detroit River was colonized with 187<br />
hectares <strong>of</strong> wetlands and the Clfnton River<br />
had over 1,295 hectares <strong>of</strong> habitat at its<br />
mouth and f t s flood basln. Some coastal<br />
areas* particular1 y north <strong>of</strong> the Cl i nton<br />
River appear to have been drained for<br />
agriculture in the 1860s suggestfng that<br />
the 1868-1873 data represent rninjrnum<br />
rather than maxlmum wetland acreage.<br />
In contrast to Michigan's wetlands<br />
being lost to the ong<strong>of</strong>ng urbanizationr<br />
Ontarfo wetlands are being lost to<br />
agriculture, <strong>The</strong> wetlands from the Thames<br />
River north to Chenal Ecarte dwindled Prom<br />
about 3,600 hectares in 1965 to 2,900<br />
hectares in 1970 (McCul lough 1982). About<br />
700 hectares or nearly 20% <strong>of</strong> the<br />
prlvately owned resource base was lost<br />
over the 5-year period. Figure 59 and<br />
Table 38 reveal specific areas <strong>of</strong> wetland<br />
loss <strong>of</strong> the coastal Ontarlo wetlands<br />
excluding the Walpole Island Indian<br />
Reserve. Drai nlng for agricultural use<br />
has accounted for 91% <strong>of</strong> the wetland loss<br />
whereas marina and cottage development has<br />
consumed 9% <strong>of</strong> the resource. Clearly,<br />
urban pressure Is presently a problem In<br />
Michigan and agriculture pressure a<br />
problem in coastal Ontario,<br />
Although the Ontario shore <strong>of</strong> <strong>Lake</strong><br />
<strong>St</strong>, <strong>Clair</strong> is low lying and subject to<br />
flooding, erosion has on7 y been tdentified<br />
as a hazard from the Thames Rfver westward<br />
(Boul den 1975). This suggests that<br />
permanent loss <strong>of</strong> wetland habitat north <strong>of</strong><br />
the Thames River 1s due t o human<br />
interventton. During the record high lake<br />
l eve1 fn the early 1970~~ about lrOOO<br />
hectares <strong>of</strong> emergent shore1 f ne marsh f rom<br />
Mitchell Bay southward to the Thames River<br />
were temporarily lost (McCullough 19821.<br />
This loss was tempered in part by the<br />
flooding <strong>of</strong> transitjon vegetation which<br />
occurred on the upland (east) margfn <strong>of</strong><br />
the wetlands, Slmflarly the delta and<br />
Anchor Bay are subject to flooding, though<br />
erosion ds not identified as a serfous<br />
problem (Great <strong>Lake</strong>s Basfn Commission
Table 38. -<br />
a<br />
Michigan and Ontario wetland losses on <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />
Mlchigan<br />
1 ocat i on<br />
1868-73<br />
(ha)<br />
1973<br />
(ha) Loss (ha)<br />
<strong>St</strong>. <strong>Clair</strong> Flats 5,473<br />
Swan Creek 75<br />
Marsac Point 6 1<br />
New Baltimore 2 1<br />
Salt River 162<br />
Cl i nton River 1,295<br />
Gaukler Polnt 187<br />
Total 7,274 2 r 022 5,252<br />
Ontar lo 1965 to 1973 Wet1 and<br />
1 ocat i on Area loss (ha) type Cause<br />
Thames Rfver 59<br />
Thames Rlver mouth 7 4<br />
Bradley Marsh 327<br />
Balmoral Marsh 11<br />
Snake Island Marsh 156<br />
<strong>St</strong>. Lukes Bay 22<br />
Patricks Cove 5 9<br />
Mitchell Bay 7<br />
Mud Creek Marsh 167<br />
Tota 1 882<br />
diked<br />
open<br />
diked<br />
diked<br />
diked<br />
diked<br />
diked<br />
d I ked<br />
d l ked<br />
agriculture<br />
mari ne/cottage<br />
construction<br />
agriculture<br />
agriculture<br />
agriculture<br />
agriculture<br />
agriculture<br />
agriculture<br />
agriculture<br />
a Data sources: Jaworski and Raphael (19761, McCullough (1982).<br />
1975). <strong>The</strong> recent losses here are due to<br />
dlktng and/or ffllfngr fn most Instances,<br />
Par urban growth.<br />
<strong>The</strong> wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> hav~<br />
been mapped from older navfgatton charts<br />
and recent topagraphlc maps CFlgure 60).<br />
Thfs figure illustrates signiffcant losses<br />
between 1873 and 1968 on the periphery <strong>of</strong><br />
the lake, However, the most apparent<br />
1 mpacts are evident in the Clinton River,<br />
the <strong>St</strong>. Clai r Delta* and the eastern shore<br />
<strong>of</strong> the lake. It is ev4dent that the<br />
wetlands in all three areas have been<br />
modified along their margins. <strong>The</strong><br />
wetlands on the eastern shore1 ine were<br />
approximately 2.5 km wide. Since 1873<br />
they have been impacted from the landward<br />
side and are now about 0.8 km in width.<br />
<strong>The</strong> progression <strong>of</strong> losses Jn the <strong>St</strong>. <strong>Clair</strong><br />
Delta follows a similar geographic<br />
pattern. <strong>The</strong> losses in both areas were<br />
fnf tial ly stimulated by agriculture. In<br />
the Cl inton River area, wetl and losses<br />
occurred from both the landward as well as<br />
the 1 akeward boundary. Fundamental lyr an<br />
1 sol ated wetl and has been created.
Figure 59. Wetland losses along the Ontario shore1 ine <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. C'lair<br />
(McCul lough 1982).
Figure 60. Comparison <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r<br />
1968.<br />
Table 39 represents a summary <strong>of</strong><br />
wetland losses in <strong>Lake</strong> <strong>St</strong>. Clalr. Since<br />
the topographic quadrangles and the<br />
navigation charts only dlsplay emergent<br />
wetlands, the area values <strong>of</strong> submergent<br />
aquatics were estimated. <strong>The</strong> re1 atlve<br />
1 osses between the historical perf ods are<br />
evldent. <strong>The</strong> losses on Table 39 are<br />
greater than that noted by Jaworski and<br />
Raphael (1976) because thais data did not<br />
include what is now surm!sed to be<br />
submersed vegetation, <strong>The</strong> percentage <strong>of</strong><br />
wetland area lost fn Michfgan is greater<br />
than In Ontario (45% vs. 34X)r but the<br />
actual area lost in Ontario exceeds that<br />
lost in Michigan. In Michigan as well a5<br />
fn Ontario* more <strong>of</strong> the wetland base was<br />
l ost along the shore1 ines than in the<br />
delta. Slnce the coastal zone wetlands<br />
are parallel to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, their<br />
accesslbilfty plays an faportant role in<br />
thefr modification. <strong>The</strong> <strong>St</strong>. Clafr Delta<br />
conversely projects into <strong>Lake</strong> <strong>St</strong>. Clafrr<br />
limfting accessibi7ity and hence adverse<br />
LAKE<br />
38.968<br />
coastal wetlands distribution in 1873 and<br />
impacts. It should not be inferred from<br />
the data that the wetland quality and<br />
diversity have not changed over the 95-<br />
year period. In sum, 9r 136 hectares <strong>of</strong><br />
the resource base have been lost.<br />
Both natural disturbances, such as<br />
water-level fluctuations (as the Great<br />
<strong>Lake</strong>s water levels oscill ate) as we1 7 as<br />
cultural disturbances ( ~ . g . diking) ~<br />
affect the wetland ecosystems. A s<br />
reflected In Eugene Odurnts qlpulse<br />
stability concept," some <strong>of</strong> these<br />
disturbances can be beneficial. In this<br />
section, however, the lmpacts <strong>of</strong> adverse<br />
cultural effects have been reviewed. Of<br />
particular concern are the following: 1)<br />
wet1 and loss* 2) dikl ng, fragmentationr<br />
and loss <strong>of</strong> hydrologic connectivity and<br />
3) changes in the environmental gradient<br />
and plant communities.
Table 39. Comparison <strong>of</strong> 1873 and 1968 <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetland areas.<br />
Location<br />
<strong>St</strong>. <strong>Clair</strong> Delta 5,414 3,077 9,641 7,234 15,055 10,311<br />
Cl 1 nton River 19192 248 -- -- 1,192 248<br />
Remaining Mich.<br />
shore1 I ne 1,900 806 -- -- 1,900 806<br />
Remai njng Ontario<br />
shore1 1 ne -- -- 4,219 1,862 4,219 1,862<br />
-- -- -_IL<br />
Total 7,506 4,131 13,860 9,096 22,366 13.227<br />
As discussed in this section, during<br />
the period 1873-1968? approximately 9,136<br />
hectares <strong>of</strong> coastal wetlands have been<br />
lost. This amounts to a 41% reductlon in<br />
the original coastal wetl and resource base<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> greatest<br />
percentage loss has been on the American<br />
side and the greatest area loss on the<br />
Canadian side. More wetlands have been<br />
destroyed along the shorel ines than in the<br />
delta. In additionr except for Dickinson<br />
Island and <strong>St</strong>. John's Marsh* the upper<br />
zones (i.e.# the sedge marsh-shrub-swamp<br />
fringe) were converted to other land uses.<br />
<strong>The</strong> juxtapositfon <strong>of</strong> corn fields and diked<br />
cattail marshes on <strong>St</strong>. Anne Island is a<br />
case in point. At present, nearly half <strong>of</strong><br />
these extant wetl ands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
consist <strong>of</strong> cattail communities.<br />
Loss <strong>of</strong> coastal wetlands along the<br />
Michigan sfde <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr results in<br />
a loss <strong>of</strong> wetland function and value. For<br />
example, public drains installed to<br />
improve run<strong>of</strong>f now occupy former creek<br />
channels, which no longer benefit from the<br />
flood water storage, sedfment trapping,<br />
and nutrient uptake afforded by the<br />
natural wetlands. Nor do the remalnlng<br />
wetlands along the river mouths and<br />
shorel ines, which have been reduced f n<br />
size, partially developed (especi a1 1 y on<br />
the lakeward sfdeIo and otherwfse<br />
impacted, have the fish and wildlife<br />
valves they once had.<br />
hectares* fn the <strong>St</strong>. Clafr Delta are now<br />
diked. This includes <strong>St</strong>. John's Marsh,<br />
which is isolated from <strong>Lake</strong> <strong>St</strong>. Clafr by<br />
earthen berms, but is not subject to water<br />
level controls. With the exception <strong>of</strong> <strong>St</strong>.<br />
John's Marsh, the upper part <strong>of</strong> the <strong>St</strong>.<br />
<strong>Clair</strong> Flats <strong>St</strong>ate Game Area, and a few<br />
other small parcels, over 75% <strong>of</strong> the diked<br />
wetlands are colonlred by cattails and<br />
associated submersed aquatic comunlties.<br />
<strong>The</strong> phrase "diked cattail marshesn is<br />
indeed appropriate. What is important<br />
here is that it fs the remainfng open-<br />
system wetl ands which are contributing to<br />
flsh production, wild1 lfe habitat, and<br />
dlving duck feeding <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. In<br />
contrast, the diked wetlands provide more<br />
<strong>of</strong> a single purpose - that <strong>of</strong> attracting<br />
dabbling ducks, especially mallards for<br />
waterfowl hunters.<br />
<strong>The</strong> adverse impact <strong>of</strong> isolatfng and<br />
fragmenting wetlands by means <strong>of</strong> roadbeds,<br />
canals, earthen dikes, and other<br />
developments appears not to be fully<br />
recognfzed. For example, many conser-<br />
vat1 onists 1 n Michigan are advocating the<br />
preservation <strong>of</strong> <strong>St</strong>. John's Marsh, but feb<br />
call for an increase In Its hydrologfc<br />
connectfvfty to <strong>Lake</strong> <strong>St</strong>, Clafr. more over^<br />
unless a wetland fs physically destroyed,<br />
not mere1 y f ragmentedr most wet1 and<br />
researchers would not refer to an Is01 ated<br />
wetland as be'lng gtlost.gt<br />
Approxlmately one-half <strong>of</strong> the Current coastal development not on1 y<br />
remafnfng coastal wetlands, f .e., 5,280 results .fn fragmentation and loss <strong>of</strong><br />
119
hydrologic connectivity, but also<br />
frequently Is assocfated with the loss <strong>of</strong><br />
As indtcated in Appendix M and on<br />
Figure 61r most <strong>of</strong> the ownership <strong>of</strong> the<br />
upper p1 ant communities (Jaworski and larger* extant coastal wetlands 1 ies in<br />
Raphael 1976). <strong>The</strong>refore, most <strong>of</strong> these public hands. Exceptions are <strong>St</strong>, Johnts<br />
extant wetlands conslst <strong>of</strong> just cattail<br />
and submersed aquatfc communities, rather<br />
Marsh, whlch the <strong>St</strong>ate <strong>of</strong> Michigan is<br />
attempttng to purchase from the remal n l ng<br />
than a compl eta wet1 and communfty private ownersr and Walpole Is1 and,<br />
contfnuum. Changfng <strong>of</strong> water levels In Ontarior which forms an Indian Reserve.<br />
the Great <strong>Lake</strong>s w i l l t then, result in the Many <strong>of</strong> the undfked coastal wetlands on<br />
loss <strong>of</strong>the current functfon fn a given<br />
wetland as opposed to the shift <strong>of</strong> this<br />
theCanadian side <strong>of</strong> the <strong>St</strong>. CJaSr Delta<br />
are owned by the Province <strong>of</strong> Ontario, wlth<br />
function laterally in accord with the the exception <strong>of</strong> Walpole Island, Canada's<br />
vegetatfon movements. In contrast, as<br />
exemplified by Dickinson Island, wetlands<br />
Department <strong>of</strong> the Environment (DOE) owns<br />
the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> National Wildlife Areap<br />
which are connected directly to <strong>Lake</strong> <strong>St</strong>. whereasDucks Unlimited has ownership OF<br />
<strong>Clair</strong> and sxhlbit a full environmental<br />
gradient, tend to experience lateral<br />
diked marshes south <strong>of</strong> Mitchell Bay.<br />
shifts In function and values.<br />
Furthermore, they are mafntained at little<br />
<strong>The</strong> land ownership <strong>of</strong> the Michigan<br />
portion <strong>of</strong> the delta is shown in Figure<br />
or no cost to the pub1 ic. 62. Two open water parcels, f.e.* Units A<br />
and B <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Waterfowl Refuge,<br />
Small, isolated wetlands near<br />
developments such as suburban housing*<br />
marinas, etc., oxhibit proximity and <strong>of</strong>fsite<br />
Impacts. Proximity impacts include<br />
ambfent noise levels as well as human and<br />
pet intrusions. Off-site affects center<br />
on nutrfent and sediment loading resulting<br />
f rem wlnd and water transport mechanisms.<br />
Xf the extant parcel <strong>of</strong> wetland is zoned<br />
for resldentlal or some other intensive<br />
use, there are pressures for filling and<br />
development. Fl rep as a disturbance,<br />
seams to be limfted to the cattafl and<br />
sedge marshes <strong>of</strong> Dickinson Island<br />
(Jaworskl et a1, 1979).<br />
5.2 WETLAND OWNERSHIP AND<br />
MANAGEMENT<br />
Wetland ownarship Js in the hands <strong>of</strong><br />
individuals as we91 as <strong>St</strong>ate, provincial<br />
and Federal agencies. <strong>The</strong> management <strong>of</strong><br />
the wetlands by these groups follows<br />
df verse strategies. <strong>The</strong> current ownership<br />
<strong>of</strong> the coastal wetlands as well as the<br />
zoning and long-range plans wf 11 strong? y o 5 10 15 20<br />
affect the future use <strong>of</strong> these Slatule M~les<br />
envfronmenls. In thfs section, an attempt<br />
has been made to project the future land Figure 61. Location map <strong>of</strong> individual<br />
