27.04.2013 Views

The Ecology of Lake St. Clair Wetlands: A Community Profile

The Ecology of Lake St. Clair Wetlands: A Community Profile

The Ecology of Lake St. Clair Wetlands: A Community Profile

SHOW MORE
SHOW LESS

Create successful ePaper yourself

Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.

Char1 es E. Herdendorf<br />

Center for <strong>Lake</strong> Erie Area Research<br />

Department <strong>of</strong> Zoo1 ogy<br />

<strong>The</strong> Ohio <strong>St</strong>ate University<br />

Columbus, OH 43210<br />

and<br />

C. Nicholas Raphael<br />

Eugene Jaworsk j<br />

Department <strong>of</strong> Geography and Geology<br />

Eastern Michigan University<br />

Ypsil anti, MI 48197<br />

Project Officer<br />

Walter G. Duffy<br />

Natlonal Wet1 ands Research Center<br />

U. S. Fish and Wildlife Service<br />

2010 Gause Boulevard<br />

S1 idell, LA 70458<br />

Prepared for<br />

National <strong>Wetlands</strong> Research Center<br />

Fish and Wild1 ife Service<br />

U. S, Department <strong>of</strong> the Interior<br />

Washington, DC 20240<br />

Fur salr by the Superizitt.ndrn: U! I)~c~rnenf,<br />

U.S. Guvernmrn!<br />

Printing Offrrc. U'ashrngton. D.C. 20102<br />

Biological Report 85(7.7)<br />

September 1986


tierdcndorf, Charle5 T.<br />

<strong>The</strong> ccolnqy <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Ila~r wetldnd~~.<br />

(R~oloqical rcport ; 8!7( J. i))<br />

"Sep tcarirer 1911b."<br />

Supt. <strong>of</strong> Ilocc;. no.: I 49.R0/i':ti'~(i.?)<br />

I, tictlanci t~ology--Saint (,lair, I.crke (Micii. arid Oiit.)<br />

?. Ndn--1rif luent e on ndture--Sdint ("l~flr, I d/\~ (Mi~h.<br />

drld 011t.) 3. Lic)tlar~d ccoloc~y--Great i ~ ikrs Rccjion.<br />

4, Mdrr--1trfI~ic~ntc on rratut-c--Gr-edt I crkcs Krcjl'on.<br />

1, Ral)t?~tcl , t. Nictlolds, lQ'37- . l i. ,Icrwor-ikr,<br />

Eutjc.rie, i i l. Ndtiondl Wctfdr~rls iie$c-'rdt ti C~nter (11.4.)<br />

IV. T~tlc. V. Titlc: icolarjy <strong>of</strong> 1 akr 'Liint (Idi.<br />

wctlilndi. VI. C;ilt.ics: fiiolor~it~tl t*i")or+t (W(~rlshincjl.:~r~,<br />

l1.L.) ; !3!>-7* 7*<br />

fhii report may be cited as:<br />

Herdendnrf, C. t, , C. N. Rdyhac? , and E. Jaworskl 1986. <strong>The</strong> ecology <strong>of</strong> iake<br />

<strong>St</strong>. Ciair-weliarlas: d~~~ii~lilfr~it~~i~;i:~.<br />

3.5. Fi~f;W:id!. Ser-t. Sjc?. Rep.<br />

85(7.1) 187 pa,


This report on <strong>Lake</strong> S t , <strong>Clair</strong><br />

wetlands is one <strong>of</strong> a series <strong>of</strong> community<br />

pr<strong>of</strong>iles that deal with marine and<br />

freshwater habitats <strong>of</strong> ecological impor-<br />

tance. <strong>The</strong> delta marshes, estuaries,<br />

lagoons, and channel wetlands which fri nge<br />

the Michigan and Ontario shores <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>, and the <strong>St</strong>. <strong>Clair</strong> and Detroit<br />

rivers, are among the most productive<br />

areas in the Great <strong>Lake</strong>s. Because they<br />

occur in the proximity <strong>of</strong> a densely<br />

populated, heavily industrial ized, and<br />

intense1 y farmed region, the marshes have<br />

suffered losses in both area and qua1 ity.<br />

However, the remaining marshes are vital<br />

habitats for migratory waterfowl, fur-<br />

bearers, and fish, and perform many<br />

important hydrological and ecological<br />

functions. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta is unique<br />

in the Great <strong>Lake</strong>s and fosters some <strong>of</strong> the<br />

finest coastal wetlands in the region.<br />

<strong>The</strong> biological resources <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> have attracted man since prehistoric<br />

time. Not only were fish and wlldlife<br />

consumed but the wetland plants were used<br />

for crafts, cures, and kitchen utensils by<br />

the early Indian settlers. Archaeological<br />

sites and early explorers <strong>of</strong> the region<br />

have recorded the close interactJon <strong>of</strong><br />

early man and the coastal environment.<br />

<strong>Lake</strong> <strong>St</strong>. Clafr was lldiscoveredN by<br />

Europeans on Saint Clalreqs Day, August<br />

12, 1679, and was named in honor <strong>of</strong> the<br />

saint. Early fur traders were consls-<br />

tently overwhelmed by the bountiful fish<br />

and wildlife stocks <strong>of</strong> the lake and <strong>of</strong>ten<br />

remarked in their journals about the<br />

tranquil beauty <strong>of</strong> the marshesc flora. A<br />

colorful episode <strong>of</strong> the regfon was l inked<br />

to the prohibitton era. During this time,<br />

local residents claim that the <strong>St</strong>. <strong>Clair</strong><br />

Flats, or delta, was a hub <strong>of</strong> activity for<br />

the importation <strong>of</strong> i 1 legal 8 but excel 1 ent<br />

quality* whiskey from Canada to the United<br />

<strong>St</strong>ates. Today, <strong>Lake</strong> <strong>St</strong>. Clais is one <strong>of</strong><br />

PREFACE<br />

the most intensively utilized lakes in<br />

North America with its International<br />

wetlands continuing to be significant<br />

contributors to recreational activities<br />

throughout the year. Without these wet-<br />

lands, the value <strong>of</strong> the lake would be<br />

greatly diminished.<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is noted for its<br />

storms and rapid water level changes. <strong>The</strong><br />

moderate energy produced by these storms<br />

1 imits the existence <strong>of</strong> coastal wetl ands<br />

to places where some type <strong>of</strong> natural or<br />

artificial protection is available.<br />

Corresponding1 yr the coastal marshes <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fall Into four categorf es:<br />

1) delta wetlands, 2) coastal lagoons<br />

behind protective barrier beaches, 3)<br />

estuarine tributary mouths, and 41 managed<br />

marshes protected by earthen and rip-rap<br />

dikes. According to Gowardin et al,<br />

(1979) these wetl ands would include<br />

elements <strong>of</strong> riverine, 1 acustrf ner and<br />

palustrine systems <strong>of</strong> the recently<br />

developed National Wet1 and Cl ass T f ication.<br />

<strong>The</strong> li.Jossary Geol09y (Bates and<br />

Jackson 1980) def f nes '"freshwater estu-<br />

aries" for the Great <strong>Lake</strong>s as, '%he lower<br />

reach <strong>of</strong> a tributary to the lake that has<br />

a drowned rtver moutht shows a zone <strong>of</strong><br />

transition from stream water to lake<br />

water, and is influenced by changes fn<br />

lake level as a result <strong>of</strong> sefches or wind<br />

tides." Brant and Herdendorf ( 19721 were<br />

among the f f r st investigators to descrf be<br />

the characterfstics <strong>of</strong> Great <strong>Lake</strong>s<br />

estuaries. Such estuarfes are important<br />

wetland habitats in western <strong>Lake</strong> Erie.<br />

<strong>The</strong> definitfon gfven above provides an<br />

adequate physical decrfption for the<br />

purposes sf thf s report.<br />

<strong>The</strong> information in this report Is<br />

intended to provide a baslc understandfng<br />

<strong>of</strong> the ecological relatfonships wfthln the


<strong>Lake</strong> <strong>St</strong>. CY air wetlands and the impact <strong>of</strong><br />

natural and man-i nduced disturbances on<br />

the coastal marsh community. References<br />

are provided for those seeking more<br />

detai 1 ed treatment <strong>of</strong> speci f l c aspects <strong>of</strong><br />

the coastal marsh ecology. An appendix 4s<br />

included which lists the dimensions and<br />

ownership <strong>of</strong> the major marshes, and the<br />

important biologfcal species <strong>of</strong> a1 yae,<br />

macrophytes, invertebrates, fish,<br />

amphi bians, reptiles, birds# and mammals<br />

occurring in the coastal marshes,<br />

Any questions or comments about, or<br />

requests for this publication should be<br />

addressed to:<br />

Informat-ion Transfer Speci a1 ist National Wet1 ands Research Center<br />

U. S. Fish and Wildlife Service<br />

NASA-Sl idell Computer Complex<br />

1010 Gause Boulevard<br />

Sl idell, LA 70458<br />

(50% i 546-73 10 FTS 680-73 20


FOR NlETRlC (SI) UNITS TO U.S. CUSTOMARY UNITS OF MEASUREMENT<br />

Metric (SI) units <strong>of</strong> measurement used in this report can be converted to U.S.<br />

customary units as follows:<br />

Linear measurements;<br />

mill imetres (mm)<br />

cent imetres (cm)<br />

metres (m)<br />

k i 1 ometres (km)<br />

square metres (m2)<br />

hectares (ha)<br />

square kilometres (km2)<br />

Yo1 ume measurements;<br />

cublc metres (dl<br />

cubic metres (dl<br />

cubic kilometres (kd)<br />

Mass measu r emenk<br />

grams (g)<br />

kilograms (kg)<br />

metric tons (m ton)<br />

Bate measurements;<br />

0.039 inches (in)<br />

0.394 inches (in)<br />

3.281 feet (ft)<br />

0.621 miles (mi)<br />

10.764 square feet (ft2)<br />

2.471 acres<br />

0.386 square miles (mi21<br />

35.318 cubic feet (ft3)<br />

1.308 cubic yards iyd3)<br />

0.240 cubic miles (mi31<br />

0.035 ounces (ozl<br />

2.205 pounds (lbf<br />

1.102 U.S. tons (ton)<br />

centimetres per second (cm/sec) 0.394 inches per second I1 n/sec)<br />

metres per second (m/sec) 3.281 feet per second (ft/sec)<br />

metres per second (m/sec) 1.943 nautical miles per hour (knot)<br />

cubic metres per second (m3/sec) 35.318 cubic feet per second (cfs)<br />

metres per hour (m/hr) 3.281 feet per second (ft/secf<br />

kilometres per hour (km/hr) 0.621 miles per hour (mphl<br />

degrees Celsius (oC) 9/jOC+?2 oegreesFahrenheZt (OF1


CONTENTS<br />

PREFACE ............................................................. i i i<br />

CO?4VERSX01! TABLE.. ..........................................*........ V<br />

FIGURES ............................................................. i x<br />

..............................................................<br />

TABl.ES xiii<br />

ACKNOWLEDC&lENTS ..................................................... x vi<br />

Cbdpter 1 . IN?KODUI:TION ............................................ 1<br />

1.1 Coastal Wctldnds ot the (iredt <strong>Lake</strong>s ..................... 1<br />

1.1 Compari sori <strong>of</strong> b'udstal and Inland Marshes ................ 4<br />

1.3 C~inctron drrll Valur <strong>of</strong> Coastal Wettnntlz .................. 5<br />

1.4 lake <strong>St</strong>. Cia1 r Wpt landr .................... . .. . ..... . 6<br />

Chaptcar Y . I'IIYSIL'AL l NVIRONMENT .................................... 11<br />

7.1 (;ctoIoqy ..................................... ............ 11<br />

2.2 Climat~ and Wpdthpr ..................................... 23<br />

2.3 Hydrology ............................................... 26<br />

2.4 Wdter Quality ........................................... 35<br />

Chapter 3 . BIUTIC ENVIRONMENT ...................................... 43<br />

3.1 Plankton ................................................ 43<br />

3.7 ketland V~yutatron ...................................... 47<br />

3. 3 lnvertrkrates ........................................... 54<br />

3.4 t-lsh ................................. ......*.*.*....... 58<br />

3.5 Aniphthlans and Reptiles ..................... . ....... . 66<br />

3. b Htrds ................................................... 68<br />

3 . 7 Marnrr~als .......................++.....*......,.... 73<br />

C'haytcr 4 . FC0101;ICAL PKI)CErjSI-7 .................................... 78<br />

4.1 Or igi n and f vuiution <strong>of</strong> L.akt. <strong>St</strong> . <strong>Clair</strong> Wet lands ......... 78<br />

4.2 B~olog~ cal Product ion ................... . ............. 85<br />

4.3 Cornrriuri~ ty Procpssc. s ..................................... Yl<br />

4.4 Wctlands as Habitat to Fish dnd W~ldFlfe ................ 101<br />

Chapter 5 . HUMAN iMPACIS ANO APPLIED ECOLOGY ....................... 104<br />

5.1 Wet1cfnrfl)rsturt)anc~ ..................................<br />

104<br />

5.2 Wrt 1 d nd &nrr.~hlp ar~l Managc~ni~nt ........................ 120<br />

5.3 Prospects for tho Future 126<br />

................................<br />

A U~rnens~ons <strong>of</strong> lake <strong>St</strong> . Cla~r and <strong>St</strong> . Llair<br />

1ilvt.r Wet idnd$ by tlnit~d <strong>St</strong>at~s and Canatfa<br />

Vuadrangle Maps ......................................... 141<br />

B Mean Monthly <strong>St</strong> . <strong>Clair</strong> Klvpr Flows 19OO to 1980 ......... 144<br />

C A1 yae Occurrt ng ln Coastal W~tlands dnu Ncarshore<br />

Wdters <strong>of</strong> Lzke S? <strong>Clair</strong> ................................ 146<br />

.<br />

U looplankton Occijrrt ng I n the Loastal Wfhtlands and<br />

Nc2arsnore Wdters <strong>of</strong> <strong>Lake</strong> <strong>St</strong> . Clai r ...................... 152<br />

vii


- Page<br />

Vascular Aquatic Plants Occurring in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

<strong>Wetlands</strong> ............................................ 157<br />

Benthic Macroi nvertebrates <strong>of</strong> Nearshore <strong>Lake</strong> <strong>St</strong>. Cl ai r<br />

and Connecting Waterways .............................. 164<br />

Taxonomic Listing <strong>of</strong> Freshwater Mol lusca <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

Clai r Coastal Marshes, Nearshore Waters, and Tributary<br />

Mouths ................................................ 169<br />

Habitat Preference <strong>of</strong> Freshwater Mollusca <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

Cl ai r Coastal Marshes and Nearshore Waters .............. 173<br />

Fish Known or Presumed to Utilize the Coastal <strong>Wetlands</strong><br />

<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ............... 175<br />

Amphibians and Reptiles Occurring in the Coastal<br />

<strong>Wetlands</strong> <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong> River ...... 178<br />

Birds Occurring in the Vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r and<br />

<strong>St</strong>. Clai r River Coastal <strong>Wetlands</strong> .................... . 180<br />

Mammals Occurring in the Coastal <strong>Wetlands</strong> <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong> Delta ....................... 185<br />

Ownership <strong>of</strong> Major <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Wetlands</strong> .............. 186<br />

viii


Pap <strong>of</strong> the Great <strong>Lake</strong>s drainage bastn .................... 2<br />

Map <strong>of</strong> <strong>St</strong>. Clalr Rl'vor-Lakc <strong>St</strong>, ClaJr-Detroit Rfver<br />

Ecosystem ............................................... 8<br />

Physical factors inf lutmcing tho occurrence <strong>of</strong> coastal<br />

wetlands .............................................. 11<br />

Dupositlanal fuaturcs and 1andfor.m~ <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

Oolta ................................................. 12<br />

Lfttlo Muscornoot Day, an interdlstributary bay botwoon<br />

Middle Channel and South Chdnnol <strong>of</strong> tho <strong>St</strong>. <strong>Clair</strong><br />

r?ulta .................................................. 13<br />

Woll-developed crevasse deposits adjacent to Mlddls<br />

Channel <strong>of</strong> <strong>St</strong>. Clajr Dolta ............................... 14<br />

Deltaic landforms <strong>of</strong> I)lckfnson Island, Mfchfgan .......... 15<br />

Gaoloylc cross-section <strong>of</strong> the northern portfon <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> showlng depth to bedrock. and thjcknoss <strong>of</strong><br />

glacl al, lacustrine, and deltaic deposlts 2 7<br />

................<br />

Cross-sectlon <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta showing distrfbutlon<br />

<strong>of</strong> deltaic sedlmsnts ..................................... 18<br />

Chronology <strong>of</strong> lake levsls and lake stages 3n the<br />

t-iufon and Erie baslns .................................. 1 H<br />

................<br />

Vagetatfan transoct on Uickinsen Island, <strong>St</strong>, <strong>Clair</strong><br />

Oelta, showing character <strong>of</strong> wetland soils %2<br />

Monthly wind frequency dlagrams for Wlndsor, Ontario ..... 26<br />

Monthly wfnd froquoncy for Mt. Clomens, Mlchigan ......... 26<br />

Current patterns for <strong>Lake</strong> <strong>St</strong>. Clafr undor varjous<br />

wlnd condftlons .......................................... 28<br />

iake <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Clafr Rlver drainage basfn ............ ~ ( 7<br />

Mean monthly discharyc. <strong>of</strong> the <strong>St</strong>. Claf r River<br />

:~s-:%a ................................................ 31)<br />

<strong>St</strong>. <strong>Clair</strong> Dolta distributary channels .................... 3 1


Page<br />

Hydrograph <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air and <strong>Lake</strong> Erie water levels<br />

showing the impact <strong>of</strong> a record ice jam in the <strong>St</strong>. <strong>Clair</strong><br />

Delta in March and April 1984 ..........................,e 34<br />

Distribution <strong>of</strong> specific conductance in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.. . . 36<br />

Mean water quality characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ., , . , 38<br />

Characteristics <strong>of</strong> <strong>Lake</strong> Huron water mass and Ontario<br />

nearshore water mass ..................................... 42<br />

Temperature and dissolved oxygen relationship in a <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> coastal wetland ............................... 42<br />

Concentration gradient <strong>of</strong> algal pigments across <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> .............................................. 43<br />

Mean distribution and concentrations <strong>of</strong> zooplankton<br />

groups in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ................................. 45<br />

Distribution <strong>of</strong> total rotifers and most abundant genus,<br />

j3rachlonur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ............................ 46<br />

Ontario shore1 ine <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River at Cathcart Park . . . . 47<br />

Vegetation transects on Harsens Island* <strong>St</strong>. <strong>Clair</strong> Delta . . 48<br />

Distribution <strong>of</strong> six aquatic macrophytes in the <strong>St</strong>. <strong>Clair</strong><br />

River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-Detroit River ecosystem ............. 49<br />

Emergent stand <strong>of</strong> reed grass (Phraamites australis)<br />

along Chematogan Channel, Walpole Is1 and, Ontario . . . . . . 50<br />

Emergent stands <strong>of</strong> narrow-leaved cattail (m<br />

~ t i f oia l on Little Muscamoot Bay* Harsens Island,<br />

Michigan ................................................. 51<br />

Wetland plant communities <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta .... . . , . . 53<br />

Wetland vegetation on Dickinson Island in 1964 ........... 56<br />

Wetland vegetation on Dickinson Island in 1975 .........., 57<br />

Walleye spawning and nursery areas in the Detroit River<br />

and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ....................................... 61<br />

Yellow perch spawning and nursery areas in the Detroit<br />

River and <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> ................................. 62<br />

<strong>Lake</strong> sturgeon spawning and nursery areas in the Detroit<br />

River, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, and <strong>St</strong>. <strong>Clair</strong> River ... . . . . .. . . . . . . 63<br />

Rock bass, pumpkfnseed/bluegill, largemouth bass, and<br />

smallmouth bass spawning and nursery areas in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> ................................................ 6 4


Number<br />

.........................<br />

3 8 Northern piker muskellunge, channel catfish, and carp<br />

spawning areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> 65<br />

Sport fishing areas by specjes in the vicinity <strong>of</strong> <strong>St</strong>.<br />

<strong>Clair</strong> Delta wetlands ....................L.,............ 66<br />

Least bittern (Jkobrvchu~ exllis) with young in broad-<br />

leaved cattail (Tveha latifoliq) nest .................... 73<br />

Food chain <strong>of</strong> the great blue heron (brdea herodlas)<br />

in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................................ 74<br />

Muskrat ffhousesw i n a well populated section <strong>of</strong> Magee<br />

Marsh, western <strong>Lake</strong> Erie, Ohio ........................... 75<br />

Raccoon (procvon lotar), a common predator <strong>of</strong> duck<br />

nests in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetlands ......................... 77<br />

Twelve wet1 and habitats recognized in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ..... 81<br />

Cycling <strong>of</strong> energy and material through the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

coastal wetland ecosystem ................................ 87<br />

Energy pyramid in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal marshes ..,.,.,., 88<br />

Seasonal growth <strong>of</strong> dominant submersed macrophytes in the<br />

<strong>St</strong>. Clai r River-<strong>Lake</strong> <strong>St</strong>. Cl air-Detroit River ecosystem ... 92<br />

Successional changes on Dick i nson Is1 and in response t o<br />

water level fluctuations ................................. 94<br />

Wet1 and pl ant community displacement model for Dickinson<br />

Island ................................................... 94<br />

Fall migration corridors for dabbling ducks (Tribe<br />

Anatini) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................... 98<br />

..........................<br />

Fall migration corridors for diving ducks (Tribe<br />

Aythyini) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> 99<br />

Fall migration corridors for Canada geese (w<br />

ranadensis) across <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ........................ 100<br />

Historic trends in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetland disturbance .... 105<br />

Aerial photograph <strong>of</strong> the mouth <strong>of</strong> the Clinton River*<br />

Macomb County, Michigan .................................. 107<br />

55 Aerial photograph <strong>of</strong> <strong>St</strong>. John's Marsh on Bouvier Eay, and<br />

North Channel, <strong>St</strong>. Cl a1 r Delta (April 1949) .............. 108<br />

56 Aerial photograph <strong>of</strong> North Channel, <strong>St</strong>. <strong>Clair</strong> Delta* and<br />

<strong>St</strong>. John's Marsh on Bouvier Bay (May 1980) ............... 109<br />

57 Commercial development in Marsens Island wetland ......... 11 3


Mercury concentration in sediment core from western <strong>Lake</strong><br />

Erie near mouth <strong>of</strong> Detro'it River ......................... 114<br />

Wetland losses along the Ontario shore1 ine <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> ................................................ 117<br />

Comparfson <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air coastal wetlands dfstrib-<br />

ution in 1873 and 1968 ................................... 118<br />

Location map <strong>of</strong> individual coastal wetlands <strong>of</strong> the <strong>St</strong>.<br />

Cl ai r Rfver-<strong>Lake</strong> <strong>St</strong>. Cl ai r ecosystem ..................... 120<br />

Ownership map <strong>of</strong> the Michigan portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

Delta ............................................... 121<br />

Locatlon map <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> diked coastal wetlands .... 122<br />

Aerial photograph <strong>of</strong> Dickinson Island wetlands, <strong>St</strong>.<br />

Clafr Delta (May 1980) ................................... 124<br />

xii


Number<br />

TABLES<br />

1 Comparison <strong>of</strong> emergent coastal wetlands for the<br />

Laurentian Great <strong>Lake</strong>s ................................... 3<br />

Economic value <strong>of</strong> Michigan's coastal wetlands, 1980 ...... 6<br />

Wild1 i fe use and other functions <strong>of</strong> coastal wet1 ands<br />

at low and high water levels ............................. 7<br />

Estimated dimensions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and <strong>St</strong>. Clafr<br />

River wet1 ands . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8<br />

Interpretation <strong>of</strong> the events related to the origin <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> River Delta . . . . . .. . . . . ., .. . . . . .. . . , ,. . .. .. . 19<br />

Sediment discharge rates and characteristics in the<br />

<strong>St</strong>. <strong>Clair</strong> River .......................................... 21<br />

Mean monthly temperatures for three selected stations .... 24<br />

Mean monthly precipitation for three selected stations .. , 25<br />

<strong>St</strong>. <strong>Clair</strong> River flow time and velocity from <strong>Lake</strong> Huron to<br />

<strong>Lake</strong> <strong>St</strong>. Cl air . . . . . . . . . , . . . . . . . . . . . . . . . . . . . . , . . . . . . . . . . . . 30<br />

Record storm surge levels <strong>of</strong> Canadlan stations on <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and connecting channels . . . . . . . . . . . . . . . . . . . . . . . . 32<br />

Average monthly water transfer factor parameters . . . . . . . . . 35<br />

Water level data for <strong>Lake</strong> <strong>St</strong>. Clai r . . . . . . . . . . . . . . . . . . . . . . 35<br />

<strong>Lake</strong> <strong>St</strong>. Cl air water qua1 ity measurements dur'ing the 1973<br />

growing season ........................................... 37<br />

Chemical and physical characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

and adjacent waters ...................................... 39<br />

Geochemlcal composition <strong>of</strong> sediments In Big Point Marsh,<br />

Ontario on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> , . . . . . . . . . . . . . . . . . . , . . . . . . . . 40<br />

Chemical composition <strong>of</strong> water in Big Point Marsh, Ontario<br />

on <strong>Lake</strong> <strong>St</strong>. Clafr .,.......,......... e L O a I I a I e e * r * + e e a * L e L 40<br />

Comparison <strong>of</strong> chemical characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, C1 af r<br />

and associated waters ......................ee.ss.L).ee*il*e 41<br />

xiii


3 1<br />

3 2<br />

Macrozoobenthos <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r and the lower <strong>St</strong>. <strong>Clair</strong><br />

River and Delta ..........................................<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> unionid bivalves and their glochidial<br />

host fish ..... ....*.............................."e**...e<br />

...............<br />

...............<br />

Average annual sport fishing harvest for Michigan water<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and connecting waterways<br />

Birds observed in <strong>St</strong>. <strong>Clair</strong> Delta wetlands<br />

Productivity and seasonal abundance <strong>of</strong> waterfowl in<br />

Bouvier Bay and Harsens Island wetlands ..................<br />

Species composltfon <strong>of</strong> ducks harvested in the <strong>St</strong>. <strong>Clair</strong><br />

Flats <strong>St</strong>ate Game Area (1974-75) ..........................<br />

Muskrat harvest in Michigan and Ontario bordering <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and connecting waterways .......................<br />

Factors influencing lake and marine delta morphology ..,..<br />

Morphology and re1 ative physical relationships <strong>of</strong> the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands ..................................<br />

Concentrations <strong>of</strong> nutrients and metals in wetland plants<br />

at the stage <strong>of</strong> maximum development, Big Creek Marsh*<br />

Long Point, Ontario .....................................<br />

Biomass and productivity <strong>of</strong> emergent wetl and pl ants<br />

<strong>of</strong> the Great <strong>Lake</strong>s region ................................<br />

Estimates <strong>of</strong> standing crop biomass for submergent<br />

vegetatfon in Anchor Bay, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> .................<br />

Seasonal estimates <strong>of</strong> standing crop biomass for<br />

submersed macrophytes at several locations in the <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> system .........................................<br />

Domlnant submersed aquatic macrophyte taxa <strong>of</strong> the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> system ................................... 92<br />

Vegetation changes on Dickinson Island in response to<br />

lake level fluctuatfons .................................. 95<br />

33 <strong>St</strong>atus <strong>of</strong> wildlife in the vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

and connecting waterways, 1970 ........................... 102<br />

3 4<br />

Land use change at <strong>St</strong>. Johnrs Marsh ...................... 106<br />

3 5 Adverse impacts to the various morphology types <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands ................................. 111<br />

36 Historical dredg'ing totals <strong>of</strong> Corps <strong>of</strong> Engineers<br />

projects in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, 1920-72 ...............a=.*... 112<br />

xiv


37 Metals concentrations in 1 Jvfng plant tissues from<br />

Big Point Marsh. Ontario. on <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> .............. 115<br />

38 Michigan and Ontario wetland losses on <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> .... 116<br />

39 Comparison <strong>of</strong> 1873 and 1968 <strong>Lake</strong> <strong>St</strong> . <strong>Clair</strong> wetland<br />

areas .................................................... 119


In preparing a comprehensive report<br />

such as this pr<strong>of</strong>ile, it is not possible<br />

to mention a11 the individuals who pro-<br />

vided information, gufdance, and stimulus.<br />

Much <strong>of</strong> the credit goes to colleagues at<br />

<strong>The</strong> Ohio <strong>St</strong>ate University and Eastern<br />

Michigan University, including Ronald<br />

<strong>St</strong>uckey, Milton Trautman, Cl arence Taft,<br />

David Johnson, and Loren Putnam. <strong>St</strong>udents<br />

and staff <strong>of</strong> the F. T. <strong>St</strong>one Laboratory at<br />

Put-in-Bay, and the Ohfo Sea Grant Program<br />

who helped In researching various aspects<br />

<strong>of</strong> the marsh scology, are Alan Riemer,<br />

Andrea W i l son, Gary Kfrkpatrick, Mark<br />

Wageman, Laura Fay, and Suzanne Hartley.<br />

Tim Donelly <strong>of</strong> Eastern Michigan University<br />

assisted with data collection.<br />

ACKNOWLEDGMENTS<br />

Ted Ladewski <strong>of</strong> the University <strong>of</strong><br />

Michfgan, <strong>St</strong>an Bolsenga <strong>of</strong> the Great <strong>Lake</strong>s<br />

Environmental Research Laboratory (NOAA) ,<br />

Karl Bednarik <strong>of</strong> the Ohio Division <strong>of</strong><br />

xvi<br />

Wildlife, Susan Crispin <strong>of</strong> the Michigan<br />

Natural Features Inventory, Brooks<br />

Williamson <strong>of</strong> the U.S. Army Corps <strong>of</strong><br />

Engineers, Gary McCullough <strong>of</strong> the Canadian<br />

Wild1 ife Service, Joseph Leach <strong>of</strong> the<br />

Ontario Ministry <strong>of</strong> Natural Resources, and<br />

Bernard Griswol d, Thomas Edsall , Bruce<br />

Panny, and Don Schloesser <strong>of</strong> the Great<br />

<strong>Lake</strong>s Fishery Laboratory, U.S. Fish and<br />

Wildlife Service, all provided useful<br />

informat ion and encouragement.<br />

Wiley Kitchens and Walt Duffy <strong>of</strong> the<br />

National Wet1 ands Research Center provided<br />

guidance and advice throughout the<br />

project. Funding for this project was<br />

provided by the Office <strong>of</strong> the Chief and<br />

the DetroTt District, U.S. Army Corps <strong>of</strong><br />

Engineers. And we especially want to<br />

thank Sandra Herdendorf for typing and<br />

editlng the manuscript. To all those who<br />

he1 ped, we are grateful.