use <strong>of</strong> these areas, and by usinpa those coastal wetlands <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />
p r ogect 1 ans address the issues <strong>of</strong> River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ecosystem; character-<br />
managemnts restorationl and arti f i ci a1 istics and ownership <strong>of</strong> each wetland are<br />
construct don <strong>of</strong> wet1 ands, given in Appendix M.<br />
728
Figure 62. Ownership map <strong>of</strong> the Michigan portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta.<br />
are administered by the Michigan DNR.<br />
Approximately 40% <strong>of</strong> Dickinson Island is<br />
in private ownership, especially the<br />
northern and eastern shores as well as a<br />
block in the south-central area. In<br />
addition, the U,S. Army Corps <strong>of</strong> Engineers<br />
manages two disposal sites on Dickinson<br />
Is1 and along North Channel, and there are<br />
private bottom1 and patents along the<br />
distributary channels <strong>of</strong> both Dickinson<br />
and Harsens Islands. Although the<br />
Department <strong>of</strong> Natural Resources is slowing<br />
the residential and summer home<br />
development along these dfstributary<br />
channels, the land remains in private<br />
ownership. At present, the lower part <strong>of</strong><br />
Harsens Is1 and, and much <strong>of</strong> Dickinson<br />
Is1 and, and those unpatented wet1 ands and<br />
adjacent shall ow-water bottomlands would<br />
be considered part <strong>of</strong> Michigan's <strong>St</strong>. <strong>Clair</strong><br />
Flats <strong>St</strong>ate Game Area. Ownership <strong>of</strong> the<br />
small, undj ked wet1 and parcels scattered<br />
along Anchor Bay and along the westerns<br />
eastern, and southern shores sf <strong>Lake</strong> <strong>St</strong>.<br />
Cl air is generally private.<br />
At present, in the delta there are<br />
two basic management stradagjes: 1)<br />
natural and 2) managed for waterfowl<br />
hunting and perhaps breeding as well. <strong>The</strong>
pub1 icly owned* open wetland systems are<br />
natural responding to seasonal and 1 ong-<br />
term water level fluctuatlons. In<br />
comparf son, the diked wetl ands consf st<br />
largely <strong>of</strong> cattail marshes that are<br />
managed by means <strong>of</strong> electric pumps, flood<br />
gates and weirs. In several managed<br />
marshes, crops are grown, then flooded to<br />
facll itate waterfowl feedfng.<br />
No sf gnif fcant changes in the wetland<br />
management practices are antlclpated. It<br />
fs expected that the status quo w i l l<br />
generally continue. Further diking <strong>of</strong><br />
wetl ands is strong1 y d 1 scouraged because<br />
<strong>of</strong> the adverse impacts discussed earller,<br />
With regard to the management <strong>of</strong> the<br />
currently dlked areas, It i s recommended<br />
that waterfowl breedlng be encouraged In<br />
a1 1 <strong>of</strong> the contained marshlands by means<br />
<strong>of</strong> nesting platforms and areas. Moreover,<br />
to facilitate flsh use, perhaps the<br />
wetlands could be opened up during the<br />
spring spawning season or at least during<br />
those years when no water-level<br />
manipulat 1 on is necessary for waterfowl<br />
management. Creating artlf icial islands<br />
and allowing better-drained areas to<br />
revert to shrub or maadow communities may<br />
also add dlverslty.<br />
DFTRO/T RIVER<br />
---*.<br />
.,. - - ~.<br />
KILOMt lHf S<br />
Figure, 63. Location map <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
diked coastal wetlands (see text for<br />
explanation <strong>of</strong> numbers).<br />
Other practices, such as experimental<br />
marsh burning* have improved the waterfowl<br />
loafing population in Ontario's diked<br />
wetlands (Ball 1984). Fertflfration<br />
however, is not recomnended at this time.<br />
Wherever possible, hydrologic connectivity<br />
should be increased.<br />
<strong>St</strong>. Clalr Delta wetlands are diked: <strong>St</strong>.<br />
Johnrs Marsh (no. l)r lower Harsens Island<br />
(no. 2)s lower Walpole Island near Goose<br />
<strong>Lake</strong> (no. 31, lower <strong>St</strong>. Anne Island (no.<br />
4 ) ~ and Mitchell Bay-Thames Rtver mouth<br />
regfon (no. 5). Nearly one-ha1 f <strong>of</strong> the<br />
total delta wetlands, some 5,280 hectares,<br />
Dlcklnson xsland~ because <strong>of</strong> Its 'pen<br />
system nature and nt act ronment<br />
gradients. must be managed se~aratel~ as<br />
it may be the<br />
wetland in the Great <strong>Lake</strong>s region,<br />
Keepfng P t an open system, with only<br />
passive recreation permitted, w i l l Insure<br />
are separated from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> by<br />
earthen dfkes. Most <strong>of</strong> these djked<br />
environments occur .fn zones where cattai 1s<br />
natural 1 grow and el evat range<br />
between 175 and 176 m above Internatfonal<br />
<strong>Lake</strong>s datum*<br />
Its multiple functions for fish, Except for the wetlands in <strong>St</strong>, John's<br />
waterfowl, other wildlffe* and food chain Marsh and those along western <strong>Lake</strong> <strong>St</strong>.<br />
support, Once fil ledr the confined <strong>Clair</strong>, most <strong>of</strong> these dfked marshlands are<br />
disposal sitesrnay be used for wildlife, managed for waterfowl purposes. Dikjng<br />
but not for a theme park as suggested by and water-level manlpulatlon <strong>of</strong> wetlands<br />
some local residents, <strong>The</strong> provfslon <strong>of</strong> for the benefit <strong>of</strong> waterfowl hunters is a<br />
public utilfties, on either Dickinson or traditional practice in the Great <strong>Lake</strong>s<br />
Harsens fslands* would Jnitiate reglon among both private and public<br />
uncontrol 1 ed development pressures. shooting clubs. <strong>The</strong> earthen dfkes Impound<br />
As indfeated<br />
water insfde the marshlands during towwater<br />
perfods in the Great takes, but also<br />
in Figure 53# several large areas <strong>of</strong> the f~ncti~n to protect the marsh from erosion<br />
12.2<br />
A
and excessive high water during above-<br />
average lake level conditions, <strong>The</strong> <strong>St</strong>.<br />
<strong>Clair</strong> Flats <strong>St</strong>ate Game Area, Canada Club*<br />
and the <strong>St</strong>e, Anne Club marshes are among<br />
the venerable diked wetlands managed for<br />
waterfowl huntlng during the fall<br />
migration,<br />
<strong>St</strong>. Johnfs Marsh, whlch straddles<br />
Hjghway M-23 in Clay Township9 is included<br />
as a diked wetl and because the l i ne <strong>of</strong><br />
houses and access road along Bouvier Bay<br />
(<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. C1 air) hydrologically<br />
separate it from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Figure<br />
56). Approximately 217 hectares <strong>of</strong> dfked<br />
marshlands are sftuated west <strong>of</strong> hlghway M-<br />
299 and about 478 hectares lie east <strong>of</strong><br />
this roadway. Interior dikes and the<br />
roadbed with only one bridge (or culvert)<br />
further fragment this wetl and and preclude<br />
water exchange, Although there are five<br />
openfngs to Bouvier Bay, water exchange<br />
and fish access are limited.<br />
Much <strong>of</strong> lower Harsens Island is<br />
contafned within the <strong>St</strong>. <strong>Clair</strong> Flats <strong>St</strong>ate<br />
Game Area (1,085 hectares) and is managed<br />
by Michtgan DNR as a public waterfowl<br />
hunting facility. In the upper section,<br />
which consists <strong>of</strong> 515 hectares, crops such<br />
as corn are grown and flooded in fall to<br />
attract migratory waterfowl, <strong>The</strong> more<br />
lakeward portion <strong>of</strong> the game area is<br />
essential1 y a water-level control led<br />
cattail marsh which provides cover and<br />
some breeding habitat for the waterfowl.<br />
During the 1974-75 waterfowl hunt, 76% o f<br />
the harvest from the 60 blinds on the game<br />
area consisted <strong>of</strong> mallard along with<br />
lesser numbers <strong>of</strong> black duck, pintail,<br />
green-winged teal and American wigeon<br />
( Jaworski and Raphael 1978).<br />
In Ontario, diking i s commonly<br />
practiced (Table 38). As shown on Figure<br />
63, diked area no. 3 consists <strong>of</strong> three<br />
diked systems located on lower Walpole<br />
Island. <strong>The</strong> area <strong>of</strong> these dfked<br />
marsh1 ands totals 1,097 hectares.<br />
Cattails and submerged aquatfcs appear to<br />
dominate the vegetation. Some <strong>of</strong> the<br />
wetl ands <strong>of</strong> the northernmost d ?ked area<br />
are now being utilized for cultivated<br />
crops.<br />
Diked area no. 4 on Figure 63 refers<br />
to two dfked systems on lower <strong>St</strong>. Anne<br />
Island. <strong>The</strong> diked area located on the<br />
west side <strong>of</strong> Chenal Ecarte consists <strong>of</strong> 660<br />
hectares, whereas the dlked system on the<br />
east side is comprised <strong>of</strong> 415 hectares.<br />
Aerfal photographs indicate dense colonies<br />
<strong>of</strong> cattails in these managed wetlands,<br />
with few canals or other human<br />
disturbances. <strong>The</strong> most 1 akeward diked<br />
wetland on lower Walpole Island also has<br />
few canals across it. Pumping stations<br />
and flood gatas or welrs are employed to<br />
manage water levels here and at the other<br />
waterfowl shooting club marshes.<br />
Along eastern <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> many <strong>of</strong><br />
the coastal wetlands are diked. Diking is<br />
done by the local farmers who wish to<br />
cultivate as close to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> as<br />
possible. In addition, at Mitchell Point<br />
there is a managed wetland, and two more<br />
exist near the mouth <strong>of</strong> the Thames River.<br />
<strong>The</strong> areas <strong>of</strong> these three dfked systems<br />
are: 194 hectares, 393 hectares, and 500<br />
hectares. <strong>The</strong> latter two are designated<br />
as Area IV and Area 111, respective1 yr on<br />
the Tflbury (Ontario) Quadrangle.<br />
Moreoverr east <strong>of</strong> Area IV, there is a 156-<br />
hectare, square-shaped diked area. A<br />
total <strong>of</strong> 1,329 hectares were planimetered<br />
on current maps <strong>of</strong> this coast. Cattails<br />
appear to be the domfnant vegetation type,<br />
<strong>The</strong> adverse impacts <strong>of</strong> diking coastal<br />
wet1 ands are poor1 y understood. Never-<br />
theless, several adverse impacts include:<br />
1) loss <strong>of</strong> fishery function, 2) blockage<br />
<strong>of</strong> water exchange, 3) loss <strong>of</strong> food cha'ln<br />
support, 4) fragmentation and isolation <strong>of</strong><br />
diked wetlands, and 5) lower diversity and<br />
water quality <strong>of</strong> diked areas. Loss <strong>of</strong><br />
fish habitat and associated fishery<br />
functions is perhaps the principal adverse<br />
impact <strong>of</strong> coastal wetland diking. Diking<br />
has long been a source <strong>of</strong> conflict between<br />
waterfowl- and f ishery-resource managers.<br />
Clearly, most fish9 with the exceptton <strong>of</strong><br />
extremely tolerant species such as carp<br />
and bullheads, do not spawn in dfked<br />
wetl ands. Furthermore, l arval and juve-<br />
nile fish cannot easily disperse Into nar<br />
tolerate the f 1 uctuatlng water temper-<br />
atures and water qua1 Jty <strong>of</strong> these<br />
contained systems. <strong>The</strong> authors have<br />
noticed that predator fish tend to forage<br />
only fn those coastal wetlands where lake<br />
water masses f nvade the open wetlands,<br />
For example, on Dickfnson Island one may<br />
observe 'l argemouth bass and bl uegi 71 s on? y
Figure 6$1 i'ier ial jir~otoornijii <strong>of</strong> !)i~kin*<strong>of</strong>~ Xs'lana wctlancis, <strong>St</strong>, <strong>Clair</strong> DPI ta {May 19R0je<br />
Molr, df~+jn,%rl~ fr-rrn top IC,;I'.;TA! f1t2r~hf"~ :n ~?F~E>'I' Ray (rrsff ~irie!, Vpv.fh ch;?nne'l ic +t<br />
t8?c?i!rwl Fq iti tilf2 tv~tt~)rli ~f LOP pirotagraph,<br />
the ?ti$, Il'lfj~ll?~<br />
324
in cl rcul ating marsh waters, particularly<br />
near culverts* or where lake water masses<br />
prevai 1,<br />
Di k i ng prevents water exchange<br />
between the wetland and the adjacent water<br />
body. Thusp the waters <strong>of</strong> the canals and<br />
open-water areas <strong>of</strong> the contafned sites<br />
stagnate, cover with f f lm, and exhibit<br />
l arge diurnal temperature ranges (Mudroch<br />
and Capobianco 1978). Although the<br />
dissolved oxygen levels In the dlked<br />
wet1 ands remain above 75% satu rat1 on<br />
(Figure 22)r in response to atmospheric<br />
dfffusion and photosynthetic activityr<br />
excessive daytime water temperatures in<br />
summer reduce the dissolved oxygen<br />
concentrations to critical levels. During<br />
the cold winter months with abundant snow<br />
cover, oxygen depletion may occur under<br />
the ice cover, resulting in fish kf 11s and<br />
die-<strong>of</strong>f <strong>of</strong> invertebrates.<br />
zooplanktonr and ye1 low perch caught over<br />
macrophyte-free areas contained hfgher<br />
numbers <strong>of</strong> these invertebrates than did<br />
fish caught over macrophyte areas. <strong>The</strong><br />
paradox in these data may be explafned by<br />
marsh drafnage which 4s associated with<br />
large numbers <strong>of</strong> f I? ter-feeding zooplankton<br />
and yellow perch regardless <strong>of</strong><br />
the occurrence <strong>of</strong> vegetated substrates.<br />
Only two areas<br />
<strong>of</strong> coastal wetland are jdentifjed for<br />
restoration. First, the function and<br />
value <strong>of</strong> <strong>St</strong>. John's Marsh could be<br />
increased with 1 mproved hydro1 og ic<br />
connectivity (Figure 56). It fs<br />
recommended that the culverts connecting<br />
this wetland to Bouvier Bay be en1 arged to<br />
f aci 11 tate greater water mass exchange,<br />
Also, water exchange across M-29 could be<br />
Improved with the addltion <strong>of</strong> another<br />
larqe culvert close to Pearl Beach.<br />
~dditionally, the water quality near the<br />
definitive research has Perch Isle Point subdivision should be<br />
been conducted which the monitored9 and some <strong>of</strong> the fnterfor dikes<br />
exceptional fish production in <strong>Lake</strong> <strong>St</strong>. west <strong>of</strong> N-29 should be breached or<br />
<strong>Clair</strong> (Hatcher 1982) r it is be1 ieved that<br />
the wetlands are important in the food<br />
webs. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta wetlands are<br />
important sources <strong>of</strong> nutrients,<br />
particul ate organic matter, phytop? ankton,<br />
small zooplankton, and drift organisms,<br />
Zoopl ankters probably f $1 ter the preferred<br />
food items out <strong>of</strong> the marsh drainage.<br />
Figure 64 illustrates drainage from the<br />
delta wetlands. Leach (1980) found that<br />
the waters from the main channels <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> River are lower in nutrients<br />
than the waters derived from the<br />
tributaries on the Canadian side <strong>of</strong> the<br />
delta. Because Chenal Ecarte and the<br />
other Ontario tributaries Incorporate<br />
considerable flow from the adjacent<br />
wetlands, it is felt that these waters<br />
contain important quantities <strong>of</strong> nutrients,<br />
detrftus, and living matter for the food<br />
webs <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />
Poe ( 1983 attempted to demonstrate<br />
the relationship <strong>of</strong> the aquatic<br />
macrophytes and associated perlphyton to<br />
the fish production <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, In<br />
spite <strong>of</strong> some positive relatfonships<br />
establ ished i n some sampl l ng stationsp<br />
Poets data for Muscanoat Bay seemed<br />
contradictory. Sampling <strong>of</strong> Muscamoot Bay<br />
revealed that macrophyte-free areas had<br />
higher numbers <strong>of</strong> macrophytoplankton and<br />
removed.<br />
Second1 y the fragmented wetland<br />
complex south <strong>of</strong> the Cllnton Rfver Estuary<br />
should be restored (Figure 543, Perhaps<br />
this area should be comprehensively<br />
planned since a private developer has<br />
already attempted t o construct<br />
subdivisions in that area. <strong>The</strong> Detro<strong>St</strong><br />
District Corps <strong>of</strong> Engineers refers to this<br />
threatened wetland as Verchev9s Marsh. A<br />
comprehensivs plan which fncludes the<br />
funct-ion <strong>of</strong> the Black Creek* the impact <strong>of</strong><br />
the Metropol i tan Beach development# and<br />
connectivtty to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is<br />
suggested. Restoratfon <strong>of</strong> a wetland on<br />
this western side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clajr could<br />
partially mitigate the historic wetland<br />
losses in this coastal stretch, Moreover*<br />
certaln fdsh such as northern pike and<br />
1 argemouth bass require spawning habitat<br />
ddstributed every 8 to 10 km along a coast<br />
because they are not Isng-dqstance<br />
migrants (Jaworski and Raphael 1978).<br />
No al-ti-fi-<br />
cjaf construction <strong>of</strong> wetlands is<br />
recommended at this time because the<br />
current wet1 ands appear adequate and <strong>Lake</strong><br />
<strong>St</strong>, <strong>Clair</strong> Js very productfve, If the<br />
current wetland resource base can be
conserved and/or Improved, then the important ecological cornmunlt~es, <strong>The</strong>se<br />
present ecology <strong>of</strong>: the lake may well be coastal wotlandst par-tfcularly the <strong>St</strong>.<br />
assured. ClaJr Delta wetlands, have had and should<br />
continue to have signSPIcant functions 3n<br />
perform<br />
va'uable functions for<br />
munfcf~alft'es* <strong>The</strong>y ~rovlde habjtats for<br />
desfred plants and animals, they filter<br />
pollutants and sedfments from storm<br />
the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ecosystem. Conclusions<br />
fn thls sectlon are based an physical.<br />
blologfcal econamfc, and poi lt-jcal<br />
factors.<br />
drainage, and they protect coastal <strong>The</strong> rocent history <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>,<br />
structures from wave attack, On the other ~1 a and fs cha ratter zed by<br />
handr wetlands are <strong>of</strong>ten barriers to lake rel stabf 1 as wel 1 as h<br />
access and are susce~tibfe to function and value. Gjven the general<br />
filng* and which<br />
in turn lmpaf rs thef r natural functfons.<br />
degradatfon whlch has taken place the<br />
lower Great <strong>Lake</strong>s durfng the past several<br />
<strong>The</strong> fmportant functfons <strong>of</strong> wetlands the future prospscts for the<br />
are <strong>of</strong>ten dfsregarded by those wishing to<br />
are<br />
develop an area, <strong>St</strong>ate <strong>of</strong> Michfgan reasonably brfght* As coastal wetlands fn<br />
has recogn'lzed this and has forrnul ated<br />
adjacent lakes the re' st've<br />
pol cy to wetlands.<br />
~h~ 'Importance <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. Clalr wetlands<br />
Shore1 ands Protection and Management Act is increasfng* there has a<br />
<strong>of</strong> 1970. provfdss programs whlch include 4X' loss In the <strong>Lake</strong> <strong>St</strong>* wet'ands8<br />
the protection <strong>of</strong> wetlands in the Great<br />
and Of the extant wet'ands are dfkedr<br />
<strong>Lake</strong>s shorelands. At the local level* a<br />
special problsm facfng declslon-makers is<br />
the delta wetlands have not suffered<br />
degradation comparable to that <strong>of</strong> Green<br />
Saginaw Bay, and westerr1 <strong>Lake</strong> Erie.<br />
the tdentf f fcatjon and regulation <strong>of</strong><br />
adequate buffer areas surrounding the,<br />
desf gnated wetlands,<br />
Tha physfcal framework <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />
Cl alr wstl ands partla1 ly explafns the high<br />
In Mlchlgan, once "areas <strong>of</strong> concernw productf vl ty <strong>of</strong> these wet1 ands. Firstr<br />
have bean establ lshed~ the level <strong>of</strong> the total area <strong>of</strong> coastal wetlands Is<br />
control must be determined. Speclal use large In comparfson to the relatively<br />
psrmlts and 1 lsts <strong>of</strong> acceptable<br />
csnstruetton near wetlands provfde a<br />
small sfze <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr. Second,<br />
these deltafc wetlands are dlstrfbuted<br />
method <strong>of</strong> mlnlmizfng wetland dlsturbance over a large area <strong>of</strong> the northern part <strong>of</strong><br />
by permitting only those %ctlvltfes which the 1 ake, Thf rd, physical dlvorsity<br />
contrfbute least to change. Wet1 ands are fact 1 ltatss mu1 tfple subsystems. This<br />
unsuitable for most klndg <strong>of</strong> development. creates numerous aquatic habttat and<br />
D'isruptfve dredging and fdlf fng are <strong>of</strong>ten coastal wet1 and interfaces which are<br />
necessary to create a usable area for the located in remote areas and thus buffered<br />
constrirctfon af bulld.fngsr parkfng areas, fr~n cultural 4mpacts. In addition, the<br />
and access foads. On-sfte sanitary sewage <strong>St</strong>. Clalr Delta Is geologlcal1y stable.<br />
treatment is dffficu'ft because <strong>of</strong> poor Subsldenca Is not occurring and delta<br />
percoIat4on and the high water table.<br />
Recregtfori usas can be developed around<br />
extensi on or erosion have not been<br />
slgnificant over the last century.<br />
wetlands in a manner compatfble wfth<br />
natural systems, Camping# canoeing, and Qn the ecosystem level, as well as an<br />
nature tratls make use <strong>of</strong> the water a local habltat scalet the hydrology and<br />
recrezrtfonel value <strong>of</strong> wet1 ands. with such hydrolog ic connectlvl ty <strong>of</strong> the <strong>St</strong>, Clafr<br />
precautionsr construc9lan need not. cause Delta wetlands are most favorable.<br />
excessive damage, Re? atlvel y cool, hfghly-oxygenated, and<br />
5.3. PROSPECTS FOR THE FUTURE<br />
non-turbld waters enter from <strong>Lake</strong> Huron<br />
and pass through the deltaic wetlands,<br />
<strong>The</strong> df stributaries and crevasse channels<br />
(1 ocally celled "hfghwaysn) dl stribute<br />
this high-qua1 f ty tnput water throughout<br />
<strong>The</strong> freshwater wetlands along <strong>Lake</strong> the wetlands, For example, an abandoned<br />
<strong>St</strong>, CT a1 r w $1 1 canttnue to corratltuts dlstrftsutery channel permits water flow<br />
7%
across the entire length <strong>of</strong> Djckinson<br />
Island. Given a residence time <strong>of</strong> on1 y 7<br />
to 9 days in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r circulatlon<br />
is well established and little stagnation<br />
occurs. Water exchange between the lake<br />
and coastal wetlands is a1 so f aci 1 itated<br />
by seiches.<br />
<strong>The</strong> we? 1-known turbidity problems <strong>of</strong><br />
other coastal wetland systems? such as<br />
Green Bay and Saginaw Bay, are not a major<br />
adverse impact in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
wetlands. SiltatJon and turbidity are low<br />
in the deltaic wetl ands because the <strong>St</strong>.<br />
Cl ai r River has a very low suspended load<br />
and the bedload consists <strong>of</strong> fine sand.<br />
Some local turbidity occurs at the mouth<br />
<strong>of</strong> the Cl inton and Thames Rivers durlng<br />
spring run<strong>of</strong>f and flooding. Also, within<br />
<strong>Lake</strong> <strong>St</strong>. Clai r, resuspension <strong>of</strong> bottom<br />
sediments, as In Anchor Bay during storms,<br />
can discolor the water over 1 arge areas.<br />
Wetland erosion is not a problem<br />
along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> due to the relatively<br />
short fetch and shallowness <strong>of</strong> the lake<br />
basin. Coastal erosion resulting from<br />
eustatically rising water levels and f ram<br />
long-term lake level fluctuations is not a<br />
significant problem as it is in <strong>Lake</strong> Erie,<br />
Because the deltaic wetlands are largely<br />
open systems that slope lakeward, lake<br />
level oscillations do not result in<br />
wetl and die-back or senescence ' as is true<br />
in many other coastal wet1 and systems.<br />
Rather, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands shift up<br />
or down the gradient, maintaining their<br />
productivity at various 1 ake levels.<br />
Dickinson Is1 and is an excellent example.<br />
Biological 1 y, these coastal wet1 ands<br />
exhibit relatively high habitat and<br />
species diversity, Productivity in <strong>St</strong>.<br />
<strong>Clair</strong> Delta wetlands is also high. Such<br />
conditfons manifest themselves in surplus<br />
waterfowl foods and Impressive flsh<br />
catches. Open wetlands systems, with<br />
complete envi ronmental gradients and<br />
numerous ecotones, exhibit "pulse"<br />
stability and high productivity. Although<br />
further research is needed to define the<br />
importance <strong>of</strong> the flux <strong>of</strong> particulate<br />
organic matterr drift organisms, and other<br />
organic foods from the wetlands into <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>r there i s an apparent<br />
association <strong>of</strong> waterfowl r game fish, and<br />
other wildlife with these wetland<br />
discharges. <strong>The</strong> high dissolved oxygen<br />
levels and high flushing rates,<br />
accompanied by relatively low contaminant<br />
and turbid1 ty levels* are positively<br />
correlated with the high productlvfty.<br />
From an economic vjewpoi nt these<br />
coastal wetlands exhibit high fish and<br />
wildlife values. Because these functions<br />
and values are linked to the ecology <strong>of</strong><br />
<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, they cannot easil y be<br />
replaced or mitigated. Wfth regard to<br />
human uses, passive recreation and open<br />
space designation appear to be more<br />
suitable than intensf ve activl ties. Uses<br />
involving hydro1 ogic modification <strong>of</strong> the<br />
wetlands and permanent intrusions <strong>of</strong>ten<br />
create large and irreparable<br />
disturbances. It i s expected that urban<br />
development w i l l result in the filling or<br />
destruction <strong>of</strong> most coastal wetlands along<br />
western <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, northward to <strong>St</strong>.<br />
John's Marsh. Moreover, south <strong>of</strong> Mitchell<br />
Bay, agricultural encroachment may is01 ate<br />
all but the cattail marsh fringe. It is<br />
unlikely that the Clinton River mouth and<br />
the <strong>St</strong>. John's Marsh wetlands w i l l be<br />
restored. Neverthelessr because <strong>of</strong> the<br />
recognized value <strong>of</strong> the remaining<br />
wetlands, we believe that most <strong>of</strong> the<br />
extant wetlands w i l l be preserved in their<br />
present state.<br />
<strong>The</strong> future impacts on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
wetlands are dependent upon leg is1 ative<br />
efforts and public awareness in the Great<br />
<strong>Lake</strong>s basin <strong>of</strong> the value <strong>of</strong> the resource<br />
base. Legislatively, the <strong>St</strong>ate <strong>of</strong><br />
Michigan has been a leader i n<br />
environmental protection <strong>of</strong> wet1 ands. For<br />
example, the followfng laws have been<br />
enacted :<br />
--<br />
Public Act 247 <strong>of</strong> 1955, as amended*<br />
prohibits constructing or dredgi ng any<br />
artificial body <strong>of</strong> water that would<br />
ultimately connect with a Great <strong>Lake</strong>.<br />
This act also requires a permit from<br />
the Department <strong>of</strong> Natural Resources to<br />
f i 11 any submerged 1 ands in a Great<br />
<strong>Lake</strong>, including <strong>Lake</strong> <strong>St</strong>, Clalr.<br />
ment Act -- PublJc Act 245 <strong>of</strong> 1970t as<br />
amended* designates wetlands adjacent<br />
to a Great <strong>Lake</strong> or a connecting<br />
waterway such as the <strong>St</strong>. <strong>Clair</strong> River as<br />
envf ronnental areas necessary to<br />
preserve flsh and wildl $fee
connectivity Is good and the wetland<br />
diversity Is well displayed, Converse1 yr<br />
regulates wet1 ands through several many wetlandsp especially In Ontarfor have<br />
important laws relating to shorelands<br />
and submerged 1 ands.<br />
been diked and are less diverse s'lnce<br />
there Ss a dominance <strong>of</strong> cattail. It %s<br />
<strong>of</strong>ten pointed out that <strong>Lake</strong> <strong>St</strong>.,Cla%r is<br />
In Ontarfo, wetlands are viewed as located tn waterfowl flyways and is aw<br />
recreational land and are thus taxed as important staging area for migration, a<br />
such. Taxation <strong>of</strong> agricultural land is fact which gustlffdes dfking to some<br />
less than recreational 1 and. This degree, Howevers as noted earlier, the<br />
unbalanced tax structure ls an incent-fve economlc value <strong>of</strong> sport fishing greatly<br />
to drain and dlke wetlands In Ontario for<br />
agrlcul tural purposes (G. B. McCulloughr<br />
exceeds waterfowl hunting and trapping<br />
combined, More specifically, the economic<br />
Canadian Wildlife Service; pers. comm.1, value <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> to sport flsh'ing*<br />
Several wetlands north <strong>of</strong> the Thames River including coldwater species, has a value<br />
including huntlng clubs lands have <strong>of</strong> $59641 per wetland hectare/year<br />
succumbed to the tax pressure and have (Jaworski and Raphael 197131. To detach<br />
been converted to agriculture. the sportfish habltat from the wetland<br />
ecosystem may well decrease wetland<br />
It is most fortunate that large dlversity and the economic value <strong>of</strong> the<br />
tracts <strong>of</strong> wetlands are owned by <strong>St</strong>ate or <strong>St</strong>. Clalr Delta. Also the cost for dike<br />
Federal agencies. Assuming that the construction and maintenance in view <strong>of</strong><br />