CHAPTER 1. INTCRODeQCTUOM<br />

1.1 COASTAL WETLANDS OF THE GREAT wetland communityr a portion <strong>of</strong> the Great<br />

LAKES <strong>Lake</strong>s - the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Clai r River<br />

system.<br />

In recent years there has been an<br />

increasing awareness <strong>of</strong> the resource value<br />

<strong>of</strong> our coastal wetlands and the urgent<br />

need to protect and conserve these<br />

ecosystems. <strong>The</strong> wetlands <strong>of</strong> the Lauren-<br />

ti an Great <strong>Lake</strong>s have been greatly altered<br />

by natural processes and cultural prac-<br />

tfces, <strong>The</strong> impacts to coastal wetlands in<br />

the Great <strong>Lake</strong>s region has become a<br />

subject <strong>of</strong> particular concern for the<br />

emerging coastal management programs in<br />

the eight states and the one Canadian<br />

province bordering the lakes (Figure 1).<br />

Traditionally, wetl and conservation<br />

efforts along the Great <strong>Lake</strong>s have been<br />

aimed at protecting waterfowl breeding, or<br />

to a lesser degree, fish spawning and<br />

nursery habitat. More recent efforts<br />

toward preservation are based on the<br />

know1 edge that wetl ands provide additional<br />

benefits? including flood control, shore<br />

erosion protection, water management*<br />

control <strong>of</strong> nutrient cycles, accumulation<br />

<strong>of</strong> sediment, and supply <strong>of</strong> detritus for<br />

the aquatic food web. Although the<br />

intrinsic value <strong>of</strong> Great <strong>Lake</strong>s wetland<br />

areas are being more fully recognized, no<br />

comprehensive studies have been undertaken<br />

to characterize the ecological relationships<br />

within them. <strong>The</strong> U.S. Fish and<br />

Wildlife Service (Herdendorf et al,<br />

1981ar bsc) inventoried the existing<br />

l fterature <strong>of</strong> physical, biological, and<br />

cultural aspects <strong>of</strong> the coastal wetl ands<br />

associated with each <strong>of</strong> the Great <strong>Lake</strong>s.<br />

<strong>The</strong>ir study pointed out many gaps in our<br />

knowledge <strong>of</strong> the resources found In Great<br />

<strong>Lake</strong>s wetl ands, partlcul arl y si te-spec1 f ic<br />

fnformation on marshes, This report is<br />

f ntended as a contrfbution toward ff 11 ing<br />

these voids by presenting a pr<strong>of</strong>ile <strong>of</strong> the<br />

For the purposes <strong>of</strong> this report,<br />

wetlands are defined as areas which are<br />

peri odi call y or permanent1 y inundated w 1 th<br />

water and which are typically character-<br />

ized by vegetation that requires saturated<br />

soil for growth and reproduction. This<br />

definition i n cl udes areas that are<br />

commonly referred to as bogs, fens,<br />

marshes, sloughsr swamps, and wet meadows.<br />

<strong>The</strong> coastal wetlands <strong>of</strong> the Great <strong>Lake</strong>s<br />

are further defined as all wetlands<br />

1 ocated within 1 km <strong>of</strong> the lake shore or8<br />

if farther from the shore, those dl rectl y<br />

influenced by water level change <strong>of</strong> the<br />

lakes or their connecting waterways.<br />

Great <strong>Lake</strong>s coastal wetl ands are<br />

highly productive, diverse communities<br />

which interface between terrestrial and<br />

aquatic envf ronments. <strong>The</strong> most obvious<br />

and unique feature <strong>of</strong> these wetlands is<br />

thelr characteristic vegetationr whlch<br />

provides a diverse community structure<br />

<strong>of</strong>fering cover and food for the animal<br />

components <strong>of</strong> the system. Because <strong>of</strong> the<br />

ability <strong>of</strong> this vegetation to siow the<br />

flow rate <strong>of</strong> water passing through?<br />

wetl ands are valuable for erosion control,<br />

trapping sediments before they reach the<br />

open lake# and attenuating the force <strong>of</strong><br />

waves to lessen the4r destructive power,<br />

<strong>The</strong> same vegetation provides a natural<br />

pol 1 ution abatement mechanism by serving<br />

as a filter for coastal trlbutaries<br />

through the reduction <strong>of</strong> the quantity <strong>of</strong><br />

nutrients and toxic pol 1 utants belng<br />

washed into the Great <strong>Lake</strong>s.<br />

Coastal wet1 ands are high1 y valued as<br />

recreational sites for activitfes such as<br />

hunting, trappfngr fishing, boatfng access<br />

to larger bodies <strong>of</strong> water, bdrdwatching?


and general aesthetic enjoyment. <strong>The</strong><br />

combination <strong>of</strong> recreatlonal desi rab ll ity,<br />

agricultural and residential potential,<br />

and the proximity <strong>of</strong> coastal wetlands to<br />

larger bodfes <strong>of</strong> water have contributed to<br />

thei r status as endangered environments.<br />

<strong>The</strong>ir unique properties are susceptible to<br />

numerous natural and human-caused factors<br />

that are now causing coastal wetlands to<br />

disappear at an alarming rate.<br />

<strong>The</strong> Laurentian Great <strong>Lake</strong>s system<br />

within the United <strong>St</strong>ates (Figure 1)<br />

extends from Duluth, Minnesota* at the<br />

western end <strong>of</strong> <strong>Lake</strong> Superior, to Massena,<br />

New York on the <strong>St</strong>. Lawrence River. It<br />

possesses a shoreline length <strong>of</strong> over 6,000<br />

km and a water surface area <strong>of</strong> 158,000<br />

km2. Herdendorf et a1 . (1981a) enumerated<br />

a total <strong>of</strong> lr370 coastal wetlands for the<br />

Great <strong>Lake</strong>s and thei r connect1 ng channel Sr<br />

for a combined wetland area <strong>of</strong> lr209 k d<br />

(Table 1). <strong>The</strong> greatest number and area<br />

<strong>of</strong> coastal wetlands ring <strong>Lake</strong> Michigan,<br />

the only Great <strong>Lake</strong> entirely wfthin the<br />

United <strong>St</strong>ates. <strong>Lake</strong> Super+or has the<br />

second highest number <strong>of</strong> wetlands, but<br />

they are relatively small in size. On the<br />

average* the 1 argest wet1 ands are found<br />

along <strong>Lake</strong> Huron and its discharge channel<br />

(the <strong>St</strong>. Clai r River); for example, the<br />

emergent wetlands <strong>of</strong> the <strong>St</strong>. Clalr Delta<br />

alone cover 36 km2. <strong>The</strong> highly<br />

industrialized <strong>Lake</strong> Erie shore has the<br />

smallest number and area <strong>of</strong> wetlands while<br />

<strong>Lake</strong> Ontario has the smallest average size<br />

<strong>of</strong> wetlands, 1 argely due to is01 ated<br />

marshes In the Thousand Islands area <strong>of</strong><br />

the <strong>St</strong>. Lawrence River. <strong>The</strong> presence or<br />

absence <strong>of</strong> coastal wetl ands is 1 argely<br />

Table 1. Cogparison <strong>of</strong> emergent coastal wetlands for the U.S. Laurentian<br />

Great <strong>Lake</strong>s .<br />

<strong>Lake</strong><br />

Total Mean Percent<br />

Shore Number <strong>of</strong> area <strong>of</strong> area <strong>of</strong> <strong>of</strong> total<br />

1 ength wet1 ands wet1 ands wet1 ands area<br />

<strong>Lake</strong> Superior and 1598 348 267 0.77 22.1<br />

<strong>St</strong>. Marys River<br />

<strong>Lake</strong> Michtgan and 2179 417 490 1.18 40.5<br />

<strong>St</strong>r. <strong>of</strong> Mackinac<br />

<strong>Lake</strong> Huron 832 177 249 1.41 20.6<br />

<strong>St</strong>. <strong>Clair</strong> River, 256 20 36 1.80 3 .O<br />

<strong>Lake</strong> <strong>St</strong>. Cl airr<br />

and Detroit River<br />

<strong>Lake</strong> Erie and 666 96 83 0.86 6.9<br />

Niagara River<br />

<strong>Lake</strong> Ontario and 598 3 12 84 0-27 6.9<br />

<strong>St</strong>. Lawrence<br />

River<br />

TOTAL/MEAN 6129 1370 1209 0.88 100.0<br />

a~ata source: Herdendorf et a?. (1981a1,<br />

3


dictated by the geomorphology <strong>of</strong> a given<br />

shoreline and the recent history <strong>of</strong> water<br />

level fluctuations. Each lake has a<br />

particular set <strong>of</strong> geomorphic features<br />

which exert control on the development <strong>of</strong><br />

coastal wet1 ands.<br />

1.2 COA4PARISON OF COASTAL AND<br />

INLAND MARSHES<br />

<strong>The</strong> coastal wetlands <strong>of</strong> the Great<br />

<strong>Lake</strong>s differ in several ways from inland<br />

wetl ands. <strong>The</strong> coastal wetl cnds are<br />

subject to temporary short-term water<br />

level changes. <strong>Lake</strong> <strong>St</strong>. Claf r wetlands<br />

are subject to inundation and dewaterfng<br />

by seiches, Long-term cycl ic water level<br />

changesr related to water budgets <strong>of</strong> the<br />

lake basins, also affect the coastal<br />

wetlands. Such fluctuationsr occurring<br />

over a period <strong>of</strong> approximately seven to 10<br />

years, may cause vegetation dieback5<br />

erosion <strong>of</strong> the wetlands* or lateral<br />

dtsp? acements <strong>of</strong> the vegetative zones <strong>of</strong><br />

wetlands. Many coastal wetlands, such as<br />

those along western <strong>Lake</strong> Erie, are exposed<br />

to relatively high wave energy.<br />

Coastal wetl ands a1 ong the Great<br />

<strong>Lake</strong>s do hot appear to exhibit the aging<br />

process associated with ini and freshwater<br />

wetlands. Because <strong>of</strong> the fluctuating<br />

water. levels <strong>of</strong> the Great <strong>Lake</strong>s, constant<br />

rejuvenation <strong>of</strong> wet1 and communi t5es<br />

occurs. As a consequencer diagrams 5n<br />

textbooks illustrating the gradual<br />

senescence <strong>of</strong> freshwater wetlands are more<br />

applicable to inland wetlands <strong>of</strong> the<br />

glaclated Midwest than to the Great <strong>Lake</strong>s<br />

coastal wetlands. Many inland freshwater<br />

wetlands undergo senescence and converslon<br />

to terrestrial environments as a result <strong>of</strong><br />

dnorganic and organic depositlon (e.g.,<br />

formation <strong>of</strong> peat deposits).<br />

Coastal wetl ands <strong>of</strong>ten display a<br />

diversity <strong>of</strong> 1 andforms not normal 1 y<br />

encountered in other wetland environments.<br />

Owing to changes in the water levels <strong>of</strong><br />

the Great <strong>Lake</strong>s since the retreat <strong>of</strong> the<br />

Pl e i skocene ice sheetsr 1 and forms such as<br />

coastal barrfersr deltas, and natural<br />

levees have been deposited. <strong>The</strong>se<br />

geamaspholagf ca? features have f n f 1 uenced<br />

the formatf on <strong>of</strong> coastal wetlands. <strong>The</strong><br />

continuing fluctuatiloc <strong>of</strong> water levels f n<br />

the Great <strong>Lake</strong>s 1s also an important<br />

varfable In determining many <strong>of</strong> the<br />

distingulshfng characteristics and the<br />

diversity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />

wetlandsr as well as those <strong>of</strong> the<br />

1 andforms the wetl ands occupy.<br />

Coastal wetlands exhi b f t topograph-<br />

ical pr<strong>of</strong>lles which are more gentle than<br />

what is generally observed in inland<br />

wet1 ands. Many l nl and wet1 ands are formed<br />

in confined glacial features such as<br />

kettle lakesp which exhibit abrupt slopes<br />

on their perimeters. Because <strong>of</strong> the low<br />

angle slope conditionss vegetation zona-<br />

tion Is more dfstlnct and better displayed<br />

In coastal wetlands. As described in this<br />

report, the <strong>St</strong>. Clafr Delta is colonized<br />

by several broad submersed and emersed<br />

wet1 and zones. In1 and wetl ands* howeverr<br />

are more conftned and do not display the<br />

extensive zonation observed in the coastal<br />

zone.<br />

Water levels in coastal wetlands are<br />

determined by climatic factors over<br />

several years. Generallyr annual higher<br />

water levels in the Great <strong>Lake</strong>s occur in<br />

late summer and lower levels occur in<br />

January. Water level changes in inland<br />

wetlands respond to immediate or short<br />

term run<strong>of</strong>f condftions. High water most<br />

<strong>of</strong>ten, occurs in the spring when run<strong>of</strong>f 1s<br />

highest; low water levels occur in early<br />

fall.<br />

Furthermore* the contribution <strong>of</strong><br />

groundwater flow is more significant to<br />

Inland wetlands, During the dry summer<br />

months with high evapo-transpiration<br />

rates, the survival <strong>of</strong> such wetlands may<br />

depend on groundwater i nf 1 ow. Great<br />

<strong>Lake</strong>s' controlled wetlands, however, shift<br />

1 andward or lakeward depending upon water<br />

level conditions over several years.<br />

Clearly, the relationship between<br />

groundwater and wetlands is intimate in<br />

inland areas, but this relationship in the<br />

coastal zone is less significant.<br />

In the Great <strong>Lake</strong>s, with few<br />

exceptions, the substrate is composed <strong>of</strong><br />

mineral sediments. Because <strong>of</strong> the<br />

f 1 ushing action <strong>of</strong> seiches and periodic<br />

water level changes, organic deposits are<br />

not common. Conversely, the hydrolsgic<br />

connect1 vfty <strong>of</strong> in7 and wetl ands is poorer.<br />

Fine mineral soils, with high organic<br />

content f n the inland marshes, contri bute


to thefr gradual senescence and account one may refer to the dollar value <strong>of</strong> a<br />

for more ac-ld-tolerant vegetation than waterfowl hunting dayl as well as to<br />

that found 5n the coastal wetl ands. Peat concepts such as margfnal lty (or scarcfty<br />

fs not typicalfy found along the coast <strong>of</strong> value) and opportunf t y costs.<br />

<strong>Lake</strong> <strong>St</strong>. Cfafr. Nevertheless, not a1 3 wetland functions<br />

(e.g,, flood water storage) are vendable<br />

In the marketplace.<br />

1.3 FU NCTlON AND VALUE OF COASTAL<br />

WETLANDS<br />

Coastal wetlands fn the Great <strong>Lake</strong>s<br />

are mu1 t f -f unct ional because these<br />

environments are part <strong>of</strong> both the up1 ands<br />

and the open-water ecosystems, It is the<br />

fnterface with the Great <strong>Lake</strong>s that<br />

mu1 t ip1 ies the wetland functtons and<br />

contr-ibutes to the "open system" dynamics.<br />

<strong>St</strong>reams and coastal waterways enhance the<br />

ecosystem connectlvf ty, whil a obstacles<br />

such as earthen berms and dikes result in<br />

coastal wetl and f ragmantation and 1 oss <strong>of</strong><br />

function. Functional loss then, can<br />

result from both bank-derlved and lake-<br />

derfved forces. An awareness <strong>of</strong> the<br />

ef f s ct <strong>of</strong> 1 ong-term lake level changes on<br />

the functlon <strong>of</strong> the coastal wetlands is<br />

begfnning to emerge, and the concept <strong>of</strong><br />

pulse stablllty fs being ascribed ta these<br />

envf ronments,<br />

Durlng the past decade, considerable<br />

research was carried out regardlng the<br />

functfon and value <strong>of</strong> wetl ands. Important<br />

general sources Include Tfner (19841,<br />

Greeson et al, (19791, and Messman et al.<br />

(19771, In reference to the Great <strong>Lake</strong>s,<br />

slgnl f icant contr7butlons were made by<br />

Herdendorf et a1 . ( 1981al b,c), Raphael and<br />

Jaworski (19791, Jaworskf t 1981) r Jaworskf<br />

and Raphael (19781, and f f 1 ton et al,<br />

(1978). According to the U, S. Fish and<br />

WfldJ f fe Service Classification (Cowardin<br />

et a1. 1979), most Great <strong>Lake</strong>s wetlands<br />

would be classified as lacustrine or<br />

palustrine,<br />

Wet1 and functions are those processes<br />

occurrf ng fn wetlands whfch are associated<br />

with the functioning <strong>of</strong> the ecosystem or<br />

with the hydrosystem. Examples <strong>of</strong> such<br />

functions are primary production and water<br />

storage, In contrast, wetland values are<br />

those wetland products or servfcss whfch<br />

satfsfy a human need. Comonly, economic<br />

values are employed to such wetland<br />

commodities and services. As a result,<br />

In response to Sectjon 404 and other<br />

permitting authority, the U.S. Army Corps<br />

<strong>of</strong> Engineers (Reppert and Sigleo 1979)<br />

developed the following list <strong>of</strong> wetland<br />

functions and ssrvlces:<br />

1. Natural biological functions<br />

a. net primary productivity<br />

b. food chain (web) support<br />

2. Habitat for aquatic and wetland<br />

speci es<br />

3. Aquatic study areas, sanctuaries<br />

and refuges<br />

4. Hydrologic support functlons;<br />

a. shoreljne protection from wave<br />

attack<br />

b. storage <strong>of</strong> storm and flood waters<br />

c. water purlflcatlon through<br />

natural filtration, sediment<br />

trapping* and nutrient cycling<br />

uptake<br />

d. groundwater recharge<br />

5. Cultural or auxll fary values<br />

fncluding consumptive and noncon-<br />

sumptfve recreation as well as<br />

aesthetic value.<br />

Except for item 5, these wetland benef fts<br />

tend to accrue to the public, and are not<br />

general 1 y vendable by a private 1 andowner.<br />

Quantlta;fve economlc value data for<br />

Great <strong>Lake</strong>s wetlands are avaflable, A<br />

study by Jaworski and Raphael t19781<br />

basicall y addressed the cultural values<br />

using a gross annual economic return<br />

method01 ogy, whereas TIlton et al, (1978)<br />

focused on ecosystem replacement values fn<br />

regard to f fsh productton, waterfowl<br />

feed fng/breedIng, nutrfent removal and<br />

control# and water supply. By combinf ng<br />

the results af these two studfes and<br />

updatfng the data to 1980# a mare complete<br />

account <strong>of</strong> the economfc value <strong>of</strong><br />

Mfchf gan's coastal wetl ands can be<br />

presented (Table 2).


Although methodologies are mlxed,<br />

Table 2 f s useful in establ ishing economic<br />

values within an order <strong>of</strong> magnitude and in<br />

priority. Notf ce that nutrient control J<br />

sport fishing, and fish production are<br />

clearly <strong>of</strong> more value than the traditfonal<br />

uses, i .e. , waterfowl huntingp trappfng*<br />

water supply, and commercial fishing,<br />

Conversely, these data are presented as<br />

average economic values, even though not<br />

all wetlands are a1 ike. Moreover, the<br />

methodologies did not consider surpf us<br />

value* as would be assessed by the<br />

wfllingness to pay technique.<br />

Table 2. Economic vaJue <strong>of</strong> Michigan's<br />

coastal wetlands, 1980.<br />

Economic sector<br />

Run<strong>of</strong>f Nutrient Control<br />

Sport Fishing<br />

Fish Production<br />

Waterfowl Breedbng/Feedi ng<br />

Nonconsumptl ve Recreation<br />

Waterfowl Hunting<br />

Trapping <strong>of</strong> Furbearers<br />

Water Supply<br />

Commercial FishIng<br />

a~ata source: Jaworski ( 1981).<br />

-<br />

(per (per<br />

hectare) acre)<br />

Wetland functions and values vary<br />

from wetl and t o wetland (f .e., spatially)<br />

as well as from year to year (i.e.,<br />

temporally), particularly as a result <strong>of</strong><br />

precipitation and hydrologic changes.<br />

Jaworskf et ale (19791, studied the effect<br />

<strong>of</strong> water level fluctuatfons on seven<br />

wetland complexes in the Great <strong>Lake</strong>s<br />

reg3 on, including Dickinson Island <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> Delta. As fllustrated in Table<br />

3, wild1 ife use and wetland functjons<br />

change as water levels fluctuate, As<br />

discussed in Section 2.3, law water levels<br />

occurred in 1964-65 whereas record high<br />

levels took place during the 1972 to 1974<br />

$nterlrn.<br />

very dense and the water table lies<br />

beneath the mean level <strong>of</strong> the marsh<br />

surface, It Is at such low water<br />

conditf ons that waterfowl management calls<br />

for diking and flooding <strong>of</strong> these emergent<br />

communities. Converselyr during hfgh<br />

water periods the hydrologic clrculat~on<br />

Is facilitated and lake water can enter<br />

the wetland via inlets, canals, and<br />

breaches in barrier beaches. Excessive<br />

high water is associated wlth plant<br />

community retrogression as we1 1 as erosion<br />

<strong>of</strong> the nearshore vegetation and beaches.<br />

<strong>The</strong>refore, the function and value <strong>of</strong> a<br />

coastal wetland is most dynamic, and<br />

short-term studf es <strong>of</strong> wetl and values<br />

should receive careful interpretation.<br />

In summary, a multi-functioning<br />

wetland tends to have a higher value than<br />

those with fewer functions, Converse1 y,<br />

coastal wetlands which are isolated by<br />

barriers or degraded by factors such as<br />

land drainage* exhibit fewer functions and<br />

have 1 ower- values. Moreover, wet1 ands<br />

vary in value from place to place and<br />

temporal 1 y as we1 1 r especi a1 1 y in response<br />

to lake level changes. In addition,<br />

continual wetl and acreage loss results in<br />

a marginality value <strong>of</strong> those extant<br />

wetlands. <strong>The</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />

have exceptional value due to their<br />

ecosystem relationship wfth the lake and<br />

connect1 ng channel s. <strong>The</strong> decline <strong>of</strong><br />

western <strong>Lake</strong> Erte marshes has a1 so<br />

enhanced the re1 ative value <strong>of</strong> the <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> wetlands.<br />

In recognitfon <strong>of</strong> the specfa1<br />

environmental val ue <strong>of</strong> wetl ands, Federal<br />

and <strong>St</strong>ate leglslatlon has recently been<br />

enacted to pratect these areas. In<br />

concert with the National Wetland<br />

Inventory programr each <strong>St</strong>ate in the<br />

Unlted <strong>St</strong>ates has begun the process <strong>of</strong><br />

inventorying and mapping thei r wetl and<br />

resources. Wet1 ands whose hydro1 ogic<br />

regime has been altered or destroyed may<br />

be mitigated.<br />

1.4 LAKE ST. CLAlR WETLANDS<br />

Durlng low water levels* the emergent <strong>The</strong> outlet <strong>of</strong> <strong>Lake</strong> Huron is through<br />

communftles <strong>of</strong> cattails and sedges are the <strong>St</strong>. <strong>Clair</strong> River, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, and


the Detroit River to <strong>Lake</strong> Erie (Figure 21,<br />

a distance <strong>of</strong> 139 km with a fall <strong>of</strong> 2.4 m.<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is an expansive* shallow<br />

basin having a distinctive heart shape.<br />

Although not one <strong>of</strong> the world's largest<br />

lakes, f t does rank 122 on the basis <strong>of</strong><br />

surface area (Herdendorf 1982). <strong>The</strong> lake<br />

has a surface area <strong>of</strong> 1,110 km2 and a<br />

drainage basin <strong>of</strong> 16,900 km2. A 29 km-<br />

long navigation channel, dredged to a<br />

depth <strong>of</strong> 8.2 m below low water datum<br />

( LWD) , extends f rom the mouth <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> River to the head <strong>of</strong> the Detroit<br />

River. <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has a shoreline<br />

length <strong>of</strong> approximately 272 km. <strong>The</strong> shore<br />

<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River, including main<br />

distributary channels, adds 192 km and the<br />

head <strong>of</strong> the Detroit Rfver, plus Belle<br />

Isle, add 32 km <strong>of</strong> shoreline to the study<br />

area. Of this total shoreline length <strong>of</strong><br />

496 km, approximately 188 km or 38% can be<br />

classified as coastal wet1 ands (Table 4<br />

and Appendix A).<br />

<strong>The</strong> delta is the most characteristic<br />

feature <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />

Table 3. Wildlife use and other functions <strong>of</strong> coastal wetlands at low and high<br />

water 1 eve1 s.<br />

Use/Functions <strong>of</strong> wetlands<br />

Mildlffe use<br />

Blue-winged teal (breeding)<br />

Red-w i nged b1 ackb i rd<br />

Ma1 1 ard (breeding)<br />

Sedge wren<br />

Muskrat<br />

Black tern<br />

Ye1 lowheaded b1 ackbird<br />

Great blue heron<br />

Belted kingfisher<br />

Crayfish<br />

Frogs and turtles<br />

Fish spawning (pikes)<br />

Forage fish<br />

Dabbl ing ducks (feeding)<br />

Diving ducks (feeding)<br />

tionq<br />

Peat accumul at1 on<br />

Sediment trapping<br />

Hemi-marsh<br />

Water ci rcul atf on<br />

Dominance <strong>of</strong> l and drainage<br />

Dom,inance <strong>of</strong> lake water masses<br />

Turbidity levels<br />

Export <strong>of</strong> detritus<br />

Re-suspension <strong>of</strong> In slf;ra clay<br />

a~ata source: Jaworski et a1 . (1979).<br />

Water levelb<br />

Low Water High Water<br />

b~se or function relative to water level fndi cated by 1 $nee<br />

7


<strong>Clair</strong>system. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Flats 4s a<br />

massive deposit formed by a series <strong>of</strong><br />

active and inactive distributaries at the<br />

mouth <strong>of</strong> the river. This is the largest<br />

delta in the Great <strong>Lake</strong>s system. <strong>The</strong><br />

Michigan portion <strong>of</strong> the delta has been<br />

urbanized to some extent* but the Ontario<br />

portion has been set aside as an Ind'ian<br />

reservation. <strong>The</strong> Ontario side <strong>of</strong> the<br />

delta, along with <strong>St</strong>ate wildlife areas on<br />

the Michigan side, represent the finest<br />

coastal wetlands in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

system.<br />

<strong>The</strong> shore1 lne <strong>of</strong> the <strong>St</strong>. Cl ai r River<br />

has been intensively developed for<br />

residential, commercial, industrial, and<br />

recreational use. <strong>The</strong> only extensive<br />

natural areas are on the delta wetlands<br />

(<strong>St</strong>. <strong>Clair</strong> Flats), formed where the river<br />

enters <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> vegetation <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> Delta occurs in broad<br />

arcuate zones which extend from the apex<br />

near Algonac, south into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

<strong>The</strong> zones show a progresslon from hardwood<br />

MICHIGAN<br />

Figure 2. Map <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>-Detroit River ecosystem.<br />

forests near Algonac to cattail and<br />

bulrush marshes at the lake.<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has a shallow#<br />

expansive basin with low marshy shores.<br />

Other than the delta, the best coastal<br />

marshes are found on the east shore <strong>of</strong> the<br />

lake between Mitchell Bay and the mouth <strong>of</strong><br />

the Thames River (Ontario) and in Anchor<br />

Bay between the delta and the mouth <strong>of</strong> the<br />

Cl lnton River (Michigan). <strong>The</strong> only other<br />

notable wetlands are submerged beds cf<br />

aquatjc plants in the vicinity <strong>of</strong> Peach<br />

Island and Belle Isle at the head <strong>of</strong> the<br />

Detroit Rf ver.<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, the smallest lake Jn<br />

the Laurentian Great <strong>Lake</strong>s system, has a<br />

mean depth <strong>of</strong> only 3.0 m, a maximum<br />

natural depth <strong>of</strong> 6.4 m, and a volume <strong>of</strong><br />

Table 4. Estimated dimensions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> and <strong>St</strong>. Clail; River emergent and<br />

submergent wetl ands,<br />

Coastal length Area <strong>of</strong><br />

Jurisdiction <strong>of</strong> wet1 ands wet1 ands<br />

(km) (krn2)<br />

MICHIGAN<br />

Wayne County 4.0 0.7<br />

Macomb County 6.6 11.2<br />

<strong>St</strong>. <strong>Clair</strong> County 75.2 140.7<br />

Michigan Total 85.8 152,6<br />

ONTARIO<br />

Essex County 11.5 6.8<br />

Kent County 26.5 62.8<br />

~arnbton County 64.4 161.7<br />

Ontario Total 102.4 231.3<br />

TOTAL 188.2 383.9<br />

a~ata sources: United <strong>St</strong>ates Geological<br />

Survey, Dept. Interior, 7.5-minute Quad-<br />

rangle Maps (1:24,000 scale) ; Canada<br />

Department <strong>of</strong> Energy, Mines and Resources,<br />

7.5-minute Quadrangle Maps (1:25,000<br />

scale); Direct fie1 d and aircraft obser-<br />

vations, 1983 and 1984.


4,O km3. Its deepest waters are found<br />

along a dredged navigation channel which<br />

extends from the mouth <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

River to the head <strong>of</strong> the Detroit Rfver.<br />

Approximately 98% <strong>of</strong> the water flowing<br />

into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> enters through the <strong>St</strong>.<br />

<strong>Clair</strong> Delta. <strong>The</strong> Detroit River, the on1 y<br />

outlet, drains <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> at a rate <strong>of</strong><br />

5,044 m3/sec. This yields a retention<br />

time for the lake basin <strong>of</strong> only 9.2 days.<br />

<strong>The</strong> <strong>St</strong>. Clzir River is one <strong>of</strong> the<br />

four major connecting channels on the<br />

Great <strong>Lake</strong>s, fl owl ng between <strong>Lake</strong> Huron<br />

and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> two principal<br />

cities on the river are Port Huron,<br />

Michigan, and Sarnia, Ontario, both at the<br />

head <strong>of</strong> the river at <strong>Lake</strong> Huron. <strong>The</strong> <strong>St</strong>.<br />

<strong>Clair</strong> River has two characteristic<br />

sections, the upper channel, and the<br />

lower, or delta, portion. <strong>The</strong> upper<br />

channel runs from <strong>Lake</strong> Huron to the head<br />

<strong>of</strong> Chenal Ecarte, a distance <strong>of</strong> 43 km.<br />

<strong>The</strong>re are two islands in the upper portion<br />

<strong>of</strong> the river, <strong>St</strong>ag Island and Fawn Island.<br />

At Chenal Ecarte, the river begins to<br />

branch out i n t o a number o f<br />

distributaries, forming a vast pattern <strong>of</strong><br />

islands, marshes, and waterways that are<br />

known as the <strong>St</strong>. Clalr Flats.<br />

<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River receives most <strong>of</strong><br />

Its flow from <strong>Lake</strong> Huron. Prfncipal<br />

tributaries in Michigan are the Belle,<br />

Black, and Pine Rivars. Water quality<br />

throughout the <strong>St</strong>. <strong>Clair</strong> River is<br />

genera1 1 y excel lent. Some degradation<br />

occurs i n localized areas where<br />

tributaries join the river. <strong>The</strong> width <strong>of</strong><br />

the <strong>St</strong>. Cl air River ranges from about 250<br />

to 750 m, and the maximum depth ranges<br />

between 11 and 15 m. <strong>The</strong> river's annual<br />

average discharge is about 5,070 m2/sec.<br />

Because the river's main water source is<br />

<strong>Lake</strong> Huron rather than surface run<strong>of</strong>f9<br />

seasonal f 1 ow variations are minimal .<br />

Herdendorf et al. (1981~1 1 isted four<br />

wetland complexes that are located in the<br />

lower portions <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River.<br />

<strong>The</strong> Pointe Aux Tremble Wetland (15<br />

hectares), the Pointe Aux Chenes Wetland<br />

(7 hectares), and the Russell Island<br />

Wetland (5 hectares) are all located in<br />

the North Channel <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River.<br />

<strong>The</strong> Algonac Wetland (64 hectares) is<br />

1 ocated 0.3 km f rom the shore1 f ne <strong>of</strong> the<br />

river. <strong>The</strong> Cuttle Creek Area Wetland (1,s<br />

hectares) and the Port Huron Area Wetland<br />

(1.2 hectares) are both located along the<br />

upper portion <strong>of</strong> the river.<br />

A1 though the 1 argest area <strong>of</strong> wetl ands<br />

1s found i n the <strong>St</strong>. <strong>Clair</strong> Delta, there are<br />

fragments <strong>of</strong> wetl ands around the perimeter<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Beginning at the mouth<br />

<strong>of</strong> the Detroit River, small patches <strong>of</strong><br />

rfveri ne, pal ustri ne, and 1 acustri ne open-<br />

water wetlands exist along Belle Isle,<br />

Aquatic beds <strong>of</strong> submersed vascul ar pl ants<br />

dominate as a park and other developments<br />

havs resulted in the removal <strong>of</strong> the upper<br />

communities.<br />

Along the southwestern shore <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>9 from Grosse Pointe to the<br />

Clinton River* the shoreline is inten-<br />

sively developed. Much <strong>of</strong> the shoreline<br />

is protected by seawalls, and pjers and<br />

marinas are scattered a1 ong th i s stretch.<br />

Although the shoreline is classified as<br />

1 acustrine open-water wet1 ands9 there is<br />

l i ttle wetland vegetation except for<br />

pollution-tolerant submersed aquatics such<br />

as coontail and Eurasian water-mil f<strong>of</strong>l<br />

growing in the boat channels and slips<br />

(see Appendix E for common and scientiftc<br />

names <strong>of</strong> vascular plants).<br />

Where the Clinton River empties Into<br />

<strong>Lake</strong> <strong>St</strong>. Cl airr along the western shore <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>9 there are two coastal<br />

wetland areas. South <strong>of</strong> the riverr in the<br />

Metropolitan Beach area, over 4 hectares<br />

<strong>of</strong> palustrine emergent and pal ustrine<br />

open-water wetlands exist. North <strong>of</strong> the<br />

Metropol itan Parkway highway, i .e, , along<br />

Black Creek, which is an old tributary <strong>of</strong><br />

the Cl inton River, there are sedge meadow*<br />

cattaf 1 and open-water (aquae lc bed 1<br />

communities. Because the connectyon wfth<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is poor, the water<br />