governling agencies wf 11 preserve the decreasing diversity ought to be<br />
resource9 most <strong>of</strong> the Michigan portion <strong>of</strong><br />
the <strong>St</strong>, <strong>Clair</strong> Delta w j l l be protected from<br />
considered.<br />
future development. In Ontario the Coastai erosion is modest on the<br />
Wal pole Island Indian Reserve encompasses shore1 lnes <strong>of</strong> the laker and selches and<br />
Seaway, Squirrel, Wal pole and <strong>St</strong>. Anne flooding are beneficial flushing agents<br />
Islands. <strong>The</strong> preservation prospects for and nutrient transport mechanisms to the<br />
this portion <strong>of</strong> the modern delta are open-water bodies <strong>of</strong> the bays and lake.<br />
reasonably bright. D i k 1 ng preserves a rather monoiyp fc<br />
wetland9 prohibits adequate hydrologfc<br />
<strong>The</strong> MichZgan DNR has made a concerted connectlvityr and decreases wetland<br />
and sustained effort to inform the publlc utlllty In a low wave energy environment.<br />
<strong>of</strong> the value, in terms <strong>of</strong> economic Dikfng--to preserve wetlands in higher<br />
returns, <strong>of</strong> the <strong>St</strong>ate's wetlands. Through wave energy environments or in coastal<br />
funded wetland research and literatures areas dominated by fine clastic sediments<br />
the assets<strong>of</strong> the wetland resources have or where geological subsidence fs<br />
been dissemfnated to the public, occurring such as western <strong>Lake</strong> Erie-appears<br />
to have some justiflcation and<br />
In spite <strong>of</strong> legislative an3 public trade-<strong>of</strong>f value. In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> the<br />
awareness efforts# there are still areas envf ronmental (i .eeP hydrological and<br />
sf concern. Both in Michigan and Ontario, geological 1 considerations ought to<br />
bul kheading and f i 11 i ng for small encourage wetlands preservatlon to be<br />
residential development continuer particu- dfrected t o optlmurn hydrologic<br />
larly along the natural levees <strong>of</strong> South connectivity between the resource and the<br />
and M-lddle channels and the shelf areas, open lake,<br />
In spite <strong>of</strong> Michfgan PA 247 and PA 24Sr<br />
dfked disposal sftes for polluted dredge In November 1984, a "Great <strong>Lake</strong>s<br />
spoil were const rlscted on the premodern Coastal Wet1 ands Col loqui um, " co-sponsored<br />
def tr? <strong>of</strong> Dickinson Island in the mid- by the Sea Grant Program and Envfronment<br />
f97Ds, Although Michigan@s efforts to Canada* was held on the campus <strong>of</strong> Michigan<br />
preserve these coas;tal wetlands have been <strong>St</strong>ate University to explore natural and<br />
posltfve, alteratton is continuing. manipulated water levels in Great <strong>Lake</strong>s<br />
wetlands, Resides pr~vidjng a forum for<br />
Dick;nson Island pr~vides an example the exchange <strong>of</strong> current research<br />
<strong>of</strong> a coastal wetland which has not been linformationr the rneetjng resulted In the<br />
signfficantly altered, <strong>The</strong> hydrological establ lshment <strong>of</strong> a network <strong>of</strong> wet1 and<br />
1 28
ecolog~sts and managers In the Great <strong>Lake</strong>s<br />
region and plans for the pub1 fcation <strong>of</strong> a<br />
quarterly newsletter for such workers,<br />
Although the conference p<strong>of</strong>nted out the<br />
need for more research djrected toward<br />
understanding processes and wild1 ife<br />
utllizatjon <strong>of</strong> the coastal marshes* an<br />
equally important consSderatlon was the<br />
alarming loss <strong>of</strong> wetlands. If there was a<br />
central theme <strong>of</strong> the conference* it would<br />
have to be that the conferees were in<br />
agreement that the existing coastal<br />
wetlands must be vigorously preserved if<br />
these valuable habitats are to survjve<br />
Intense development pressures.<br />
Gary B. McCullough <strong>of</strong> the Canadian<br />
Wild1 i fe Service documented the extensive<br />
conversion <strong>of</strong> coastal marshes to<br />
agricultural 1 ands along the Ontario shore<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> which has taken place in<br />
the past several decades. He estimated<br />
that 36% <strong>of</strong> the original wetlands have<br />
been lost. Much <strong>of</strong> the remaining wetland<br />
area south <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta is<br />
protected by dikes and water level control<br />
structures. John Ba7 1 <strong>of</strong> the University<br />
<strong>of</strong> Guelph reported that in these marshes,<br />
such as the ones owned by Ducks Unlimited<br />
or within the <strong>St</strong>. Clalr National Wildlife<br />
Area (Canada)# waterfowl utilization can<br />
be enhanced by clearing open areas in the<br />
cattail marshes (Ball 1984). To attract<br />
migrating ducks the optimal size <strong>of</strong> the<br />
opening appears to be approximately 50 m.<br />
<strong>The</strong> col 1 oqu ium concluded with the remarks<br />
<strong>of</strong> Harold H. Prince, underscoring the<br />
"signaturew which coastal wetlands impart<br />
to the fish and wildlife resources <strong>of</strong> the<br />
region. He pointed out that the<br />
significance <strong>of</strong> these marshes can be<br />
appreciated by recognizing: 1) that so<br />
many animals have embraced wet1 and<br />
habitats as part <strong>of</strong> their 1 ife strategy<br />
and 2) that wetlands are indicators <strong>of</strong><br />
llttoral and upland environmental quality.<br />
<strong>The</strong> ideas put forth at this conference are<br />
1 ikely to shape the direction <strong>of</strong> wetlands<br />
research in the Great <strong>Lake</strong>s region for the<br />
next decade.
Allen, H.L. 1971. Primary productivity,<br />
chemo-organotrophy, and nutritional<br />
interactions <strong>of</strong> epiphytic algae and<br />
bacteria on macrophytes in the<br />
1 ittoral <strong>of</strong> a 1 ake. Ecol . Monogr.<br />
41(2): 97-127.<br />
Andrews, R. 1952. A study <strong>of</strong> waterfowl<br />
nesting on a <strong>Lake</strong> Erie marsh. M.S.<br />
<strong>The</strong>sis. Ohio <strong>St</strong>ate Univ., Columbus.<br />
85 PP.<br />
Assel, R.A., F.H. Quinn, G.A. Leshkevich,<br />
and S. J. Bolsenga. 1983. Great <strong>Lake</strong>s<br />
ice atlas. U.S. Dept. Commerce, Great<br />
<strong>Lake</strong>s Envir. Res. Lab. Ann Arbor, MI.<br />
115 pp,<br />
Ayres, J.C. 1964. Currents and related<br />
problems at Metropol itan Beach, <strong>Lake</strong><br />
<strong>St</strong>. <strong>Clair</strong>. Great <strong>Lake</strong>s Res. Div.<br />
Spec. Rep. No. 20, Univ. Michigan,<br />
Ann Arbor. ,55 pp.<br />
Bajorunasr S.J. 1968. River ice jams; a<br />
literature revlew. U.S. Army Corps <strong>of</strong><br />
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Baumgartner, L . L. 1942, Ecological survey<br />
<strong>of</strong> muskrat populations and habitats.<br />
Mich. Dep. Nat. Resour., Proj. Rep.<br />
14-R. 128 pp.<br />
Bayly, I. 1975. Preliminary study <strong>of</strong> the<br />
nutrient regime <strong>of</strong> marshland <strong>of</strong> the<br />
<strong>St</strong>. <strong>Clair</strong> National Wildlife Area -<br />
Dover Township, Ontario. Rep. to<br />
Can. Wildl. Serv., London, Ontario,<br />
27 PP*<br />
Bednarik, K.E. 1956. <strong>The</strong> muskrat in Ohio<br />
<strong>Lake</strong> Erie marshes. Ohio Dep. Nat.<br />
Resour., Ohio Div. Wildl. 67 pp.<br />
Bednar i kr K. E. 1975. Environmental impact<br />
assessment report, Magee Marsh<br />
Wildlife Area dike construction. Ohio<br />
Dep. Nat. Resour,, Crane Creek Wildl.<br />
Exp. <strong>St</strong>a., Oak Harbor. 13 pp.<br />
Behler, J.L. and F.W. King. 1979. <strong>The</strong><br />
Audubon Society field gufde to North<br />
American reptiles and amphibians.<br />
Alfred A. Knopf, New York. 743 pp.<br />
Bellrose, F.C. 1968. Waterfowl migration<br />
Baker, DeG.r and J.G. <strong>St</strong>rub, Jr. 1965. corridors east <strong>of</strong> the Rocky Mountains<br />
Climate <strong>of</strong> Minnesota. Part 111:<br />
In the United <strong>St</strong>ates. Illinois Nat.<br />
temperature and its appl ication. History Survey Biol, Notes. No. 61.<br />
Unfv. Minnesota, Agr. Expr. <strong>St</strong>a.<br />
Tech. Bull. 254. 43 pp.<br />
24 PP-<br />
Ball, J.P. 1984. Habitat selection and Bellrose, F.C* 1976- Ducks* geese* and<br />
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Geol. Instit.# Falls Church,<br />
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VA. 749 taxonomy. Unfv. Michigan, Mus. Zool.<br />
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<strong>of</strong> Cladocera, mostly from take <strong>St</strong>.<br />
Gfafrr Mtchfgan, Pagss 43-47 fn J. E.<br />
Food and feeding In fresh-water<br />
mussels. Bull, U.S. Bur. FSsh.<br />
Welghard, ed., A bfologScal 39:439-471.<br />
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Mfch. flsh Corn. No, 4. Clarke, A.M. 1901. Tho freshwater molluscs<br />
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Pagss 710-718 & Proc. 15th Conf.<br />
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wetlands and deepwater habitats <strong>of</strong><br />
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103 pp.<br />
Crowderr L.B., and W.E. Cooper, 1979.<br />
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<strong>The</strong>sis* Univ. Michiganr Ann Arbor.<br />
124 pp.<br />
Dennis, D.G., and R.E. Chandler. 1974.<br />
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and R.K. Ross. 1981. An update<br />
assessment <strong>of</strong> migrant waterfowl use<br />
<strong>of</strong> the Ontario shorelines <strong>of</strong> the<br />
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Serv. Rep. Ser. (manuscript rep. 1<br />
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Waterfowl use <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
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Goodyear, C.D., T.A. Edsallr D.M. Dempsey,<br />
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Atlas <strong>of</strong> the spawning and nursery<br />
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Erie, Ohio. Ohio J. Sci. 71(6):321-<br />
342.<br />
U*S* Fish<br />
Great <strong>Lake</strong>s Fish. Lab.,<br />
Mich., Admin. Rep. 82-7.<br />
Ann Arbor*<br />
49 pp.<br />
stuckey, R. L, 1972. Taxononly and<br />
distribution <strong>of</strong> the genus<br />
(Cruciferae) in North America. Sida<br />
Contrih. Bot. 279-430.<br />
Schloesser, D.W., and B.A. Manny. 1984.<br />
Distribution <strong>of</strong> Eurasian water-<br />
mil foil (Mvrioohvllum ~oic-1 in<br />
the <strong>St</strong>. <strong>Clair</strong>-Detroit River system in<br />
1978. J. Great <strong>Lake</strong>s Res. 10(3):322-<br />
326.<br />
Scott, S.L., ed. 1983. Field guide to the<br />
birds <strong>of</strong> North America. National Geo-<br />
graphic Soc. Washington, D.C. 464 pp.<br />
Shelemon, P.J. 1971. <strong>The</strong> Quaternary<br />
deltalc and channel system in the<br />
central great valley, Cal i forn ia.<br />
Ann, Pssoc. Am. Geogr. 61(3):427-440.<br />
Sieghardt, H. 1973. Utilization <strong>of</strong> solar<br />
energy and energy content <strong>of</strong><br />
di f ferent organs <strong>of</strong> Phra~mi tes<br />
somrnu Trin. Pol, P,rch. I-cydrobiol .<br />
20:151-156.<br />
Smith, F. 1894, List <strong>of</strong> protozoa and<br />
mollusca observed in <strong>Lake</strong> <strong>St</strong>. Clai r<br />
in the summer <strong>of</strong> 1893. Pages 42-44<br />
3. E. Heighard, ed., A biological<br />
examination <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Bull.<br />
Mich. Fish Comm. No. 4.<br />
<strong>St</strong>ansbery, I?. H. 1960. <strong>The</strong> Unionf nae<br />
(Molluscar Pelecypoda, Naiadacea) <strong>of</strong><br />
Fishery Bay, South Bass Island, <strong>Lake</strong><br />
Erie. Ph.D. Diss., Ohio <strong>St</strong>ate Univ.,<br />
Columbus. 216 pp.<br />
<strong>St</strong>ockner, J .G., and F.A. Armstrong. 1971.<br />
Periphyton <strong>of</strong> the experimental 1 akes<br />
area, northwestern Ontario. J. Fish.<br />
Res. Board Can. 28(2):215-229.<br />
<strong>St</strong>uckey, R.L. 1975. Aquatic vascular<br />
plants <strong>of</strong> the Sandusky River Basin.<br />
Pages 295-333 in D.B Baker, hl.P.<br />
Jackson, and P.L. Prater, eds. Proc,<br />
Sandusky River Basin Symposium.<br />
PLUARG/ I nt. Joint Comm., Great <strong>Lake</strong>s<br />
Regional Office, Windsorr Ontario.<br />
475 pp.<br />
<strong>St</strong>uckey, R.L. 1979. <strong>The</strong> declfne <strong>of</strong><br />
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Taftp C.E., and C. W. Taft. 1971. <strong>The</strong><br />
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Teal, J .M. 1962. Energy flow in the salt<br />
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43 : 614-624.<br />
Tilton, D.L., P.H. Kadlec, and P.R.<br />
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Tiner, R.W., Jr. 1984. <strong>Wetlands</strong> <strong>of</strong> the<br />
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U.S. Dep, <strong>of</strong> Interior 11. S. Govt.<br />
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PP<br />
U.S. Army Corps <strong>of</strong> Engineers. 1974,<br />
Proposed diked disposal area on<br />
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National Cceanic and Atmospheric<br />
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PP-<br />
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Walters, L.J., T.L.. Kovack, and C.E.<br />
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Role <strong>of</strong> habitat in the distribution<br />
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Home Econ. Exp. <strong>St</strong>a. Iowa <strong>St</strong>ate<br />
Univ,, Spec. Rep. No. 43. Ames. 31<br />
PP *<br />
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58 PP*<br />
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M.S. <strong>The</strong>sis, Univ. Western Ontario,<br />
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Wilson, J., and L.J. Walters. 1978.<br />
Sediment-water-biomass interaction <strong>of</strong><br />
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<strong>Lake</strong> Erie. Ohio <strong>St</strong>ate Univ., Cent.<br />
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113 pp.<br />
Winner, J.M,, A.J. Oud, and R.G. Ferguson.<br />
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Proc. 13th Conf. Great <strong>Lake</strong>s Res.,<br />
Int. Assoc. Great <strong>Lake</strong>s Res.<br />
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Microsc. Soc. 24:85-107.<br />
Wolfertr D.R,, W.N. Busch, and C.T. Baker.<br />
1975. Predation by fish on walleye<br />
eggs on a spawning reef in western<br />
<strong>Lake</strong> Erie. 1969-71. Ohio J. Sci.<br />
75(3) :118-125.<br />
Wolverton, C. 1981. Manual for wetland<br />
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draft. Div. Land Resour. Prog.# Mich.<br />
Dep. Nat. Resour., Lansing. 30 pp.<br />
Wright, L. D., and J .M. Coleman. 1972.<br />
River delta morphology: wave cl imate<br />
and the role <strong>of</strong> the subaqueous<br />
pr<strong>of</strong>ile. Science 176(4032):282-284.
DIMENSIONS OF LAKE ST. CLAIR AND ST. CLAIR RIVER WETLANDS BY UNITED STATES<br />
AND CANADA QUADRANGLE MAPSa<br />
Jurisdiction and<br />
quadrangle map<br />
MICHIGAN :<br />
Wayne County<br />
Be1 1 e Is1 e Quadrangle<br />
Detroit Quadrangle<br />
Subtotal<br />
Macomb County<br />
Grosse Point Quadrangle<br />
Mt. Cl emens Quad rangl e<br />
New Haven Quad rangl e<br />
Subtotal<br />
<strong>St</strong>. C1 air County<br />
A1 gonac Quad rangl e<br />
Marine City Quadrangle<br />
New Baltimore Quadrangle<br />
Port Huron Quadrang1 e<br />
<strong>St</strong>. Cl ai r Quadrangle<br />
<strong>St</strong>, <strong>Clair</strong> Flats Quadrangle<br />
Subtotal<br />
MICHIGAN TOTAL<br />
ONTARIO:<br />
Essex County<br />
Be1 le River Quadrangle<br />
Riverside Quadrangle<br />
<strong>St</strong>oney Point Quadrangle<br />
Ti1 bury Quadrangle<br />
Subtotal<br />
Coastal length Area <strong>of</strong><br />
af wet1 ands wet1 ands<br />
(km) (km2)
Jurisdiction and<br />
quad rangl e map<br />
Kent County<br />
Mi tche11 Bay Quadrangle<br />
Ti 1 bury Quadrangle<br />
Wall aceburg Quadrangle<br />
APPENDIX A (CONTINUED)<br />
Subtotal<br />
Lambton County<br />
Courtright Quadrangle<br />
Johnston Channel Quadrangle<br />
Mi tchel 1 Bay Quadrangle<br />
Sarn i a Quad rangl e<br />
Seaway Is1 and Quadrangle<br />
Sombra Quadrangle<br />
Wall aceburg Quadrangle<br />
Wal pol e Is1 and Quad rangl e<br />
Coastal 1 ength Area <strong>of</strong><br />
<strong>of</strong> wet1 ands wet1 ands<br />
( km? ( km2 I<br />
Subtotal 64.4 161.69<br />
ONTARIO TOTAL 102.4 231.33<br />
--<br />
LAKE ST, CLAIR/ST. CLAIR RIVER TOTAL 188.2 382.91<br />
%ata sources: United <strong>St</strong>ates Geological Survey, Dept. Interior, 7.5-minute<br />
Quadrangle Maps (1:24,000 scale?; Canada Department <strong>of</strong> Energy, Mines? and<br />
Resourcest 7.5-minute Quadrangle Maps (1:25,000 scale),
APPENDIX A (CONCLUDED)
MEAN MONTHLY ST. CLAIR RIVER FLOWSp 1900 TO 1980a<br />
Flaw In cubic meters per second<br />
WGTfl<br />
YEAR JAN FED MAR &PR M&Y JUN JUL AUO SEP OCT NOV DEC MERN<br />
1900 4190 3960 3960 5130 5240 5300 5440 5470 5530 5530 5640 5440 5077<br />
1901 4700 3570 4360 3600 3499 5720 5780 5780 5660 5610 5550 5300 5105<br />
1902 4130 4300 5180 5270 5320 5440 5410 5490 5380 5270 5300 5150 9141<br />
1903 3990 3880 4000 5240 5240 5320 5440 5440 5490 9610 5440 5150 3102<br />
1904 4330 4190 4360 5300 5520 3720 5780 5800 5780 5800 5490 5350 5304<br />
190% 3570 3940 4560 5550 566.0 5780 5930 5860 5360 5330 5720 5610 5321<br />
1306 5470 4250 4810 5460 5780 5780 5830 5800 5720 5610 5550 5150 5458<br />
1907 4420 4110 4900 5580 5640 5720 5830 5800 5330 5720 5810 5490 5395<br />
1908 3360 3740 4730 5440 5610 57YO 5330 5830 5630 '5550 5440 5380 3258<br />
1909 4810 3450 4360 25210 5380 5520 5L;SC) 5520 3490 f;3E:0 5210 5(:)40 5E1'3(1
APPENDIX B (CONCLUDED)<br />
Flow in cubfc meters per second<br />
WGTD<br />
YEW AN FEB I"ICIR APR MAY JUN JUL CSUG SEB OCT NOV DEC MEAN<br />
a Data source: U,S, Army Carps <strong>of</strong> Engineers.
ALGAE OCCURRING IN COASTAL WETLANDS AND NEARSHORE WATERS OF LAKE ST. CLAIRa<br />
Taxa Form and habitat<br />
Phylum Cyanophyta (blue-greens)<br />
Class Myxophyceae<br />
Order Chroococcal es<br />
Family Chroococcaceae<br />
Order Harmogonal es<br />
Famlly Osclllatoriacaas<br />
Family Nostocacaae<br />
SP *<br />
Fa<br />
Phylum Euglsnaphyta Cauglenoids)<br />
C7 ass Eugl enophyceas<br />
Order Eugl snal es<br />
Family Euglenaceae<br />
246<br />
Colonial; planktonic<br />
Colonial; planktonlc<br />
Colonial; planktonic<br />
Colonial; planktonic<br />
Colonial ; pl anktonfc<br />
Colonial ; p1 anktonic<br />
Colonial; gl anktanic<br />
Colonfal; planktonic<br />
Colonfal; pl anktonic<br />
Fi l amentous; pl ankton ic<br />
Fil amentous; sessile 8<br />
pl anktonfc mats<br />
Filamentous; planktonlc<br />
Colonial; among marsh p1 ants8<br />
sessile & planktonic<br />
Sol itary; pl anktonic<br />
Solftary; planktanic<br />
Sol i tary; pl ankton jc<br />
Sol ftary; p1 anktonic
APPEND SX c ( CrMlf NUED 1<br />
f axa Form and Rabftat<br />
Phylum Chrysophyta (golden-browns)<br />
Class Xanthophyceae (yellow-greens)<br />
Ordor Hoterococeales<br />
Family Chlorothaclacsae<br />
Qrdar kleterotrichal es<br />
Famlly Trlbonematacsaa<br />
SP *<br />
Order Hatoroslphonalas<br />
Farnfly Vauchorlacaae<br />
Class Chrysophyceas (goldan algae1<br />
Qrdcs Chrysomonadalas<br />
Famfly khromnadacsao<br />
Class Bacl llarlophycaa Idlatms3<br />
Order Centrales (centrfc dlatomsl<br />
Famlly Corcfnadfscacsas<br />
w<br />
Order Psnoal es ( psnnate d 1 atoms)<br />
Fanlly Frayllarlaceae<br />
Faml l y Achnanthacaae<br />
sP<br />
Family Navicul aceas<br />
Sol<strong>St</strong>ary; among marsh plants<br />
8 planktanfc<br />
Filamentous; planktonlc<br />
Filamentous; among marsh plants<br />
Filamentous; among marsh plants<br />
Sol ltary~ planktonlc<br />
So1 ftary; pl anktanic<br />
Sat ftary; planktontc<br />
$01 ftary; planktonfc<br />
Sot i tary; plankton f c<br />
$01 1 tary; planktanf c<br />
Sol 1 taryt planktonfc<br />
sol ltary; pl anktonds<br />
Sol I tary; p1 anktonfc<br />
Sol ftary; pf anktanfcr &<br />
attached to aquatic plants<br />
Sol ltary; ptanktonlc
f axa<br />
Famfly Gomphonsmaceas<br />
Family Cymbellaceas<br />
Phylum Pyrrhsphyta (fire algae)<br />
Class Dfnaphycsas (din<strong>of</strong>lagellates)<br />
Qrdar Psridiniales<br />
FnmZ f y Per 4 dl n 1 aceat3<br />
Phylum Chlarophyta (greens)<br />
Class Chlorophycsaa<br />
Ordar Volvoealas<br />
Famfly Ghlamydmanadacea~<br />
SP *<br />
BrQas Tetrasporal @s<br />
Fa<br />
Order Mf crssporal ss<br />
Fanfly Mlcrosporacsao<br />
spa<br />
Order Chaetopharaf ss<br />
5%<br />
Farm and habitat<br />
Sol itary; yl anktanfc<br />
So1 1 tasy; p7 anktonic<br />
Sol Ztary; planktonlc<br />
Sol l tary; planktonlc<br />
$01 1 tery; planktonic<br />
Calonlal; planktonfc<br />
Colonlal; planktonlc<br />
Colonial; planktonlc<br />
Colanfal; planktonic<br />
Colonlal; glanktanfc<br />
F$lamsntous; planktonic<br />
Filamentous; attached to<br />
aquatJc plants & debrfs
APPENDIX C (CONTINUED)<br />
Taxa Form and habitat<br />
Fami l y Protococcaceae<br />
Family Coleochaetaceae<br />
Order Cl adophoral es<br />
Fami 1 y Cl adophoraceae<br />
SP<br />
--<br />
SP 0<br />
--<br />
m b o r v a n u m<br />
Ped.i_arj.truml2ahsmanhtusum<br />
Order Oedogonial es<br />
Fami 1 y Oedogon i aceae<br />
Order Chl orococcal es<br />
Family Hydrodictyaceae<br />
ped i astrm -taturn<br />
-m-<br />
-mtetras<br />
Sorastrums<strong>of</strong>nulasuol<br />
Fami 1 y Coel astraceae<br />
Family Botryococcaceae<br />
,l2uulu<br />
Solitary; planktonic attached<br />
to organic debris<br />
Filamentous; attached to<br />
organic debris<br />
Fil amentous; attached to<br />
aquatic p1 ants<br />
Fi1 amentous; p1 anktonic<br />
Filamentous; attached to<br />
aquatic plants<br />
Colonial ; pl anktonic<br />
Colonial; planktonic 8<br />
among marsh plants<br />
Colonial; planktonic 8<br />
among marsh plants<br />
Colonial ; p1 anktonic<br />
Colonial ; pl anktonic<br />
Colonial; planktonic<br />
Colon i a1 ; pl ankton i c<br />
Col on i a1 ; pl ankton ic<br />
Colonial ; planktonic<br />
Colonial ; planktonic<br />
Colonial ; p1 anktonic 8<br />
among marsh plants<br />
Colon i a1 ; pl ankton i c<br />
Colon i a1 ; among marsh pl ants<br />
Colonial ; planktonic<br />
Colonial; planktonic
Taxa Form and habitat<br />
Famt 7 y Scenadesmaceae<br />
Order Zygnsmatales<br />
Family Zygnematacsae<br />
Family Ossmfdiateas Cdasmlds)<br />
So1 ltary; pl anktonfc<br />
Sol l tary; p1 anklon lc<br />
Colon4al; pl anktonfc<br />
Colonial; amang marsh plants<br />
Salltary; planktonlc<br />
Colonf a1 ; pl anktonlc<br />
Sol ltary; among marsh pl ants<br />
Colonial; planktonjc<br />
Colonlal; planktonlc<br />
Colonlal; planktonlc<br />
Colonial ; among marsh plants<br />
Colonial; amng marsh plants<br />
Colonf a1 ; pl anktsnic<br />
Colonfal; planktonlc<br />
Filamentous; planktonic<br />
Filamentous; marsh pools<br />
Filamantous; plantonfc<br />
Solitary; among marsh plants<br />
Sol i tary; amang marsh plants<br />
Solitary; among marsh plants<br />
Solltnry; among marsh plants<br />
Sulftary; among marsh plants<br />
Sol ftary; amang marsh plants<br />
Saldtary; among marsh plants<br />
Solitary; attached Lo debrfs<br />
Salltary; among marsh plants<br />
Solitary; among marsh plants<br />
5cZltaryy among marsh plants<br />
Sol l tary; amang marsh plants<br />
Sclftary; among marsh plants<br />
Solttary; among marsh plants<br />
So? f tary; among marsh pl ants<br />
Salltary; planktanfc & sessile<br />
Sol 4 tary; among marsh plants<br />
Solitary; among marsh plants<br />
Solltary; among marsh plants<br />
Sot ttargr; air*orzy marsh pl atits<br />
Sol f fary; among marsh plants
WPENDIX C (CONCLUDED)<br />
Taxa Form and habitat<br />
Sol i tary; among marsh p1 ants<br />
Sol i tary; among marsh plants<br />
Sol i tary; among marsh plants<br />
So1 i tary; among marsh pl ants<br />
Sol i tary; among marsh pl ants<br />
Fil amentous; among marsh pl ants<br />
Fil amentous; among marsh pl ants<br />
Filamentous; among marsh plants<br />
Sol i tary; among marsh plants<br />
Sol 1 tary; among marsh pl ants<br />
Sol i tary; among marsh pl ants<br />
So1 itary; among marsh pl ants<br />
Fi 1 amentous; among marsh pl ants<br />
Filamentous; among marsh plants<br />
Sol 1 tary; among marsh plants<br />
Solitary; among marsh plants<br />
So1 i tary; among marsh p1 ants<br />
Sol i tary; among marsh pl ants<br />
Sol i tary; among marsh pl ants<br />
Filamentous; among marsh plants<br />
Solitary; among marsh plants<br />
Sol i tary ; among marsh pl ants<br />
Sol i tary; among marsh plants<br />
Sol itary; p1 anktonic &<br />
among marsh plants<br />
So1 i tary; p1 anktonic &<br />
among marsh plants<br />
Sol itary; among marsh plants<br />
Sol itary; planktonic<br />
Sol itary; among marsh pl ants<br />
So1 dtary; p1 anktonic &<br />
among marsh plants<br />
Sol itary; among marsh pl ants<br />
Sol itary; among marsh pl ants<br />
So1 itary; among marsh pl ants<br />
So1 i tary; among marsh plants<br />
Sol i tary; among marsh pl ants<br />
Solitary; among marsh plants<br />
Sol itary; among marsh plants<br />
Sol itary; among marsh pl ants<br />
Sol i tary; among marsh plants<br />
a Data sources: Pfeters (189311 Winner et al. (19701, Taft and Taft (19713s<br />
Michigan Water Resources Commdssion (1975).