frequently stagnates resulting in lower<br />

fish and benthic invertebrate values,<br />

North <strong>of</strong> the Clinton River, located<br />

between Sand Point and Self ridge Air Force<br />

Basep is an open wetland. This wetland 7s<br />

largely a palustrine system with submersed<br />

aquatic plant communities.<br />

<strong>The</strong> shoreline <strong>of</strong> Anchor bay^ sftuated<br />

in northwest <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r is a150


largely urbanized. Residential develop-<br />

ment, marinasr and boat sljps are common<br />

in the mouths <strong>of</strong> streams as we11 as in<br />

lakeshore points. Very 1 ittle undisturbed<br />

wetland exists and much <strong>of</strong> the shorel ine<br />

is elther bulkheaded or piled with rip-rap<br />

to reduce erosion assocjated with the<br />

current high water levels. Although the<br />

entire shoreline i s classifled as<br />

lacustrine open-water wetl ands, on1 y<br />

scattered patches <strong>of</strong> emergent cattails and<br />

submersed aquatics exist along the creek<br />

mouths and behf nd bulkheads. Clumps <strong>of</strong><br />

bulrushes occur along the shorel ine9 along<br />

with strips <strong>of</strong> wild celery and other wave<br />

tolerant submersed aquatics.<br />

<strong>The</strong> first large area <strong>of</strong> deltaic<br />

wetland occurs along Bouvier Bay, and Is<br />

locallv referred to as <strong>St</strong>. Johnqs Marsh.<br />

~~~roximatel~ 1,000 hectares in sJze, this<br />

traditional 1 y p? anted in corn and other<br />

artificial waterfowl foods. Most <strong>of</strong><br />

Dickinson Island and the lower half <strong>of</strong><br />

Harsens Island together make up the <strong>St</strong>.<br />

Clafr Flats Wildlife Area, which is<br />

managed by the Michigan Department <strong>of</strong><br />

Natural Resources for waterfowl hunting.<br />

With reference to the Ontario side <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> Delta, that wetland complex<br />

i n cl udes port ions <strong>of</strong> Bassett Is1 andt<br />

Squirrel Island, Walpale Island# and <strong>St</strong>.<br />

Anne Is1 and. Bassett Is1 and consi sts<br />

mostly <strong>of</strong> cattail marsh with open-water<br />

ponds and a thin beach colonized by shrubs<br />

and herbaceous pl ants. Squi rrel Is1 and is<br />

also largely a natural cattail marsh* but<br />

the northern part, where sedge marsh would<br />

naturally occur, i s diked <strong>of</strong>f and<br />

cu1 tivated.<br />

Canada's major wetland complex in the<br />

is found both 'Ides delta focuses on Walpole Island where<br />

Of M-29? and extends from Fair<br />

Goose <strong>Lake</strong> and Johnston Bay are located.<br />

Haven to Pearl Beach. Plant communities<br />

Deciduous woodlands, shrubs, and sedge<br />

in this are by meadows form the upper part <strong>of</strong> this<br />

and sedge<br />

meadows cattaf1s8 bul and<br />

leaved and submersed aquat icr.<br />

CDnstructi on Of access roads and<br />

f'll jng Iror residential as<br />

"11 as the Of highway M-29 have<br />

fragmented this wetl and.<br />

island* but much <strong>of</strong> the natural shrub and<br />

meadow zones have been cleared and owed<br />

into row-crop agrlculture, A diked and<br />

water-level regulated cattail marsh occurs<br />

north <strong>of</strong> Goose <strong>Lake</strong>. South <strong>of</strong> Goose <strong>Lake</strong><br />

the wetlands are dominated by cattails and<br />

sedqe communities along the distributary<br />

~ i ~ 1 ~ and k1 js i the ~ 1 argest ~ ~<br />

chaknels<br />

~<br />

and shorel ine beaches. <strong>The</strong><br />

natural undeveloped, and functioning <strong>Wetlands</strong> near Johnston are open to<br />

wetland complex along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>Lake</strong><strong>St</strong>*<strong>Clair</strong>*<br />

Approximately lr200 hectares, this wetland<br />

has a continuum <strong>of</strong> environments. To the<br />

north, where elevations average 1.5 to 3 m<br />

above North Channel* swamp forest and<br />

dogwood shrub communities occur. In the<br />

mfddle <strong>of</strong> this deltaic island, several<br />

abandoned river channels bisect vast<br />

stretches <strong>of</strong> sedge meadow and cattail<br />

communit f es. Along the shore1 ine, the<br />

emergent wetlands give way to various<br />

<strong>The</strong> wetlands on <strong>St</strong>. Anne Island<br />

appear much more managed. Large corn<br />

fields on the upper and middle portions <strong>of</strong><br />

the island extend to the limit <strong>of</strong> the<br />

cattail marsh. At this point* the marshes<br />

are diked for the benefit <strong>of</strong> private<br />

waterfowl clubs. On1 y the shorel f ne<br />

fringe <strong>of</strong> the wetlands is open to <strong>Lake</strong> <strong>St</strong>.<br />

C1 air.<br />

open-water aquatic bed-type wetlands,<br />

<strong>The</strong> last major wetland area along the<br />

Ontario side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr Is in the<br />

Another major wetland complex along Mitchell Bay area. Because <strong>of</strong> the<br />

the American side <strong>of</strong> the <strong>St</strong>. Clai r Delta proximity <strong>of</strong> the managed marshes on <strong>St</strong>.<br />

is Harssns Island. Only the lower quarter Anne Island and the cultivation by<br />

<strong>of</strong> thls deltaic island remains a wetland, Canadian farmers rtght up to the lakeshore<br />

consisting largely <strong>of</strong> hybrid cattail<br />

(Typha x ~lauca) colonies along with the<br />

assocfated floating-1 eaved and submersed<br />

canal and shore1 Sne communities. <strong>The</strong><br />

center <strong>of</strong> Harsens Island is part <strong>of</strong> the<br />

<strong>St</strong>. Clafr Flats Wildlife Area and is<br />

in Kent and Lambton counties, the wetlands<br />

<strong>of</strong> Mitchell Bay consist mainly <strong>of</strong> openwater<br />

ar submerged cammunlties. Moreover,<br />

considerable diking and waterfowl nesttng<br />

improvements have taken place along<br />

Mitchell Point.


This section contains a description<br />

<strong>of</strong> the geographic location <strong>of</strong> the study<br />

areao including a discussion <strong>of</strong> topog-<br />

raphy, morphometry, and coastal geomor-<br />

phology, Although no bedrock or glacial<br />

features are directly related to the<br />

distribution <strong>of</strong> wetlands there are bio-<br />

geographical associations with more recent<br />

landforms such as natural levees, crevasse<br />

deposits, interdistributary bays, and<br />

other del talc deposits.<br />

<strong>The</strong> occurrence and distribution <strong>of</strong><br />

wetlands is determined, to a significant<br />

degree, by a combination <strong>of</strong> physical<br />

factors. In the Great <strong>Lake</strong>s, coastal wet-<br />

lands are at the interface <strong>of</strong> land, water,<br />

and atmosphere. <strong>The</strong>se three components <strong>of</strong><br />

the biosphere are delicately balanced to<br />

produce and maintain the biologi.ca1<br />

resource. In this chapter the physical<br />

parameters <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />

are exam1 ned.<br />

<strong>Wetlands</strong> occur in basins or topo-<br />

graphical depressions whlch are wet or<br />

flooded for at least several months <strong>of</strong> the<br />

year. <strong>The</strong> occurrence <strong>of</strong> the resource is<br />

therefore in part determined by water<br />

depth, which is in turn determined by the<br />

submarine topography, sed imentationr and<br />

character <strong>of</strong> 1 ake level oscillations.<br />

Another factor responsible for wetl and<br />

distribution is wave action in open water<br />

bodies and current characteristics in<br />

riverine and 1 ittoral areas. Many <strong>of</strong><br />

these factors are in turn directly related<br />

to climatic conditions. A simplified<br />

diagram illustrating these relationshjps<br />

appears in Figure 3.<br />

<strong>The</strong> more extensive wetlands <strong>of</strong> the<br />

Great <strong>Lake</strong>s are located in ancient lake<br />

bottoms or in areas <strong>of</strong> h5gh sedi mentation<br />

CHAPTER 2. PHYSICAL EIUIVOIRBWMENT<br />

I CLIMATE I<br />

1<br />

BOTTOM TOPOGRAPHY<br />

Figure 3. Physical factors influencing<br />

the occurrence <strong>of</strong> coastal wetl ands.<br />

rates. In the geological past, water<br />

levels in the Great <strong>Lake</strong>s were higher and<br />

progl aci a1 1 akes inundated the lower<br />

terrain, As the water levels <strong>of</strong> the lakes<br />

droppedr a series <strong>of</strong> clay lake plains were<br />

exposed which were colonized by vast<br />

wetl ands. Western <strong>Lake</strong> Erie, includfng<br />

the Maumee River embaymentp for examplep<br />

was an extensive wetland known as the<br />

Black Swamp. <strong>The</strong> margins <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />

Clai r, especially in Ontarf or are composed<br />

<strong>of</strong> a similar deposit.<br />

<strong>The</strong> rivers flowing f nto take <strong>St</strong>.<br />

Clai r deposited sediments a1 ong valley<br />

floors to create floodplains which were<br />

colonized by wetland communities. At the<br />

same time deposition in the lake created<br />

the <strong>St</strong>. Clafr Delta. Although most <strong>of</strong><br />

Michigan has a glacfal herttage, coastal<br />

wetlands are a product <strong>of</strong> fluvial,<br />

deltaic? and lacustrine sedimentation.<br />

<strong>The</strong> <strong>St</strong>, Clai r DeTta has been depos-<br />

ited* in past because <strong>of</strong> the shallowness<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Compared to marfne<br />

shore1 inesp ongolng coastal deposit ion<br />

which produces subaeri a1 l andforms such as<br />

the delta is not common in the Great


<strong>Lake</strong>s. In fact, no other signff icant<br />

deltas occur in the Great <strong>Lake</strong>s, <strong>The</strong><br />

combination <strong>of</strong> a shallow receivfng basfn<br />

and an abundance <strong>of</strong> sediments fs largely<br />

responsible for the deposttion <strong>of</strong> the <strong>St</strong>.<br />

Clafr Delta which fs colonized by the most<br />

extensive wetlands in the coastal Great<br />

<strong>Lake</strong>s.<br />

<strong>The</strong> progradatton or recession <strong>of</strong> the<br />

coastal zone is 1 argely influenced by wave<br />

and currents. High energy environments<br />

discourage the establ i shment <strong>of</strong> pal ustri ne<br />

and lacustrine wetlands. Also high wave<br />

energies hinder delta deposition and<br />

growth. In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wave and<br />

current energy are not excessive. Thls<br />

physical condition has allowed the<br />

development <strong>of</strong> the delta with its diverse<br />

wetland habjtats. Also the low wave<br />

energies have encouraged wetland coloniza-<br />

tion north <strong>of</strong> the Thames River along the<br />

eastern shore1 ine <strong>of</strong> the 1 ake.<br />

<strong>The</strong> climatic elements Including<br />

temperature, precfpi tation* evaporation,<br />

and wfnd velocities and dire~tions exert a<br />

control on water levels and wave<br />

parameters, <strong>The</strong>se in turn dictate wetland<br />

occurrence and types. With changes In<br />

water levels the geographfcal di stributlon<br />

<strong>of</strong> wetlands shfft or "pulse stabi71tyw<br />

occurs. Also the character <strong>of</strong> sedimentation<br />

is altered to a point where water<br />

quality (i.aer turbidity) levels are<br />

impacted.<br />

It has been determfned that wetlands<br />

in the Great <strong>Lake</strong>s colonize a diversity <strong>of</strong><br />

geomorphic settings such as backbarrfer<br />

flats, alluvial flood plains and deltas.<br />

<strong>The</strong> <strong>St</strong>. C1 air Delta (Figure 4) is rnade up<br />

<strong>of</strong> a suite <strong>of</strong> landforms which support a<br />

diversity <strong>of</strong> wet1 and types, Deltaic<br />

deposits in the Great <strong>Lake</strong>s are rare<br />

features, suggesting that fluvial sedirnen-<br />

tation is not abundant and/or wave<br />

energies are too high thus not permfttlng<br />

delta development to occur. <strong>The</strong>refore the<br />

<strong>St</strong>. Cl ai r Delta Is a unique Great <strong>Lake</strong>s'<br />

Figure 4. Depositional features and landforms <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta.<br />

12


coastal feature supporting abundant and<br />

diverse wet1 and communities not found on<br />

other Great <strong>Lake</strong>s' shore7 i nes.<br />

Since the classical study <strong>of</strong> <strong>Lake</strong><br />

Bonneville by Gilbert (18901, deltas in<br />

lakes* with only a few exceptions, have<br />

been largely ignored by geomorphologists.<br />

<strong>The</strong> major deltas <strong>of</strong> the world are located<br />

on contlnental shelves and have held the<br />

interest <strong>of</strong> geologists and geographers<br />

because <strong>of</strong> petroleum resources, agricul-<br />

tural production, and waterborne activi-<br />

ties. A1 though 1 ake deltas are smaller,<br />

they share most <strong>of</strong> the processes<br />

characteristic <strong>of</strong> marine deltas. However,<br />

a unique aspect <strong>of</strong> lake deltas is their<br />

dynamic response to re1 atively rapid<br />

changes <strong>of</strong> base level that not only<br />

produce special 1 andforms, but dl sti nctive<br />

vegetation zonation as we1 1.<br />

<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta is the largest<br />

delta in the Great <strong>Lake</strong>s basfn. Despite<br />

the delta's recreational sign1 f icance and<br />

commercl a1 importance with regard to<br />

navigation, literature on the area has not<br />

been abundant, <strong>The</strong> first significant<br />

investigation was done several decades ago<br />

by Cole (19031 who determined that the<br />

delta was beina deposited atop deep water,<br />

proglacial 1 &e clays. p ore recent1 yr<br />

Wfghtman (1961) attempted to establish a<br />

1 ate Quaternary chronology for the del tats<br />

formation. Detailed investigations by<br />

geol og 5 sts have part1 ally documented the<br />

sediment01 ogical composition <strong>of</strong> the<br />

deltaqs surface, parti cularly in Muscamoot<br />

and Goose bays (Pezzetta 1968, Mandelbaum<br />

1969). During the 1950~8 engineering<br />

studies by the U.S. Army Corps <strong>of</strong><br />

Engineers preceded construction <strong>of</strong> the 8-m<br />

deep <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel through the<br />

delta. Although not published, much <strong>of</strong><br />

these data, in the form <strong>of</strong> bore records*<br />

are on file at the Detroit District<br />

Office. Recent environmental studies<br />

associated with flooding problems on the<br />

shores <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and dredged spoil<br />

disposal s<strong>St</strong>e locations have also<br />

contributed some useful data for this<br />

outlets <strong>of</strong> the Great <strong>Lake</strong>s were changing<br />

or even blocked, causing the lake levels<br />

to oscillate several meters. <strong>Lake</strong> Erie<br />

established its present level some 4,000<br />

years ago and <strong>Lake</strong> Huron shortly<br />

thereafter. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River and its<br />

delta came to existence during these<br />

changing lake levels.<br />

As a lake delta, the <strong>St</strong>. <strong>Clair</strong><br />

exhibits several <strong>of</strong> the landform<br />

characteristics <strong>of</strong> marine deltas* such as<br />

active and inactive distributaries,<br />

interdfstrfbutary bays (Figure 5)s and<br />

crevasses or wlde breaches in the channel<br />

banks which lead into interdistributary<br />

bays (Figure 6). <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta has<br />

a classical bi rd-f oot morphology , as does<br />

the Mississippi River Delta. However,<br />

significant landform differences are also]<br />

apparent. Atypical landforms include a<br />

premodern surface at the apex <strong>of</strong> the delta<br />

and unusual ly wide distributary channels.<br />

section <strong>of</strong> the study (U.S. Army Corps <strong>of</strong><br />

Engineers 19741,<br />

Wfth tke retreat <strong>of</strong> the Late<br />

Figure 5. Little Muscamoot Bay, an interdistri<br />

butary bay between Middle Channel<br />

and South Channel <strong>of</strong> the <strong>St</strong>. Cfair 3e7ta<br />

W i sconsin ice sheet some 13,000 years ago, (August 1984). Note emergent cattails<br />

a series <strong>of</strong> moraines and till plains were (Typha an ustifolia) and solitary<br />

deposited in the Great <strong>Lake</strong>s basln. <strong>The</strong> pied-bili&~ddilymbus podiceps).


Figure 6. We1 1-developed crevasse deposits adjacent to Middle Channel <strong>of</strong> <strong>St</strong>. <strong>Clair</strong><br />

Delta. <strong>The</strong>se crevasses have been intermittently active for over 100 years. Dickinson<br />

Island is to the upper left and Harsens Island is to the lower right.<br />

<strong>The</strong> active dJstributaries--Norths<br />

Middlet and South C,hannel s--average some<br />

500 m in width and 11 m in depth.<br />

Howevert widths <strong>of</strong> 675 m and depths <strong>of</strong> 26<br />

m are not uncommon. At the mouths <strong>of</strong> the<br />

dfstrfbutarfes, channel depths decrease<br />

abruptly Indicating the presence <strong>of</strong> river<br />

mouth bars 2 to 4 m below mean lake level.<br />

As a deposftional basin, <strong>Lake</strong> <strong>St</strong>. Clafr fs<br />

relatfvely small with a maximum natural<br />

depth <strong>of</strong> 6.4 m and width <strong>of</strong> 40 km.<br />

North, Middle* and South Channels<br />

exhibit shoulder-1 i ke features or shoals<br />

along both the cutbank and point-bar sides<br />

(Ffgure 71, A sfmjlar morphology has been<br />

attrfbutsd to periodic cut and ffll<br />

associated wfth slfght base level<br />

oscillations (Butzer 1971). Borings<br />

obtained from the Corps <strong>of</strong> Engineers<br />

reveal that the distributary channels <strong>of</strong><br />

the <strong>St</strong>. ClaSr Delta are entrenched in<br />

laeustrine c1 ays which 1 ie beneath a 3.5<br />

to 5 m veneer <strong>of</strong> coarser deltaic<br />

sediments. Qn the cutbank side, where<br />

flow velocftfes may be relatlvely high<br />

during high water condltlons, an erosfonal<br />

berm usually less than 2.5 m below the<br />

water surface slopes gently from the<br />

cutbank toward the channel center, <strong>The</strong>se<br />

features may be caused by lateral erosion<br />

<strong>of</strong> the fine, sandy deltaic sedtments which<br />

over1 ie the 1 acustrfne clays. On the<br />

inside bank, especially along Middle<br />

Channel , point bar deposits characterized<br />

by ridge and swale topography are<br />

conspicuous. Here the distributary<br />

shoulders probably represent a fill<br />

deposit which is colonized by emergent<br />

vegetati on during 1 ow-water periods.<br />

Because the water level fluctuates<br />

only 45 to 60 cm seasonally, spring floods<br />

are not a normal occurrence within the<br />

delta, hence natural levees are scarcely<br />

dfscernable adjacent to modern distrib-<br />

utaries. Even though levees are poorly<br />

developeds averaging 10 to 45 cm in<br />

elevationr flooding and subsequent<br />

overbank flow does occur. Overbank flow<br />

is assocfated with breaching <strong>of</strong> levees in<br />

abnormally low areas along a levee. As a


.................<br />

...............<br />

MODERN SURFACE<br />

SUBMERGED RIVER<br />

RELICT CHANNEL<br />

Figure 7. Deltaic landforms <strong>of</strong> Dickinson Island, Michigan,


levee fs breachedr crevasse deposits are<br />

Introduced into the interdistributary bays<br />

at right angles to the channels (Figure<br />

71. With contf nued depositionp the open-<br />

water bay w i l l be filled with crevasse<br />

deposits and colon 1 zed by sedges and<br />

emergent aquatics.<br />

Crevasse channel s9 1 ocal l y known as<br />

"highwaysw (see Figure 57 in Section 5,1Ir<br />

are operative for several years.<br />

Nevertheless, deposition into the bays i5<br />

not rapid, A comparison <strong>of</strong> navigation<br />

maps reveals zhat such features may be<br />

part <strong>of</strong> the delta landscape for over a<br />

century. Pezzetta 11968) graphical1 y<br />

compared the western portton <strong>of</strong> the delta<br />

front between 1903 and 1961. Over the 58-<br />

year perjod crevasse fills were minor. In<br />

fact human-mads a1 terati ons such as<br />

dredging and bulkheadlng dominated the<br />

1 andscape. This suggests that crevasse<br />

channels are active intermittently and<br />

transport l i t tle sediment into the<br />

interdistributary bays.<br />

During the winter and early spring<br />

when <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is frozen, pack ice<br />

accumulates at the mouths <strong>of</strong> distributarjes<br />

forming ice jams. Channel flow is<br />

then dfverted lnto crevasses and some<br />

overbank flow may occur, Because the<br />

dominant grain size is sand, 1 ittle bed<br />

load sedlment is transported fr~m the deep<br />

distributaries Into the interdistributary<br />

bays. Thus, In the <strong>St</strong>, <strong>Clair</strong> Delta the<br />

filling <strong>of</strong> interdistrSbutary bays and<br />

delta growth Is a slow process.<br />

In contrast, the Mississippi Delta<br />

crevasse depos f ts rapidly convert open<br />

interdistributary bays into mud flats<br />

which are subsequently colonized with<br />

marsh grasses. During flood stage, the<br />

crevasse channels are scoured deep enough<br />

to be maintained and rapid depositlon<br />

occurs, In the past 135 years, crevasse<br />

deposf t s have transformed the open<br />

interdistributary bays <strong>of</strong> the modern<br />

Mississippi Rfver Delta into a complex <strong>of</strong><br />

marshy subdeltas (Coleman 1976).<br />

Beaches on the present delta<br />

shorel ine are poorly developed and appear<br />

transgress'ite in orf~fn, On t he Canadian<br />

side, where the beaches are somewhat<br />

better developed* the berms may reach I t o<br />

1.5 m In height and are colonfzed with<br />

sumac and small trees, Borings reveal<br />

that the principal constituents <strong>of</strong> these<br />

beaches are coarse sand or f lne gravel<br />

(0.3 m or -L.5 phi unJts) separated by<br />

layers <strong>of</strong> sand and organic materials<br />

including rafted logs, bulrush stems, and<br />

other debris. Coarse sands and fine<br />

gravels are not evident, and storm berms<br />

seldom exceed 0.7 m In elevation.<br />

Characteristic vegetation <strong>of</strong> these<br />

shorel ine features are either sedge marsh<br />

or a complex community <strong>of</strong> grasses,<br />

thistles8 and other nonwoody species.<br />

Within the interd9stributary marshes,<br />

especf a1 1 y on lower DiC~inSon and Harsens<br />

is1 ands, are arcuate-shaped features<br />

resembling beach ridges. Borings through<br />

one <strong>of</strong> these ridges revealed up to 3 m <strong>of</strong><br />

flne sand (Raphael and Jaworski 1982).<br />

<strong>The</strong> absence <strong>of</strong> washover deposits and<br />

organlc sediments indicate that these<br />

features may be regressive beaches and<br />

represent shorelines as delta accretion<br />

took place.<br />

A comparison between the eastern and<br />

western portions <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta<br />

illustrates two other distinct dif-<br />

ferences. On the CanadSan side, Chenel<br />

Ecarte and Johnson Channel are narrow,<br />

shall ow distributaries which do not carry<br />

a significant portion <strong>of</strong> the volume <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> River. Moreover9 open Inter-<br />

distributary bays are few and are<br />

colonized by marsh vegetation. Delta<br />

extension has ceased and maximum delta<br />

accretion ls now occurring to the west as<br />

evidenced by the active dlgital<br />

dtstributaries <strong>of</strong> North, Middle, and South<br />

Channels, and Chenal A Bout Rond. In the<br />

past, Chematogan and Bassett Channels were<br />

approximately 500 m wide comparable to the<br />

modern distributaries, but have been<br />

alluviated and colonized with aquatfc<br />

plants as abandonment occurred.<br />

In most deltas with large fluvial<br />

systems, lateral migration <strong>of</strong> the delta<br />

occurs because <strong>of</strong> the changes in the<br />

course <strong>of</strong> the river upvalley. <strong>The</strong><br />

premodern 1 ateral displ acements <strong>of</strong> the<br />

Mississippi River Delta originated within<br />

the alluvial valley several miles from the<br />

Gulf <strong>of</strong> Kexico% coast. Such a dlversfon<br />

process is evident in some lake deltas as<br />

well. Several older deltas <strong>of</strong> the Omo<br />

Rlverr for example, have been identi f fed


and related to the former positlon <strong>of</strong> the<br />

rfver within its alluvial valley (Butzer<br />

1971).<br />

<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta# however, has no<br />

comparable a1 luvial uall ey, and de1 ta<br />

migration has occurred from east to west<br />

at the shoreline <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>. As<br />

the Canadian distributaries degeneratedr<br />

new distributaries were created on the<br />

western side <strong>of</strong> the delta.<br />

A series <strong>of</strong> borings, cores, and<br />

exposures indicates that i n cross sect i on<br />

the <strong>St</strong>. <strong>Clair</strong> Delta is a thin and sandy<br />

deposit. An east-west cross section<br />

reveals that above the shale bedrock,<br />

1 acustrine c1 ays have been deposited over<br />

a thin deposit <strong>of</strong> glacial till (Figure 8).<br />

<strong>The</strong> coarse deltaic deposits, having a<br />

maximum thickness <strong>of</strong> 7 m rest upon blue<br />

lake clays.<br />

<strong>The</strong> north-south cross section from<br />

the apex <strong>of</strong> the delta into <strong>Lake</strong> <strong>St</strong>. Cl air<br />

S<strong>of</strong>t<br />

Clay<br />

f 11 ustrates the near-surface stratigraphy<br />

(Figure 9). Topographically however, this<br />

cross section reveals two distl nct 1 evels,<br />

a modern and the equally obvious premodern<br />

surface. <strong>The</strong> premodern surface,, standing<br />

about 1.5 rn above take <strong>St</strong>. <strong>Clair</strong>, consists<br />

<strong>of</strong> coarse* 0x1 dized sand and is confined<br />

to the apex <strong>of</strong> the delta complex. It has<br />

been dissected by long, sinuous channels<br />

which have been alluviated. Occasionally<br />

during high 1 ake levels these channels<br />

have been reoccupfed~ particularly in<br />

areas such as Dickinson Island, where<br />

human interference has been minimal.<br />

Because this hfgher surface i s<br />

topographically and sedimentologically<br />

distinct, it must be a surface which was<br />

deposited during a pre-existing higher<br />

lake level. <strong>The</strong> modern delta with its<br />

fine sand sediment (0.25 to 0.125 mn or +2<br />

to 3 phi units) is located at present mean<br />

lake level and is represented by the<br />

active crevasses and interdistributary<br />

marsh deposits.<br />

Based on the chronology <strong>of</strong> the<br />

progl acial Great <strong>Lake</strong>s*. field data, and<br />

Figure 8. Geologic cross-section <strong>of</strong> the northern portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> showing<br />

depth to bedrock, and thickness <strong>of</strong> glacial, lacustrine, and deltaic deposits,<br />

S<strong>of</strong>t<br />

Clay


61 inkc<br />

<strong>St</strong>. <strong>Clair</strong><br />

I<br />

Modern Delta Premodern Delta<br />

Bassett Harrens<br />

Island Island<br />

Walpole<br />

Island<br />

I North<br />

1 2 3 4 5 6 7 8 9 10 11 12 Miles<br />

I I I --LL--l I I I I<br />

Deltaic Sediments<br />

a ORGANIC<br />

SILTY SAND<br />

SANDY CLAY<br />

Figure 9. Cross-secti on <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta showi<br />

L .~ - A-2<br />

.- . --.-- - _I<br />

stribution <strong>of</strong> deltaic sediments.<br />

available Carbon-14 data, the relative materfals range in age from 6,100 + 80 to<br />

geomorphological events <strong>of</strong> the <strong>St</strong>. Clafr 98300 + 200 years. <strong>The</strong>se organic<br />

Delta may be determined. Sfnce the materials do not appear to behsifad and<br />

retreat <strong>of</strong> the Late Wisconsin ice sheet,<br />

the outlets <strong>of</strong> tho Great <strong>Lake</strong>s, and hence<br />

lake levels in the <strong>St</strong>. <strong>Clair</strong> basin, have<br />

oscillated sufficient1 y to construct two<br />

deltas. Compared to other proglacial<br />

Great <strong>Lake</strong>s, <strong>Lake</strong> <strong>St</strong>. Clafr was a less<br />

permanent feature as the lake bottom was<br />

exposed to subaerial modlf ication due to<br />

the f 1 uctuatf ng ice sheet which determl ned<br />

lake level conditl~n~ (Figure 10).<br />

One C-14 date retrieved from the<br />

upper portion <strong>of</strong> the lake claysr but<br />

beneath the premodern delta, f ndicates<br />

that both deltas are less than 7,300 + 80<br />

years old, Based upon older proglacial<br />

Great <strong>Lake</strong>s chronology, the premodern<br />

delta may have been deposited during the<br />

. . .580'<br />

T-- - 7<br />

12000 9000<br />

Years belore the present<br />

ALGONQUIN STANLEY NIPISSING<br />

latter stage <strong>of</strong> Algsnquf n time (Wightman Figure 10. Chronology <strong>of</strong> lake levels and<br />

19611, Additfonal C-14 dates have been lake stages in the Huron and Erie basins<br />

obtained by Mandelbaum (1969). <strong>The</strong> dated (Darr and Eschman 1970).


they have not been jncluded in our<br />

interpretation, However, even if the<br />

latter dates were used they would not<br />

detract from our argument. Pezzetta<br />

(1968) also has radiogenic data that<br />

reveal that the minimum age <strong>of</strong> ,the delta<br />

sediment is 4,300 years before present<br />

(B.P. 1.<br />

With the subsequent retreat <strong>of</strong> the<br />

ice during the <strong>Lake</strong> <strong>St</strong>anley low-water<br />

stager an outlet for the <strong>Lake</strong> Huron basin<br />

to the <strong>St</strong>. Lawrence was created via North<br />

Bay, Ontario. During this stage the pre-<br />

modern sands <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta may<br />

have been exposed, oxidized, and dissected<br />

(Figure 10). Following the low-water<br />

stage, <strong>Lake</strong> Huron once again rose to the<br />

Nipissing stage and drained into the lower<br />

Great <strong>Lake</strong>s via the <strong>St</strong>. <strong>Clair</strong> River.<br />

As determined by the more recent<br />

chronology, the premodern delta may have<br />

been deposited during Nipissing time some<br />

3,500 to 5,000 B.P. and not during<br />

Algonquin time (Dorr and Eschman 1970).<br />

<strong>The</strong> premodern delta was deposited during a<br />

higher than present lake level and the<br />

subsurface sediments range In age from<br />

7,300 to 9,300 B.P. It i s suggested that<br />

fol lowf ng the <strong>Lake</strong> <strong>St</strong>anley stage (i.e.,<br />

Nipissing stage), the <strong>St</strong>. <strong>Clair</strong> River once<br />

again flowed south into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />

the premodern delta was deposited at an<br />

elevation sl ightly higher than the modern<br />

delta (Table 5).<br />

On many coasts, evidence for lower<br />

than present sea levels is determined by<br />

depositional surfaces which dip beneath<br />

younger sediments (Shelemon 1971). Had<br />

the premodern delta been deposited du r i ng<br />

the Algonquin stage and the subsequent<br />

early <strong>Lake</strong> <strong>St</strong>anley stage, evidence <strong>of</strong> that<br />

event as a depositional surface beneath<br />

the modern delta or as a paleosol should<br />

be apparent. Numerous borings suggest<br />

that the premodern delta does not plunge<br />

beneath the modern sediment. Since the<br />

stratigraphy imp1 ies that lake levels were<br />

lower than present only once, a Nipissing<br />

stage is favored for the depositton <strong>of</strong> the<br />

Table 5. Interpretation <strong>of</strong> the events related to the origin <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> Delta. a<br />

Approximate <strong>Lake</strong> stage Event<br />

Dates B.P.<br />

3,500 to Present Modern <strong>Lake</strong> <strong>St</strong>. Cl air Flow <strong>of</strong> <strong>Lake</strong> Huron continues<br />

and Algoma Phase southward; deposition <strong>of</strong><br />

modern <strong>St</strong>. Cl air River Delta<br />

and dissection <strong>of</strong> premodern<br />

surface<br />

3,500 to 5,000 <strong>Lake</strong> Nipissing Deposltion <strong>of</strong> premodern delta<br />

approximately 1.5 m (5 ftl<br />

above present mean 1 ake level<br />

5,000 to 10,500 <strong>Lake</strong> <strong>St</strong>anley <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin exposed;<br />

outlet for upper Great <strong>Lake</strong>s<br />

via North Bay* Ontario<br />

101500 to 12,500 <strong>Lake</strong> A1 gonqui n Val ders Maximum<br />

and Post-Algonquin<br />

Phases<br />

a~ata sources: Dorr and Eschman (19701, Raphael and Jaworski 619825,


premodern delta. Organic deposits, com-<br />

monly composed <strong>of</strong> large wood fragments<br />

(probably red ash) 1le beneath both<br />

deltas. Assuming that the peat accum-<br />

ulated from organic growth above the water<br />

table, it probably was deposited during a<br />

low-water period prior to the Niplssing<br />

stage ( possl bl y <strong>Lake</strong> <strong>St</strong>an1 ey 1.<br />

Following the <strong>Lake</strong> Nipissing high<br />

level, <strong>Lake</strong>s <strong>St</strong>. <strong>Clair</strong> and Huron fell to<br />

their present levels. <strong>The</strong> premodern<br />

channels were sl lghtly entrenched and the<br />

premodern surface oxidized during the last<br />

3,500 years. Towards the apex <strong>of</strong> the<br />

premodern delta, beyond the areas <strong>of</strong><br />

flood1 ng, a mix <strong>of</strong> hardwoods has colonized<br />

the oxidized soils and evidence <strong>of</strong><br />

drowning <strong>of</strong> a pre-<strong>Lake</strong> <strong>St</strong>anley delta by<br />

Niplssing high water levels is lacking.<br />

With the fa11 to the approximate present<br />

lake level, the modern delta was deposited<br />

in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Deltaic landforms occur where<br />

substantial quantities <strong>of</strong> clastic sediment<br />

are Introduced and deposited into a<br />

receiving basin. Sediment is normally<br />

eroded from a drainage basin and<br />

transported down an alluvia1 valley by a<br />

master stream and its tributaries.<br />

<strong>The</strong>refore the rate <strong>of</strong> delta development is<br />

dependent upon the flow regime and the<br />

avaf 1 abil ity <strong>of</strong> sediments.<br />

<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River maintains a<br />

relatively stable channel and i s not an<br />

alluvlal stream In the common connotatlon.<br />

Fundamentally. the river is a strait<br />

connecting two large water bodies and<br />

therefore does not have a high variability<br />

<strong>of</strong> flow. Flow measurements during the<br />

ice-free season in 1959 revealed that<br />

discharge ranged from a low <strong>of</strong> 3,900<br />

m3/sec to a high <strong>of</strong> 5,700 m3/sec. By<br />

contrast the Mississippi River flow<br />

between Baton Rouge and the Gulf <strong>of</strong> Mexico<br />

varies from a minimum <strong>of</strong> 1,400 d/sec to a<br />

maxJmum <strong>of</strong> 44,400 m3/sec (Duane 1967).<br />

<strong>The</strong>refore* for the <strong>St</strong>. <strong>Clair</strong> River, low<br />

flow was 68% <strong>of</strong> the high flow as compared<br />

to the Mississippi River where low flow<br />

was only 31% <strong>of</strong> the high flaw. <strong>The</strong><br />

discharge <strong>of</strong> the <strong>St</strong>. Clatr River is<br />

relatively constant and averages about<br />

5~097 m3/sec (International Joint<br />

Commissfon 19811, Instead <strong>of</strong> a spring<br />