Phylum Protozoa fpsotoroansl<br />
C1 ass Sarcad l na<br />
Qrdar<br />
Class @Illopksre leflfalosf<br />
Order HaleP~lckEda<br />
FamSly Tracheldnldaa<br />
Order 925g(st,rlchlda<br />
Famfly Tlnttnfitda6<br />
ZWPLANKVQN OCGURRING Xt\B THE COASTAL WETLARDS<br />
AW NEARSHORE WATERS QF LAKE ST, CtAXWa<br />
1 52<br />
Abundance status<br />
Rare<br />
Cornon<br />
Camn<br />
Cornon<br />
Abundant<br />
Cornon
APPENDIX 0 (~oNTIMUED)<br />
Spec 1 es Abundance status<br />
Order Psri trichida<br />
Family Vorticallldae<br />
Family Ophrydiidae<br />
Phylum Rotatoria (rotifsrs)<br />
Class Monogononata (sfngle ovary rotifers)<br />
Order Flosculariacea<br />
Famf 1y Conochil idae<br />
unicorniq<br />
Family Tostudinellidae<br />
Order Co? 1 othecacea<br />
Famfly Collothscidae<br />
Order Ploimacea<br />
Family Notommatldae<br />
Family Synchaetidae<br />
Family Gastropodidae<br />
Occasional<br />
Common<br />
Rare<br />
Uncomnon<br />
Common<br />
Abundant<br />
Common<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Occasional<br />
Occasional<br />
Abundant<br />
Abundant<br />
Rare<br />
Rare<br />
Occasional<br />
Common
Spec 1 os Abundance status<br />
Rare<br />
Bcca5fonal<br />
ff are<br />
Rare<br />
Rare<br />
Occasional<br />
Rare<br />
Rara<br />
Rare<br />
Rare<br />
Abundant<br />
Rare<br />
Csmn<br />
kcas tonal<br />
kcas f anal<br />
Rare<br />
Ris re<br />
Rare<br />
Raro<br />
Rare<br />
Abundant<br />
Abundant<br />
Cornan<br />
Cmon<br />
Raro<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Ram<br />
Ra ra<br />
Rare<br />
Rare
APPENDIX D (CONTINUED)<br />
Species Abundance status<br />
Family Tylotrochidae<br />
--<br />
llxxEaG<br />
eriorlslghniq auadranu<br />
Phylum Arthropoda (joint-legged animals)<br />
Cl ass Crustacea<br />
Order Cl adocera (water fleas)<br />
Family Daphnidae<br />
aaPhnfa-<br />
DnDhn_aw-<br />
DaD.hnia-<br />
DaPbn.la-<br />
DaDhnLsaul_ex<br />
BaBhnlsret;racurvn<br />
--<br />
Fami 1 y Leptodoridae<br />
Le~todoril hindtii<br />
Family Sididae<br />
Uahmxam -era1 anum<br />
smd-<br />
Fami 1 y Hol oped i dae<br />
LlmQCm<br />
Family Bosminidae<br />
Bosmlna-<br />
EubPsminm.coreaon5<br />
Fami 1 y Chydoridae<br />
Alona-<br />
Alona-<br />
Blneanuadranaularis<br />
81snella SPP*<br />
-a<br />
Gdmmaas-<br />
Family Macrothrici dae<br />
Rare<br />
Uncommon<br />
Fare<br />
Rare<br />
Common<br />
Common<br />
Occasional<br />
Uncommon<br />
Common<br />
Occasional<br />
Uncommon<br />
Uncornmon<br />
Rare<br />
Occasional<br />
Abundant<br />
Abundant<br />
Uncommon<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Rare<br />
Occasional<br />
Rare<br />
Rare<br />
Rare
Spec 3 es<br />
Abundance status<br />
Unc mion<br />
hhc oarnor?<br />
SuBardtrr. bZ&rpaet Jc~Zda 4 hat-past lc<strong>of</strong>d cogspr~ds)<br />
e drrrf 1 y blasrpacticldao<br />
rn Rare
APPEMDlX E<br />
VASCULAR AQUATIC PLANTS KCURRING IN LAKE ST, CLAIR WETLANDSa<br />
Ecologicaf niche and species Abundance statusb<br />
Class Angiospermae (flowering p1 ants)<br />
Subclass Monocotyledonae (monocots)<br />
Qrdsr Pandanal es<br />
Family Typhaceae (cattails)<br />
(narrow-leaved cattail)<br />
road-1 saved cattail<br />
(hybrid cattail)<br />
Family Sparganium (bur reeds)<br />
(giant bur reed)<br />
Order A1 ismales<br />
Famlly Alismataceae (water plantains)<br />
(water plantain)<br />
mmon arrowhead)<br />
Family Butomaceae (flowering rushes)<br />
(flowering rush)<br />
Order Grami nal es<br />
Fami 1 y Grami neae (grasses) [bluejoint grass1<br />
tlesnake grass)
Ecological ndche and specles Abundance status<br />
Famf 1 y Cyperaceae (sedges)<br />
(tussock sedge)<br />
(redrooted cyperusl<br />
(rusty cyperus)<br />
f redfooted spiks-rush)<br />
unt spike-rush)<br />
1 (American bulrush,<br />
(dark green bulrush)<br />
(river bulrush)<br />
sat bul rush)<br />
Order Xyrldalas<br />
Fsmif y Pontedsriaceao (ptckeral weads)<br />
(water stargrass, mud plantain)<br />
(p fckarel wwd)<br />
Qrder ifllalaa<br />
Famlly Juncaceae (rushes)<br />
(s<strong>of</strong>t rush)<br />
(knotted rush)<br />
CTorreyrs rush1<br />
Ordnr arch Idalss<br />
Family Orchidaceae (orchf ds)<br />
(prairfe fringed orchfdl<br />
5ubcIass Olcatyladanas tdfeotsl<br />
Order Sal fcales<br />
Famfly Sallcacaae (willaws)<br />
C eastern ca.t;tonwood)<br />
(quaking aspen)<br />
y wlSSaw9
WPENDXX E (CONTINUED)<br />
Ec<strong>of</strong> ogical niche and species Abundance status<br />
Order Myrical es<br />
Family Jug1 andaceae (walnuts)<br />
(shagbark hickory)<br />
Order Fagal es<br />
Family Fagaceae (beeches)<br />
bicolar (swamp white oak)<br />
(burr oak)<br />
(pin oak)<br />
Order Urtical es<br />
Fami 1 y Ul maceae (elms 1<br />
Y.lnrus mericana (American elm)<br />
Famlly Urticaceae (nettles)<br />
Ilrtica dioica (stinging nettle)<br />
Order Polygonal es<br />
Family Polygonaceae (smartweeds)<br />
Pol va- am~hi b ium (water smartweed)<br />
Po1 vgfau~~ ~occine~ (swamp persicaria)<br />
Po1 v m ~onvol- (black bindweed)<br />
Pol vaow Mi f o l u (nodding smartweed)<br />
& d y g p~nsvl ~ ~ ~ vanica (Pennsylvania smartweed,<br />
pi nkweed)<br />
pol vgonum punctat~ (water smartweed)<br />
R j 3 vertici,ll atus (swamp dock)<br />
Order Ranales<br />
Family Nymphaeaceae (water lilies)<br />
Jutes (American water-lotus)<br />
Nu~hu & v e ~ ( spatterdock, ye1 1 ow water 1 $1 y 1<br />
Order Papaverales<br />
Family Cruciferae (mustards)<br />
Wamine pensvlvania (bitter cress)<br />
IWiQJB i2-1 (marsh cress)<br />
Order Rosa1 es<br />
Family Crassulaceae (orpines1<br />
Pent- (ditch stonecrop)<br />
Fami 1 y Rosaceae ( roses)<br />
(sf 1 verweed)<br />
mp rose)
Order SapSnd&lea<br />
F emf 3 y Anacardf aceao I sum&< b b<br />
(btaghorn sumac)<br />
family Acoraceact Imaplus)<br />
MS % tsf3ver maple)<br />
Ut-4c.r hbal alas<br />
FamfXy eorna~sae (woyruodsl<br />
gray dqiwsrjda<br />
# r ran a5 4er d~xjhwjd<br />
1
AQPEHDIX E ( CONTINUED 1<br />
Ecological niche and species Abundance status<br />
Family Boragfnaceae (borages)<br />
(common comfrey)<br />
Family Solanaceae (nightshades)<br />
(nightshade)<br />
Order P<strong>of</strong>emoniales<br />
Family Labiatae (mints)<br />
(water horehound)<br />
Family Acanthaceae (acanthids)<br />
(water wfllow)<br />
mad-dog sku1 lcap)<br />
Order Rub i a1 es<br />
Family Rubiaceae (madders)<br />
~ccidentali~ (buttonbush)<br />
Order Campanul a1 es<br />
Family Lobelliaceae (lobellas)<br />
(blue lobelia)<br />
Order Asteral es<br />
FamSly Composftae (composites)<br />
(nodding beggar tick)<br />
(purple-stemmed swamp beggar tick)<br />
(Hi11@s thistle)<br />
(bonset)<br />
Cl ass Ang i ospermae ( f 1 ower i ng pl ants 1<br />
Subclass Monocotyledonae (monocots9<br />
Order Na j adal es<br />
Fami 1 y Potamogatonaceae ( pondweeds)<br />
(knotty pondweed)<br />
Subclass Dicotyledonae Cdicots)<br />
Order Po1 ygonal es<br />
Family Polygonaceae (buckwheats)<br />
(water smartweed)<br />
(swamp smartweed)
APPENDIX E (CONCLUDED)<br />
Ecological niche and species Abundance status<br />
Order Xyridales<br />
Family Pontederiaceae (pickerel weeds)<br />
(mud plantain, water stargrass) C<br />
Subclass Dicotyledonae (dicots)<br />
Order<br />
-<br />
Ranal ss<br />
Family Ranunculaceae (buttercups)<br />
Jonairastris (water crowfoot)<br />
Order Myrtal es<br />
Family Ha1 oragidaceae (water mi 1 foils)<br />
Mvrf0~h~l1t,@ a1 terniflor_urn (1 Jttle water mil foil<br />
(water mil foil 1<br />
~~riophvllwm s~icatw (water milfoil)<br />
Family Onagraceae (evening primroses)<br />
(water primrose)<br />
Class Angiospermae (flowering plants)<br />
Subclass Monocotyledonae (monocots)<br />
Order AraZ es<br />
Family Lemnaceae (duckweeds)<br />
Lemnq $rI scu1 q (star duckweed)<br />
Subclass Dicotyledonae (dicots)<br />
Order Ranales<br />
Family Ceratophyllaceae (hornworts)<br />
Cerato~hvllace_ae demersw (hornwort, coontail) A<br />
Order Polemon fa1 es<br />
Family Lentibulariaceae (bladderworts)<br />
(bl adderwort)<br />
"0ata sources: <strong>St</strong>uckey (1968, 1972, 1975, 19791, J aworski and Raphael<br />
( 1978) Herdendorf et a1 . (1981~).<br />
b~bundance<br />
status: A = abundant, c = common, 0 = cccasional, R = rare,<br />
U = uncormnon.