flood typical <strong>of</strong> most rivers, discharge in<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fs s1 ightly Increased In<br />

late-summer when lake levels In <strong>Lake</strong> Huron<br />

are highest, Thus, overland flow and<br />

flooding Is not an annual event and -its<br />

occurrence is dependent in part upon<br />

seasonal 1 ake l eve1 condi tions.<br />

Greatest delta extension occurs In<br />

areas <strong>of</strong> highest flow and sediment yield.<br />

Flow di stribution data make it clear that<br />

the most active portlon <strong>of</strong> the delta is<br />

presently confined to the western side <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Much <strong>of</strong> the flow <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> River is carried by the large<br />

distributaries on the Amerlcan side <strong>of</strong> the<br />

delta. North, Middle, and South Channels<br />

account for approximately 95% <strong>of</strong> the flow<br />

volume whereas the principal Canadian<br />

distributary, Chenal Ecarte, accounts for<br />

5%. Although North Channel appears to<br />

have been the main channel a century agoj<br />

-. -<br />

construction and continual dredsins <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel has fncreased the<br />

f1 ow <strong>of</strong> South Channel.<br />

Because the <strong>St</strong>. <strong>Clair</strong> River is not a<br />

true river system and has few tributary<br />

streams the source area for the deltaic<br />

sediments is not solely <strong>of</strong> fluvial origin.<br />

Rather, the principal source appears to be<br />

the shorelines <strong>of</strong> southern <strong>Lake</strong> Huron.<br />

Although some organic matter is carried In<br />

suspension, the clastic fraction is<br />

dominant. In terms <strong>of</strong> composftion, quartz<br />

sand Is the most commonr but fragments <strong>of</strong><br />

feldspar, chert, carbonate minerals and<br />

igneous and metamorphic rock fragments<br />

also occur. According to Duane (19671,<br />

the mean diameter <strong>of</strong> the suspended river<br />

sediment ranges from 0.17 to 3.16 KWI while<br />

the mean diameter <strong>of</strong> the bed load varies<br />

from 1.98 to 2.64 mm. Detailed sedimen-<br />

tological studies suggest that the shallow<br />

bays <strong>of</strong> the delta are composed <strong>of</strong> fine<br />

sand (0.115 mm) deposits and the sorting<br />

Is fair (Mandelbaum 1966). It may be<br />

concluded that the <strong>St</strong>. <strong>Clair</strong> Delta is<br />

composed <strong>of</strong> sand-sized sediments. In<br />

contrast most marine deltas consist <strong>of</strong> a<br />

finer sediment fraction. Also where<br />

organic deposlts occur on the delta<br />

surface or subsurfacer they were formed in<br />

place and not transported into the de1 t a<br />

comp l ex.<br />

Estimates <strong>of</strong> the sediment discharge<br />

rates were compiled by Pezzetta (1968) and


are summarized on Table 6. <strong>The</strong> high<br />

variability is probably a reflection <strong>of</strong><br />

the time <strong>of</strong> sampling and <strong>of</strong> refinements in<br />

sampling and analytfc techniques.<br />

Furthermore, the load is not distributed<br />

uniformly from one reach to anotherr a<br />

fact which accounts for the variable<br />

transport rates,<br />

Under storm conditions there is a<br />

marked increase in the suspended load <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> River. <strong>The</strong> suspended sed-<br />

iment load is directly related to wind<br />

velocities and duration over southern <strong>Lake</strong><br />

Huron. During periods <strong>of</strong> northerly wfnds<br />

the suspend load is decreased. River<br />

velocities are competent to transport<br />

material comprised <strong>of</strong> the beach sediment<br />

from southern <strong>Lake</strong> Huron. Sediment<br />

deposits in the nearshore zone <strong>of</strong> <strong>Lake</strong><br />

Huron have a mean diameter <strong>of</strong> 0.125 mm to<br />

0.25 mm; and according to Duane (19671,<br />

most clastic sediments suspended in the<br />

<strong>St</strong>. <strong>Clair</strong> Delta have mean diameters <strong>of</strong><br />

slightly finer sizes. Particle size data<br />

suggest that the sediments <strong>of</strong> the delta<br />

are derived from reworked nearshore<br />

sediments in <strong>Lake</strong> Huron.<br />

<strong>The</strong> soil types <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

wetlands ref 1 ect the late Quaternary<br />

history <strong>of</strong> the regfon and may be divided<br />

into two broad categor'les. <strong>The</strong> soils <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> Delta proper are derived<br />

from sources adjacent to <strong>Lake</strong> Huron<br />

whereas the soil types along the eastern<br />

shore <strong>of</strong> the 1 ake are derived from lake<br />

pl a? n sediments i n southwestern Ontario.<br />

As such the delta soils general 1 y have a<br />

coarser texture than the soils <strong>of</strong> eastern<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Table $5. Sediment discharge rates and characteristics in the <strong>St</strong>. <strong>Clair</strong><br />

River.<br />

Transfer rate Location Load type Sediment<br />

grade<br />

54,700 1n3/~ear Shore <strong>of</strong> <strong>Lake</strong> Huron at Tract ion Sand<br />

head <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River<br />

61,600 d/year <strong>St</strong>. <strong>Clair</strong> River at Bay Point Suspended Sf1 t-cl ay<br />

Light (after storm)<br />

37,500 m3/& <strong>St</strong>. <strong>Clair</strong> River bottom 12 km Traction Cl ay<br />

between <strong>St</strong>. <strong>Clair</strong> 8 Marine<br />

City (after severe storm)<br />

40,100 <strong>St</strong>. <strong>Clair</strong> River Channel Tract ion ---<br />

wall between <strong>St</strong>. Cl a3 r<br />

8 Marine City<br />

19,900 d/year <strong>St</strong>. <strong>Clair</strong> River bank between Traction ---<br />

<strong>St</strong>. <strong>Clair</strong> & Marine City<br />

15,100 d/&y Surface and bottom <strong>of</strong> Chenal Total Clay<br />

A Bout Rond, 3 km below North<br />

Channel (after a storm)<br />

15,200 m3/year Entire <strong>St</strong>. <strong>Clair</strong> River Total Sand<br />

a~ata sources: Cole ( 1903 1 r Duane ( 19671, Pezzetta ( 1968).


<strong>The</strong> soi.78 <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta were<br />

derived from the shoreline and nearshore<br />

zone <strong>of</strong> <strong>Lake</strong> Huron. <strong>The</strong> sediment was<br />

transported,by the <strong>St</strong>. <strong>Clair</strong> River and now<br />

forms the delta proper. Two s<strong>of</strong>l types<br />

are evident in the delta. At the apex <strong>of</strong><br />

the landform, 1.5 to 2 m above mean lake<br />

level, Sanilac loam (Col wood fine sandy<br />

loam in Ontario) is prevalent. This soil<br />

is a very fine sandy loam and occurs on<br />

moderate (0-8) slopes. <strong>The</strong> soil formed<br />

in limey, water-laid sediment <strong>of</strong> very fine<br />

sandy loam and loamy very fine sand (U.S.<br />

Department <strong>of</strong> Agriculture 1974). A1 though<br />

the surface layer is dark-brown (0-15 cm),<br />

sand pits reveal that the deeper soil is<br />

ye1 lowish-brown (Munsell color code = 10<br />

YR 5/61, a fact suggesting slight<br />

oxidation. This soil type represents a<br />

premodern delta that was formed at higher<br />

lake levels in the geologic past.<br />

<strong>The</strong> wetlands proper are characterized<br />

by the Bach loam ("marshw soils in<br />

Ontario). <strong>The</strong> sol1 has a very fine sand<br />

Y., 1<br />

loam texture which farmed in limey<br />

lacustrine sediments (Figure 11).<br />

Occurring on land with a slope <strong>of</strong> 2% or<br />

less, the soil is <strong>of</strong>ten water l ogged, In<br />

a typlcal pr<strong>of</strong>ile the surface layer is<br />

black* calcareous, very fine sand loam, 18<br />

cm thick. <strong>The</strong> subsoil is 75 cm th9ckv and<br />

is made up <strong>of</strong> several layers <strong>of</strong> gray,<br />

calcareous, very friable* very fine sand<br />

loam. It represents the modern delta<br />

front, local depresslons~ and ancestral<br />

channels <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> dlstributaries.<br />

<strong>The</strong> Clyde loam is a poorly drained<br />

loam occupying the eastern shoreline <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. It is characterized by 0<br />

to 20 cm <strong>of</strong> very fine, very s<strong>of</strong>t, very<br />

dark gray silt-cl ay with large quantities<br />

<strong>of</strong> organic debris. From 20 to 30 cm deep.<br />

Clyde loam is dark gray, fine, firm silt<br />

and clay (Mudroch 1981). This s<strong>of</strong>l type<br />

developed on older, more f i ne-grained<br />

sediments <strong>of</strong> a lake plain and i s therefore<br />

finer and generally has a higher organic<br />

content than the delta soils.<br />

6rrvsmad Jvlm 26, 1977<br />

re.1 *.t.r,<br />

I -- 1 I 1 1 1<br />

r-"<br />

100<br />

---<br />

400 600 800<br />

--- -T -,<br />

1000 I200<br />

1<br />

1400 1600<br />

I<br />

1800 ZOO0 2200 2100 2600 2800 IOOOFI<br />

0 :a0 ZOO 100<br />

100 100 600 I00 800 900 1<br />

tread R(4p.<br />

$and and Peat alack<br />

Srrvr#ad Ocl 7, 1977<br />

' 5-7-<br />

Ebst<br />

115.5<br />

115.0<br />

171,s<br />

-----I a<br />

3000 3200 1100 1600 3804 4040 4200 1400 4600 4800 5000 5200 5400 5600 5100 6000fl<br />

-- I-- -<br />

I--<br />

(st*) 1000 i:5t IZJO lion ICOG !fn@ !do0 1300 1800 8<br />

Figure 11. Vegetation transect on Dickinson Island, <strong>St</strong>. <strong>Clair</strong> Delta, showing character<br />

<strong>of</strong> wetland soils (Jaworski et a7. 1981).<br />

22<br />

116 0<br />

115 1<br />

11s 0<br />

li4 5


<strong>The</strong> soils in the <strong>Lake</strong> <strong>St</strong>. Clafr<br />

region are basically derived from<br />

unconsolidated sediments deposited over<br />

bedrock in early Holocene time (1 .ear past<br />

20,000 years). <strong>The</strong>se s<strong>of</strong>ls contribute to<br />

the geochemical composition <strong>of</strong> bottom<br />

sediments <strong>of</strong> the wetlands. Typically, the<br />

wetland sediments investigated by Mudroch<br />

f 1981) are submerged mineral soils, with a<br />

pH ranging between 6.9 and 7.2. <strong>The</strong><br />

wet1 and sediments from the Walpole Island<br />

sector <strong>of</strong> the delta are coarser: more than<br />

half <strong>of</strong> the particles are at least sand<br />

size whereas the eastern shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> silt size particles are dominant.<br />

<strong>The</strong> absence <strong>of</strong> extensive peat or<br />

organic rich soils is noteworthy. Peat<br />

accumu1ations are poor and were probably<br />

formed in place rather than transported<br />

into the wetlands. Deltaic wetlands are<br />

<strong>of</strong>ten characterized by blanket peat<br />

deposits especially in sheltered local-<br />

ities such as bays and backbarrler flats.<br />

In the Great <strong>Lake</strong>s coastal wetlands, peats<br />

are with few exceptions (e.g.$ western<br />

Saginaw Bay) uncommon. This suggests<br />

several possibil itles: 1) wetlands during<br />

recent geologlc time were not abundant, or<br />

2) peat deposlts decomposed rapidly Or9 3)<br />

water exchange between the wetlands and<br />

lake was efficient in exporting organic<br />

material. In any caser clastic sediments<br />

are dominant in the wetland soils <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>.<br />

2.2 CLIMATE AND WEATHER<br />

<strong>The</strong> variable weather elements such as<br />

windr precipitation, air temperature,<br />

humidity* and growing season influence the<br />

physical and biological character <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. A unfque aspect <strong>of</strong> this water<br />

body Is rapid water level change due to<br />

seiches or wind tides. Also; spring ice<br />

jams can significant1 y a1 t er the water<br />

level <strong>of</strong> the lake.<br />

<strong>The</strong> climatic variables such as<br />

temperature, wind, atmospheric pressure,<br />

and precipitatfon play a stgnjf icant role<br />

in the terrestrial and lacustrfne<br />

environment <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r. <strong>The</strong>se<br />

elements are primarily responsible for the<br />

water budget <strong>of</strong> the lake as well as wave<br />

activityt the extent <strong>of</strong> the growing season<br />

in the wetlands, the distribution and<br />

length <strong>of</strong> ice cover, and the water level<br />

conditions on a daily, monthly, and<br />

seasonal bas 1 s.<br />

<strong>The</strong> climate <strong>of</strong> the lake i s<br />

characterized by cold wlnters (coldest<br />

month below 0% and warm summers (warmest<br />

month above 220C) with precipitation<br />

uniformly distributed through the year<br />

(climate type Daf based on the Koppen<br />

system). Cyclonic storms occur through<br />

the yearr but decrease in frequency durfng<br />

late June, Julys and August. Durlng this<br />

time convectional up1 ift is frequent and<br />

thunderstorms are common. Most <strong>of</strong> the<br />

moisture that falls on the Great <strong>Lake</strong>s<br />

basin as precipitation originally<br />

evaporated from the tropical At1 antic<br />

Ocean or the Gulf <strong>of</strong> Mexico (Phil1 ips and<br />

McCulloch 1972). On an average day, 10<br />

billion m3 <strong>of</strong> water in the form <strong>of</strong> water<br />

vapor are advected across the Great <strong>Lake</strong>s<br />

basin. <strong>The</strong> efficiency <strong>of</strong> precipitation<br />

mechanisms is such that only about 15% or<br />

1.4 billion m3 fa11 as rain on a given<br />

day.<br />

<strong>The</strong> frost-free season is defined as<br />

the interval in days between the last<br />

occurrence <strong>of</strong> frost in spring and the<br />

first occurrence in fall and is thus an<br />

indicator <strong>of</strong> the length <strong>of</strong> the growing<br />

season. <strong>The</strong> mean annual frost-free period<br />

for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is 160 days. With the<br />

exception <strong>of</strong> the south shore <strong>of</strong> <strong>Lake</strong> Erie,<br />

<strong>Lake</strong> Ontario* and the south shore <strong>of</strong> <strong>Lake</strong><br />

Michigan; which experience 180 f rost-f ree<br />

days annually, <strong>Lake</strong> <strong>St</strong>. Clafr has the<br />

longest frost-free period in the Great<br />

<strong>Lake</strong>s basin. <strong>The</strong> spring cooling exerted<br />

by the lake prevents premature growth <strong>of</strong><br />

the vegetation thus lessening the chances<br />

<strong>of</strong> crop loss due to late spring frosts<br />

(Eichenlaub 1979). Conversely, the slow<br />

cooling in autumn retards the occurrence<br />

<strong>of</strong> the first fall frost thus extending the<br />

growing season.<br />

C1 osel y re7 ated to the length <strong>of</strong> the<br />

frost-free season is the accumulation <strong>of</strong><br />

growing degree days as an index <strong>of</strong> the<br />

amount <strong>of</strong> heat available d~ring the<br />

growing season. <strong>The</strong> index is usually<br />

defined as the number <strong>of</strong> degrees <strong>of</strong> mean<br />

daily temperature above a base <strong>of</strong> 5,SoC.


<strong>The</strong> growing degree-day concept has been<br />

applied to delimit areas suitable for<br />

particular vegetation types and as a means<br />

<strong>of</strong> predicting the hatching date <strong>of</strong> various<br />

insects (Baker and <strong>St</strong>rub 1965). On the<br />

basis <strong>of</strong> using mean monthly temperature<br />

and the number <strong>of</strong> days in a month, the<br />

accumulated number <strong>of</strong> degrees above a base<br />

<strong>of</strong> 5.60C in a normal year is 2,340 OC in<br />

the southern portion <strong>of</strong> take <strong>St</strong>. <strong>Clair</strong> and<br />

29200 OC in the northern sector <strong>of</strong> the<br />

1 ake.<br />

Table 7 reveals that the mean annual<br />

average air temperature for three selected<br />

stations is 9.4OC. Mean monthly<br />

temperatures in January and February are<br />

well below freezing (-3.50C) but summer<br />

means exceed 220C. Such continental<br />

conditions commonly occur in regions which<br />

are remote from maritime air masses.<br />

One <strong>of</strong> the characteristics <strong>of</strong> the<br />

climate <strong>of</strong> the Great <strong>Lake</strong>s is the lack <strong>of</strong><br />

any marked seasonal i t y <strong>of</strong> prec i p i t at i on.<br />

Mean monthl y precipitation values for<br />

three stations (Table 8) average 77.83 cm.<br />

It should be noted that the Detroit and<br />

Windsor stations are located in highly<br />

urbanized areas compared to the Mount<br />

Clemens station, As noted on the table*<br />

the precipitation is 1 owest at Mount<br />

Clemens. Research in urban precipitation<br />

patterns indicates that the presence <strong>of</strong> a<br />

1 arge city such as Detroit does cause an<br />

increase in precipitation in the urban<br />

areas (Sanderson 1980). <strong>The</strong>refore9 Mt.<br />

Clemens i s more representative <strong>of</strong><br />

temperature conditions over <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>.<br />

Wind direction and frequency are<br />

significant with regard to waves, ice jams<br />

and short term (i.e., seiches) water level<br />

changes in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. According to<br />

Ayres (19641, the circulation in <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> is strongly influenced by prevail ing<br />

wind direction as are longshore currents.<br />

Figures 12 and 13 illustrate the average<br />

monthl y di rectfonal frequency <strong>of</strong> winds for<br />

Windsor and Mt. Clemens respectively. <strong>The</strong><br />

more spherical wind roses (e.g. March,<br />

April) have the greatest variabil ity in<br />

wind direction. <strong>The</strong> more skewed wind<br />

Table 7. a Mean monthly ternperatures for three selected<br />

stations.<br />

Mount Cl er,~ens, Detrolt City W i ndsor<br />

Michigan Airport Airport<br />

1940- 1969 1940-1969 1941-1970<br />

Month OF OC OF OC OF OC<br />

January<br />

February<br />

March<br />

Aprll<br />

May<br />

June<br />

July<br />

August<br />

September<br />

October<br />

November<br />

December<br />

Annual average 48.3 9.0 50.1 10.1 48.6 9.2<br />

a<br />

Data sources: Sanderson (1980); U.S. Dept. <strong>of</strong> Commerce, Weather<br />

Service.<br />

24


Table 9. a Mean monthly precipitation for three selected<br />

stations.<br />

Mount Cl emens Detroit C<strong>St</strong>y Windsor<br />

Michigan Airport Airport<br />

- -<br />

Month inches cm inches cm inches cm<br />

January<br />

February<br />

March<br />

April<br />

May<br />

June<br />

July<br />

August<br />

September<br />

October<br />

November<br />

December<br />

Annual average 28.07 71.29 30.95 78.60 32.91 83.59<br />

a<br />

Data sources: Sanderson (1980); U.S. Dept. <strong>of</strong> Commerce, Weather<br />

Service.<br />

roses (e.g., June, July, August) reveal a<br />

more prevailing wind. It is evident that<br />

predominant winds In the region are from<br />

the westerly direction# especially from<br />

the southwsst quadrant. Additional wind<br />

data for Windsor airport suggests that<br />

January, February, and March have the<br />

highest wind speeds averaging 20 km/hr<br />

while June, July, and August have the<br />

lowest at 13 km/hr. Furthermore, winds<br />

from the west-northwest have the highest<br />

speeds at 21 km/hrr and the lowesfwind<br />

speeds, 13 km/hro are from the southeast.<br />

Ice Cover<br />

A major physical feature on <strong>Lake</strong> <strong>St</strong>.<br />

Cl air is the occurrence <strong>of</strong> ice. Using 20<br />

years <strong>of</strong> datar Assel et a1 . (1983 1<br />

documented and mapped the distribution <strong>of</strong><br />

ice on the Great <strong>Lake</strong>s. <strong>The</strong> concentration<br />

<strong>of</strong> ice varies over the period <strong>of</strong> record<br />

mainly because <strong>of</strong> the shallowness <strong>of</strong> the<br />

lake. Although synoptic data on ice<br />

thickness especial 7y over the centrai<br />

portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> are poor#<br />

maximum thickness during severe winters is<br />

approximately 49 cm and maximum thickness<br />

during mild winters is about 5 cm,<br />

Normal 1y during mi d-December through<br />

February the 1 ake i s ice covered with the<br />

exception <strong>of</strong> Its exit--the Detroit River.<br />

During mild winters the only persistent<br />

ice cover in January and February occurs<br />

along the eastern shore1 ine <strong>of</strong> the 1 ake.<br />

However, during severe winters persistent<br />

ice will cover 100% <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from<br />

mid-December to mid-Apri 7.<br />

<strong>The</strong> impact <strong>of</strong> ice and ice jams on<br />

wet1 and vegetation is poorly understood,<br />

and can have negative as well as positSve<br />

effects, Ice jams migrating down<br />

distributary channels uproot aquatic and<br />

persistent vegetation from the substrate.<br />

Conversely, ice in the nearsh~re tone<br />

protects 1 acustrine wetlands by acting as<br />

a buffer and absorbing the impact from<br />

incoming waves. <strong>The</strong> coastal protectfon<br />

<strong>of</strong>fered by the ice In the nearshore zone<br />

is particularly significant during mild<br />

winters when most <strong>of</strong> the lake is not<br />

frozen and also durin~ the sprfng whe~<br />

storms frequently occur and wave energies<br />

are higher.


Jenuery February March , FEBRUARY<br />

/<br />

October I<br />

-- '$<br />

August<br />

November<br />

'i (/<br />

/ ' C 5<br />

June<br />

September<br />

December<br />

Figure 12. Monthly wi nd frequency diagrams<br />

for Windsor, Ontario, 1955-1972 (Sanderson<br />

1980).<br />

OCTOBER NOVEMBER DECEMBER<br />

Figure 13. Monthly wind frequency for Mt.<br />

Clemens, Michigan (Sel fridge Air Force<br />

Base) for period 1936-1953 (Ayers 1964).<br />

Note only velocities greater than 4 mph<br />

plotted; winds less than 4 mph shown as<br />

percent time.<br />

2.3 HYDROLOGY CirculatI~n in <strong>Lake</strong> <strong>St</strong>* Clajr<br />

n'storically water levels<br />

Of <strong>Lake</strong><br />

Investigations reveal that two major<br />

water masses occur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

C1air ranged apprOximate'y in Ayres (1964) modeled field data before and<br />

response changJng water budgets= Water after the construction <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

budgets In<br />

turn are to jnf l Ow and Cut<strong>of</strong>f Channel and found that the current<br />

Outflow as as evaporation and structure in the lake rnainta.lned semi-<br />

precipitation* <strong>The</strong> impact Of changing permanent patterns. <strong>The</strong> distributfon<br />

water levels within the basin On the pattern was altered according to changing<br />

shorelJne has been wind patterns, Leach (1980) also<br />

signi jcante HistorScal identified two discrete water masses in<br />

displacements have been documented with the 1 ake on the basis <strong>of</strong> cluster analysis<br />

the use <strong>of</strong> aerial photography over time. <strong>of</strong> physical and chemical data,<br />

Wetland vegetation transects and<br />

geographical dlstributions over time have <strong>The</strong> main flow <strong>of</strong> water entering <strong>Lake</strong><br />

a? so been determined . <strong>St</strong>. <strong>Clair</strong> is from the <strong>St</strong>. <strong>Clair</strong> River.


Thfs river contrfbutes about 98% <strong>of</strong> the<br />

total Inflow t o the lake. <strong>The</strong><br />

constructfon~ <strong>of</strong> the <strong>St</strong>. Lawrence Seaway<br />

and the opening <strong>of</strong> the South Channel<br />

Cut<strong>of</strong>f i n 1962 have increased the<br />

proportfon <strong>of</strong> <strong>St</strong>, <strong>Clair</strong> River water<br />

arriving directly into the main lake<br />

basin. <strong>The</strong> greater cross-sectional area<br />

<strong>of</strong> the Cut<strong>of</strong>f Channel for outflow which<br />

now exists and the straightening that the<br />

arti f ici a1 channel provides have decreased<br />

the frlctlonal effects and promoted a<br />

greater discharge <strong>of</strong> the st.-<strong>Clair</strong> River<br />

water directly into the main basin,<br />

In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> the extent and<br />

geometry <strong>of</strong> the ci rculatfng water masses<br />

are related to wind direction. <strong>The</strong><br />

predominant winds in the region are from<br />

the southwest quadrant (Sanderson 1980).<br />

Figure 14-D illustrates the two principal<br />

patterns in the eastern and western<br />

portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> basin derived<br />

from prevailing southwest winds. Figure<br />

14-A revsaxs a more djstinctive<br />

circulatory pattern associated with a<br />

north wind, Based upon Ayresl (1964)<br />

model, water in Anchor Bay originates from<br />

two distinct sources. Wl th southwest<br />

w'inds there are both a net diminution <strong>of</strong><br />

outflow through North Channel and the<br />

creation <strong>of</strong> a discrete gyre in Anchor Bay<br />

composed <strong>of</strong> Clinton River water. With a<br />

north wind, though, Anchor Bay becomes<br />

dominated by water from the North Channel<br />

<strong>of</strong> the <strong>St</strong>. Claf r River as the Clinton<br />

River water is reduced in the bay.<br />

<strong>The</strong> significance <strong>of</strong> the two main<br />

water masses is their relationship to the<br />

productivity <strong>of</strong> the lake. <strong>The</strong> water mass<br />

adjacent to Ontario is more influenced by<br />

nutrient load1 ngs f rom the Sydenham and<br />

Thames rivers and the smaller tritrutaries<br />

from Ontario. <strong>The</strong>se waterways drain an<br />

intensive1 y cultivated lake plain. Urban<br />

development on the south shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> also contributes to the loading.<br />

Because this water mass is more enriched<br />

and more stable than the mass associated<br />

with <strong>St</strong>. <strong>Clair</strong> River water, It has yielded<br />

greater productivity (Leach 1972, 1973 1.<br />

Due in part to the flushing time <strong>of</strong><br />

lake <strong>St</strong>. <strong>Clair</strong>, which is theoretically<br />

about 9 days, the fish communities have<br />

changed 1 fttle, with the exception <strong>of</strong> some<br />

col dwater species. According to Johnston<br />

(19771. the walleye and the yellow perch<br />

stocks over the past century have remained<br />

reasonably stable in spite <strong>of</strong> more<br />

intensive agricultural practices in the<br />

watershed, increased settlement in the<br />

coastal zoner and exploitatio~ <strong>of</strong> the<br />

resource from commercial f isheries. <strong>The</strong><br />

flushing action <strong>of</strong> relatively clean water<br />

from <strong>Lake</strong> Huron has prevented concentratlon<br />

<strong>of</strong> nutrients and eutrophication in<br />

most <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />

ater Discharae Into l ake <strong>St</strong>. <strong>Clair</strong><br />

In addltlon to the principal source<br />

<strong>of</strong> water from the <strong>St</strong>. <strong>Clair</strong> River,<br />

several other river basins contribute<br />

water to the lakes including the Blackr<br />

Belle9 Clinton* Thames, and Sydenham<br />

Rivers (Figure 15). <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River<br />

is not a true fluvial system but rather a<br />

strait connecting two large water bodies<br />

(<strong>Lake</strong> Huron and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>). It<br />

therefore does not exhibit typical river<br />

discharge characteristics. Furthermore,<br />

depositional processes associated with<br />

rivers are normally directly linked to<br />

extreme flow regjmes. But <strong>St</strong>. Clafr River<br />

flows and ultimate sediment distribution<br />

in the <strong>St</strong>. <strong>Clair</strong> Delta are more closely<br />

linked to ice conditions or high lake<br />

level condftfons.<br />

Of the total average fa11 <strong>of</strong> 2.5 m<br />

from the <strong>Lake</strong> Huron level to <strong>Lake</strong> Eriet<br />

1.5 m occur in the <strong>St</strong>. <strong>Clair</strong> River<br />

(Korkigian 1963). Its banks and bed have<br />

been relatively stable except for<br />

navigation improvementsr and sand and<br />

grave1 dredging over several decades.<br />

Figure 16 illustrates mean monthly<br />

discharge from 1900 through 1980 (Quinn<br />

and Kelley 1983). Average monthly<br />

discharge is 5,121 &/see and the range <strong>of</strong><br />

discharge is 4,254 m3/sec (February) to<br />

5,483 m3/sec (September). Detailed<br />

monthly data are included in Appendix 8,<br />

With flow velocities up to 3.2 km/hr the<br />

travel time through the system is<br />

re1 atively high, As determined by Dereckl<br />

(1983)~ the flow time <strong>of</strong> water from the<br />

international bridge at Port Huron to <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> is 21,1 hours (Table 9). <strong>The</strong><br />

flow time through the Detroit River (Belle<br />

Isle south to Geleron IsSandE fs 20.9<br />

hours, <strong>The</strong> initial travel timetable was<br />

developed for normal flow condltfons.


Based on the geomorphologic history South channels, which in turn each spllt<br />

Of the <strong>St</strong>* C1aBr Delta? the active before entering <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>e <strong>The</strong> flow<br />

portion <strong>of</strong> the landform at the present<br />

time 1s to the west. As the <strong>St</strong>. <strong>Clair</strong> is therefore proportional 1 y distributed<br />

River enters the Sk. <strong>Clair</strong> basin f t through the channels as illustrated In<br />

bifurcates near Algonac Jnto the North and Figure 17,<br />

. . . * . WATER LEFT LI PRLVIOUJ NlHb<br />

NORTH WINO<br />

-. cn~nlt EtA*Te (I JW**fOn CHI)IXEL<br />

-' ST CLSIR RIVER<br />

IU- * CLINIOW nivcn<br />

- * -WATER LLFY EY PREVIOU5 WlWU<br />

-----, - ~~lijvu~hci: OUTFLOW P ISCIPCOEHI<br />

WEST WINO C<br />

-' CUENhL fCI\RTE 4 JOHNSTON CIllNntL<br />

I--".<br />

ST. CLIlR RIVER<br />

I-- CLIITOR RIVER<br />

WATER LEFT aY PUEVIOUJ WINO<br />

NORTHWEST WIND<br />

--<br />

CIIENIL CCARTE 8 JOMNSTOW CHANNEL<br />

------L. ST. CLllR RIVER<br />

-+r - CLINTOM RIVER<br />

- ........ -L . NITER LEFT BY PREVIOUS WIND<br />

-----r . SUISURFACE CSCAPIYLnT<br />

SOUTHWEST WIND D<br />

Figure 14. Current Patterns for <strong>Lake</strong> <strong>St</strong>. Clafr under various wind conditions (Ayers<br />

1964).


Artificial modif icatlon In the past and 1925# and uncompensated navigation<br />

85 years has altered river levels and the improvements for the 7.7-m and 8.3-rn<br />

water volume i n <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. projects completed in 1933 and 1962<br />

Art.fficia1 channel changes 'In the <strong>St</strong>, respective1 y (Derecki 1982). <strong>The</strong>se<br />

<strong>Clair</strong> River s'lnce 1900 include dredging channel changes increased the discharge<br />

for commercial gravel removal between 1908 through the <strong>St</strong>. Cl air River and caused a<br />

---L. CHENAL LCARTE O JOHNBTOH CHANNEL<br />

-----* - 51. CLAIR RIVER<br />

--.CZ. CLINTON RIVER<br />

. , . , . , , . . . r WhTER LEFT BY PREVIOUS WIND<br />

SOUTH WlND<br />

--C. CHENAL CCARTe 6 JOUNSTON CHANNEL<br />

-----r- 1 ST. CLAIR RIVER ---u. CLINTON RIVER<br />

....... . W.PTER LEFT BY PREVIOUS WINO<br />

EAST WlND<br />

G<br />

E<br />

Figure 14. (Continued)<br />

29<br />

- ST.<br />

-----H.<br />

-r CRENAL ECARTE 6 JOHNLTON CHANNEL<br />

CLAlR WVER<br />

CLINTON RIVER<br />

WATER LEFT BY PREVIOU8 WlND<br />

-----+ BUSSURFACE OUTFLOW<br />

SOUTHEAST WlND<br />

--<br />

CHENAL LCARTE 6 JOHNSTOH CtiAhlNEL<br />

-- ST CLAIR RIVER<br />

--M r CLfNTOH RIVER<br />

- *=WATER LEFT BY PREVIOUS WIND<br />

NORTHEAST WIND H


Figure 15. <strong>Lake</strong> <strong>St</strong>. Cl ai r-<strong>St</strong>. Clai r River drainage basin show-<br />

ing location <strong>of</strong> precipitation and water level gages.<br />

Table 9.<br />

veloci$y<br />

<strong>Clair</strong>.<br />

<strong>St</strong>. <strong>Clair</strong> River flow time and<br />

from <strong>Lake</strong> Huron to <strong>Lake</strong> <strong>St</strong>.<br />

<strong>St</strong>atlon Tf me Distance Velocity<br />

(hr) (km) (km/hr)<br />

<strong>Lake</strong> Huron 0.0<br />

Blue Water Brldge 0.1<br />

Black River 0.9<br />

<strong>St</strong>ag Island 4.4<br />

<strong>St</strong>. <strong>Clair</strong> City 6.0<br />

Marine City 9.8<br />

Roberts Landing 12.0<br />

Algonac Lt, 12,5<br />

<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f 17.9<br />

J F M A M J J A S Q N O 1.1<br />

<strong>Lake</strong> <strong>St</strong>, Gfair 21.1 60.5<br />

Figure 16. Mean monthly discharge sf the<br />

<strong>St</strong>, <strong>Clair</strong> River 1900-1988. a~ata source: Derecki (1983).


Figure 17. Location map <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta distributary channels<br />

showing average discharge (cubic meters per second) and flow distri-<br />

bution (percent <strong>of</strong> total ) .


permanent lowering <strong>of</strong> the lake's level.<br />

<strong>The</strong> impact <strong>of</strong> these channel changes<br />

resulted in the l ~ ~ ~ r <strong>of</strong> i nthe g <strong>Lake</strong><br />

Michigan and <strong>Lake</strong> Huron water levels by<br />

0.27 m. Thfs depth superf mposed on the<br />

combined areas <strong>of</strong> <strong>Lake</strong>s Mfchlgan and Huron<br />

represents a permanent water loss <strong>of</strong> 32<br />

km3 or nine times greater than the volume<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air.<br />

<strong>Lake</strong> level changes and thef r duration<br />

play an important role in the character <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, its shoreline and its<br />

wetland connnunities. In general r high and<br />

stable water levels are most beneficial to<br />

the fish stock <strong>of</strong> the lake whereas low and<br />

stable or unstable conditions are least<br />

desf rable from an ecological standpoint.<br />

Converse1 y hf gher water levels when<br />

combined w l th storm events encourage<br />

higher flood f requencles and coastal<br />

erosion. <strong>The</strong> wetland communities <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>, Clalr respond to longer term water<br />

changes and hence adjust to such changes<br />

over time.<br />

All the Great <strong>Lake</strong>s includfng <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> experience changes in water<br />

levels due to short-term and long-term<br />

weather and cl imat ic conditions. Short-<br />

term changes are due to steep barometric<br />

gradients which may alter water levels for<br />

a period <strong>of</strong> several hours. During the<br />

winter months ice jams on the <strong>St</strong>. Claf r<br />

River cause abrupt water level changes<br />

whlch may span a period <strong>of</strong> 60 days. Long-<br />

term oscillations are related to water<br />

lnput and water output to and from <strong>Lake</strong><br />

<strong>St</strong>, <strong>Clair</strong>. Such water level changes,<br />

al though non-cycl lcr occur over a period<br />

<strong>of</strong> several years. Pre-1946 storm surges<br />

on <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and its connecting<br />

channels have been documented by Murty and<br />

Polavarapu (197.5 1 armd are summarlzed in<br />

Tab1 e 10. As noted* the water levels at<br />

Tecumsshp Ontario dropped 0,5 m and rose<br />

again over a perfod Qf approximately 48<br />

hours. Short-term depressions <strong>of</strong> water<br />

levels due to storm surges provide some<br />

flushing action and the fntroduction <strong>of</strong><br />

nutrfents fraorn the marshes fnto the bays<br />

and nearshore waters <strong>of</strong> the lake.<br />

However, the event is probably Loo short-<br />

1 ived to have signif posltfve impact,<br />

Table 118, Record storm surge levels <strong>of</strong><br />