BENTHIC MACROINVERTEBRATES OF NEARSWORE LAKE ST, CLAIR AND CONNECTING<br />
WATERWAYS (EXCLUDING MOLLUSKSIa<br />
Classfflcatfun and spscfss<br />
Phylum Porffera fsgongas)<br />
Class 0mosponglae (horny spangas)<br />
Famll y Spang11 l ldae ( fresh-water spanyes)<br />
SPP *<br />
Phylum Costante& ata (61 ass Cnf darfal<br />
Class Wydrozea (kydrozoans)<br />
Famtly Wydrldas<br />
SPP<br />
Phylum Platyhslmlnthes (flatworms)<br />
Class Turbel? aria t p1 anarl ansl<br />
Ordar Trfcladlda (triclads?<br />
Frtmlly Plannrlldae (planasfans1<br />
Phylum Namatoda (roundworms)<br />
Gl asa Adsnaphorao<br />
Family Trlpylfdae<br />
SP *<br />
Phylum Annelids (sagmented worms?<br />
GI ass Pal ychaeta<br />
FamE 1 y Sabal l idae ( Fan warmsr feather-duster worms1<br />
Class Bllgochaata laquatic aarthwarms)<br />
f emf l y Lun&r?cul idae
WPENDZX F (CONTINUED)<br />
Cl assi f ication and species<br />
Famdly Tubificidae (sludge worms)<br />
Famf ly Naididae<br />
sj2ik.h-<br />
Unc;.f- -<br />
Fami 1 y Gl ossoscol eci dae<br />
i2zIEE&<br />
Class Hirudinea<br />
--<br />
(leeches)<br />
Family Glossiphoniidae<br />
He1 &dell a $riserial is<br />
Order Pharyngobdellida<br />
Family Erpobdellidae
Classffjcatlon and spscfes<br />
Phylum Arthrapada (joint-lsggod anlmalsl<br />
C1 ass Arachndda<br />
Order Acarina (mites and tIcks1<br />
Superfamily Mydracarfna dwatgr rnftss)<br />
Famfly Limnssifdao<br />
Fami l y Wygrabatidaa<br />
sp,<br />
Family Plonidae<br />
Cl ass Crustacea<br />
Subclass Bstracoda (ssod shrimps)<br />
Famfly Cypridae<br />
SPP *<br />
Subcl ass Ma1 acast raca<br />
Qrder Isopoda (aquatlc sowbugs1<br />
Family Assllldas<br />
Ordsr Amphfpoda fstdeswfmrs, scuds)<br />
Family Gamaridae<br />
Order Becapoda tcrayff shesr shrimps)<br />
Subardsr Raptantfa (crayfJskss1<br />
Famlly Gambarldas (crayffshosb<br />
SP *<br />
Class Xnsacta (a Msxapodal /Insects)<br />
Order Ephemroptera (may fl 3 @r,!<br />
Fsmlly Baetfscfdas<br />
5P *
APPENDIX F (CONTINUED)<br />
Classiffcat4on and species<br />
Family Ephemeridae (burrowing mayflies)<br />
Fami 1 y Pol ymi tarci dae<br />
Order Odonata (dragonflies and damselflies)<br />
Suborder Anisoptera (dragonfl ies)<br />
Family Gomphidae<br />
SP<br />
Suborder Zygoptera (damselflies)<br />
Family Coenagrionidae (narrow-winged damselflies)<br />
SP *<br />
Order Hemiptera (bugs)<br />
Family Corixidae (water boatmen)<br />
SP<br />
-<br />
Family Hydropsychidae (net-spinning caddfsflles)<br />
SP.<br />
SP.<br />
Order Trjchoptera (caddi sf1 ies)<br />
Family Hydroptilidae (micro-caddisflies)<br />
HyGm@m SP.<br />
- SP *<br />
Family Polycentropodidae (tube-making caddisflies)<br />
SP.<br />
Family Psychomylldae (trumpet-net caddisflies)<br />
l2admuh SP*<br />
Family Mol annidae<br />
SP *<br />
Family leptocerldae (long-horned caddisfl les)<br />
!&xiz&a SP.<br />
SP *<br />
SP *
Cl asst f icaP3on and speclas<br />
Order Coleaptera (bsotles)<br />
Farnlly Hallplldaa (crawling water bsotles)<br />
Famtly Elmfdae (rlffla beetles)<br />
sp.<br />
Order DIptera ltrus flles)<br />
Faml l y Chaobor ldaa (phanton midges)<br />
SP *<br />
Family Culicidas (mosquftaesf<br />
SP *<br />
'8ata saurcos: Wo3cotf ( 1903 1 @ Edmondsan ( 1959), Hunt ( 19621, Hf 1 tunen<br />
tE971r 1BBQ)r Modlin and Gwnnon 6197311 Pennak (1S78Is Mrdsndarf et al.<br />
il$8lcIt Mfltunen and Manny (1982)r Hardsndorf and Hardsndorf (1983).
TAXONOMIC LISTING OF FRESHWATER MOLLUSCA OF LAKE ST. CLAIR<br />
COASTAL MARSHES, NEARSHORE WATERS, AND TRIBUTARY MOUTHSa<br />
Specf es<br />
Cl ass Gastropoda (snai l s)<br />
Subclass Prosobranchia (gill-breathing snails)<br />
Order Mesogastropoda<br />
Superfamily Viviparidae (mystery snails)<br />
Famlly Vivfparidae (mystery snails)<br />
816) brown mystery sna?l<br />
r 1834) banded mys<br />
(Gray, 1834) (= Y,<br />
oriental mystery snail or Japanese snail<br />
Superfamily Valvatacea<br />
Family Valvatidae (valve snails)<br />
Valvat3 w e s s q Walker, 1906 flat valve snail<br />
Valvata piscf nal is (Mu1 ler, 1774) European valve snail<br />
slncer~ sincera Say, 1824 ribbed valve snail<br />
trlcarinata (Say, 1817) three-keeled valve snail<br />
Superfamily Rissoacea<br />
Family Hydrobiidae (spire snails)<br />
clncinnatiensis (Anthony, 1840) campeloma spire<br />
snail<br />
(Baker, 1928) fl at-ended spi re snail<br />
r, 1928) Pilsbryps spire snail<br />
Say, 1817) ordinary splre snail<br />
Pilsbry, 1898 small spire snail<br />
(Say, 1825) deep water spire snail<br />
Family Truncatellidae (looplng snails><br />
(Say, 1817) river bank looping snail<br />
Family Bfthynjidae (faucet snails)<br />
l2mULb (Lfnnaeus, 1767) faucet snafl<br />
Superfamily Cersithiacea<br />
Family Pleuroceridae (horn snai 1s)<br />
Rafinesque* 1831 fl at-sided horn snaS9<br />
(Menke, 1830) Great <strong>Lake</strong>s horn snaS3<br />
--
Spsc .i tss<br />
APPENDIX C; (CONTINUED)<br />
Subclass Pulmansla Qfuny-breathing snafls)<br />
Ordsr Basomatophosa<br />
Superfamlly Lymnaeacea<br />
Famlly Lymnaefdea (pond snaf 1s)<br />
(<strong>St</strong>reng, 18961 shouldered northern fossarfa<br />
sa, laall graceful fossarja<br />
(Say, 18251 modest fassaria<br />
(Lea, 1841) amphlblaus fossarfa<br />
er, 1907) small pond snall<br />
{Say, 1817) AmerScan ear snail<br />
nney, 18671 slsnder pond snall<br />
y pond snall<br />
1817) great pond snafl<br />
(Say, 1817) lake stagnjcola<br />
sr, 19069 mini aturg<br />
(Sayt 18211 comon stagnfcola<br />
(Say, 1821) striped staynlsola<br />
Superfamlly Physacea<br />
Family Physfdaa (tadpals snaffs)<br />
Say, 1821 tadpols snafl<br />
eman, 1841 solid lake physa<br />
(Llnnaeu~~ 1758) polished tadpole snafl<br />
Superf amll y Pl anorbacoa<br />
Family Pl anarbldae (ramshorn snaj 1 s)<br />
ETyron, 1866) flatly colled gyraulus<br />
Baker, 1932 great carinate<br />
(Sayc 1816) larger eastern ramshorn<br />
Fa e f reshwatsr l impets9<br />
.G. Adams, 18411 dusky lily-pad lfmpet<br />
(Tyron, 1863) oval lake lfmpet<br />
Cl ass Pel ecypoda<br />
Order Eul amel l ibranchia<br />
Superf am1 1 y Un i onacea<br />
Family Unionidae (pearly mussels)<br />
Subfamily Ambleminae (button shells and relatives)<br />
(Say, 1817) three-ridge<br />
Rafinesque, 1820) plg-toe<br />
(Rafi nesgue, 1820) maple-leaf<br />
Lea, 1831) warty-back<br />
(Rafinesque, 1820) purple pimple-back<br />
inesque, 1820) spike, or lady-finger<br />
(Conrad, 1836) false pig-toe<br />
Subfamily Anodonti nae ( f l oater mussels)<br />
Raf inesque, 1820) brook wedge mussel<br />
Say, 1819 ridged wedge-mussel<br />
(Barnes, 1823) white heel-splitter<br />
29) brook l asmigona<br />
er fluted shell<br />
1 (Say, 1825) mudpuppy<br />
(Lea, 1834) cyl lndrical floater<br />
ay, 1829 common floater<br />
paper pond-she1 l<br />
SugAxku ur,ululaUs (Say, 1817) squaw-foot<br />
Subfamily Lampsi1 inae (1 amp-mussels)<br />
(Rafinesque, 1820) kidney shell<br />
gue, 1820 three-horned warty-back<br />
Jruncilla maciformis (Lea, 1828) fawn%-foot<br />
Truncilla fsunc- Rafinesque, 1820 deer-toe<br />
U (Say, 1817) pink heel-spl itter<br />
(Barnes, 1823) lilliput mussel<br />
(Raf inesque, 1820) round hickory-nut<br />
Raffnesque, 1820) olive hickory-nut<br />
Rafinesque* 1820) fragile paper-shell<br />
(Barnes, 1823) mucket<br />
, 1817) polnted sand-shell<br />
(Lamarck, 1819) bl ack sand-she1 1<br />
r 1820 wavy-rayed lamp-mussel<br />
(Barnes, 1823) fat mucket<br />
f Barnes, 1823 > pocket-book<br />
, 1831) bean villosa<br />
(Leal 1838) northern rfffle shell<br />
sque, 1820) trieorn pearly mussel
Spec l sf;<br />
APPENDIX &; 6.<br />
SuperfamDly Sphaarlacsa<br />
Famlly Corblculfdae (71ttle basket clams)<br />
lMullor, 2774) Aslatls clam<br />
Famlly Sphaerifdae (ffngornafl clams and poa clams)<br />
Subfamily Sphasrilnaa (fingarnail clams)<br />
nnaous, 17583 European flngernafl clam<br />
ssfn, 1876 Arctic-alpine Flagernall clam<br />
(Say, 18221 rho&<strong>of</strong>d ffngernall clam<br />
y, 18161 grooved ffngernatl clam<br />
f lannarck, liWl8) strfated flngernaf I clam<br />
Mullsr, 27743 faks fingarnail clam<br />
(Say, 18221 swamp flngernafl clam<br />
fme, 18511 pond ffngernadl clan<br />
(Say, 1829) long ffngsrnafl clam<br />
Subfamtly Pi%ldl%nao Epsa clams or pill clams)<br />
(Muller, 17741 greater European pea clam<br />
Roper, 1890 giant northern pea clam<br />
me, 1852 Adamt% pea clam<br />
(Poli, 139%) ubdqultous poa clam<br />
Prfme, 1852 rfdged-beak pea clam<br />
Primer 1852 rsufid pea clam<br />
d, 1898 rfver pea clam<br />
Prlmsr 1852 rusty poa clam<br />
(Sheppardr 1895) Hsnslowts paa clam<br />
Clersint 1886 Lilljeborgqs pea clam<br />
1836 quadrangul ar plll cl am<br />
fisr L832 shlny pea clam<br />
rlme, 1852 fat pea clam<br />
Malm, 1855 shart-sndsd pea clam<br />
Primer 1852 triangular poa clam<br />
Primat 1851 globular pea clam<br />
rkll 1895 Walkorqs pea clam<br />
@lasainl 1877 Arctic-alplns poa clam<br />
SLerkf r 1895 perforatad pea clam<br />
--<br />
& OaLa sources: Gosdrlch and Van dor fehalla (&93211 Berry (19431, <strong>St</strong>ansbary<br />
I196OT* Clarke (X98l1,
Species<br />
APPENDIX H<br />
HABITAT PREFERENCE OF FRESHWATER MOLLUSCA OF LAKE ST. CLAIR<br />
COASTAL MARSHES AND NEARSHORE WATERS<br />
Shallow Deep Gravel<br />
((2 m) (>2 m) Mud Sand & rock Plants Comments<br />
Eutrophf c<br />
Eutrophic<br />
Rare<br />
Rare
Spec t as<br />
APPENDIX W ICQNIINUED)<br />
Shall sw Deep Gravel<br />
((2 m) f >2 m) Mud Sand 8 rock PI ants Cornants<br />
x Eutrophic<br />
x Eutraphlc<br />
x Rare<br />
X<br />
x EutrophSc<br />
X<br />
x Eutrophic<br />
X<br />
X<br />
X<br />
X<br />
X<br />
X<br />
X<br />
X<br />
X<br />
Rare
Species<br />
Shall ow Deep Gravel<br />
((2 m) (>2 n) Mud Sand d rock Plants Comments<br />
X X X<br />
x x Rare<br />
X X<br />
X X<br />
X X X<br />
X X X X<br />
X X X<br />
X<br />
Rare<br />
a~ata<br />
sources: Goodrich and Van der Schal ie (1932) Berry (19431, Clarke<br />
( 1981 1.