Canadian stations o$ <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />

connecting channel s.<br />

Maximum<br />

Date <strong>of</strong> Observat l on surge<br />

storm stat i on (m) (ft)<br />

November1913 Fighting Islandb 0.48 1.56<br />

Isle Aux Pechesc 0.28 0.91<br />

October 1916 Fighting 1s1 andb 0.71 2.34<br />

Isle Aux Pechesc 0.51 1.67<br />

November 1940 ~ecumsehd 0.54 1.76<br />

a~ata source: Murty and Pol avarapu (19751,<br />

b~ower Detroit River.<br />

'upper Detroit River.<br />

d~ake <strong>St</strong>. <strong>Clair</strong>.<br />

Ice jams are a common occurrence on<br />

the <strong>St</strong>. <strong>Clair</strong> River and <strong>of</strong>ten detafn<br />

Interlake shipping. Ice blockades usually<br />

form in February and March in the <strong>St</strong>.<br />

<strong>Clair</strong> River and occasionally extend into<br />

April. On the average, ice retards the<br />

flow by 16%. Ice-jamming reduces the<br />

outflow from <strong>Lake</strong> Huron and raises its<br />

level and at the same time reduces inflow<br />

to <strong>Lake</strong> <strong>St</strong>. Clai r and lowers its level.<br />

During the winters <strong>of</strong> 1904-1905 and 1905-<br />

1906, average inflow from <strong>Lake</strong> Huron to<br />

<strong>Lake</strong> Erie decreased 1,369 m3/sec over 4<br />

months and 904 m3/sec over 5 months,<br />

respective1 y (Bajorunas 1968).<br />

During the spring <strong>of</strong> 1984, strong<br />

winds from the east and north caused a<br />

massf ve fce jam south <strong>of</strong> <strong>Lake</strong> Huron.<br />

Brash ice clogged the 62.5 km-course <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> River until the end <strong>of</strong><br />

April, and jams <strong>of</strong> 3 m or more in depth<br />

developed at the apex <strong>of</strong> the delta. This<br />

situation led to a decrease in discharge<br />

from the monthly average <strong>of</strong> 5,096 m3/sec<br />

for April, to about 2,50Om3/sec below the<br />

projected mean volume for that month.<br />

Also there was a subsequent drop fn the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> water level by more than<br />

0.76 m from March 21 to April 19 t 175.40<br />

to 174.59 m above mean sea level). In the<br />

closing days <strong>of</strong> Aprtl a convoy <strong>of</strong> Canadian<br />

and Amerfcan ice breakers, alded by<br />

unseasonably mild temperatures broke the<br />

ice jam at the upper end <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong>


Delta allowing 70 vessels anchored In the Although non-cyclicr the water levels<br />

Detrolt River and western <strong>Lake</strong> Erje to appear to have a high water period and a<br />

enter the upper Great <strong>Lake</strong>s. low water period every 7 to 10 years.<br />

Figure 18 records the impact <strong>of</strong> this<br />

ice jam in the <strong>St</strong>, Clafr River on the<br />

water level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Water<br />

level began its drop in mid-March 1984 and<br />

did not recover until the end <strong>of</strong> May 1984.<br />

<strong>The</strong> change (March 15-April 15) in the<br />

water level was 0.48 m. Ice jams<br />

temporarily a1 ter the hydrology <strong>of</strong> the <strong>St</strong>,<br />

Cl af r Delta, As the distributary channels<br />

are jammed, river water is diffused<br />

through crevasses into the bays (e.g.,<br />

Muscamoot and Goose Bays), This process<br />

is responsible for subaqueous<br />

sedimentation in the bays and the<br />

subsequent colonFzatfon by emergent<br />

wet1 ands.<br />

February 1984 was relatively mild and<br />

<strong>Lake</strong> Huron was largely ice freer but<br />

exceptionally cold weather followed and an<br />

ice cover formed over the entire lake by<br />

about March 10th (Great <strong>Lake</strong>s Commfssion<br />

1984). By the outset <strong>of</strong> April, <strong>Lake</strong> Huron<br />

again was nearly Ice free except at the<br />

head <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River. Unusually<br />

strong and persistent westerly winds<br />

reduced the movement <strong>of</strong> drifting ice down<br />

the river by blowing much <strong>of</strong> it from the<br />

south end <strong>of</strong> <strong>Lake</strong> Huron eastward.<br />

Following this event? winds from the east<br />

and north forced the jumbled ice into the<br />

<strong>St</strong>. <strong>Clair</strong> River over a 3-week period.<br />

Water level data for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

have been continuously recorded since<br />

1899. <strong>The</strong> long-term level <strong>of</strong> the 1 ake is<br />

determined by the inputs <strong>of</strong> water derived<br />

from run<strong>of</strong>f or inflow <strong>of</strong> connecting<br />

channels and tributaries to the lake, and<br />

precipitation over the lake basin. <strong>The</strong><br />

outputs are due to evaporatfon and outflow<br />

via the Detroit River. Long-term lake<br />

level oscfllations are usually thought <strong>of</strong><br />

as volumetric changes which are<br />

cl imatically controlled. Changes in 1 ake<br />

storage may be sumnarlzed as follows:<br />

Water voTume and lake level <strong>of</strong> a<br />

given 1 ake bast n are dependent upon water<br />

inflow and 0utf10wr which in turn is<br />

determined by factors such as overwater<br />

precipitation, evaporationr and run<strong>of</strong>f<br />

fnto and out <strong>of</strong> the basin. <strong>The</strong>se factors<br />

are interrelated and may be expressed as a<br />

transfer factor. <strong>The</strong> transfer factor is<br />

defined as the sum <strong>of</strong> the monthly<br />

precip itati on and run<strong>of</strong>f minus the<br />

evaporatfon and change in storage. QuInn<br />

( 1976 1 computed transfer factors for <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> (Table 11). <strong>The</strong> run<strong>of</strong>f from the<br />

land into <strong>Lake</strong> <strong>St</strong>. Clai r was determined<br />

independently for the four river basins<br />

draining fnto the lake. <strong>The</strong>y are the<br />

Be1 1 e, Cl i nton, Thames, and Sydenham<br />

Basins (Figure 15). On an annual basis<br />

and during most <strong>of</strong> the year, the transfer<br />

factor is most sensitive to run<strong>of</strong>f. <strong>The</strong><br />

evaporation however, becomes the most<br />

important factor during the high<br />

evaporation months <strong>of</strong> August to October.<br />

Figure 18 records the water level<br />

history <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r for 1900 to<br />

1984. <strong>The</strong> lake has a mean monthly<br />

elevation <strong>of</strong> 174.87 m above sea level at<br />

Father P<strong>of</strong>nt, Quebec in the Gulf <strong>of</strong> <strong>St</strong>*<br />

Lawrence. However, as the figure<br />

demonstrates, the water level in recent<br />

years has been we1 1 above this 1 eve1 .<br />

Table 12 indicates that the lake has had a<br />

record high <strong>of</strong> 175,64 m (June 1973) and a<br />

record low <strong>of</strong> 173.71 m or a range <strong>of</strong> 1.93<br />

m. Low water or chart datum is 174,25 m.<br />

This level is a fixed reference plane<br />

selected by the United <strong>St</strong>ates and Canada<br />

so that the majority <strong>of</strong> time during the<br />

navigation season the Great <strong>Lake</strong>s actual<br />

levels w i l l be above that plane. Water<br />

levels are lowest in February; based on a<br />

1900-1983 mean, the level for this month<br />

i s 174.47 m. HIghest water levels Bccur<br />

in July; the 1900-1983 average for thi s<br />

month is 174.94 m.<br />

S = (P-E) + (1-0) <strong>Lake</strong> level fluctuations occur<br />

naturally on <strong>Lake</strong> <strong>St</strong>. CSair and are the<br />

where: S = change in storage volume result <strong>of</strong> prolonged periods <strong>of</strong> precip-<br />

P = precipitation itation, drought* storm-related events, or<br />

E = evaporation ice conditions, <strong>The</strong> lake level depends on<br />

I = inflow the balance between the total water<br />

0 = outflow supplied to the lake from precipftation on


LEGEND<br />

I 1<br />

LAKE LEVELS<br />

Hydrographs are In feet above (+)<br />

or below (-) Chart Datum, the plane<br />

-'<br />

REGORDED on each lake to whrch navigation<br />

chart depths and Federal navigatton<br />

ImprOvement depths are re-<br />

PROBABLE<br />

ferred.<br />

1900-1 983<br />

Chart Datum and all other eleva-<br />

AVERAGE<br />

tions sire in feet above mean water<br />

level at Father Point, Quebec (In-<br />

MAXIMUM m,E 1973 ,973 ternattonai Great <strong>Lake</strong>s Datum - 1955). To convert feet to meters,<br />

MINIMUM 1936 1934 1926 multiply feet by o 30480.<br />

-<br />

Figure 18. Hydrograph <strong>of</strong> iake <strong>St</strong>, <strong>Clair</strong> and <strong>Lake</strong> Erie water levels, showing<br />

the impact <strong>of</strong> a record ice jam in the <strong>St</strong>. <strong>Clair</strong> Delta in March and April 1984<br />

(U,S, Army Corps <strong>of</strong> Engineers), Note the slight dip in iake Erie water levels<br />

in May in response to the <strong>St</strong>. Clai r Be1 ta ice jam.


Table 11. take <strong>St</strong>, <strong>Clair</strong> average monthly water transfer factor parameters. a<br />

Cubic &rs oar ntglrth<br />

Parameter J F M A M J J A S O N D<br />

Precipitatfon 23 20 25 34 25 34 31 31 25 23 28 28<br />

Tributary run<strong>of</strong>f 156 190 342 277 136 62 37 28 25 42 79 153<br />

<strong>Lake</strong> evaporation 25 25 11 8 48 37 34 45 59 51 42 20<br />

Change in storage -76 42 79 42 17 17 -6 -17 -37 -48 -25 14<br />

a Data source: Quinn (1976).<br />

Table 12. Water level data for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. a<br />

Parameter Meters Feet Remarks<br />

Low water datum 174.25 571.70<br />

Month1 y water levels<br />

Average 174.87 573.72 1900-1983 data base<br />

Record high 175.64 576.25 June 1973<br />

Record low 173.71 569.90 January 1936<br />

a~ata source: U.S. Army Corps <strong>of</strong> Engineers.<br />

the lake and run<strong>of</strong>f from the surrounding<br />

drainage area and the amount <strong>of</strong> water<br />

discharged from the Detroit River or by<br />

evaporation. <strong>The</strong> relationship between<br />

long-term changes in lake level and the<br />

climatic events which cause the changes is<br />

usually quite evident, even though there<br />

is generally a time lag in the response <strong>of</strong><br />

1 ake levels to imbalances in the system.<br />

<strong>The</strong> high levels <strong>of</strong> 1973 on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

as well as the other Great <strong>Lake</strong>s were<br />

attributed to a 16% increase In precip-<br />

itation and a 24% decrease in evaporation<br />

during 1972 compared to the long-term<br />

average for both precipitation and<br />

evaporat I on.<br />

2.4 WATER QUALITY<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> waters are usually<br />

we11 mSxed by both horizontal and vert-ical<br />

currents. <strong>The</strong> two major factors affectfng<br />

current patterns and water quality are the<br />

flow-through current <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

River (Figure 19) and the seasonal water<br />

temperature changes, Variations in water<br />

density, largely caused by changes in the<br />

water temperature and amount <strong>of</strong> suspended<br />

sol ids, and winds affect the 1 ateral<br />

movement at the lake surface, while tur-<br />

bulence created by storms, and rec-<br />

reational and commercfal boatfng actlv-<br />

ities affect the vertical movement (U.S.<br />

Army Corps <strong>of</strong> Engineers 1974).<br />

Water quality informatfon fs presented<br />

here f n terms <strong>of</strong> nutrlents and<br />

contaminants which either enhance or 1 imit<br />

wetland productivlty. Of particular concern<br />

are phosphorusr nltrogenr df ssol ved<br />

oxygen, alkal lnftyt pH, major fans, trace<br />

elementst and toxic organic compounds.<br />

Tharmaf structure, ! fght penetration" and<br />

sediment load are also addressed f n terms<br />

<strong>of</strong> Impacts to wetland stab4llty and<br />

habftats,


Figure 19. Distribut.ion <strong>of</strong> specific conductance in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

(Bricker et al. 1976). Note low values from the <strong>St</strong>. <strong>Clair</strong> River and<br />

high values at the mouths <strong>of</strong> the Thames River and Clinton River.<br />

In March 1970 the Canadlan government<br />

announced that 5,500 kg <strong>of</strong> commercially<br />

caught walleye I L !LjAB@)<br />

from <strong>Lake</strong> <strong>St</strong>. Cl afr were destroyed because<br />

<strong>of</strong> mercury contamination. In the early<br />

19705s mercury concentrations in fish in<br />

<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> ranged from 0.3 ppm to over<br />

5 ppm, the higher values were usually<br />

found in larger fish and in predacious<br />

species. <strong>The</strong> United <strong>St</strong>ates and Canada<br />

adopted a concentration <strong>of</strong> 0.5 ppm in fish<br />

muscle as a maximum for commercial fish,<br />

Both governments closed commerci a1 fishing<br />

in the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>,<br />

as well as walleye fishing in western <strong>Lake</strong><br />

Erie,<br />

<strong>The</strong> mercury pollution was largely<br />

attributed to waste discharges from chlor-<br />

alkali processing plants on the <strong>St</strong>. <strong>Clair</strong><br />

and Detroit Rivers which began operation<br />

in the 1950s. Several <strong>of</strong> these plants<br />

ceased operation in the early 1970s and as<br />

a result, mercury contamination has<br />

diminished in the environment. Levels <strong>of</strong><br />

total mercury In walleye collected in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> have decl jned from a mean <strong>of</strong><br />

over 2 ppm in 1970 to 0.5 ppm in 1980. In<br />

western <strong>Lake</strong> Erie, 1968 levels <strong>of</strong> mercury<br />

were 0.8 ppm as compared to only 0.3 pprn<br />

in 1976. <strong>The</strong> rapid environmental response<br />

subsequent to the cessation <strong>of</strong> major point<br />

source d fscharges at Sarnia, Ontario, and


at Wyandotte, Mfchlgan* can be attributed<br />

to rapid flushing by the <strong>St</strong>. <strong>Clair</strong>-Detrolt<br />

Rivers system and rapid burial <strong>of</strong><br />

contaminated sediments by the high load <strong>of</strong><br />

suspended sediment delivered to western<br />

<strong>Lake</strong> Erie.<br />

Few comprehensive water quality<br />

investigations have been conducted on <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and measurements in the coastal<br />

wetlands are rare. <strong>The</strong> Michigan Water<br />

Resources Commission (1975) conducted a<br />

thorough study <strong>of</strong> the open lake and bays<br />

along the American shore in 1973 (Table<br />

13 1 and Leach (1972, 1973, 1980) reported<br />

on conditions in CanadIan waters (Figure<br />

20). <strong>The</strong> U.S. Envl ronmental Protection<br />

Agency operates a national water qua1 i ty<br />

data storage and retrieval system<br />

(STORET). Pub1 ished and unpubl i shed <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> data from several Federal,<br />

state, provincial, and local agencies have<br />

been entered into this STORET for the<br />

period 1967-1982. Table 14 represents a<br />

retrieved STORET summary (mean values) for<br />

22 parameters measured in the <strong>St</strong>. Cl air<br />

Rfver, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Detroit River, and<br />

western <strong>Lake</strong> Erie during this period.<br />

Mudroch and Capobianco (1978) determfned<br />

the geochemical composition <strong>of</strong> the<br />

surface sediments (0 to 20 cm) and the<br />

overlying rnarshwater in Big Point Marsh on<br />

the Ontario shore <strong>of</strong> <strong>Lake</strong>-<strong>St</strong>. ClaOr<br />

(Tables 15 and 16). Mineralogically the<br />

sed irnents are composed <strong>of</strong> ill ite, quartz,<br />

calclte, dolomite, K-fe1 dspars,<br />

pl agi ocl ose, and mi nor amounts <strong>of</strong> chl ori te<br />

and kaol inite. <strong>The</strong> water can be<br />

characterized as hard with high levels <strong>of</strong><br />

nutrients. <strong>The</strong> pH <strong>of</strong> the marsh sediment<br />

is about 7.0, while the overlying water<br />

ranges between 8.1 and 8.4.<br />

Two important features which<br />

determine the water temperature <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> are the lake's shallowness and<br />

the high flow rate from <strong>Lake</strong> Huron.<br />

Because <strong>of</strong> the shallow depth <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>, it warms quickly in the spring and<br />

cools rapidly In the fall. <strong>The</strong> lake is<br />

too shallow to stratify thermally;<br />

therefore the water is almost always<br />

isothermal from surface to bottom, <strong>The</strong><br />

high inflow from the <strong>St</strong>. Clalr River<br />

Table 13. <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> water quality measurements for 1973 growing season, a<br />

Parameter<br />

-lake--<br />

July Aug Sept July Aug Sept July Aug Sept July Aug Sept<br />

Secchi depth (rn) 1.9 1.6 2.2 2.0 1.3 1.7 2.0 1.4 2.3 0.6 0.9 1.2<br />

Conductlvlty(umohs/cm) 184 203 202 181 213 205 183 214 210 340 273 274<br />

Temperature OC 19.1 22.3 21.2 19.4 23.5 22.0 20.2 22.9 21.9 23.3 24.6 21.1<br />

pH (units) 8.6 8.6 8.8 8.5 8.7 8.8 8.6 8.6 8.7 8.5 8.8 8.5<br />

DIssolvedoxygen(mg/l) 8.1 8.2 7.2 8.3 8.7 7.7 6.9 8.0 7.6 8.5 7.9 7.1<br />

Suspendedsollds (mg/l) 7.3 11.6 1.8 4.0 12.0 15.0 4.3 6.9 12.0 35.7 11.7 26.3<br />

Dlssolved solids (mg/lf 127 129 -- 127 120 -- 127 124 - 218 153 --<br />

Alkalfnlty (mg/l) 80 81 90 78 80 88 80 80 90 114 90 108<br />

Hardness (mg/l) -- -- 95 -- -- 95 -- - 96 -- -- 119<br />

Sodium (t~~/l) 5.1 6.1 5.2 5.1 5.5 5.3 5.3 5.6 5.1 15.8 9.7 12.7<br />

Magnesium (mg/l) 7.8 -- 8.0 8.0 - 7.9 8.0 - 8.0 12.1 -- 9.8<br />

Calclurn (rng/l) -- 26.1 24.9 -- 26.0 25.0 -- 26.9 25.3 -- 31.0 31.7<br />

Potassfurn (mg/l) -- -- 0.8 -- -- 0.9 -- -- 0.8 -- -". 1.6<br />

Sulfate (mg/l) -- 18.8 17.7 -- 17.5 17.5 -- 18.7 17.9 -- 19.1 22.9<br />

Phosphorus, total (ug/l) 29 60 15


Figure 20. Mean water quality characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from April<br />

through November 1971 (Leach 1972). Note increased concentrations near the<br />

southeast shore in the vicinity <strong>of</strong> the Thdmes River mouth.


Table 14. Chemical and physical characgeristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />

adjacent waters (mean record 1967-1982).<br />

Parameter<br />

Temperature OC<br />

Di ssolv%fed oxygen mg/l<br />

D,O, saturation %<br />

Conductivity (250C) umohs/cm<br />

Df ssol ved so1 f ds mg/ 1<br />

Suspended sol ids mg/l<br />

Secchi depth m<br />

Alkalinity mg/l<br />

pH<br />

Calcium, total<br />

units<br />

mg/l<br />

Magnes i um, total mg/ 1<br />

Potassi um, total mg/l<br />

Sod 1 urn, total mg/l<br />

Chloride, total mg/l<br />

Sulfate, total mg/l<br />

Fl uori de, total mg/l<br />

Silica, dissolved ug/1<br />

Ammonia, dissolved<br />

Nitrate + Nitrite, diss.<br />

ug/l<br />

ug/l<br />

Phosphorus, total ug/l<br />

Phosphorusr dissolved ~g/l<br />

Chlorophyll p ug/l<br />

<strong>St</strong>. ~lairb <strong>Lake</strong> Detrolt Western<br />

Uni t s River <strong>St</strong>. <strong>Clair</strong> River <strong>Lake</strong> Erie<br />

a ~ ta a source: U.S. Environmental Protection Agency, Large <strong>Lake</strong>s<br />

<strong>St</strong>ation, Grosse Ile, Michigan, STORET Data System.<br />

b~eadings do not reflect mean concentration at the <strong>St</strong>. <strong>Clair</strong> River due<br />

to location <strong>of</strong> sampling stations near major outfall s.<br />

(approxlmatel y 5,000 m3/sec) quick1 y<br />

rep1 aces the water in the lake with water<br />

from <strong>Lake</strong> Huron (in about 9 days). Thus,<br />

the temperature <strong>of</strong> Laks Huron has a strong<br />

influence on the temperature <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>. This is especially true in Anchor<br />

Bay and along the Michigan shore because<br />

most <strong>of</strong> the water from the <strong>St</strong>. <strong>Clair</strong> River<br />

enters the lake via the North Channel<br />

(33%) a t the east side <strong>of</strong> the bay.<br />

Ljke <strong>Lake</strong> Erie, highest temperatures<br />

are reached in August. On the Ontarto<br />

shore, water temperatures average about<br />

240C in July and August. On the western<br />

side <strong>of</strong> the lake, which is influenced more<br />

by the cool water from <strong>Lake</strong> Hurcn,<br />

temperatures are usually 2 to 40C lower.<br />

Temperatures are highest in the shallow<br />

coastal wetlands, reaching over 28OC, <strong>The</strong><br />

typfcal summer thermocl ine <strong>of</strong> deeper 1 akes<br />

in the Great <strong>Lake</strong>s system is non-exlstent<br />

in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />

In the fa1 1, the 1 ake Is warmest on<br />

the western sfde* a reversal <strong>of</strong> the summer<br />

pattern. Water is coolest in the shallows<br />

near shore and warmest near the mouth <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> River. Normally <strong>Lake</strong> <strong>St</strong>,<br />

<strong>Clair</strong> Ss almost completely ice covered by<br />

mid-January. Ice break-up usually occurs<br />

in early March.<br />

Mid-summer water transparency (Secchf<br />

dfsc) In <strong>Lake</strong> <strong>St</strong>. @lair normally ranges<br />

from 0,7 m along the Michigan shore to 2,6


Table 15. Geochemical composition <strong>of</strong><br />

sediments in B+g Point Marsh, Ontario, on<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Parameter Concentration ( ppm)<br />

Sf02<br />

A1 203<br />

Fe2~3<br />

CaO (%I<br />

Na2o<br />

K20<br />

Ti02<br />

MnO<br />

'205<br />

C r<br />

Ni<br />

Cd<br />

Co<br />

Cu<br />

Organic C (%I<br />

63.7<br />

93.7<br />

%ata source: Mudroch and Capobianco<br />

(19781,<br />

km near the center <strong>of</strong> the lake to 0.6 m<br />

along the Ontario shore (Herdendorf 1970).<br />

In the nearshare reglons the water color<br />

is brownish-green. Water in the vfcinfty<br />

<strong>of</strong> the navigation channel Is a d$stincttva<br />

cloudy-green but near the center <strong>of</strong> She<br />

lake away from the channel is a cfear-<br />

green.<br />

Table 16. Chemical composition <strong>of</strong> waterd<br />

Big Point Marsh, Ontario, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Parameter<br />

Ammonia-Nitrogen<br />

Nitrate-Nitrogen<br />

Nitrogen, Kj<br />

Phosphorus, total<br />

Potass 1 urn<br />

Cal cl um<br />

Magnes f urn<br />

Sod i um<br />

Alkalinity, total<br />

Lead<br />

Zinc<br />

Chromi um<br />

Copper<br />

Cadmi um<br />

Min. Max,<br />

(mg/l) (mg/l)<br />

a Data source: Mudroch and Capobianco<br />

(1978).<br />

<strong>The</strong> minimum intensity <strong>of</strong> subsurface<br />

light that permits photosynthesls has been<br />

set at 1.0% <strong>of</strong> incident surface light<br />

(Cole 1983). <strong>The</strong> zone <strong>of</strong> a lake from the<br />

surface to a depth at which 99% o f the<br />

surface light has been absorbed is known<br />

as the euphotic zone. Below this depth,<br />

primary productivity is considered n i 1.<br />

<strong>The</strong> relationship between Secchi disc<br />

transparency and the thickness <strong>of</strong> the<br />

euphotic zone provides a simple way to<br />

estimate the photosynthetically active<br />

region <strong>of</strong> a lake. A ratio <strong>of</strong> 3.0 to 3.5<br />

has been empirically determined for this<br />

relationship, yielding a thickness <strong>of</strong> the<br />

euphotfc zone rangfng from about 2 m along<br />

the shores to over 9 rn near the center <strong>of</strong><br />

the lake,<br />

Nitrogen* phosphorus, and sil ica are<br />

the primary lfmfting nutrients in <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>. Bicarbonate alkalintty is in such


great abundance (>80 mg/l) that a source<br />

carbon for primary productivity is never<br />

lackfng. Relatively high nitrate (>400<br />

vg/1It phosphorus (>ZOO ug/l) and silica<br />

(>2 mg/7) in the nearshore waters <strong>of</strong> the<br />

wetland bays (Table 13) reflect high<br />

watershed inputs, Leach (1972) also found<br />

high nutrient loading from the Thames<br />

River (Table 17). <strong>The</strong> mid-lake lower<br />

concentrations reflect low nutrient<br />

loading from <strong>Lake</strong> Huron water. <strong>The</strong> mean<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> concentrations <strong>of</strong> nutrients<br />

lie between values for tha upper Great<br />

<strong>Lake</strong>s and those <strong>of</strong> western <strong>Lake</strong> Erie<br />

(Schelske and Roth 197318 yielding a<br />

mesotrophic classification for this lake.<br />

On the basis <strong>of</strong> water chemistry,<br />

Leach (1980) identified two distinct water<br />

masses i n <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>: 1) a<br />

northwestern mass consisting primari 1 y <strong>of</strong><br />

<strong>Lake</strong> Huron water flowing from the main<br />

channels <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River and 2) a<br />

southeastern mass <strong>of</strong> more stable water<br />

enriched by nutrient loadings from Ontarlo<br />

tributaries and shoreline urban devel-<br />

opment. <strong>The</strong> margins <strong>of</strong> the water masses<br />

shift according to the wind di rectf on and<br />

characteristics <strong>of</strong> the two water masses<br />

for the period 1971-1975 is shown in<br />

Figure 21.<br />

Because thermal stratification does<br />

not occur* due to the lake's shallowness<br />

and rapid flow <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> River,<br />

dissolved oxygen concentrations are<br />

usually at saturation. In Bradley Marsh,<br />

north <strong>of</strong> the Thames River, Mudroch and<br />

Capobianco (19781 found that the<br />

temperature and dissolved oxygen<br />

saturation in marshwater during the<br />

growlng season are inversely related<br />

(Figure 22), but the 0, saturatian did not<br />

fa1 l below 75%. <strong>The</strong> Lcoastal marshes are<br />

shallow and well mixed by wind, and<br />

consequent1 y the water temperature Is<br />

affected di rectly by air temperature. <strong>The</strong><br />

diurnal temperature changes are 1 arge8 for<br />

example in July 1976, they found<br />

marshwater temperature at 0600 hours to be<br />

20°C and at 1500 hours* 27%. <strong>The</strong>se<br />

changes coupled with wind maintain high 0<br />

levels In the marshwater during thg<br />

summer.<br />

speed, but the-overall discreetness <strong>of</strong> the<br />

distribution 1s maintained during the ice-<br />

free periods <strong>of</strong> the year ( April-December) . <strong>The</strong> Michigan Water Resources<br />

A compression <strong>of</strong> the water quality Commission (1975) reported that<br />

Table 17. Comparison <strong>of</strong> cJemica1 characteristics <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> and associated waters.<br />

De1 t a <strong>Lake</strong> Thames<br />

Parameter (units) Channels <strong>St</strong>. <strong>Clair</strong> River<br />

Chlorophyll 3 (vg/l) 1.2<br />

Pheopigments (ug/ll 0.4<br />

Part. org. carbon (mg/1) 0.7<br />

Sol. react. phosphate (pg/1) 0.6<br />

Total phosphate (vg/l) 2.5<br />

Nitrate (pg/l) 17 8<br />

Nitrite (vg/l) 1.6<br />

Sol. react. silicate (mg/l) 0.3<br />

Total a1 kal tnity (mg/l) 87<br />

Sul fate (mg/l) 15.3<br />

Chloride (mg/l) 8.1<br />

'Flat3 source: Leach ( 19721,


sooi<br />

1 ". . .." . . ---- - - .<br />

' CHLOROPHYLL 5? (ug/L) 1<br />

Figure 21. Characteristics <strong>of</strong> <strong>Lake</strong> Huron<br />

water mass (A) and Ontario nearshore water<br />

mass (B) in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> from 1971 to<br />

1975 (Leach 1980). Solid line is mean<br />

water qua1 i ty for each water mass and<br />

dashed line is prediction based on a<br />

central station in each water mass.<br />

chlortnated hydrocarbon concentrations in<br />

sedfments are low throughout the lake.<br />

Lindane, aldrin, endrin, heptachlor<br />

(detectfon limits 0.001 mg/kg)r and<br />

chlorodane (detectton 1 imi t 0.005 mg/kg)<br />

concentrations were below detection, and<br />

dfeldrfn (detection limlt 0.001 mg/kg) was<br />

only detected at a few stations, then at<br />

maximum levels <strong>of</strong> only 0.006 rng/kg, DOE*<br />

TDE, and DDT (detection 1 Smit 0.002 mg/kgl<br />

were below detection at most stations, but<br />

high concentrations (0.039 mg/kg) were<br />

found In L'Anse Creuse Bay near the<br />

Clinton Rfver splllway. PCB (detection<br />

limits 0.05 mg/kg) levels were below<br />

detectfon in most nearshore areas, but<br />

concentrations ranged from 0.50 to 1.10<br />

mg/kg near the splllway, indicating that<br />

the Cl inton River is an important source<br />

<strong>of</strong> PCBfs.<br />

#<br />

o Temperature<br />

Apr ~ 6 y , I .lul Aug Sep 1976<br />

Figure 22. Temperature and dissolved<br />

oxygen relationship in a <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

coastal wetland, near the mouth <strong>of</strong> the<br />

Thames River, Ontario (Mudroch and<br />

Capobianco 1978).


3.1 PLANKTON<br />

Phvto~l Sessf 1 e A-<br />

Planktonic and sessfle algae,<br />

although not as important as macrophytes<br />

in the coastal marshes9 are signif icant<br />

primary producers <strong>of</strong> organfc matter in<br />

wetlands, converting the sun's energy into<br />

chemical compounds that in turn are used<br />

as food by animals and nonphotosynthetic<br />

mtcro-organisms. In the nearshore and<br />

open waters <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, however9<br />

alga productivity dominates. Phyto-<br />

plankton production and distribution are<br />

fnfluenced by sun1 ight, temperature,<br />

nutrients, morphometry, water movementsr<br />

grazing by zooplankton, and other factors.<br />

Many inorganic elements are required for<br />

algal cell growthp incl uding nitrogens<br />

phosphorus, potassium, calcium, and iron.<br />

Algae reproduce rapidly when phosphorus is<br />

added to the water9 and continue to<br />

reproduce as more phosphorus is added.<br />

However, nitrogen and other nutrients must<br />

also be present if algal production is to<br />

contlnue, Recycling <strong>of</strong> nutrients within a<br />

marsh may be sufficient to promote algal<br />

blooms for several years after the initial<br />

loadfng (Sawyer 1954).<br />

Pieters (1893) was the first botanist<br />

to describe the algae <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r<br />

largely from the marshes <strong>of</strong> Anchor Bay.<br />

He noted that over 60 species <strong>of</strong> green<br />

algae, largely desmids, were associated<br />

with the vascular aquatlc plants <strong>of</strong> the<br />

coastal wetlands. W i nner et ale (19701<br />

and the Michigan Water Resources<br />

Commission (1975) have made more recent<br />

studies <strong>of</strong> phytoplankton populations in<br />

Ontario and Michigan nearshore waters<br />

respectively. <strong>The</strong> results <strong>of</strong> these in-<br />

vestigations are summarized in Appendix C.<br />

Diatoms are the most abundant forms <strong>of</strong><br />

pl anktonlc algae in the surface waters <strong>of</strong><br />

the lake. r mr and<br />

are important genera throughout<br />

the water column. Filamentous forms such<br />

as -r are common where sol id substrate<br />

is available, whjle Sp$roqaa is<br />

found in open waters <strong>of</strong> the marshes.<br />

<strong>The</strong> distribution <strong>of</strong> the algal<br />

pigment, chlorophyll gr in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

can be used as an indicator <strong>of</strong> the<br />

re1 ati ve abundance <strong>of</strong> phytoplankton.<br />

Leach (1972) found that starting at the<br />

delta channels, the concentrations <strong>of</strong><br />

chlorophyll g and pheopi gments increase<br />

toward the southeast (Figure 231, reaching<br />

the highest levels (20 pg/l) fn the<br />

marshes at the mouth <strong>of</strong> the Thames River.<br />

Figure 23. Concentration gradient <strong>of</strong><br />

algal pfgments across <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> from<br />

the <strong>St</strong>. <strong>Clair</strong> Delta to the mouth <strong>of</strong> the<br />

Thames River (Leach 1372).


<strong>The</strong> strongest positfve correlat~on was<br />

Found during the spring plankton blooms.<br />

SIl ica also has a strong relationship<br />

to the phytoplankton composftion. De-<br />

pletion <strong>of</strong> sil lca toward the end <strong>of</strong> the<br />

summer greatly reduces the percentage <strong>of</strong><br />

df atoms in the phytoplankton community.<br />

In the southern portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>,<br />

diatom concentration falls from 95% <strong>of</strong> the<br />

total phytoplankton population in May to<br />

10% <strong>of</strong> the population in July. Con-<br />

verse1 y, cyanophytas (b1 ue-greens) i n-<br />

crease from 0% <strong>of</strong> the total population in<br />

May up to 25% <strong>of</strong> the population in July<br />

(Leach 1972).<br />

<strong>The</strong> term periphyton generally refers<br />

to micr<strong>of</strong>loral growth, particuTar1 y<br />

sessile algae, on submerged substrate.<br />

Modifiers are normally used to indicate<br />

type <strong>of</strong> substrate: epipelic - growfng on<br />

sediment; epilithic - growing on rock;<br />

epiphytic - growing on macrophytes; and<br />

epfzooic - growing on animals (Wetzel<br />

1975). <strong>The</strong> periphyton <strong>of</strong> western <strong>Lake</strong><br />

Erie consists <strong>of</strong> predominant1 y 1 lttoral<br />

communitfe~r most commonly associated with<br />

wet1 and vegetation, A greater number <strong>of</strong><br />

species occurs in the littoral zone than<br />

in the llmnetlc zone <strong>of</strong> the lake due to<br />

the greater diversity <strong>of</strong> habitats<br />

available in the nearshore region,<br />

Although no detalled studies have been<br />

done in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlandsr<br />

Millie (1979) examined the epiphytlc<br />

diatom flora <strong>of</strong> aquatic macrophytes fn<br />

marshes along the south shore <strong>of</strong> western<br />

<strong>Lake</strong> Erie. Three common species <strong>of</strong><br />

wetland macrophytes were studied as hosts,<br />

narrow-leaved cattail (w -usti -<br />

white water lily (<br />

and swamp smartweed (<br />

a), 157 were new<br />

ds. Centrlc forms,<br />

were the most common taxa, but each marsh<br />

possessed a dlstfnct flora and<br />

suecessJona1 pattern. Millie attributed<br />

this heterogeneity to the diversity <strong>of</strong><br />

littoral habitats in the marshes*<br />

particularly differences in chemical and<br />

physical factors, Because <strong>of</strong> the nearness<br />

and similarity <strong>of</strong> the <strong>Lake</strong> Erie marshes to<br />

the wetlands along the Ontario shore <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. Claj r, comparable peri phyton<br />

communlties would be expected for both<br />

1 akes,<br />

In Lawrence <strong>Lake</strong>, Michi ganr A1 1 en<br />

( 1971) measured the annual net production<br />

<strong>of</strong> aquatic macrophytes and their attached<br />

algal forms. He found that the epiphytic<br />

algae were responsible for approxf matel y<br />

31% o f the total 1 ittoral production in<br />

the lake. Brock (19701, working in the<br />

Everglades, observed that epiphytf c algae,<br />

rather than macrophytes were responsible<br />

for the majority <strong>of</strong> the primary production<br />

<strong>of</strong> bl adderwort ( sp. 1 com-<br />

munities. Diatoms were found to be the<br />

most abundant form <strong>of</strong> 1 ittoral periphyton,<br />

both in number and biomass, In several<br />

Ontario 1 akes (<strong>St</strong>ockner and Armstrong<br />

1971). Likewiser it is anticipated that<br />

epiphyttc diatoms are an important<br />

component <strong>of</strong> the energy base in <strong>Lake</strong> <strong>St</strong>.<br />

Cl a1 r marshes.<br />

By definition, plankton are floating<br />

organisms whose movements are more or less<br />

dependent on currents. However# some<br />

zoopl ankters exhibit active swimming<br />

movements that aid in maintaining vertical<br />

posltion. Zoopl ankters are diverse in<br />

their feeding habits. Herbivores graze on<br />

phytoplankton, periphyton, and macrophytes<br />

within the coastal marshes. Carnivores<br />

prey on attached protozoans and other<br />

zooplankters, while omnivores feed at all<br />

trophlc levels. In turn, zooplankton are<br />

important fish and waterfowl food. Every<br />

fish species and many duck species<br />

urilizing the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands eat<br />

zooplankters during some portion <strong>of</strong> the1 r<br />

lffe cycle.<br />

<strong>The</strong> anlmal components <strong>of</strong> the wetland<br />

plankton (Figure 24) consist <strong>of</strong> three<br />

groups: protozoans, rotifers (Figure 251,<br />

and microcrustaceans (primarily<br />

cl adocerans and copepods) . Common zoo-<br />

pl ankters, including sessile or periphytic<br />

protozoans# occurring fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

wetlands are listed in Appendix D.<br />

<strong>The</strong> first study <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

zooplankton, and one <strong>of</strong> the earl lest<br />

1 imnol ogical investigations in the Great<br />

<strong>Lake</strong>s, was organized by Refghard (1894)<br />

and included work by many <strong>of</strong> the leading


Figure 24. Mean distribution and concentrations <strong>of</strong> zoo-<br />

plankton groups in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> for April through Novem-<br />

ber 1972 (Leach 1973). Note correlation between total<br />

zooplankton population and a1 gal pigments.


CARFA ENLARGED /<br />

Figure 25. Distribution <strong>of</strong> total roti fers and most abundant genus,<br />

Brachionus, in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> during July-September 1973 (Bricker et al.<br />

" m N b t e high concentration in the vicinity <strong>of</strong> the Clinton River Cut<strong>of</strong>f<br />

Channel.<br />

specfalists <strong>of</strong> that perlod. Species lists<br />

<strong>of</strong> Protozoa (Smfth 18941, planktonic<br />

Rstlfera (Jennings 18941s Gladocera (Bi rge<br />

1894)r and Copepoda [Marsh 1895) were<br />

compiled from samples collected among<br />

aquatic plants fn Anchor Bay and at other<br />

nearshore stations along the west shore <strong>of</strong><br />

take <strong>St</strong>, Clai r, Planktonic organ1 sms<br />

remained negl scted untll Winner et a1 .<br />

(2970) obtained zooplankton data from two<br />

estuary mouths on the Canadian shore.<br />

Leach ( 1973 ) col lected zooplankton samples<br />

from the South Channel <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

Daita* Mitchell Bay* the estuary mouth <strong>of</strong><br />

the Thames Rivsrr and other nearshore and<br />

<strong>of</strong>fshore stations on <strong>Lake</strong> <strong>St</strong>. Clalr.<br />

Brfcker et a7, (1976) documented the zoo-<br />

plankton populatfons <strong>of</strong> Anchor Bay, the<br />

nearshore region in the vicinity <strong>of</strong> the<br />

Clinton River estuary* and <strong>of</strong>fshore re-<br />

gions <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> results <strong>of</strong><br />

these collections are included in Appendix<br />

0.<br />

<strong>The</strong> distribution <strong>of</strong> zooplankton in<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is largely dependent on the<br />

prevailing currents <strong>of</strong> the lake. Other<br />

inf l oences, such as temperature, algal<br />

abundance, and chemical characteristics<br />

a1 so determine the concentratlons <strong>of</strong><br />

zooplankton. Because these factors vary<br />

greatly throughout the lake, the<br />

distributions <strong>of</strong> zooplankton also vary<br />

consf derabl y in different portlons <strong>of</strong> the<br />

lake (Bricker et al. 1976). A persistent<br />

eddy structure In the southeastern portion


<strong>of</strong> the lake (Figure 141 promotes<br />

entrajnrnent <strong>of</strong> the waters and allows<br />

nutrient and algal levels to increase<br />

above other portions <strong>of</strong> the lake (Leach<br />

1972). Because <strong>of</strong> the high phytoplankton<br />

levels in this area, the zooplankton<br />

biomass is greater here than in any other<br />

regton, <strong>The</strong> rapid flow conditions <strong>of</strong><br />

water through the delta and the Michigan<br />

portion <strong>of</strong> the lake inhibits any great<br />

concentration <strong>of</strong> zooplankton; therefore,<br />

the biomass is greater on the Ontario side<br />

(Figure 24). Despite rapid flow rates,<br />

zooplankton biomass is elevated near the<br />

Cl lnton Rfver Cut<strong>of</strong>f Canal (Figure 25) due<br />

to nutrient loading from the watershed<br />

(Bricker et al, 1976).<br />

Because <strong>of</strong> the rapid flushing rate <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (9 days) and the dense<br />

subrnergent macrophyte populations competing<br />

for available nutrients, the total<br />

biomass <strong>of</strong> plankton in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is<br />

low. Zooplankton that succeed the best in<br />

this environment are those that have a<br />

short 1 ife span and a fast turnover rate.<br />

For that reason* rotlfers are the most<br />

abundant forms <strong>of</strong> zooplankton found. Mean<br />

numbers <strong>of</strong> rotifers ranged from<br />

approximately 62/1 in July and 69/1 In<br />

August down to 10/1 in September (Leach<br />

1973 1 . <strong>The</strong>se organisms are thirty times<br />

more abundant at the Clinton River Cut<strong>of</strong>f<br />

Canal (Bricker et a1. 1976). Crustaceans<br />

have a consistently low abundance in the<br />

lake ranging from a high <strong>of</strong> around 6/1 to<br />

a low <strong>of</strong> 0.3/1. As with rotifers, the<br />

peak abundance occurs in July and the low<br />

occurs in September. Cyclopoid and<br />

ca1 anoi d copepods, and cl adocerans (water<br />

fleas) are the predominant microcrustaceans<br />

found in the lake (Figure 24).<br />

Protozoans vary fn abundance but have the<br />

second highest occurrence following the<br />

rot1 fers. <strong>The</strong>y compose an average <strong>of</strong> 37%<br />

<strong>of</strong> the total zooplankton communlty.<br />

3.2 WETLAND VEGETATltON<br />

Although wetland communities exist in<br />

the <strong>St</strong>. <strong>Clair</strong> River (Figure 261, Clinton<br />

RJver, Marsac P<strong>of</strong>ntt Swan Creakp and <strong>St</strong>,<br />

<strong>Clair</strong> Delta complexes, only the latter<br />

area has been w'ell studied. <strong>The</strong>refore?<br />

the description <strong>of</strong> the pl ant communities<br />

Figure 26. Ontario shore1 ine; <strong>of</strong> <strong>St</strong>. <strong>Clair</strong><br />

River at Cathcart Park (August 1984),<br />

herein is based on the <strong>St</strong>, <strong>Clair</strong> Delta<br />

wet1 andsr particular7 y those on DJck f nson<br />

Is1 and. Important 1 iterat~re sources, by<br />

wetland area, are: <strong>St</strong>. John" Marsh<br />

(Roller 19761, Dfckinson Island (Jaworski<br />

et al. 1979, 19813, Canadian sfde <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> Delta (Eayly 1975)? and the<br />

enti re del ta complex ( Jaworski and Raphael<br />

1978; Raphael and Jaworskf 1982). Lyon<br />

(1979) mapped the wetlands communfties on<br />

the Michigan side <strong>of</strong> the <strong>St</strong>. C1 alr Delta<br />

using the wetlands classfficatfon <strong>of</strong> the<br />

U, S, Fish and Wildlife Service and a map<br />

scale <strong>of</strong> 1:24b000, Hayes (1964) surveyed<br />

the plant communities <strong>of</strong> a wet prairle on<br />

Harsens Is1 and,


Several distinct environmental types*<br />

or pl afit zones, have been identified<br />

wfthin the <strong>St</strong>, <strong>Clair</strong> Delta wetland complex<br />

located in northeastern <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

<strong>The</strong>se zones are part <strong>of</strong> a vegetation<br />

continuum (Figure 27). For example, on<br />

the 11.3 km2 Dickinson Island* the plant<br />

comrnuniPles grade from cattafl marsh along<br />

the southwest shore to upland hardwoods on<br />

the northeast point. <strong>Lake</strong> level<br />

fluctuations control the development and<br />

areal extent <strong>of</strong> the various zones<br />

(Jaworski and Raphael 1978). <strong>The</strong><br />

important aquatic macrophytes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> are 1 isted in Appendix Er and<br />

distribution maps <strong>of</strong> several species are<br />

presented In Figure 28,<br />

1.5 me Certain species, such as the<br />

cattails and the bulrushes can withstand<br />

moderate wave energy. Common vascular<br />

pl ants Include the following specfes:<br />

(eel grass,<br />

(pickerel weed)<br />

(yellow water Illy)<br />

water smartweed)<br />

(Eurasian<br />

(hybrid cattail)<br />

(hard-stem bul rush)<br />

(three-square<br />

(sago<br />

pondweed)<br />

Char9 sp. (muskgrass or stonewort).<br />

This type includes plant communities<br />

such as those present in Goose Bay,<br />

Gooseneck Pond <strong>of</strong> Fisher Bay, and Little<br />

Muscamoot Bay. Species are largely sub- LOW* narrow beaches <strong>of</strong> fine sand and<br />

merged and emergent aquatic macrophytes. stands <strong>of</strong> emergent vegetation (Ff gu r e 29)<br />

Habitats consist <strong>of</strong> low-wave energy lake are found on the Canadian sfde <strong>of</strong> the<br />

bottoms and nearshare environments <strong>of</strong> bays delta. <strong>The</strong> sand extends down less than 1<br />

where the water depths range from 0.3 to m and exhibits alternatqng layers <strong>of</strong> clean<br />

.3<br />

SW.<br />

July 1972 dlarcs~ooi Bay<br />

DOGWQOR<br />

MCAROW SE THREE<br />

E SQUARE BULRUSH<br />

-IY 0<br />

0<br />

5<br />

-5 +-- T-. r-- ?--I<br />

0<br />

ASPEN<br />

2 00<br />

V € r 20 V+(l.letbw notfc rtal*<br />

--r---7. - - - ,-<br />

8<br />

400 LOO 800 1W0 3600 3800<br />

"71-__7__7,-- T---- --<br />

FEET<br />

October 1974<br />

FEET<br />

I<br />

40W<br />

BULRUSH<br />

- --- 0<br />

---..<br />

---- 5<br />

1000 3600<br />

Figure 27. Vegetation transects on Harsens Island, <strong>St</strong>. <strong>Clair</strong> Delta, showing impact <strong>of</strong><br />

rising water levels (Jaworski and Raphael 1976).<br />

4%


MICHIGAN<br />

ANCHOR BAY<br />

PORT<br />

HUROiL<br />

MICHIGAN<br />

ANCHOR BAY<br />

ONTARIO<br />

/ ST. CLAlR S<br />

MICHIGAN<br />

PORT<br />

HURON<br />

Potarnogeton crispus Potanlogeton ri chardsoni i Butornus umbel 1 atus<br />

(curly pondweed) (Richardson's pondweed) (flowering rush)<br />

PORT HURsEU<br />

MICHIGAN hllCHlGAN MICHIGAN<br />

ANCHOR BAY<br />

Val 1 i sneria americana<br />

(wil d celery)<br />

ANCMOR BAY<br />

Heteranthera dubia<br />

(water stargrass)<br />

Myriophyl lum spicatum<br />

(water-mil foil )<br />

Figure 28. Distribution <strong>of</strong> six aquatic macrophytes in the <strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>-Detroi t River ecosystem (Schloesser and Manny 1982, unpubl ished data supplied by<br />

R. L. <strong>St</strong>uckey).


Fiaura 29. Emeraent stand <strong>of</strong> reed qrass<br />

(fhragmi tes austbl is) a1 ong he ma tog an<br />

Channel, Wal pole Island, Ontario (August<br />

sand and organic-rich sand or peat.<br />

<strong>St</strong>randed or transgressive, beach ridges<br />

are low, sandy features on the delta<br />

Islands that can be up to 100 m wide.<br />

Normally these ridges are only slightly<br />

above the water level. Characterfstfc<br />

vegetation <strong>of</strong> these sandy habltats<br />

Includes the following vascular species:<br />

bindweed).<br />

( staghorn sumac)<br />

(touch-me-not,<br />

(swamp thistle)<br />

This marsh type is prevalent within<br />

abandoned channels and fn cattail marshes<br />

where open water and sandy sediments<br />

occur. Bulrushes are also common In<br />

shallow water along the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />

shore and in bays, as in Anchor Bay.<br />

Water depths range from 0.6 to 1.2 m.<br />

Representative plants include the<br />

followfng species:<br />

Scir~us acut~ (hard-stem bul rush)<br />

Ce~ha1 anthyg occidental I 5<br />

(buttonbush)<br />

Char3 sp. (muskgrass or stonewort)<br />

Various emergent and submergent<br />

species.<br />

Cattai 1 Marshes<br />

<strong>The</strong>se marshes are broad zones located<br />

on the lower portions <strong>of</strong> the delta islands<br />

where water depths exceed 15 cm. <strong>The</strong>y are<br />

also found on inundated shoulders <strong>of</strong> river<br />

channels and bottoms <strong>of</strong> shallow ponds and<br />

bays. Pure colonies <strong>of</strong> cattails (lyebq x<br />

are associated with peaty or<br />

clayey sediments (FJgure 30). In small,<br />

shall ow open i ngs in the cattall marshes,<br />

duckweeds (hmna minor and SDI rode1 q<br />

~glyrhlzg), water-mil foils (Nvrio~t).ylJm<br />

al_tecniflorum and d % ~icatum), and<br />

bl adderwort (Ib;triculariq. vulgari~) are<br />

abundant.<br />

Sedae Marshes<br />

Thfs mdian type comprises a narrow<br />

zone between the cattail marsh and the<br />

grassy meadow or the dogwood shrub zones.<br />

Sedge mavshes are also found along river<br />

channels* at the base <strong>of</strong> old shore1 ines<br />

now stranoed in cattail marshes, and along<br />

the edge <strong>of</strong> eroding lake and bay<br />

shorelines. <strong>The</strong>se marshes form on<br />

occasional 1 y flooded wet sites where<br />

permanent water depths do not exceed 15<br />

cm. Common species present include the<br />

fol lonring:<br />

ana ad ens is (bluejoint<br />

mex strictq (tussock sedge)<br />

!2E.S% (tussock sedge)<br />

!a% (tussock sedge)<br />

1 anuainosq (tussock sedge)<br />

Carex sartwell i I (tussock sedge)


Figure 30. Emergent stands <strong>of</strong> narrow-<br />

leaved cattail (Typha angustifolia) on<br />

Little Muscamoot Bay, Harsens Island,<br />

Michigan (August 1984).<br />

So1 a w<br />

gfffcinale$ (common comfrey)<br />

(nightshade).<br />

iN&&u@m<br />

<strong>The</strong>se habitats include artificial<br />

canals and openings in the marshes<br />

(ponds), Water depths are general 1 y<br />

shallow, 0.3 to 0.6 m, and sediments are<br />

usually clayey or organic. Communities<br />

can be quite variable. Typical species<br />

fnclude the following:<br />

ellow water lily)<br />

(white water lily)<br />

Lemna minor (duckweed)<br />

Wederi g 5;ordata (pickerel weed)<br />

,- ~anadens-ls (waterweed)<br />

mjgm (water smartweed)<br />

Potamoael;on 5:ris~uq (curly pondweed)<br />

Potamq&&Qn sp~. (oondweeds)<br />

Ce~hal anthus o~cid&ntal is<br />

(buttonbush)<br />

W sp. (muskgrass or stonewort)<br />

C1 ildo~hora sp. ( f il amentous green<br />

algae).<br />

Abandoned distributary channels in<br />

the delta are generally shallow, less than<br />

1 m deep, and have silty or peaty<br />

sediments. <strong>The</strong> variable plant communities<br />

include the following species:<br />

Nuohar advena (yellow water Illy)<br />

$uberosa (white water lily)<br />

M~rioptydJm<br />

--<br />

tr!lterniflor~ (little<br />

water-mll foil<br />

Wttaria latifolfa (common<br />

arrowhead)<br />

,cutus (hard-stem bulrush)<br />

Sci rpus americanus (three-square<br />

bulrush)<br />

(buttonbush).<br />

This type 4s a zone <strong>of</strong> transition<br />

vegetation or ecotone, between the sedge<br />

marshes and the hardwood community. This<br />

zone 1 fes sl ightly above the water table<br />

and is infrequently flooded. <strong>The</strong><br />

communities consist <strong>of</strong> a mixture <strong>of</strong><br />

grasses, herbs and shrubs, and water-<br />

tolerant trees, including the fallowing<br />

species:<br />

po~ul uz Qemuloide~ (quaking aspen)<br />

(red ash)<br />

d osier<br />

~alustrSs (swamp rose)<br />

Sol i &gg spp. (goldenrods)<br />

sp. [panic grass)<br />

(rattlesnake


Carey strictp (tussock sedge)<br />

wamp mi 1 kweed)<br />

ush)<br />

(marsh fern)<br />

sf 1 verweed) ,<br />

This type also represents a tran-<br />

sition into up1 and hardwood vegetatfon.<br />

<strong>The</strong> depth to the water table is normally<br />

0.5 to 1.0 m in this zone. Communities<br />

are mlxed shrubsr water-tolerant treesr<br />

and some understory plants typical <strong>of</strong> the<br />

meadows. Prominent plants include the<br />

following species:<br />

1- (eastern<br />

cottonwood<br />

PODU~Y+ tremu181das. (quaking aspen)<br />

Fraxiw Pennsvlrnnca (red ash)<br />

Gornus ~tolonifera (red osier<br />

dogwood<br />

Qxnuj racemosa (gray dogwood)<br />

oalmata (wlld grape)<br />

sp. (hawthorn).<br />

During low-water periods, these shrubs and<br />

swamp trees invade the sedge marshes.<br />

However, during high water condi tions, as<br />

fn the 1970~~ flooding and dle back <strong>of</strong><br />

woody plants in the shrub zone are common.<br />

<strong>St</strong>ands <strong>of</strong> oak-ash hardwoods occur in<br />

the upper portion <strong>of</strong> Dickinson, Harsens,<br />

<strong>St</strong>. Anne, Squirrel, and Walpole is1 ands at<br />

elevations ranging from 1 t o 3 m above<br />

mean lake level. <strong>The</strong> major species <strong>of</strong><br />

these stands are Eraxinus eann+~lvanica<br />

(red ash) and (swamp white<br />

oak), Assocfated hardwoods occurring In<br />

this type include the following species:<br />

American elm)<br />

<strong>The</strong> wetland plant communities <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> Delta occur in broad, arcuate<br />

zones which extend from the vlcinity <strong>of</strong><br />

Algonac* Mlchfgan, southward into <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> (Figure 311, In spfte <strong>of</strong> extensive<br />

cultural mod1 fications, sufficient natural<br />

wetland vegetation remains to reconstruct<br />

the communlty zonation, On the various<br />

islands <strong>of</strong> the delta, five major<br />

macrop hyte communities can be mapped.<br />

<strong>The</strong>se communitfest from the driest to the<br />

wettest habitats, include: oak-ash<br />

hardwoods, dogwood meadow9 sedge marsh,<br />

cattail marsh, and bulrush marsh. Other<br />

fdentlfiable communities which are 1 imited<br />

In extent include: canal-pond-abandoned<br />

channel, lakeshore and bay bottom8 and<br />

reed grass communities.<br />

<strong>Lake</strong>ward <strong>of</strong> the hardwoods i s a<br />

complex <strong>of</strong> communfties referred to as the<br />

dogwood meadow. This vegetation zone<br />

occurs as a transftion zone between the<br />

hardwood forest and the wetter sedge<br />

marsh. It appears that periodic high lake<br />

level and human disturbance (such as<br />

burning and mowing) prevent woody species<br />

from dominating this zone. Major plant<br />

species herein are bluejoint grass<br />

(ww -1, s<strong>of</strong>t rush<br />

(m gffusus) , gray dogwood, red os i er dogwood, and quakfng aspen.<br />

<strong>The</strong> sedge marsh forms a narrow zone<br />

between the dogwood meadow and the cattail<br />

marsh, as well as along eroding lake and<br />

bay shorel ines, particularly on the<br />

Michigan side <strong>of</strong> the delta. This<br />

community also occurs along lower dlstributary<br />

channels and on old shorel i nes.<br />

<strong>The</strong> sedge marsh is associated with the<br />

alkaline, inorganic soils which are either<br />

saturated or occasional ly flooded.<br />

However, permanent Inundation <strong>of</strong> the sedge<br />

tussocks results in dieback <strong>of</strong> the sedges.<br />

Major plant species are sedges <strong>of</strong> the<br />

genus Carex and bluejoint grass. Carex<br />

stricta is particularly abundant on<br />

Dickfnson Island, whereas L lacustri~ is<br />

common on Walpole Island,<br />

Located on the lakeward margin <strong>of</strong> the<br />

delta, where water depths exceed 15 to 30<br />

cm, is the broad cattail marsh zone.<br />

Zones <strong>of</strong> cattail also occur on inundated<br />

shoulders <strong>of</strong> the river channels, on lower<br />

ends <strong>of</strong> distributary channels, and on<br />

crevasses. Pure stands <strong>of</strong> cattail are<br />

assocfated with peaty or clayey sediments.<br />

<strong>The</strong> hybrid cattail (TyDha x glauc(a)r which<br />

attafns 2 to 3 m in height and grows in


ANCHOR BAY<br />

a OAK-ASH<br />

HARDWOOD<br />

DOGWOOD MEADOW<br />

0 DREDGED DISPOSAL<br />

0 CULTIVATED LAND<br />

Figure 31. Wetland plant communities <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta (Raphael and Jaworski<br />

1982).<br />

53


circular colonlesP is the domlnant<br />

species. Some narrow-leaved (L,<br />

and broad-1 eaved cattall s<br />

1 can also be identified.<br />

<strong>The</strong> bulrush marsh i s found in<br />

abandoned river channels, in 1 arge open-<br />

water areas <strong>of</strong> the cattafl marsh* and on<br />

sand bars in low-wave energy areas <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and Anchor Bay. Colonies <strong>of</strong><br />

bul rush are associated with sandy<br />

substrates and water depth <strong>of</strong> 0.5 to 1,25<br />

m. <strong>The</strong> prfncipal species <strong>of</strong> the marsh is<br />

hard-stem bulrush. Buttonbush i s<br />

associated with the bulrush community in<br />

abandoned distributary channels where some<br />

water flow occurs.<br />

In canals, abandoned river channel sr<br />

marsh open1 ngs, and other stagnant water<br />

environments, a variety <strong>of</strong> floating and<br />

submersed aquatic communities occur.<br />

Water depths range from 0.24 to 1.0 m and<br />

water circulation is minimal except for<br />

wind-driven currents. Organic substrates<br />

are common along with turbidity due to<br />

humlc acids and suspended particulate<br />

organic matter. Whereas species diversity<br />

fs the ruler abundant plant species<br />

include duckweedsp pickerel weed,<br />

waterweed, water-mllfoll, and water<br />

smartweed. Other associated species are<br />

whfte water lily<br />

<strong>Lake</strong> and bay-bottom communities occur<br />

along low wave energy bottoms and<br />

nearshore anvi ronments such as Little<br />

Muscamoot Bay. Water depths generally<br />

range between 0.3 and 2.0 m, Major<br />

species in this zone include wild celery,<br />

muskgrass or stonewort (w sp,), and<br />

flexed naiad. Brown (1983) reported the<br />

dominance <strong>of</strong> along with pondweeds,<br />

Eurasian wat foil, waterweed* and<br />

varfous emergenks growfng about Sand<br />

Island In Anchor Bay. Because <strong>of</strong> higher<br />

wave energy, lake and bay-bottom<br />

communities are less widespread on the<br />

Canadian slde <strong>of</strong> the delta. In <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>, Schloesser and Manny (1982) found<br />

few macrophy%esp which excludes<br />

, beyond 1,25 km from shore.<br />

Although conspicuous where pressnt,<br />

the feed grass community I s <strong>of</strong><br />

insufficient dJstrlbutfon to map in the<br />

<strong>St</strong>. C1air Delta. Dense colonies <strong>of</strong> reed<br />

grass ( )I usually less<br />

than 0.5 hectares in extent* are located<br />

In scattered patches on the drowned<br />

shoulders and natural levees <strong>of</strong> North,<br />

Middle, and South channels.<br />

Wetland vegetation is not static from<br />

year to year (Section 4.3) . Dynamic<br />

change occurred In the plant cornrnunitles<br />

during the interlm 1964 and 1975 as water<br />

levels In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> fluctuated from<br />

very low levels to extreme high water<br />

conditions, Dicklnson Island, which has a<br />

full complement <strong>of</strong> wetland zonest and<br />

except for the dredged material disposed<br />

in the early 1970~~ had 1 i ttle development<br />

(Figures 32 and 33). In particular, large<br />

shifts occurred among the bulrushsubmersed<br />

commun ! ties and the emergent<br />

sedges and cattai 1 s.<br />

A literature search by Herdendorf et<br />

a1. (1981~) revealed no plant species in<br />

the <strong>St</strong>. Clatr Delta complex which appeared<br />

on Federal or <strong>St</strong>ate lists <strong>of</strong> endangered<br />

species. However, there Is a very small<br />

patch <strong>of</strong> the rare plant, wild rice<br />

) located on lower Hareover,<br />

Wagner (1977)<br />

reported the fo1 low1 ng threatened vascul ar<br />

plant species (Michigan) for Harsens<br />

Island: marsh sedge ( ~ i s t v l i s<br />

auberul.a) frtnged orchid<br />

(Habenarln 1, Hill's thi stle<br />

(CIrs1~ hi11 ii) , panrc grass (Panicurn<br />

JelbergjJ,), and sand grass (m<br />

ourouraa) *<br />

In 1976 and 1977, the U. S. Fish and<br />

Wildlife Servfce (Hfltunen 1980; Hfltunen<br />

and Manny 1982) conducted surveys <strong>of</strong> the<br />

macrozoobenthos <strong>of</strong> the lower <strong>St</strong>. Clafr<br />

River and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>se surveys<br />

Jndicatad that the <strong>St</strong>. <strong>Clair</strong> Rfver and<br />

Delta support a dfverse and abundant<br />

macrozoobenthos. <strong>The</strong> number <strong>of</strong> taxa and<br />

densfty found fn the river system was


signJficantly greater than that found in<br />

Anchor Bay or the open portions <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>, Cl afr (Table 181,<br />

In the <strong>St</strong>. <strong>Clair</strong> River and Delta,<br />

oligochaete worms are the most abundant<br />

macrozoobenthos (55%). Sixteen species<br />

are commons with the tubif icid genera<br />

and<br />

d the naidld genera and<br />

dominating. Dfpteran 1 arvae<br />

are the second most abundant group (26%).<br />

By number, the chironomids compose 95% <strong>of</strong><br />

the dipterans. Crustaceans are among the<br />

less abundant forms and are represented by<br />

the amphipods <strong>of</strong> the four genera,<br />

Gamm.ar_us* my !Zmgaux, and<br />

Ponto~oreia. <strong>The</strong> first three are<br />

permanent residents <strong>of</strong> the river wetlands;<br />

the fourth populates the deep waters <strong>of</strong><br />

the Great <strong>Lake</strong>s and is probably transient<br />

from <strong>Lake</strong> Huron. <strong>The</strong> insect orders,<br />

Trichoptera and Ephemeroptera together<br />

compose on1 y 1% to 2% <strong>of</strong> the total number<br />

<strong>of</strong> macrobenthos. Caddisfl ies are not as<br />

numerous as the mayflies, but are<br />

represented by a greater number <strong>of</strong> genera.<br />

<strong>The</strong> mayfl fes are 1 argel y Nexaaen l a (90% 9.<br />

Although the density <strong>of</strong> mayflies and<br />

caddisfl ies Is low, these organ isms are a<br />

significant part <strong>of</strong> total fauna because<br />

the biomass <strong>of</strong> their populations is<br />

greater than that <strong>of</strong> other insect groups<br />

(Hi 1 tunen 1980).<br />

<strong>The</strong> richness <strong>of</strong> the benthic fauna <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is evident in the high<br />

diversity <strong>of</strong> taxa present in the wetlands<br />

<strong>of</strong> Anchor Bay and elsewhere in the lake<br />

(Table 181. <strong>The</strong> moderate abundance <strong>of</strong><br />

01 igochaetes as compared to other groups<br />

indicates the quality <strong>of</strong> the benthic<br />

environment is high. Hi ltunen and Manny<br />

(1982) found the mean percent composition<br />

<strong>of</strong> the pol lution-tolerant 01 igochaetes<br />

ranged from 25% in Anchor Bay to 49%<br />

elsewhere in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> values<br />

are far below the standard <strong>of</strong> 60%<br />

01 igochaetes in pol 1 uted environments<br />

proposed by Goodnfght and Whit1 ey (1960).<br />

<strong>The</strong> presence <strong>of</strong> pollution-intolerant<br />

Ephemeroptera and Trichoptera in<br />

substantial numbers (300/m2) at a<br />

nearshore station in Anchor Bay 3 supports<br />

the conclusion that the benthic<br />

environment <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr is not<br />

severe1 y impai red by pollutjon (Hiltunen<br />

and Manny 1982).<br />

Macrozoobenthos constitutes an<br />

important source <strong>of</strong> food for fish and<br />

waterfowl in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-<strong>St</strong>. Claf r<br />

River wetlands. Price (19631, working In<br />

western <strong>Lake</strong> Erier showed that rainbow<br />

smel tr alewives~ glzzard shad, spottail<br />

shiners* trout-perch, yellow perch,<br />

channel catf ishr white bass, and walleye<br />

feed heavily at different life stages on<br />

01 igochaetesr leeches* cladocera, ostra-<br />

cods, amphlpodsr mayfl less caddisfl iesr<br />

rnf dges, snails, and bivalves. Although<br />

1 fttle published work exists on the food<br />

habits <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> River fishes, the<br />

aforementioned fish species and the<br />

invertebrates that they feed upon are all<br />

abundant In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>.<br />

<strong>Clair</strong> River (Hiltunen 1980). Dawson<br />

(1975) has observed that many <strong>of</strong> the<br />

macrozoobenthos reported for Anchor Bay<br />

are also important sources <strong>of</strong> food for<br />

waterfowl inhabiting that bay. A list <strong>of</strong><br />

the important benthic invertebrate species<br />

<strong>of</strong> the <strong>St</strong>. Clai r River-<strong>Lake</strong> <strong>St</strong>. Cleir<br />

ecosystem is presented in Appendix F.<br />

Duri ng four different stages <strong>of</strong> the<br />

Pleistocene Epoch, which itself lasted<br />

from about one million to about ten<br />

thousand years agor the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />

basin was covered with glacial ice.<br />

Within the glaciated region all benthic<br />

organ1 sms were destroyed, but each time<br />

the ice receded they reinvaded the<br />

previously ice-covered region. <strong>The</strong><br />

Pleistocene mollusks <strong>of</strong> the intsrgl acial<br />

lakes and their paleoecology are well<br />

documented by La Rocque (1966). Many<br />

species, particularly the unioni d bivalves<br />

and the 1 arge prosobranch snails8 require<br />

a continuous waterway for mfgration.<br />

Glochidia may be carried over long<br />

distances while attached to their fish<br />

host. <strong>The</strong> rich molluscan fauna <strong>of</strong> the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> region was largely<br />

repopulated from the Ohfo-Mississippi<br />

system when glacf a1 meltwaters in this<br />

region flowed south. Additf onal ly, Clarke<br />

(1981) speculated that most small snails<br />

(pulmanates) and small clams (sphaeri fds)<br />

are carried about imbedded fn the feathers<br />

<strong>of</strong> water birds, in mud attacked to their<br />

feet or clamped to the appendages <strong>of</strong><br />

1 argep flying aquatlc insects. Appendix G<br />

provides a lysting <strong>of</strong> the mollusks


0 5000 Feet<br />

t=TFL-r-e 7--rlf=74<br />

0 1500 Meters<br />

Figure 32. Wetland vegetation on Dickinson Island in 1964.<br />

L-3 Developed<br />

. .<br />

assoclated with the coas.ta1 wetlands and descended from land snails, must come to<br />

nearshore waters. Append fx H contains the surface periodf cally for air.<br />

Information on the hab 1 t at preference <strong>of</strong><br />

these spectes.<br />

Most aquat tc snalls are vegetarians.<br />

<strong>The</strong> veneer <strong>of</strong> livfng algae which covers<br />

most submerged surfaces is their chief<br />

We1 ?-vegetated portions source <strong>of</strong> food, but dead plant and animal<br />

<strong>of</strong> unpolluted embayments* marshes, beach materjal is frequently ingested.<br />

ponds, and slugglsh tributary mouths are Dissolved oxygen is an Important Ifmiting<br />

the most productJve localities for factor; most gilled species require high<br />

freshwater snails Jn <strong>Lake</strong> <strong>St</strong>. Clafr. <strong>The</strong>y concentratfons. w<strong>St</strong>h limpets such as<br />

Ilve an submerged vegetation* on rocks and mrissb being found only where the water<br />

on the bottom at the water's edge and out remains near saturation (Pennak 1978).<br />

to a depth <strong>of</strong> several meters. Two However, -1omp dec.ls_umr and Amnlico'ia<br />

subclasses, pro sob ranch^ and Pulmonata, l.imasa have been col lected in water with<br />

are we1 1 represented In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> less than 2 ppm oxygen (Harman 1974). <strong>The</strong><br />

coastal marshes. <strong>The</strong> former group 1s concentration <strong>of</strong> dissolved sol Ids in <strong>Lake</strong><br />

characterized by internal re y <strong>St</strong>. <strong>Clair</strong>, particular1 y bicarbonate<br />

gJlls (~tenidia)~ or as in , alkalinity at over 80 mg/l, provfdes<br />

external gflls, and an oP@rcuIum used to adequate essential materials for shell<br />

seal the shell aperaturee <strong>The</strong> latter construct;Son. Appendix W shows the<br />

group does not have gfllst but obtajns habitat preferences for the 47 species <strong>of</strong><br />

oxygen through a riIung-like" pulmonary gastropods which have been reported for<br />

cavjty, Pul monate snaf 1 s p whlch have the coastal marshes and Rearshore waters.<br />

56


Shore1 ine<br />

- 0 5000 Feet<br />

D<br />

0 1500 Meters<br />

Bul rush-Submersed (sparse)<br />

Cattails (some die back)<br />

a Dogwood Meadow<br />

Woodlands<br />

0 Developed<br />

Disposal Site<br />

Figure 33. Wet1 and vegetation on Dicki nson Island in 1975. Previous shore1 ine refers<br />

to 1964 shore shown in Figure 32.<br />

Dl ec- <strong>The</strong> bivalved molluscan<br />

fauna <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> consists <strong>of</strong> three<br />

families. <strong>The</strong> majority <strong>of</strong> the species<br />

belong to the Unionidae (freshwater<br />

mussels or naiades) and Sphaerlidae<br />

(fingernail clams). <strong>The</strong> third family,<br />

Corbicul fdae (1 fttle basket clams), fs<br />

represented by an f ntroduced Asiatic<br />

species, Bivalves are most abundant<br />

nearshore* especially in water less than 2<br />

m deep, <strong>St</strong>able gravel and sand substrates<br />

with a good current support the largest<br />

populations. Common1 y mussels inhabit<br />

substrates free <strong>of</strong> rooted vegetat i on, but<br />

there are numerous exceptionsv i nc1 udi ng<br />

Anodonta arandis and<br />

<strong>St</strong>omach contents <strong>of</strong> unionid mussels<br />

are commonly mud, desmidsv dfatoms, and<br />

other unicellular a1 gae, protozoans*<br />

rotifers, flagellates, and detritus. <strong>The</strong><br />

1 argest populations <strong>of</strong> mussels develop<br />

below areas where disintegration <strong>of</strong> rich<br />

vegetation is occurring (Churchill and<br />

Lewis 1924).<br />

<strong>The</strong> female pocket-book mussel<br />

(Wsflb ysntrfcc&a) is capable <strong>of</strong><br />

extending and pulsatl ng the posterior edge<br />

<strong>of</strong> the mantle fn such a way that it<br />

resembles an injured mf nnow (Clarke 1981).<br />

This activity attracts fish such as<br />

bluegill, white crappje, smallmouth bass,<br />

and yellow perch. and Increases the<br />

opportunitf es for juvenile mussels<br />

(glochidia) to attach themelves to a fOsh<br />

after they have been ejected from the<br />

parent. <strong>The</strong> larvae are released by the<br />

parent when its light rsnsftlve spots are<br />

stimulated* for example, by the shadow <strong>of</strong><br />

a passing fish. %veral unionid blvalves<br />

possess special mantle structures adapted<br />

to lure fDsh their vicinity. <strong>The</strong><br />

glochidia <strong>of</strong> each species <strong>of</strong> freshwater<br />

mussel (correspo~ds Oa vel dgsr larvae af<br />

marine bivalves) musk attach to the gills


Table 18. Macroziobenthos <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the lower <strong>St</strong>. <strong>Clair</strong> River<br />

and Delta (1977).<br />

Major<br />

taxa Compos<strong>St</strong>ion Density Composition Density Composition Density<br />

(% (no./m2) (8) ( no./mZ) (%I (no./m21<br />

Nemert i nea<br />

Nematoda<br />

Hi rund i nea<br />

01 igochaeta<br />

Polychaeta<br />

Amphipoda<br />

Isopoda<br />

Diptera<br />

Ephemeroptera<br />

Col eoptra<br />

Lepldoptera<br />

Trlchoptera<br />

Hydracarlna<br />

Gastropoda<br />

Pel ecypoda<br />

Totals<br />