Famlly and species<br />
Fsmfly Aclpenserldas<br />
FISH KNOWN OF4 F'Rt SlSMED TO U"I"TI_IZE THE COASTAL WETiADuUS<br />
OF THE ST, CLAXR RXVEM AND LAKE ST, CtAIRa<br />
( sturgeon 1<br />
Famfly kspfsostsldee (gars)<br />
l angnase gas) Coprm~n Gomon<br />
(spotted gar) Unc<strong>of</strong>nnrun Dncorman<br />
Family fso~ldas (pfkesl<br />
Cnorthsrn plksl Abundant Camn<br />
(musks1 1 ungo)<br />
(grass pfckarsl 1<br />
Camon<br />
Camon<br />
Uncomon<br />
Comn<br />
Famfl y Umbrf dae (mudm9 nnawsl<br />
(central mudmf nnow?<br />
Fa I rigs)<br />
(gfttard shad)<br />
Cyalden ~hjnerf<br />
blackchin shiner)<br />
spattat 1 shf narl<br />
femrald shingar)<br />
sand shiner)<br />
g fathead mS nhov I<br />
Camon<br />
Rare<br />
Ra rs<br />
Comn<br />
Camn<br />
Abundant<br />
Ataerndant<br />
Abtnradant<br />
Abundant<br />
Csmn<br />
Itncomn<br />
Comn<br />
Rare<br />
Rare<br />
Camn<br />
Cornon<br />
Abundart<br />
Abundant<br />
Camon<br />
C O ~ R<br />
Abundant<br />
Abundant
Famlly and species<br />
Family Catostomidea (suckers)<br />
(white sucker)<br />
e chubsucker)<br />
(spotted sucker)<br />
Family Ictaluridae (catfishes)<br />
.li&&uE (channel catfish)<br />
ma1 urus me1 as ( bl ack bull head 1<br />
Jctalurus ~ u l o s (brown u ~ bull head)<br />
Lctalurug natalis (yellow bullhead)<br />
Ngturus gvrinu (tadpole madtom)<br />
Family Cyprinodontidae (killifishes)<br />
Fundulu dlaohanur (banded killifish)<br />
Family Centrarchidae (sunfishes)<br />
Bmblo~l ites ~ e s t r i s ( rock bass)<br />
je~omi3 c~anellus (green sunfish)<br />
) I s albbosus (pumpkinseed)<br />
~ o c h i r u s (bluegill<br />
Micro~teru dolomiea (smal lmouth bass)<br />
Micro~teru salmoide~ (largemouth bass)<br />
Pomoxi~ gaaulari~ (white crappie)<br />
~1iaromacu1 atu (black crappie)<br />
Family Percidae (perches)<br />
fiheostoma exile (Iowa darter)<br />
Etheosto~ D I (johnny darter)<br />
perca f 1 avescens (ye1 1 ow perch<br />
Percina wades (1 ogperchl<br />
<strong>St</strong>izostedion vitreu~ (walleye)<br />
Family Perichthyidae (basses)<br />
Nrone &- (white bass)<br />
Family Sciaenidae (drums)<br />
A~lodinotus grunniens (freshwater drum)<br />
Family Gasterosteidae (sticklebacks)<br />
Culea inconstans (brook stickleback)<br />
Family Cottidae (sculpins)<br />
(mottled sculpin)<br />
-- --<br />
aorta source: Herdendorf et al. (1981~).<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Uncommon<br />
Common<br />
Common<br />
Common<br />
Common<br />
Common<br />
Common<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Uncommon<br />
Common<br />
Common<br />
Abundant<br />
Abundant<br />
Common<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Common<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Uncommon<br />
Common<br />
Common<br />
Common<br />
Common<br />
Common<br />
Common<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Uncommon<br />
Common<br />
Common<br />
Abundant<br />
Common<br />
Common<br />
Common<br />
Common<br />
Common<br />
Conimon<br />
Common
APPENDIX J<br />
WPWXRIANS AND REPTILES OGCURRING IN THE COASTAL WETLANDS<br />
OF LAKE ST. CLAIR AND THE ST. CLAlR RIVERa<br />
Spec 1 er Abundance status<br />
Class Amphfbfa (amphfbians)<br />
Order Gaudata (salamanders)<br />
PPY<br />
Crsd-spotted newt)<br />
Caastsrn tfger salamander)<br />
(blue-spotted sal amander)<br />
( spattad sal amanderf<br />
sd-backed salamander)<br />
(four-toed salamander)<br />
Order Saltantfa Cfroys and toads)<br />
tern chorus frog)<br />
% crfckot frog)<br />
Class Regtffla Creptflasl<br />
Ordar Squaaata (snakes and llzards)<br />
(northern water snake1<br />
(northern rlbbon snake)<br />
(northern red-ballfsd snake)<br />
brawn snake)<br />
mf dl and brown snake)<br />
f~astern hagnose snake)<br />
(northern rtngnsclk snake)<br />
rn $moth green snake)<br />
Comon<br />
Comon<br />
Common<br />
Comn<br />
Coman<br />
Abundant<br />
Rare<br />
Abundant<br />
Cam0<br />
Comon<br />
Comon<br />
Gomon<br />
Comon<br />
Comn<br />
Abundant<br />
Camnon<br />
Comn<br />
Abundant<br />
Abundant<br />
Uncomnon<br />
Abundant<br />
Uncomn<br />
Comon<br />
Comon<br />
Comn<br />
Coman<br />
Comn
Spec S es Abundance status<br />
feLd. (blue racer)<br />
eastern fox snake)<br />
rat snake)<br />
(eastern ml1 k snake)<br />
(eastern massasauga)<br />
Order Testudf nes (turtles)<br />
Z, ser~entlnq (snapping turtle)<br />
Common<br />
Rare<br />
Rare<br />
C0mm0~<br />
Uncomnon<br />
Comnon<br />
Rare<br />
Common<br />
Common<br />
Abundant<br />
Comon<br />
Rare<br />
a~ata sources: Ruthven et al. (1928) * Conant (1975). Behler and King (1979).<br />
Herdendorf et a1 . (1981~).
BIRDS KCURRING IN THE VICINITY OF LAKE ST. CLAIR<br />
AND ST. CLAZR RIVER COASTAL WETLANDS3<br />
Spec f es Resf dance statush<br />
Class Avas (Bfrds)<br />
Qrdar Anserifarms<br />
Famlly Anatfdae (swans# go@$@, and ducks1<br />
SubfamJly Anserinae (swans and ge@sa)<br />
Trfbe Cygnlni<br />
(tundra swan)<br />
Subfamily Andtinaa Iducksl<br />
Trlbe CafrZninl<br />
(wood Buck)<br />
(dabbl f ng ducksf<br />
whfte-fronted goose)<br />
(canvasback)
Tribe Oxyurini<br />
APPENDIX K (CONTINUED)<br />
fflehead) W<br />
(hooded merganser) M<br />
common merganser) M<br />
( red-breasted merganser) M<br />
(ruddy duck)<br />
Order Podicipedi formes<br />
Family Podicipedidae (grebes)<br />
orned grebe) M<br />
(pied-billed grebe) BR<br />
Order Pelecaniformes<br />
Family Pelecanidae (pelicans)<br />
(white pelican) M<br />
Residence status<br />
Order Gruiformes<br />
Famlly Ra11 idae (rails, moorhensp and coots)<br />
BR<br />
(Virginia rail BR<br />
BR<br />
(common moorhen) BR<br />
merican coot) BR<br />
Order Ciconi i formes<br />
Family Ardeidae (herons)<br />
(American bittern) BR<br />
BR<br />
YR<br />
BR<br />
BR<br />
(green-backed heron) BR<br />
f bl ack-crowned BR<br />
n-ight-heron)<br />
Order Charadrllformes<br />
Family Charadri idae (plovers)<br />
(semi palmated plover) M
- OIIJL<br />
h<br />
e-. L<br />
UI a3 CI<br />
wl Q. I<br />
a, n-<br />
F Ira -g<br />
%--QIW L.r<br />
o m a c m a<br />
c m-- m am<br />
~ o a m c c<br />
a?= a a<br />
X3rCh mtn<br />
C<br />
vlL"-rn@a at-<br />
L@P- .P m a<br />
@i).t-)@-W--- L<br />
116s XQIF m*, Q<br />
+totmwa~<br />
t +rn-mu~=<br />
a Cno 0-r 0 @F-<br />
E-=--uiQ. LP-ar-<br />
s<br />
x -<br />
u ."-<br />
at-<br />
3 3<br />
1 Cn<br />
x<br />
u w<br />
FO. c<br />
i r - F3
WPEMDXX K (CONTINUED)<br />
Spec f es Residence status<br />
Family Accipitridae
APPENDIX K (CONCLUDED)<br />
Spec l es Residence status<br />
Subfamily Emberizinae (sparrows)<br />
(swamp sparrow) Y R<br />
(savannah sparrow) BR<br />
a~ata sources: Peterson ( 19801, Herdendorf et a1 . ( 1981~1, Scott (1983).<br />
b~esi<br />
dence key :<br />
YR - Year-round resident BR - Breeding range (spring and summer)<br />
M - Migrant W - Winter range
APPENDIX L<br />
MAMMALS OCCURRING IN THE COASTAL WETLANDS OF LAKE ST. CLAIR<br />
AND THE ST. CLAIR DELTAa<br />
Species Abundance status<br />
Order Marsupialla (marsupials)<br />
Family Didelphidae<br />
(Vi rgfnia opossum)<br />
Order Lagomorpha<br />
Family Leporidae<br />
(eastern cottontail)<br />
(European hare)<br />
Order Rodentia<br />
Famlly Sciuridaa<br />
(woodchuck)<br />
ray squf rral<br />
rrel )<br />
( red squirrel 1<br />
Family Cricetl dae<br />
Order Carnivora<br />
Family Canidae<br />
Yu1pcs. Lu.lxa (red fox)<br />
Famil y Procyon idae<br />
( raccoon 1<br />
(muskrat 1<br />
Family Mustel idae<br />
frenat~ (long-tailed weasel)<br />
Order Art i odactyl a<br />
Family Cervf dae<br />
(striped skunk)<br />
(white-talled deer)<br />
'~ata sources: Hayes ( 1964) r Herdendorf et a1 . ( 1981~).<br />
Common<br />
Rare<br />
Occasional<br />
Uncommon<br />
Uncommon<br />
Occasional<br />
Abundant<br />
Qccas ional<br />
Occasional<br />
Occasfonal<br />
Occasional<br />
Camon<br />
Rare
APPENDIX M<br />
OWNERSHIP OF MAJOR LAKE ST, CLAIR WETLANDS<br />
Wetland Name Re1 a t i ve Water-1 eval Veg .<br />
Pub1 i c Private areab controlC typed<br />
-- ---<br />
WAYNE COUNTY, MICHIGAN<br />
I. Belle Isle M/ F<br />
MACOMB COUNTY* MICHIGAN<br />
2, Black Creek -<br />
3. Clinton Rtver Estuary M<br />
4. Betvidere Bay -<br />
5. Salt River Estuary -<br />
ST. CLAJR COUNTY, MICHIGAN<br />
6. Marsac Paint .<br />
7, Swan Cresk Estuary -<br />
8. <strong>St</strong>. Johnts Marsh S<br />
9. Dickinson Island S/F<br />
10. Harsens Is1 and S<br />
11. Bslle Rfvar Estuary -<br />
12. Pine River Estuary M<br />
13, Marysvi 11 e -<br />
14, Port Huron -<br />
ESSEX COUNN, ONTARIO<br />
25, Peach Island<br />
16. Ruscom River<br />
17, Thames River Estuary<br />
KENT COUNTY p ONTARIO<br />
18, Bradley Marsh<br />
19. BJg P<strong>of</strong>nt Marsh<br />
20, Tacky Marsh<br />
21. MJtchell Pafnt Marsh<br />
(Moon Island Marsh)<br />
22. Mitchell Bay<br />
CPlntal1 and Mud Creek<br />
marshes9<br />
23, Val 1 aceburg<br />
(Bear Creek and<br />
Pigeon marshes9
APPENDIX M (CONCLUDED)<br />
Wetland Name Re1 at i ve Water-1 evel Veg .<br />
Public Private area control type<br />
LAMBTON COUNTY, ONTARIO<br />
24. <strong>St</strong>. Anne Is1 and<br />
25, Wal pol e Is1 and<br />
26. Squirrel Island<br />
27. Bassett Island<br />
28. Seaway Island<br />
29. Port Lambton<br />
30. Fawn Island<br />
31. <strong>St</strong>ag Island<br />
-<br />
-<br />
32, Sarnia M<br />
a<br />
Ownership Key: b~rea Key: dVeg. Type Key:<br />
F = Federal S = Small (>500 ha) S = Submergent<br />
S = <strong>St</strong>ate or Provincial M = Medium (500-lrOOO ha) E = Emergent<br />
M = Municipal<br />
SC = Shooting Club<br />
L = Large (>l,000 ha)<br />
PM = Private, multiple owners CWater-1 eve1 Control Key:<br />
PS = Private, single owner D = Diked, managed marsh<br />
R = Indian Reservation U = Uncontroll ed wet1 and<br />
'~ote: Between Bradley Marsh (18) and Mitchell Bay (22) several shooting<br />
clubs operate small marshes, including Recess Club, Balmoral Club, <strong>St</strong>. Lukes<br />
Clubr Rex Club, Bay Loge Club* Rex/Cadotte Marsh, and Rankin/Sloan Marsh.
<strong>The</strong> <strong>Ecology</strong> <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Wetlands</strong>:<br />
A Communi ty Pr<strong>of</strong> i 1 e<br />
I R eW Dote<br />
September 1986<br />
b Ibr(.)<br />
Charles E. Herdendorf ,I C. Nicholas Raphael, Eugene ~aworski<br />
6. P<strong>of</strong>formln(l O~anlzotlon Rem. No<br />
Authors' Affiliations: la CmimctfTesk/Work Unit NO.<br />
lCenter for <strong>Lake</strong> Erie Area Research<br />
<strong>The</strong> Ohio <strong>St</strong>ate University<br />
Columbus, OH 43210<br />
U w m O~llaniatlon Nome and Address<br />
National <strong>Wetlands</strong> Research Center<br />
Fish and Wildlife Service<br />
U.S. Department <strong>of</strong> the Interior<br />
Washington, DC 20240<br />
Geography<br />
and Geology 11. Controct(C) w Grnnt(O) No.<br />
Eastern Michigan University ,<br />
Ypsilanti, MI 48197<br />
la Abstnct (Limit: loo words) I<br />
This publication reviews the ecological data and information on the wetlands<br />
<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> lake, whicn is a part <strong>of</strong> the Great <strong>Lake</strong>s ecosystem, is<br />
situated between southeastern Michigan and southwestern Ontario. <strong>Lake</strong> <strong>St</strong>.<br />
<strong>Clair</strong> contains a substantial portion <strong>of</strong> the remaining Great <strong>Lake</strong>s coastal<br />
wetlands, with extensive wetlands occurring in the vicinity af the <strong>St</strong>. <strong>Clair</strong><br />
River Delta. <strong>The</strong>se wetlands are among the most productive areas in the Great<br />
<strong>Lake</strong>s ecosystem. This publication summarizes the geologic history <strong>of</strong> the<br />
region leading to the development <strong>of</strong> wetlands and the present environment;<br />
biological production and community organization in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />
wetlands are also examined. <strong>The</strong> publication closes with a chapter on applied<br />
ecology which addresses issues such as wetland disturbance, management, and<br />
future prospects for these wetlands.<br />
LI. Mllltlfian/O$wt.Ended Terms<br />
Nutrient dynamics<br />
lrophic interactions<br />
Invertebrates<br />
Birds<br />
Great <strong>Lake</strong>s<br />
Navigation<br />
Rnthropogenic i n f 1 uences<br />
nt j 9<br />
- ---<br />
. Security Class (Thts Rrwm 21. No <strong>of</strong> Psges<br />
Unclassified<br />
Unlimited distribution -- -<br />
I Swd ANS1-339.11))<br />
1 Unclassified<br />
22. Pncc<br />
I<br />
ORIOMAL FORM 272 (4-77)<br />
(Forwrly NJIS-35)<br />
Dapartment <strong>of</strong> ComWrC4