a Data sources: Hiltunen (19801, Hiltunen and Manny (1982).<br />

b- Absent<br />

+ Present but not enumerated.<br />

and fins <strong>of</strong> a particular fish species or<br />

small group <strong>of</strong> species (Table 19) before<br />

further development can take place, Most<br />

glochidia never accomplish this, but those<br />

that do succeed remain attached for a few<br />

weeks as they metamorphose into tiny<br />

mussels. <strong>The</strong> young mussels then drop from<br />

the fish to take up an independent lSfe on<br />

the lake bottom, moving about and<br />

siphoning water for respiration and to<br />

obtain plankton as a source <strong>of</strong><br />

nourishment, Appendix H indicates the<br />

habitat preferences <strong>of</strong> the 64 species <strong>of</strong><br />

pelecypods which have been reported for<br />

coastal marshes and nearshore waters.<br />

3.4 FlSH<br />

<strong>Lake</strong> <strong>St</strong>. ClaJr, along with the <strong>St</strong>.<br />

<strong>Clair</strong> Rfver and the Detroit River* forms<br />

the connecting waterway between <strong>Lake</strong> Huron<br />

and <strong>Lake</strong> Erie, and is therefore an<br />

important fish conduit. <strong>The</strong> bays and wet-<br />

lands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, especially in the<br />

<strong>St</strong>. <strong>Clair</strong> Delta area at the mouth <strong>of</strong> the<br />

<strong>St</strong>, Cl air River, are important spawning<br />

and nursery areas for many species that<br />

support major fisheries in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

as well as <strong>Lake</strong> Huron and <strong>Lake</strong> Erie<br />

(Johnston 1977; Goodyear et al. 1982arb,c;<br />

Hatcher and Nester 1983). <strong>St</strong>udies<br />

conducted at a number <strong>of</strong> widely separated<br />

sltes in the Great <strong>Lake</strong>s indicate that the<br />

nearshore waters are spawning and nursery<br />

grounds for most fishes (Boreman 1976).<br />

More than 70 species <strong>of</strong> fish have been re-<br />

corded as residentsr or migrants, in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>; <strong>of</strong> these, 48 species are depen-<br />

dent on or known to use the marshes and<br />

shallow areas <strong>of</strong> the lake (Appendix I).<br />

Although many species <strong>of</strong> fish utilize<br />

coastal wet1 ands, comparative1 y few


Table 19. <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> unionid bivalves and their glochidial host fishea<br />

Fish hostsb AM5 FUS QUA ELL ALA LAS SIM AN0 STR TRU PRO CAR LEP ACT LIG LAM<br />

Northern pfke X<br />

Carp X X<br />

Golden shiner X<br />

Creek chub X X<br />

White sucker X<br />

N hog sucker X<br />

N Sh redhorse X<br />

01 bull head X<br />

Ye bull head X<br />

Br bull head X<br />

Ch catfish X X<br />

Bk stickleback X<br />

White bass X X<br />

Rock bass X X X<br />

Gr sunfish X X X X<br />

Pumpkinseed X<br />

Or-sp sunfish X<br />

Bluegill X X X X<br />

Sm bass<br />

Lm bass X X X X<br />

Wh crappie X X X X X X X<br />

B1 crappie X X X X<br />

Iowa darter . X<br />

Johnny darter X X<br />

Ye perch X X<br />

Wa1 1 eye<br />

Freshwater drum X X X X<br />

Mottled sculpin X<br />

Mudpuppy X<br />

(amphibian)<br />

a~ata sources: Full er (1974) r Clarke (1981)<br />

b ~ -<br />

northern CAMB - ~1ic-<br />

Sh- shorthead FUS - Fusc;onafa flpyg<br />

B1-bl ack QUA - guduh and P,<br />

Ye-ye1 1 ow ELL - Elliat_io dllatata<br />

Br-brook ALA -<br />

Gr-green LAS -<br />

Or-sp-orange-spotted SIM -<br />

Sm-smal lmouth AN0 -<br />

Lm-1 argemouth STR -<br />

Wh-white TRU -<br />

PRO -<br />

CAR -<br />

LEP -<br />

ACT -<br />

LIG -<br />

LAM -<br />

C<br />

X X X<br />

X X<br />

X X X<br />

X X X<br />

X X


specles are strongly dependent an aquatic<br />

vegetation for spawntng, feeding8 or<br />

cover. <strong>The</strong> more common wetland-dependent<br />

species found fn <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and<br />

occurring frequently in the coastal<br />

wetl ands I nc1 ude 1 ongnos<br />

-1, bowfin (m<br />

pike (&&,& luciu@, gras<br />

-) , central mudmlnnow (Y-a<br />

m), golden shiner (<br />

blacknose sh<br />

, blackchln shiner (&&roois<br />

mic shfner f<br />

1 sand shiner<br />

1 P lake chubsucker f<br />

, rock bass (<br />

1, green sunf<br />

smallmouth bass<br />

generally associated with lotic waters and<br />

clean sand or gravel bottoms8 but they are<br />

also common in coastal lake waters, where<br />

they may util ize deep lacustrine coastal<br />

wetlands with sand or gravel bottoms. <strong>The</strong><br />

bet.erod~ll)~ tadpole madtom (lig&dus<br />

-9, banded killifish (Egn&ulus<br />

green sunfish, rock bass, and smallmouth<br />

bass, are signif lcant game species. Among<br />

-haw) r pumpkinseed (l&pomi~ commercial and game species the walleye<br />

-1, Iowa darter ( F a - u ) r ($tizostedioo y. -1 and native or<br />

and brook stickleback (w inconslam). introduced salmonids apparently have<br />

Several other species are largely little direct association with coastal<br />

restricted to vegetated waters but are wetl ands, Other specles <strong>of</strong> recreational<br />

either uncommon or rare In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. or commercial Importance, including ye1 low<br />

<strong>The</strong>se are the spotted gar (-<br />

perch (parca -1, whlte bass<br />

$xuh$u), muskellunge (&&x -1 , (w -1, and freshwater drum<br />

pugnose shiner (- -1 and ( ~ i n o t u -1, r<br />

as we1 1 as<br />

pugnose minnow (m ~miliu). important forage species, including spot-<br />

However, a signif icant native, selfsustaining<br />

populatfon <strong>of</strong> muskellunge<br />

exists in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Haas 1978) and<br />

tail shiner (Notroo hudson_ius) and<br />

emerald shiner (- j&hgchm,)r<br />

are ubiquitous in the coastal zone <strong>of</strong> <strong>Lake</strong><br />

spawns in the delta wetlands. Here, it is<br />

harvested by the Chippewa Indians on the<br />

Walpole Island Indian Reserve, Ontario.<br />

<strong>St</strong>. <strong>Clair</strong> and may be locally commmon in<br />

certain coastal wetl ands,<br />

Known ffsh spawning areas fn the <strong>St</strong>,<br />

Several fish species found in the <strong>Clair</strong> system have been documented by the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin are common to U.S. Fish and Wildlife Service (Goodyear<br />

abundant in wetlands and adjacent waters, et a1. 1982arbrc1, including site specific<br />

although they are <strong>of</strong>ten associated with information on ffsh spawning and nursery<br />

non-vegetated habitats as well.<br />

-1,<br />

<strong>The</strong> gl z-<br />

goldfish<br />

areas for sturgeon (- ful\~asce&),<br />

alewife (w a s e u d o m ) , rainbow<br />

trout (SalmP -1, smelt (Osmru.s<br />

-1, white sucker (&&ostom~<br />

-1, smallmouth bass, rock bass,<br />

yellaw bull head (m na$al f S) r<br />

walleye, and ye1 low perch. Goodyear and<br />

her colleagues summarized information on<br />

a17 past and presently known spawning<br />

areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and the <strong>St</strong>. <strong>Clair</strong><br />

River. <strong>The</strong>se areas include the marshy and<br />

occur in quiet* low-gradient waters with<br />

bottoms <strong>of</strong> mud or clay. <strong>The</strong>se species are<br />

general 1 y cover-oriented and may be common<br />

in sheltered riverine and l acustrine<br />

coastal wet1 ands along <strong>Lake</strong> <strong>St</strong>. Cl air.<br />

All except the gizzard shad, goldfish,<br />

bl untnose rnf nnon, and fathead minnow are<br />

significant game spectes,<br />

shallow bay areas along the shore1 ine and<br />

the wetlands <strong>of</strong> the delta. While the<br />

relationship <strong>of</strong> submersed aquatfc<br />

macrophytes in 1 ittoral waters to f i shery<br />

productf vi ty is not fully understood,<br />

prel lminary results <strong>of</strong> recent studies<br />

indicate that the submersed littoral<br />

macarophyte habitat is important to fish<br />

production in these waters (Schl oesser and<br />

Manny 1982).<br />

Walleye migrate through <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> and use the channels <strong>of</strong> the <strong>St</strong>.


<strong>Clair</strong> Delta for spawning areas (Figure<br />

34). Tagging studles (Ferguson and Derksen<br />

1971; Wolfert st a1. 1975) indicated<br />

considerable movement <strong>of</strong> wall eye between<br />

<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> and adj<strong>of</strong>ning lakes Erie<br />

and Huron. Each lake has many known and<br />

some suspected spawni ng areas. Sf nce<br />

walleyes prefer to spawn close to their<br />

p 1 ace <strong>of</strong> or i g i n ( Eschmeyer 1950; Ferguson<br />

and Derksen 1971) there appears to be a<br />

number <strong>of</strong> semi-di screte stocks coexisting<br />

in these lakes. Tagging studies have not<br />

been sufficiently intense, however# to<br />

delimit these stocks within time and<br />

space.<br />

Yellow perch (w flayescm,sI is<br />

one <strong>of</strong> the most abundant fish in <strong>Lake</strong> <strong>St</strong>.<br />

Clalr and is known to spawn over aquatic<br />

vegetation in most inshore areas <strong>of</strong> the<br />

lake (Figure 351, but it is not known<br />

whether discrete stocks exist. Commercial<br />

fishermen belleve that <strong>Lake</strong> Huron fish can<br />

be identified by shape and color in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. Similarlys Ohio fishermen<br />

postulate that a postspawning movement <strong>of</strong><br />

large yellow perch comes into the western<br />

basin <strong>of</strong> <strong>Lake</strong> Eries presumably from <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>re is no evidence to<br />

support or reject these views, <strong>The</strong> scant<br />

tagging that has been carried out with<br />

yellow perch suggests that they are<br />

capable <strong>of</strong> moving between the lake systems<br />

(Johnston 1977). However, the maintenance<br />

<strong>of</strong> a number <strong>of</strong> age-groups in <strong>Lake</strong> <strong>St</strong>,<br />

<strong>Clair</strong> is in direct contrast to the<br />

situation in <strong>Lake</strong> Erie.<br />

<strong>St</strong>urgeons a threatened species in<br />

Mfchigan and an endangered species in<br />

Ohlor are common in the deeper portions <strong>of</strong><br />

the delta and channels. <strong>The</strong> major known<br />

spawning area for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> sturgeon<br />

is the North Channel <strong>of</strong> the <strong>St</strong>, Clalr<br />

River (Figure 36). <strong>The</strong>se sturgeon migrate<br />

up to the North# South, and Middle<br />

channels to spawn at a site in the North<br />

Channel ; young-<strong>of</strong>-the-year sturgeon<br />

migrate from this spawning site to the<br />

marsh between Bouvier and Goose Baysv<br />

where they are found among the rushes<br />

(Goodyear at al. 1982a~b). <strong>The</strong> channels<br />

are also productive fishing areas for<br />

muskellunger smallmouth bass# 1 argemouth<br />

bass* freshwater drum* and northern pike,<br />

Flgures 37 and 38 depict spawning and<br />

nursery areas for several <strong>of</strong> these<br />

species.<br />

[ /fi\ WALLEYE<br />

Spawning area<br />

2.:. Nursery area<br />

Figure 34. Walleye spawning and nursery<br />

areas in the Detr<strong>of</strong>t River and <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> (Goodyear et al. 1982b,c),<br />

<strong>Lake</strong> <strong>St</strong>. CSair has been acclaimad one<br />

<strong>of</strong> the best fishing areas in Elarth America<br />

(Figure 39). This fishery is fostered by<br />

the marsh habitats which sustain species<br />

number and diversity, Sport fishing in<br />

wetlands exhfbits high sconomlc values,<br />

Jaworski and Raphael ( 1878) calcul atad the<br />

value <strong>of</strong> Mfchigan's coastal wetlands with


Detroit River<br />

~atrolt<br />

YELLOW PERCH<br />

Spawnlnp area<br />

AY. Nuraery area<br />

Figure 35. Yellow perch spawning and<br />

nursery areas in the Detroit River and<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Goodyear et at, 1982b,c).<br />

respect to sport f l shfng at $ll6/wetland<br />

hectare/year, <strong>Lake</strong> <strong>St</strong>. Cl air exhibits the<br />

highest sport flshery value for Great<br />

lakes coastal wet1 ands because <strong>of</strong> its<br />

proximity to Detroit and the high qua1 i ty<br />

<strong>of</strong> angl i ng for wa1 l eyer smallmouth bass,<br />

and yellow perch (Table 20). Current<br />

estimates indicate that the <strong>St</strong>, Clafr<br />

Delta wetlands average 8 to 16 angler-<br />

day s/wetl and hectare/year (Werdendorf et<br />

a%. 1981~1.<br />

Major game fish species whfch furnish<br />

a recreational fishery in Bouvier Bay<br />

Wetland, known also as <strong>St</strong>. Johnts Marsh,<br />

include northern pike, 1 argemouth bass9<br />

ye1 low perch, bull heads* and various<br />

panflsh (Posplchal 1977). Recreational<br />

fishing is popular from shore sites as<br />

well as from boats In the shallow bays and<br />

channels <strong>of</strong> <strong>St</strong>. Johnrs Marsh, Within the<br />

<strong>St</strong>. <strong>Clair</strong> Flats <strong>St</strong>ate Wildlife Area on<br />

Dickinson Island and on Harsens Island,<br />

recreational fishing is popular for<br />

muskellunge, northern pike, wall eye,<br />

yellow perch, smallmouth bass, channel<br />

catfish (fctalurs punctatus), bull heads,<br />

bluegill other sunfishes, crappie, rock<br />

bass, white bass, lake sturgeonr coho<br />

salmon (Qncorhv- steel head<br />

trout and other species. <strong>The</strong> area is also<br />

a prolific producer <strong>of</strong> bait-minnows. Bow<br />

and arrow fishing for carp is a growing<br />

springtime activity, which generates 3,000<br />

to 5,000 angler-days <strong>of</strong> recreation per<br />

year (Pospichal 1977). Ice fishing<br />

(commonly by hook and line) Is also an<br />

active sport in January and February.<br />

Common species pursued include ye1 1 ow<br />

perch (85%) s sunfish (10%)r walleyer and<br />

northern pike. More than 95% <strong>of</strong><br />

recreation ice fishing is for wetland<br />

associated species.<br />

Other than species composition and<br />

recreational use <strong>of</strong> the flsh fauna <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>r knowledge <strong>of</strong> the wetland<br />

fishery <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> Is lacking in<br />

several respects. Other important con-<br />

siderations include: 1) actual relation-<br />

ships <strong>of</strong> the species to the wetlands in<br />

terms <strong>of</strong> their utilization for spawning,<br />

nursery, and feeding areas, 2) fish<br />

community structure, 3 1 niche occupation,<br />

and 4) interspecific relationships within<br />

wetlands. <strong>The</strong> coastal wetlands <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. ClaJr appear to support a mlxed fish<br />

fauna <strong>of</strong> warmwater and coolwater species.<br />

In addl ti on to wetland dependent speciesr<br />

many species common in other coastal<br />

habltats are also common in coastal<br />

wetlandsE dependlng on condition <strong>of</strong><br />

shelter, bottom type, water clarity, water<br />

depth, and density <strong>of</strong> vegetation. Because<br />

<strong>of</strong> the obvious voids in fisheries<br />

information wfthin coastal wet1 ands,<br />

research in this area ranks high priority.


MlCHlGAN<br />

LAKE STURGEON<br />

Spawning area<br />

.* .+ Nursery area<br />

Middle Channel<br />

outh Channel<br />

Figure 36. <strong>Lake</strong> sturgeon spawning and nursery areas in the Detroit River, <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>, and <strong>St</strong>. <strong>Clair</strong> River (Goodyear et a1 . 1982a,b,c).<br />

63


Figure 37. Rock bass, pumpkinseed/biuegill, largemouth bass, and smallmouth bass<br />

spawning and nursery areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Goodyear et al. 1982b).


FISHING AREAS<br />

f_l_j YELLOW PERCH<br />

SMALLMOUTH BASS<br />

MICHIGAG ONTARIO<br />

Figure 39. Sport fishing areas by species In the vicinity <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> Delta wetlands<br />

(<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Advisary Committee 1975).<br />

33 AMPHlBlANS AND REPTILES<br />

<strong>The</strong> assemblage <strong>of</strong> amphlbfans and<br />

reptlles reported For the <strong>Lake</strong> <strong>St</strong>. ClaSr<br />

regfon can be divided into four major<br />

groups: 11 those specfes almost always<br />

found In wetlands and somehow dependent on<br />

wetland habitats to the degree that they<br />

are uncommon elsawherer 2) aquatic and<br />

semiaquatic species found in a wide range<br />

<strong>of</strong> law1 and wet habf tats, including coastal<br />

wetlands. 3) terrestrial species which<br />

enter water or wetlands only fnc5dentally<br />

or to breed* and 4) aquatfc and<br />

semiaquatic specles found prSmar4l y in<br />

upland or lotic waters and not common to<br />

coastal wetlands (Herdendorf st a7.<br />

1981~). <strong>The</strong> latter are not disussed in<br />

thls report. Four species recorded In the<br />

literature as occurring fn the coastal<br />

zone <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr are largely wetland<br />

dependent: mudpuppy (i&&uxi<br />

JMGW<br />

red-spotted newt ( Y&k<br />

, eastern fox snake (w<br />

eastern massasauga<br />

1. <strong>The</strong> mudpuppy is<br />

largely an open lake species which is most<br />

sonmon around large 1 acustrine wetlands<br />

with submersed vegetation, whereas the<br />

red-spotted newt is found primarily in<br />

small ponds and sheltered palustrine or<br />

riverine wetlands. <strong>The</strong> eastern fox snake<br />

is common in large coastal emergent-type


Table 20. Average annual number <strong>of</strong> sport fish takenafrom Michigan waters <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. Clai r and connecting waterways (1975-1979).<br />

Species <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Detroi t Total<br />

<strong>St</strong>. Clafr River River individuals<br />

<strong>Lake</strong> trout<br />

Rai nbow/steel head<br />

Brown trout<br />

Coho salmon<br />

Chi nook salmon<br />

Wal leye/sauger<br />

Bass<br />

Northern pike/musky<br />

Yellow perch<br />

Sunf ish/bluegill<br />

Crappie/white bass<br />

Bull head/catf ish<br />

Rock bass<br />

Sucker<br />

Whitefish/cisco<br />

Other<br />

Angler days 1,079,031 320,701 4529376 lr 852,108<br />

Fishermen 100,622 32,523 41,674 174s 819<br />

a Data source: Michigan Department <strong>of</strong> Natural Resources (unpublished data).<br />

wetlands, but the rarer eastern massasauga<br />

is primarily restricted to swamp forest<br />

and bogs.<br />

Amphibians found in a wide range <strong>of</strong><br />

lowland wet or aquatic habitats in the<br />

<strong>Lake</strong> <strong>St</strong>. Clafr coastal zone include the<br />

blue-spotted salamander (&nbvstoma 1 atec a), spotted salamander ( A m<br />

n@&ulatu!q), four-toed salamander (Hemi-<br />

Utv1 i -1, eastern tiger sa1 a-<br />

mander t,4mbvsta =), gray treefrog<br />

(fjYLB yersIcQ&x), northern spring peeper<br />

(liyljl ~ ruci fer) western chorus frog<br />

(pseudacris triseriata), Blanchardqs<br />

crfcket frog C-5 -1 r<br />

bullfrog (m -1, green frog<br />

(m m1 ano-I, pickerel frog<br />

(Baas pa1 ustris) , and northern leopard<br />

frog (m ploiens). <strong>The</strong> salamanders may<br />

<strong>of</strong>ten occur in moist terrestrial habitats,<br />

but the1 r larvae are always aquatjc.<br />

Aquatic and seml-aquatic reptiles which<br />

probably occur in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />

wetlands or their border lands include the<br />

northern ringneck snake (Digdoahfs ~unctatlls<br />

edwar_dsl), eastern milk snake<br />

(mm -1 northern<br />

water snake (&&& SL northern<br />

brown snake (<strong>St</strong>orerb L 1, northern<br />

red-bellied snake (sore* p, ~sg@ito -<br />

1par:ulatg), Butlerts garter snake (m<br />

eastern gar<br />

Several amphfblans <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>.<br />

Clafr coastal zone are largely terres-<br />

trial, although they are found in lowland<br />

areas and are dependent on water for<br />

breeding. Consequent1 y the adults ar<br />

l arvae may occur in coastal wetlands or


their margfns at least seasonal 1 y. <strong>The</strong>se<br />

specles 1 nclude the red-backed salamander<br />

)C the Amerfcan<br />

d the wood frog<br />

trial reptf le5<br />

may occur in the margins <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

coastal wetlands or incidentally in the<br />

wetlands themselves<br />

smooth green snak<br />

m ) r blue racer<br />

1, and black rat snake (<br />

I.<br />

Although few species <strong>of</strong> reptiles and<br />

amphibians are absolutely dependent on the<br />

coastal wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> for<br />

habitat, the continued abundance <strong>of</strong> most<br />

species in the coastal tone is probably<br />

dependent on the presence <strong>of</strong> extensive<br />

coastal wet? ands. <strong>The</strong> relatively un-<br />

disturbed coastal hab 'f t at a1 ong most <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> Delta probably supports a<br />

present herpet<strong>of</strong>auna similar to that <strong>of</strong><br />

pre-settl ement ti mesr both in composition<br />

and relatf ve abundance <strong>of</strong> specles.<br />

Appendix J Ilsts the specles known or<br />

presumed to occur in or adjacent to the<br />

coastal wetlands; because <strong>of</strong> the scarcity<br />

<strong>of</strong> published records, this list is<br />

somewhat specul atlve.<br />

species 1s wetland-dependent and was once<br />

common and restricted to coastal emergent<br />

wetlands and wetland margins <strong>of</strong> western<br />

<strong>Lake</strong> Erie and the coast <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air.<br />

<strong>The</strong> black rat snake is also listed as<br />

threatened In Michfgan and may occur fn<br />

drier margins af <strong>Lake</strong> <strong>St</strong>, Clafr coastal<br />

wetlands. <strong>The</strong> eastern massasauga is a<br />

candidate for Federal listing on the<br />

threat.ened/endangered speci es 1 ist. <strong>The</strong><br />

following species, considered rare in<br />

Michigan, may also occur in these coastal<br />

wetlands: eastern spiny S<strong>of</strong>tshellr spotted<br />

turtle ( 18 and fourtoed<br />

salamander,<br />

<strong>The</strong> extensive wetlands <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> Delta are utilfzed for nesting and<br />

during migration by waterfowl wading<br />

birds, gulls, terns, and shorebirds.<br />

Appendix K lists the Important bird<br />

species which occur in the vicinity <strong>of</strong> the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands. Waterfowl<br />

commonly observed in the wetlands include<br />

mallard (BagS whvn-), black duck<br />

(81195 -1, blue-wlnged teal (m<br />

$i%cocs). wood duck (&1X -1, American<br />

wigeon (m -18 and northern<br />

shoveler (u ~lvpeata) . In addi tion,<br />

canvasback (m val f +=i-1 scaup<br />

(m spp.1, ruddy duck (Ilxylrra<br />

&gnat and hooded merganser<br />

<strong>The</strong> presence <strong>of</strong> water, food, and<br />

cover, and the relative isolation from the<br />

cultural development which occurs around<br />

most other bodies <strong>of</strong> water are among the<br />

factors that contribute to the importance<br />

<strong>of</strong> wetlands in maintaining the coastal<br />

zone herpet<strong>of</strong> auna. <strong>The</strong> recreational or<br />

(w ~ 1 1 a t u s utilize ) the<br />

habitat for nesting . Several endangered<br />

commerci a1 importance <strong>of</strong> the coastal and threatened species (Michigan) <strong>of</strong> birds<br />

herpet<strong>of</strong>auna is unknown8 but the bullfrog, have been reported In the coastal zone.<br />

green frog* eastern spiny s<strong>of</strong>tshell, and<br />

snapping turtle are edible species which<br />

may be harvested. In addition, reptiles<br />

Osprey (Pandi_nn ha1 iaet~~5.1 and the Caspian<br />

tern (<strong>St</strong>erna -1 have been reported in<br />

scattered localities along the shore1 I ne.<br />

and amphi bians such as the northern water Nest1 ng colonies <strong>of</strong> ye1 law-headed<br />

snake and the snapping turtle, serve as<br />

food sources for many species <strong>of</strong> fish,<br />

birds, and mammals, or may themselves be<br />

blackbird ( 18<br />

black tern (m -1, and common<br />

tern (Sf;arna hi run&? have been observed<br />

predators <strong>of</strong> economf tally important, fish in several wetlands. <strong>The</strong> common tern is<br />

and wild1 ife. Beyond ~m3ra1 background presently on the Michigan endangered<br />

on the occurrence and relative abundance<br />

<strong>of</strong> reptiles and amphibians in the coastal<br />

specles list.<br />

wetlands* little publfshed information<br />

exists pertaining to papu1atlon8 community<br />

structurer and dynamics <strong>of</strong> the<br />

Waterfowl are a most Important<br />

component <strong>of</strong> the bIotic envf ronment. <strong>The</strong><br />

importance <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />

herpet<strong>of</strong> auna,<br />

to ducks, geese, and other waterfowl is<br />

<strong>The</strong> eastern Cox snake is listed as<br />

threatened by +Re <strong>St</strong>ate <strong>of</strong> Michtgan, This<br />

based primarily on the location <strong>of</strong> these<br />

wetlands in relation to waterfowl flyways,<br />

the productivity (in terms <strong>of</strong> waterfowl


foods) <strong>of</strong> the wetlandso and the declfne <strong>of</strong><br />

other coastal wetland systems. Except for<br />

that <strong>of</strong> mallard, black duck and blue-<br />

winged teal, nesting appears to be a<br />

secondary use <strong>of</strong> these wetlands with<br />

feeding and resting during migratlon bejng<br />

<strong>of</strong> primary importance.<br />

Based on waterfowl feeding and<br />

resting use durfng migratfon as well as on<br />

duck harvest and production <strong>of</strong> young, the<br />

wetlands and associated open waters <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> may be the most important<br />

wetland system in the Great <strong>Lake</strong>s region<br />

with the possible exception <strong>of</strong> Long Point<br />

in <strong>Lake</strong> Erie. With reference to the Great<br />

<strong>Lake</strong>s-Chesapeake Bay migration torrider*<br />

the principal flight paths <strong>of</strong> the tundra<br />

swan, formerly whist1 ing swan, (CYQILUS<br />

-1. canvasback, bufflehead<br />

( -1, and ruddy duck in-<br />

clude a resting stopover fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

(Bell rose 1976). In reference to water-<br />

fowl harvesting in Michigan, <strong>St</strong>. Clai r<br />

County, which includes most <strong>of</strong> the wetland<br />

on the American side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr, is<br />

the leading harvest county (Carney et a1 .<br />

1975). Production data for the Bouvier<br />

Bay (including <strong>St</strong>. John" Marsh) and<br />

Harsens Is1 and wetlands reveal an average<br />

<strong>of</strong> 240 young produced per km2 EHerdendorf<br />

et al. 1981~).<br />

Important 1 lterature sources re-<br />

garding the waterfowl use <strong>of</strong> the wetlands<br />

are Jaworski and Raphael (1978) as we11 as<br />

Herdendorf et al. (1981~). <strong>The</strong> latter<br />

source is especially important in regard<br />

to the Bouvier Bay, Dickinson Island, and<br />

Harsens Island wetlands. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong><br />

Flats <strong>St</strong>ate Game Area comprises the lower<br />

portions <strong>of</strong> Harsens Island and Dickinson<br />

Island, plus the two former areas <strong>of</strong> the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> National Wfldllfe Refuge,<br />

one at the mouth <strong>of</strong> North Channel in<br />

Anchor Bay, and another in between the end<br />

<strong>of</strong> Middle Channel and Channel A Bout Rond.<br />

In Ontario, the Canadian Wildl ife<br />

Service (Dennis and Chandler 1974, Dennis<br />

et a1. 1981) have documented the util ity<br />

<strong>of</strong> wetlands with regard to waterfowl. In<br />

terms <strong>of</strong> total use durlng migration, the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands rank second to<br />

Long Point (<strong>Lake</strong> Erie), Ontario. <strong>The</strong><br />

coastal marshes are presently the most<br />

important staging areas in southern<br />

Ontario for mallard, black duck, Canada<br />

1, and tundra<br />

ons <strong>of</strong> .the North<br />

asback* redhead<br />

1, and tundra swan also<br />

util ize the <strong>Lake</strong> <strong>St</strong>. Claf r marshes during<br />

migratlon staging periods.<br />

Walpole Island possesses extensive<br />

marshes, particularly at its southern end<br />

where the shore fronts on <strong>Lake</strong> <strong>St</strong>. Cl ai r.<br />

<strong>The</strong>re are flve major islands that make up<br />

the 24,000-hectare Wal pole Is1 and Indi an<br />

Reserve. Here many nesting species <strong>of</strong><br />

birds rare in both the region and Ontario<br />

as a whole have been observed, including<br />

canvasback, redhead, ruddy duck, northern<br />

harrier (Circus little gull<br />

(m -1 , Forsterts tern (w<br />

-1, and king rail (&Uui 3<br />

(Goodwin 19821, Wooded areas e<br />

island include sections <strong>of</strong> oak savannah<br />

which accommodate a rookery <strong>of</strong> blackcrowned<br />

night-herons )<br />

and great egrets (- allus).<br />

As indicated In Table 21, a diversity<br />

<strong>of</strong> waterfowl rely on the <strong>St</strong>. <strong>Clair</strong> Delta<br />

wetlands. Waterfonl use during migration<br />

is more Important than use <strong>of</strong> these<br />

wet1 ands for breeding purposes. Moreover,<br />

ducks are much more numerous than are<br />

swans and geese. With regard t o dabbling<br />

ducks, the most abundant migrants include:<br />

mallard, black duck, American wigeon as<br />

we1 1 as pintail (&as<br />

teal, and green-winged<br />

Among the more abundant migrant diving<br />

ducks are canvasback, great6<br />

(&j&ygj -1, lesser scaup<br />

ilfflnfs) * 11 a<br />

goldeneye ( and buffle-<br />

head.<br />

<strong>The</strong> use <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta<br />

wetlands for feeding by both dabbling and<br />

diving ducks fs <strong>of</strong> prime importance (Table<br />

22). Plant foods from shallon marshes and<br />

waste gralns from nearby agricultural<br />

fields are especially important in the<br />

diet <strong>of</strong> dabbl ing ducks, parti cul arly<br />

mallards and black ducks as well as Canada<br />

geese, whereas submersed aquatic plants<br />

and thei r associated i nrertebrate fauna<br />

constitute important fwd items far diving<br />

ducks such as canvasback, redhead* and<br />

scaup (Jaworski and Raphael 1978). Corn


Table p.<br />

1977).<br />

Birds observed in <strong>St</strong>. <strong>Clair</strong> Delta wetlands (1975-<br />

Species Bouvi er Dickinson Harsens<br />

~ayb Is1 and Is1 and<br />

Tundra swan<br />

Canada goose<br />

Snow goose<br />

Ma1 1 ard<br />

Black duck<br />

Gadwall<br />

Pintail<br />

Amerlcan wigeon<br />

Green-wi nged teal<br />

Bl us-w i nged teal<br />

Northern shoveler<br />

Wood duck<br />

Red head<br />

Ring-necked duck<br />

Canvasback<br />

Greater and lesser scaups<br />

Common go1 deneye<br />

Buff 1 ehead<br />

Ruddy duck<br />

Hooded and comon mergansers<br />

Horned grebe<br />

Piad-bi 1 led grebe<br />

Be1 ted kingf lsher<br />

Sedge and marsh wrens<br />

Great blue heron<br />

Green-backed heron<br />

Great and cattle egrets<br />

Black-crowned night-heron<br />

American and least bitterns<br />

K4ng and Virginia rails<br />

Sora<br />

Common moorhen<br />

Amerfcan coot<br />

Herring gull<br />

Ring-btlled gull<br />

ForsPerss and black terns<br />

Comon tern<br />

Eastern meadow1 ark<br />

Ye1 1 ow-headed blackbird<br />

Red-w i nged bl ackbi rd<br />

Common snipe<br />

American woodcock<br />

cooperfs hawk<br />

Red-tat led hawk<br />

Rough-1 egged hawk<br />

Northern harri er<br />

'~ata source: Herdendorf at a1. ( 1981~1.<br />

b~ote: Bouvier Bay survey included <strong>St</strong>. John's Marsh; other<br />

surveys less i ntense,


- - -<br />

Summer Fa1 1 Spring<br />

Adult Young Total Duration Total Duration<br />

'$able 22. Productivity and seasonal abundance <strong>of</strong> watesfowl in Bouvier<br />

Bay marshes and Harsens Island interior wetlands (1974).<br />

Area and pai rs<br />

waterfowl ( no/ km2 pop. (wks) pop. (wks)<br />

Dabbl lng ducks 35 222 41800 4-10 4r900 3-4<br />

Divers 2 14 200 - 5~000 3-4<br />

Geese 1 4 0 0 0 0<br />

Swans 0 0 0 0 0 0<br />

--<br />

Total 38 240 5 r 000<br />

Dabblfng ducks 33 210 1,400 4-10 1~500 4-5<br />

Divers 4 22 100 10 2,400 4-5<br />

Geese 1 5 - - - -<br />

Swans 38 237 lr500 - 3 * 900 -<br />

--<br />

Total 76 474 3 r 000<br />

a~ata sources: Herdendorf et al. (1981c)r Michigan Department <strong>of</strong> Natural<br />

Resources (unpubl lshed data),<br />

and other grains grown on the Harsens<br />

Island <strong>St</strong>ate Game Area and on the upland<br />

portions <strong>of</strong> the Canadian side <strong>of</strong> the delta<br />

attract dabbl ing ducks and Canada geese.<br />

In contrast* submersed aquatic plants i n<br />

the diet <strong>of</strong> diving ducks include wild<br />

celery, pondneeds, and waterweed.<br />

<strong>The</strong> significance <strong>of</strong> the plant and<br />

invertebrate food base <strong>of</strong> the <strong>St</strong>, <strong>Clair</strong><br />

Delta wetl ands in relation to waterfowl<br />

can not be overstated. An important study<br />

by Dawson (1975) <strong>of</strong> Anchor Bay in northern<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, indfcates that only a<br />

small percentage <strong>of</strong> the preferred aquatic<br />

pl ants and invertebrates avail able to<br />

migrating waterfowl fn <strong>Lake</strong> <strong>St</strong>. Claf r<br />

wetlands are currently being utilized.<br />

Waterfowl food supplies are especially<br />

important during sprf ng migration because<br />

preferred foods are general 1 y less<br />

abundant and breeding females must arrive<br />

in good condition on the breedfng grounds.<br />

Thus the spring migration is less<br />

restricted to specific feeding areas and<br />

waterfowl can be observed in lakes and<br />

wetlands whSch they did not frequent in<br />

the fall. For exampf er note in Table 22<br />

that divfng ducks are more numerous Jn the<br />

relatively shallow Bouvfer Bay and Harsens<br />

Island wetlands in sprfng than in the fall<br />

migration season. Waterfowl migratjon is<br />

discussed in more detail in Sectton 4.3.<br />

With regard to production <strong>of</strong><br />

waterfowl In the wetlands along take <strong>St</strong>.<br />

<strong>Clair</strong>, these wetlands appear to be <strong>of</strong><br />

moderate importance on1 y. Some pre-<br />

lfmlnary data collected by JaworskS and<br />

Raphael (1978) suggested that Michigan's<br />

coastal wetl ands have a duck nesting<br />

density <strong>of</strong> 32 breedjng pafrs per km2 and<br />

that goose and swan nestlng was<br />

Insfqniffcant, Data reveal that the pairs


<strong>of</strong> dabbling ducks slfghtly exceed this<br />

prel iminary average, but that paf rs <strong>of</strong><br />

diving ducks were much less than the man<br />

(Table 22) . Eterdendorf et al. (1981~1<br />

reported that ma1 1 ard, blue-winged teal#<br />

and cinnamon teal are the most common<br />

nesters in the delta wetlands along wlth<br />

smaller numbers <strong>of</strong> black duck, wood duck,<br />

pintall, redhead, and ruddy duck. Small<br />

but signi f icant numbers <strong>of</strong> the re1 at1 vel y<br />

rare redhead duck nest in <strong>St</strong>. Johnrs<br />

Marsh, on lower Harsens Island* and along<br />

coastal Walpole Island. Both mallard and<br />

blue-winged teal are cl assif fed as meadow<br />

nesters, but the mal 1 ard exhibits great<br />

versatility In regard to selection <strong>of</strong><br />

potential nesting sites.<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has long been renowned<br />

as an excel? ent duck huntfng area.<br />

Hunting pressure from both the land and<br />

water is common on the American and<br />

Canadlan sides <strong>of</strong> the delta. <strong>The</strong> main<br />

species harvested in these coastal<br />

wetlands are the mallard, along with<br />

lesser numbers <strong>of</strong> black duck, pintail, and<br />

teal (Table 23) . However, In open-water<br />

areas <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, where hunting<br />

from boats and floating blinds prevails*<br />

the majorfty <strong>of</strong> the ducks harvested are<br />

divers# i.e., scaup and the now restricted<br />

canvasback and redhead, rather than<br />

dabbling ducks.<br />

<strong>The</strong> type <strong>of</strong> shores range from sandy<br />

beaches to mudflats. <strong>The</strong> beaches attract<br />

small flocks <strong>of</strong> shorebirds In migration.<br />

Those species commonly seen feeding are<br />

bfrds, such as herons and egrets, gather<br />

to feed in the mud flats near larger<br />

streams.<br />

Nesting species in the <strong>St</strong>. <strong>Clair</strong><br />

Flats (Figure 409 include such specIes as<br />

Table 23. Species composition <strong>of</strong> ducks<br />

harvested In the <strong>St</strong>., <strong>Clair</strong> Flats <strong>St</strong>ate<br />

Game Area (1974-1975).<br />

Annual<br />

Spechs harvest Percent<br />

(avge <strong>of</strong> total<br />

individ.)<br />

Ma1 1 ard, 4,362<br />

including domestic<br />

Black duck and hybrids 498<br />

Plntail 242<br />

Green-w 1 nged teal 173<br />

American w l geon 13 8<br />

Ri ng-necked duck 78<br />

Bl ue-w i nged teal 7 1<br />

Scaup 34<br />

Gadwall 30<br />

Wood duck 28<br />

Merganser 19<br />

Buf fl ehead 18<br />

Northern shoveler 10<br />

Common go1 deneye 9<br />

Redhead 4<br />

Ruddy duck 3<br />

Oldsquaw 0<br />

Canvasback 0<br />

Scoter 0<br />

- -<br />

TOTAL 5,717 100.00<br />

a~ata source: Jaworski and Raphael (1978).<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands support the<br />

following nesting waterbirds In wooded<br />

habitats: great blue heron (<br />

-1, black-crowned night-h<br />

green-backed heron (- .-US),<br />

great egret, American woodcock (3~01<br />

OD=<br />

-1, be1 sher (Cerv1 Q D<br />

bald eagle Jeuc-1, and<br />

osprey.<br />

Investigations in 1954 <strong>of</strong> a rookery<br />

on Dfcki nson Is1 and found approximately<br />

350 pairs <strong>of</strong> great blue herons and 20<br />

pairs <strong>of</strong> black-crowned night-herons a1 ong<br />

with seven pafrs <strong>of</strong> great egrets. <strong>The</strong>Ir<br />

numbers have declined over the past three<br />

decades. Nearby Harsens Island supports a<br />

rookery <strong>of</strong> bl ack-crowned night-herons<br />

(U.S. Army Corps <strong>of</strong> Engineers 1974).<br />

S<br />

Herons are a solitary bl ).d except<br />

when breeding. <strong>The</strong>y are colonfd nesters


Figure 40. Least bittern (Ixobr chus<br />

exilis) with young in broad-leaved __Y-r catta 1<br />

(Typha latifolia) nest (Michigan Department<br />

<strong>of</strong> Natural Resources).<br />

who requlre a habitat with an abundance <strong>of</strong><br />

twigs and sticks for nest-building. <strong>The</strong>se<br />

long-legged waders prefer a diet <strong>of</strong> f ishp<br />

but w i l l also feed on insectsr<br />

crustaceans, amphibians, reptiles,<br />

mo1 lusca, and rodents (Figure 41). <strong>The</strong>y<br />

are strong fliers and are known to fly<br />

great distances in search <strong>of</strong> food, For<br />

example in <strong>Lake</strong> Erie, herons fly 15 km<br />

from their rookery on West Sister Island<br />

to feed fn the marshes at Locust Point on<br />

the Ohio mainland (Meeks and H<strong>of</strong>fman<br />

1980).<br />

Wading birds usually forage in the<br />

shore1 i nes <strong>of</strong> the tributary streams and<br />

rfthin the coastal marshes <strong>of</strong> the <strong>St</strong>.<br />

Clatr Delta. <strong>The</strong>ir diet consists <strong>of</strong> fish<br />

for the most part, but crayfish and<br />

fnsects are also consumed in significant<br />

quantities.<br />

Short-eared owls (m<br />

red-tai 1 ed,<br />

1 have been<br />

reported fn <strong>St</strong>, John's Marsh in Bouvier<br />

Bay (Pospichal 1977). Bald eagle and<br />

osprey nest in wooded habitats assxiated<br />

with delta wetlands (Kelley et a1. 19639.<br />

3.7 MAMMALS<br />

<strong>The</strong> wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> are<br />

important habitat for several species <strong>of</strong><br />

mammals (Appendix L). <strong>Wetlands</strong> are<br />

essential for muskrat (I)nda.tra gjhethi-1<br />

and mink (w -1. <strong>The</strong> Virginia<br />

opossum (U~&J&IS 1, striped<br />

skunk (w whiti,~)~ and red fox<br />

(m u)<br />

are commonly observed in<br />

the wetlands. Furbearers are a valuable<br />

resource fn many coastal wetlands*<br />

particularly those associated with the<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Delta, including <strong>St</strong>. John's<br />

Marsh (a portion <strong>of</strong> Bouvier Bay wetlands),<br />

Dickinson Island* Harsens Island* Hal pole<br />

Is1 and, and Mitchell Bay wetlands.<br />

Fifteen species <strong>of</strong> mammals have been<br />

reported from the delta fslands (Hayes<br />

1964, U.S. Army Corps <strong>of</strong> Engineers 19749~<br />

ossum, eastern<br />

f10ridanus)~ Eur<br />

Limited deer and upland game hunting is<br />

permltted on the delta i slands.<br />

<strong>The</strong> pri nci pal mmal inhabit+ ng <strong>Lake</strong><br />

<strong>St</strong>. Claf r wetlands in terms <strong>of</strong> occurrence<br />

and economf c value is the muskrat. Based<br />

on fts feeding and lodging habitsf the<br />

density <strong>of</strong> muskrats per unit <strong>of</strong> coastal<br />

wetland can be estimated, Jaworski and<br />

Raphael ( 19781 determfnsd that Macomb


DEA'D PLANTS AND<br />

MINERAL NUTRIENTS<br />

Figure 41. Food chain <strong>of</strong> the great blue heron (Ardea herodias) in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> (U.S. Amy Corps <strong>of</strong> Engineers 1974).


County* Michigan wetlands have sustained<br />

yield <strong>of</strong> 16 muskratsfhectare. This is a<br />

relatively high yfeld for coastal wetlands<br />

which usually range from 5 to 15/hectare.<br />

<strong>The</strong> food supply <strong>of</strong> these omnivorous<br />

herbivores may be diverse, especi a1 1 y<br />

durf ng periods <strong>of</strong> stress. Baumgartner<br />

(1942) has identified 34 food items for<br />

muskrats in Michigan. <strong>The</strong> preferred food<br />

is cattail, bulrush, and bluejoint grass.<br />

<strong>The</strong>se emergent plants are common in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> wet1 ands,<br />

Little site specific research has<br />

been conducted on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> muskrats,<br />

but work done on this species in western<br />

<strong>Lake</strong> Erie can provide some insight.<br />

Several structures observed In <strong>Lake</strong> Erie<br />

coastal marshes are associated with the<br />

activity <strong>of</strong> muskrats (Bednarik 1956). <strong>The</strong><br />

most noticeable structure 1s the muskrat<br />

"house", a dome-shaped pile <strong>of</strong> emergent<br />

vegetation, primarily cattail (Figure 42).<br />

<strong>The</strong> average house varies in slze from 1 to 2.5 rn In diameter at the base and from 0.7<br />

to 3 m in height. <strong>The</strong>y are located in<br />

stands <strong>of</strong> emergent vegetation or along the<br />

periphery <strong>of</strong> such stands. <strong>The</strong>y are <strong>of</strong>ten<br />

constructed on protuberances in the marsh<br />

bottom, util izing plants in the Immedi ate<br />

area. <strong>The</strong> majority <strong>of</strong> houses are<br />

constructed during the fall, chiefly from<br />

October to November. Building activity<br />

occurs mainly during perlods <strong>of</strong> darkness.<br />

Typical densities <strong>of</strong> active muskrat houses<br />

in Sandusky Bay marshes average 4/hectare.<br />

Small houses have one living chamber and<br />

larger houses have two or three lfvfng<br />

chambers above the water 1 ine. <strong>The</strong> 1 iving<br />

Figure 42. Muskrat "houses" in a well-<br />

populated section <strong>of</strong> Magee Marsh, western<br />

<strong>Lake</strong> Erie, Ohio (Bednarik 1956).<br />

chamber is about 35 to 50 cm in height and<br />

is formed by the muskrat chewing out the<br />

vegetation. Most houses have two<br />

underwater exits or plunge holes.<br />

Other structures made by muskrats are<br />

associated with feeding activity. Rafts<br />

are constructed from stems <strong>of</strong> plants piled<br />

in a circular fashion to serve as a<br />

feed1 ng platform. "Feeding bogsw are<br />

covered, floating platforms which are<br />

smaller in dimensions than the muskrat<br />

house and are no higher than 40 cm,<br />

averaging 60 cm in diameter. <strong>The</strong>se<br />

structures are usual 1y located some<br />

distance from the larger muskrat house and<br />

serve as protected feeding sites. Itpush-<br />

upsn are built only in the winter and are<br />

small, hollowr dome-shaped shells <strong>of</strong><br />

submergent vegetatton over a plunge hole<br />

in the ice. <strong>The</strong>se protected plunge holes<br />

allow the muskrat to extend the area over<br />

which it can forage since It can travel<br />

greater dfstances under the ice.<br />

<strong>The</strong> reproductive cycle <strong>of</strong> the muskrat<br />

on Sandusky Bay <strong>of</strong> <strong>Lake</strong> Erie has been<br />

documented by Bednari k (1956) and Donohoe<br />

(1966). Reproductive activity begins in<br />

January and ends In September, with the<br />

greatest acti vl ty occurring I n February<br />

and March. <strong>The</strong> time <strong>of</strong> ff rst mating is<br />

determined in part by the time <strong>of</strong> ice<br />

breakup. <strong>The</strong> gestation period varies f ram<br />

20 to 28 days. Females usually have two<br />

1 itters <strong>of</strong> young per season although some<br />

may have three litters. Placental scar<br />

counts indicate that the mean number <strong>of</strong><br />

young per 1 itter Is l l r but Bednarik<br />

concluded that at the time <strong>of</strong> birth, the<br />

average Iltter size is eight. Sandusky<br />

Bay has habitats similar to those <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> and Is located on the south<br />

shore <strong>of</strong> <strong>Lake</strong> Erie approximately 100 km<br />

southeast <strong>of</strong> Detroit.<br />

Predation by mink occurs but is not<br />

an important influence on the muskrat<br />

population because <strong>of</strong> the low numbers <strong>of</strong><br />

mink in the marshes. Predation on juv-<br />

enlle muskrats by raccoons and Norway rats<br />

occurs In early summer and is not an<br />

Important mortal f t y factor. Some mortal -<br />

fty is attributed to hemorrhagic or<br />

"Erringtonts" disease, This disease can<br />

cause signif icant fluctuatjons in the<br />

muskrat populatfsns 3n western <strong>Lake</strong> Erie<br />

marshes (Bednarik 19561 and presumably<br />

also in lake <strong>St</strong>. CPaSr marshes,


<strong>The</strong> annual harvest <strong>of</strong> muskrat in the<br />

Michigan countfes and Ontario areas<br />

borderfng <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> is glven in Table<br />

24. <strong>The</strong> harvest from Mlchigan and Ontario<br />

marshes can exceed 100,000 individuals per<br />

year. With a pelt plus carcass price <strong>of</strong><br />

approximately $10.00 per animals muskrat<br />

trapping Is a million dollar a year<br />

Industry in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> region,<br />

Jaworskl and Raphael (1978) observed that<br />

furs from the coastal wetlands are <strong>of</strong> high<br />

quality8 hence the fur prices in these<br />

areas are usually high. <strong>The</strong> recent<br />

decline In Ontario production is<br />

attributed to roughly a 30% decline in<br />

-<br />

Table 24. Muskrat harvest in Michigan and<br />

Ontario bordering Ldke <strong>St</strong>. <strong>Clair</strong> and<br />

connect1 ng waterways.<br />

Year <strong>St</strong>. <strong>Clair</strong> Macomb Wayne Total<br />

1965 2,675 1,272 112 4,059<br />

1966 15,515 4,084 2,700 22.299<br />

1967 24,335 2,658 392 27,385<br />

1968 13,395 5,469 1,006 19.870<br />

1969<br />

-<br />

16,063 6,012 1,484 23,559<br />

1970 30,350 3,479 1,149 34,978<br />

Annual<br />

mean 17,055 3,829 1,141 22,025<br />

Year Dover Twnshp. Walpole Is. Total<br />

1974 23,512 67 880 91,392<br />

1975 22,591 73 r 796 96,387<br />

1976 21,174 31,212 52,386<br />

1977 13 r 174 14,790 31,964<br />

1978 11,658 15,385 27 1 043<br />

1979 12,715 37,340 50,055<br />

1980 15,138 26,891 42 029<br />

1981<br />

1982<br />

8,886<br />

9,215<br />

26,363<br />

29,780<br />

35t249<br />

38,995<br />

Annual<br />

~ a r 15,785 ~<br />

35,937 51,722<br />

a Data sources: Jaworski and Raphael (19781<br />

and George Ducknorth, Ontar30 Ministry <strong>of</strong><br />

Natural Resources (pers, cm. ) .<br />

wetland area during the same period IG.<br />

Duckworthv Ontarlo Ministry <strong>of</strong> Natural<br />

Resources; pers. corn.<br />

Again* little direct information is<br />

available on the ecology <strong>of</strong> raccoons in<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands# but western <strong>Lake</strong><br />

Erie studies can be helpful in<br />

understanding this ant ma1 . Several as-<br />

pects <strong>of</strong> the life history <strong>of</strong> the raccoon<br />

in Sandusky Bay marshes were investigated<br />

by a trapping and telemetry study (Urban<br />

1968, 19701, <strong>The</strong> density <strong>of</strong> the raccoon<br />

population was estimated to be 17/km2.<br />

<strong>The</strong> juvenile to adult female ratio was<br />

1.2xl.0, indfcating a moderate<br />

productivity for the raccoon population.<br />

<strong>The</strong> mean weights <strong>of</strong> both adults and<br />

juveni 1es increased from sprlng untll<br />

early winter and then decreased over the<br />

winter.<br />

<strong>The</strong> telemetry portion <strong>of</strong> the study<br />

provided information on raccoon movements,<br />

home ranger and denning. General lyr raccoons<br />

spend the daytime period in or near<br />

dens. <strong>The</strong> amount <strong>of</strong> nocturnal movement is<br />

related to the size <strong>of</strong> the home range,<br />

Raccoons w i th 1 arger home ranges move<br />

greater distances. Raccoons move at a<br />

mean rate <strong>of</strong> 162 m/hr. Marshland f s the<br />

major habitat type encompassed in an<br />

average night <strong>of</strong> travel and the habitat<br />

type in which raccoons spend the most<br />

time. Raccoons spend approximately 73% <strong>of</strong><br />

the time in the vicinity <strong>of</strong> shallow water<br />

(Figure 43). Male juvenile raccoons<br />

disperse from the marsh in the fall.<br />

Raccoons do not appear to search out<br />

waterfowl nests, since little change<br />

occurs in their movement patterns when<br />

waterfowl nesting is initiated. Dikes<br />

receive high usage in proportion to the<br />

amount <strong>of</strong> area they represent in the<br />

marsh. Movements <strong>of</strong> female raccoons<br />

encompass more wooded area per night than<br />

the male raccoons. Home range for the<br />

average raccoon is 51 hectares in size.<br />

Food items <strong>of</strong> raccoons in Sandusky<br />

Bay marshes (<strong>Lake</strong> Erie) include muskrat,<br />

crayf ishr fish, duck eggs, p1 ant materf a?<br />

seeds, and birds (Andrews 1952~ Bednarik<br />

1956, Urban 1968). Fish, crayfish, and<br />

plant material were the chlef food items


In all seasons. Muskrat fur was found in<br />

only 8% <strong>of</strong> raccoon scats collected year<br />

round, but In 47% <strong>of</strong> scats collected in<br />

the sprfng. Andrews (1952) noted that<br />

chln~ney crayfish 1 Is a<br />

favorite food <strong>of</strong> raccoons in the summer,<br />

Raccoons were responsible for the<br />

termlnat-lon <strong>of</strong> 39% <strong>of</strong> the waterfowl nests<br />

built on the Sandusky Bay dikes in 1967<br />

and 1968 (Urban 1970).<br />

Raccoons are the most signlf icant<br />

furbearer In the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetlands<br />

Sn terms <strong>of</strong> pelt value <strong>of</strong> the total<br />

harvest. <strong>The</strong> density <strong>of</strong> raccoons in <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> wetlands has been estimated at<br />

about 0.3jhectare by Jaworskl and Raphael<br />

Figure 43. Raccoon (Procyon lotor), a (1978) <strong>The</strong>y fndlcate a value per pelt<br />

common predator <strong>of</strong> duck nests in <strong>Lake</strong> <strong>St</strong>. PIUS carcass <strong>of</strong> about $40.00 for the early<br />

<strong>Clair</strong> wetlands. to middle 1970s. <strong>The</strong>refore, the 38,000<br />

hectares <strong>of</strong> wetlands wfthfn the study area<br />

has a total raccoon value <strong>of</strong> about<br />

$456 r 000.


4.1 ORIGIN AND EVOLUTlOM OF LAKE ST.<br />

CLAlR WETLANDS<br />

<strong>The</strong> relationship <strong>of</strong> water levels,<br />

topography and the geomorphic devel opment<br />

<strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal zone is<br />

significant to b fogeographlcal development<br />

and maintenance <strong>of</strong> the wetlands and<br />

their areal development. This is particularly<br />

true for the <strong>St</strong>. <strong>Clair</strong> Deltar which<br />

is constantly changing its morphology.<br />

Because <strong>of</strong> shifting channels, variable<br />

flow regimes, and sedimentatfon, the<br />

geological nature, (and hence the<br />

b~ologlcal characteristic <strong>of</strong> the 1 f ttoral<br />

zone and wetlands) has not been constant.<br />

Sfnce not a11 rivers which debouch<br />

into a basin deposft deltas, the marine<br />

processes are slgniffcant. A delta is not<br />

solely a praduct <strong>of</strong> river deposition.<br />

Basicall yp del taf c 1 andforms are a product<br />

<strong>of</strong> both river and marine processes. <strong>The</strong><br />

rfver contributes the sediments to the<br />

receiving bas1 n. Whether the sediments<br />

accumulate in the mouth is dependent upon<br />

the marine processes such as wave action<br />

and long-shore currents? and the geo-<br />

morphlc character <strong>of</strong> the submarf ne topo-<br />

graphy. If there is a dominance <strong>of</strong> the<br />

marine process the sediments may be trans-<br />

ported down coast and a delta may not be<br />

extensive, Conversely? a rlver domlnated<br />

system contributing an excess <strong>of</strong> sediment<br />

coupled wjth a low wave climate w i l l<br />

prsmote deltaic development.<br />

Several <strong>of</strong> our concepts <strong>of</strong> delta<br />

development have been obtained from the<br />

M$ssiss$ppi deltafc plain, Because <strong>of</strong> <strong>of</strong>f<br />

and gas exploratfan, rfver and marine<br />

CQmm63rCer and f he extensive wet1 ands in<br />

the arear the Missfssippi River Delta has<br />

been extensive1 y studied geologicall yr<br />

blologica~lyr and geographically. It<br />

CHAPTER 4, ECOLOGllCAL PROCESSES<br />

serves as a model by which other marine<br />

and 1 acustrjne deltas are compared.<br />

Traditlonally deltas have been<br />

classified on the basis <strong>of</strong> their<br />

morphology and descriptive terms such as<br />

arcuate, b i rd-foot, and cuspate have been<br />

applied to the various delta shapes.<br />

Using this classification the <strong>St</strong>. <strong>Clair</strong>?<br />

like the Misslssippi Delta may be<br />

described as bird-foot, It is generally<br />

assumed that a given set <strong>of</strong> processes is<br />

responsible for each deltaic morphology.<br />

Although* morphological sf mil aritles are<br />

evident, significant differences also<br />

occur (Table 25).<br />

A unfque aspect <strong>of</strong> a delta's<br />

geomorphology is related to its changfng<br />

base levels. <strong>The</strong> base level <strong>of</strong> all marine<br />

deltas appears to have been eustatf call y<br />

stable over the past 4*000 years. In the<br />

Misslssippi Delta? because <strong>of</strong> the stable<br />

sea level? the ancestral streams deposited<br />

a serles <strong>of</strong> delta lobes along the<br />

Louisiana coast. A t <strong>Lake</strong> <strong>St</strong>. Clalr,<br />

changing base levels have produced two<br />

distinct deltas at different levels, the<br />

premodern and the modern. Examples <strong>of</strong><br />

this base level change are apparently rare<br />

with regard to deltaic deposition.<br />

Marine deltas are <strong>of</strong>ten characterized<br />

by subsidence due to sediment loading.<br />

Geosyncl i nal downwarping and sediment<br />

compaction in response to sediment ac-<br />

cumulation in the Mississippi Delta has<br />

allowed up to 122 m <strong>of</strong> deltaic deposits to<br />

accumulate since the termination <strong>of</strong> the<br />

Late Wisconsin glacial ice (Morgan 1970).<br />

With continued sedimentation and<br />

subsidence, the shtftjng delta lobes<br />

overlapped# burying older deltaic<br />

envf ronments. <strong>The</strong>reforer the traditional<br />

sedimentologlcal units, top-setr fore-set,


Table 25. Factors influencing lake and marine delta morphology. a<br />

Factors Characteristics <strong>St</strong>. <strong>Clair</strong> Delta Mississippi Delta<br />

Geomorphic: Base 1 eve1 Long- and short-term <strong>St</strong>able<br />

variations<br />

Flow regime Seasonally constant Seasonally variable<br />

Channel d iversl on Downstream (spring Wfthfn alluvial<br />

ice jams) valley (sprfng floods)<br />

<strong>St</strong>ructural behavior <strong>St</strong>able<br />

<strong>of</strong> basin <strong>of</strong> deposition<br />

Sediment compaction<br />

and significant<br />

subsidence<br />

Dominant sedlment size Fine to coarse sand Clay to silt<br />

Relative rate <strong>of</strong> S1 ow<br />

sedimentat ion<br />

Marine: Relative wave energy Low Low<br />

Rapid<br />

Offshore pr<strong>of</strong>ile Flat to gently Flat to gently<br />

slopfng sloping<br />

Vegetative: Vegetatlon Deciduous trees to Predominantly fresh<br />

zonation freshwater marsh to brackish to salt<br />

marsh<br />

Control <strong>of</strong> vegetation Short-term 1 ake level Salinity, substrate<br />

distribution changes composition, and tidal<br />

range<br />

a Data source: Jaworski and Raphael (1973).<br />

and bottom-set beds, identified on the<br />

margins <strong>of</strong> <strong>Lake</strong> Bonneville by Gilbert<br />

(1890) do not occur in the Mississfppi<br />

Delta complex.<br />

Subsurface data reveal that the <strong>St</strong>.<br />

<strong>Clair</strong> Delta is stable. Sediments ac-<br />

cumulating in the delta are not <strong>of</strong><br />

sufficient thickness to allow any<br />

significant downwarping to occur, Borfngs<br />

on several beaches suggest that the base<br />

<strong>of</strong> these depositional features Is at<br />

apgroxlmately low water datum. Apparently<br />

the beaches have not subsided as the<br />

cheniers have done on the flanks <strong>of</strong> the<br />

Mississippi Delta,<br />

Isostatic rebound i n the <strong>St</strong>. Clafr<br />

basin, due to the waning <strong>of</strong> Wisconsin ice<br />

fo1 lowing the Valders maximum some 10~500<br />

to 1Zt500 years ago, has been minlmaf ,<br />

Pertinent zero isobases since the Valders<br />

maximum ice advance are the Algonquin and<br />

Nipissing hinge 1 friest which are oriented<br />

east-west and located in the central<br />

portion <strong>of</strong> <strong>Lake</strong> Huron. <strong>The</strong>refore, crustal<br />

displacement due to regional glacial<br />

rebound or 1 ocal ired sed imen%atlcn does<br />

not appear to have direct1 y influenced the<br />

vertical and horizontal morphology <strong>of</strong> the<br />

<strong>St</strong>. Claf r Dslta, <strong>The</strong> <strong>St</strong>, <strong>Clair</strong> Delta has*<br />

in fact, extended itself across an es-<br />

sentiall y rlgid platform.


Although structural stabil tty and<br />

mfnimum subsfdence characterize the <strong>St</strong>.<br />

<strong>Clair</strong> Delta, significant changes in the<br />

level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> in the past few<br />

millenla have contributed to the mor-<br />

phology <strong>of</strong> the delta. Based upon a<br />

structural framework* deltas may be de-<br />

posited in one <strong>of</strong> three geologic<br />

envlronments: a subsiding area, a stable<br />

area, or an elevating area. <strong>The</strong><br />

Mtssissippf Delta with its thick ac-<br />

cumulation <strong>of</strong> recent sediments* is a<br />

classic example <strong>of</strong> a subsiding delta.<br />

Although the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> basin is<br />

geologically stable, the effect <strong>of</strong> a "tower<br />

lake level on the delta complex is similar<br />

to elevatlng the 1 and, Since base level<br />

in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> has dropped in the 1 ast<br />

4,000 years, the focus <strong>of</strong> depostion has<br />

shifted lakeward creating the modern <strong>St</strong>,<br />

Cl ai r Delta and wetl and habitat.<br />

<strong>The</strong> configuration <strong>of</strong> deltas is<br />

related to <strong>of</strong>fshore slope" relattve wave<br />

power in the nearshore zone, and riverine<br />

Influence (Wrlght and Coleman 1972). A<br />

delta such as the Mississippi with its<br />

digltal distributaries and marshy<br />

embayments 1s dominated by Its rlver since<br />

Its <strong>of</strong>fshore slope is flat to convex and<br />

the nearshore wave power is low. Although<br />

long-term wave data have not been recorded<br />

for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, wave energy must be<br />

considered to be low since the maximum<br />

fetch for wave generation for the delta is<br />

only 40 km. <strong>The</strong> subaqueous delta front is<br />

shallow and concave, causing waves<br />

generated In the shallow lake to attenuate<br />

lakeward <strong>of</strong> the delta, Also, marshes have<br />

become establ ished on exposed <strong>of</strong>f shore<br />

areas between Johnson Bay and the -2 m<br />

contour suggesting that law wave energy<br />

conditfons prevafl, Although there may be<br />

an occasional dominance <strong>of</strong> martne<br />

pracessss as evidencad by the presence <strong>of</strong><br />

transgressive beaches, f t must be<br />

concluded that the geometry <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> Delta, like that <strong>of</strong> the Mississippi,<br />

is prfncfpally dTctated by Its rfver,<br />

<strong>The</strong> pkevtous discussion suggests that<br />

although the <strong>St</strong>, Clafr and Mississippi<br />

river deltas have similar appearances<br />

several dissfmf l ar features and p rocssses<br />

prevafl . For example, the domfnant<br />

sediment size in the Misslssfppi Delta 1s<br />

silt to clayo whereasr the <strong>St</strong>, <strong>Clair</strong> Delta<br />

Is fine t o coarse sand. Alsos the<br />

relative rate <strong>of</strong> sedfmentation in the <strong>St</strong>.<br />

Clafr Delta is slow compared to the rapid<br />

rate <strong>of</strong> the Misslssfppi Delta. <strong>The</strong><br />

wetl ands dlscussed in this community<br />

pr<strong>of</strong>ile exhibit a dfversity <strong>of</strong> form.<br />

However, thef r distributfon can be re1 ated<br />

to fundamental geomorphic features some <strong>of</strong><br />

which are unique. Jaworski et a1. (1979)<br />

examined and developed geomorp hic model s<br />

for the coastal wetlands <strong>of</strong> the Great<br />

<strong>Lake</strong>s as a step to understanding their<br />

origin, stability, and distribution.<br />

<strong>The</strong> occurrence, distrfbution, and<br />

diversity <strong>of</strong> coastal wetlands are, in<br />

part, determined by the morphology <strong>of</strong><br />

coastal <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Perhaps in no<br />

other b f ogeograp hic envi ronment i s the<br />

relationship between 1 and forms and<br />

vegetation so evident. All wetlands occur<br />

in topographical depressions created by<br />

nature or more recently by humans. <strong>The</strong>se<br />

topographical lows may be erosional fn<br />

origin such as river channels or<br />

depositional fn origin such as lagoons or<br />

floodbasins.<br />

Although elevations <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

Delta are low, a diversity <strong>of</strong> landforms<br />

exists (Sectlon 2.1) . When the entire<br />

shorel ine <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl air is considered,<br />

the habitat diversity is even greater.<br />

Twelve wetl and sett 1 ngs have been<br />

identified in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Figure 44).<br />

A l l but one s e t t i n g (f.e.,<br />

diked/disturbed) are <strong>of</strong> a natural origin.<br />

<strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta exhibits a greater<br />

diversity <strong>of</strong> wetland settings because it<br />

is a product <strong>of</strong> river as we71 as marine<br />

processes. <strong>The</strong> habitats beyond the<br />

deltaic plain are fewer in number but<br />

unfque to the lake shoreline. A third<br />

suite <strong>of</strong> wet1a:jd habltats occurs both In<br />

the delta complex and on the shorel jne.<br />

<strong>The</strong> followfng section discusses the<br />

12 wetland habitats recognized in <strong>Lake</strong> <strong>St</strong>.<br />

Claf r. <strong>The</strong> deltaic habitats are descrfbed<br />

first followed by the coastal envi ronmsnts<br />

<strong>of</strong> wetland occurrence. Finally, wetl ands<br />

occurrfng in both the <strong>St</strong>. Clafr deltaic<br />

and coastal plains are described:<br />

1, are shall ow submerged<br />

features <strong>of</strong> the df stributary channels


1. River shoulders<br />

<strong>Lake</strong><br />

4. Crevasse deposits<br />

5. Shallow interdistributary bays<br />

6. Deep water interdistributary bays<br />

7. Transgressive beaches<br />

10. Transitional areas<br />

Premodern<br />

2. Abandoned channels<br />

8. Shallow shelves<br />

9. Barrierllagoons<br />

<strong>Lake</strong><br />

Plain<br />

12. Diked areas<br />

Figure 44, Twelve wetland habitats recognized in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

<strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> De1 ta. Paralleling<br />

the shore1 ine <strong>of</strong> the channels, the<br />

submerged river shoals extend 35 m into<br />

the rlver and have maximum depths <strong>of</strong> 2<br />

m. <strong>The</strong> sediment consists <strong>of</strong> fine sand.<br />

<strong>The</strong> feature is reasonably continuous<br />

along the length <strong>of</strong> the channels but fs<br />

normally absent on the outsfde or<br />

cutbank side <strong>of</strong> the channel. <strong>The</strong><br />

feature attains a maximum width at<br />

submerged point bars. At the channel<br />

shoreline, reeds such as the robust<br />

are dense (Figure<br />

29); however, submergent aquatics are<br />

dominant. An aggressive fnvader,<br />

Euraslan water-mll fol 1 has become more<br />

abundant in the last decade in the<br />

distributary channels (Schloesser and<br />

1 1. Eustuarinelflood<br />

basin<br />

Manny 19823, It appears to be a common<br />

submersed macrophyte and a minor<br />

nuisance to recreational boating In the<br />

delta region as well as in Anchor Bay,<br />

Furthermore, the plant denslty appears<br />

to be greater in the non-commercial<br />

navigation channels (i.e., North and<br />

Middl e channels).<br />

2. dissect the<br />

up1 and surface <strong>of</strong> the delta and merge<br />

into the modern delta when traced<br />

lakeward. <strong>The</strong> channels are long and<br />

linear and <strong>of</strong> varfable width. <strong>The</strong><br />

1 argest channel on Dickfnson Island is<br />

some 500 m wjde, but usual channel<br />

wfdths are about 100 m. <strong>The</strong> sediments<br />

have a fine texture and a high organfc


content suggesting gradual decay over<br />

time, <strong>The</strong> boundary between the<br />

premodern delta upland and the<br />

abandoned channel is usual1 y distinct<br />

especial1 y at the apex <strong>of</strong> the delta.<br />

During years <strong>of</strong> low lake level several<br />

<strong>of</strong> the ancient channels are above the<br />

water table and plant stress may occur.<br />

Emergents such as hard-stem bul rush<br />

(Scfrow+ -1, arrowhead (<br />

lati f ol la), and three-square<br />

colonize the deeper water<br />

nels (1 m) as do water<br />

ccasionally buttonbush<br />

3. are generally<br />

similar to the abandoned<br />

channels discussed except that they are<br />

st111 in the process <strong>of</strong> decay.<br />

Chematogen Channel 3s a dlstrlbutary<br />

channel which contfnues to conduct <strong>St</strong>.<br />

Clalr Rlver water to <strong>Lake</strong> <strong>St</strong>. Clalr.<br />

<strong>The</strong>se channels are malnl~ on the<br />

Ontario sido <strong>of</strong> the delta ind were<br />

responslble for the depositfon <strong>of</strong> the<br />

Canadian portion <strong>of</strong> the delta. Since<br />

that time this area was abandoned and<br />

the more recent depositlonal act iv'i ty<br />

began in Michfgan. As abandonment<br />

occurred, the channels were gradual 1 y<br />

s l l t e d so that only narrow<br />

distributaries remain today. Such<br />

channels not only dlssect the premodern<br />

delta but the modern surface as well.<br />

<strong>The</strong> sedlment <strong>of</strong> the channel fflls .Is<br />

fine sand, but some silts have been<br />

deposited from the adjacent 1 ake pl afns<br />

as well. Since these channels occur on<br />

the modern delta, slopes are more<br />

gentle. In contrast abandoned river<br />

channels are on the premodern surface<br />

and slopes to the channels more<br />

distinct. <strong>The</strong> channel fills <strong>of</strong> the<br />

recent1 y abandoned channels are <strong>of</strong>ten<br />

submerged beneath 15 cm <strong>of</strong> water and<br />

colonized by sedges E- spp.1 and<br />

more sporad?call y by blue-joint grass<br />

spp. and pondweeds (e.g., curly<br />

pondwsed ~ r i s ~ colon9 u ) ze<br />

the narrowfng active channels.<br />

occur on the f7anks<br />

<strong>of</strong> dfstrlbutary bays and have a lobate<br />

form, Since they are flood or hlgher<br />

energy deposits they are composed <strong>of</strong><br />

coarser sand. Channel depths are<br />

approximatel Y 4 m and are not<br />

COI onjzed, but the sand deposits<br />

adjacent to the "highwaysn support<br />

commun itf es <strong>of</strong> bul rush ~PP. 1.<br />

Most crevasse deposits are at the lower<br />

end <strong>of</strong> the active dfstributary channels<br />

and thus exposed to wave act Ion.<br />

Because <strong>of</strong> higher wave energies and<br />

perhaps due t o Ice damage and<br />

recreational boating the emergent<br />

aquatics are not particularly dense,<br />

<strong>The</strong> landward side <strong>of</strong> the crevasse<br />

deposfts appear to be better protected<br />

from wave action and thus support a<br />

denser stand <strong>of</strong> emergent vegetation<br />

including not only bulrush fn deeper<br />

habitats but cattails (Iy&h spp.) in<br />

shallower areas <strong>of</strong> the deposit. Also<br />

crevasse deposits closer to <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> (farther down the distributary<br />

channels) are exposed to greater wave<br />

actfon and are sparsely colonized with<br />

bulrush.<br />

aallow interdlstributgILy_ bavg occupy<br />

extensive areas between the open lake<br />

and the equally obvious higher<br />

premodern deltas. On the flanks these<br />

shallow wetland hab i tats are bordered<br />

by natural levees. Occasionally,<br />

particularly in Michigan, narrow<br />

discontinuous beaches occur. <strong>The</strong><br />

sediment In the bays was derived from<br />

crevasses during high water periods and<br />

are thus silty-sand size particles and<br />

mostly mineral in character. Organic<br />

sedfmsnts are sparse and discontinuous.<br />

A t the surface the organic<br />

accumulatlonr where present, ranges in<br />

thickness from 5 to 15 cm. Water<br />

depths average 5 to 30 cm. <strong>The</strong> shallow<br />

Interdlstrfbutary bays are almost<br />

excl us I vel y col on i zed by dense stands<br />

<strong>of</strong> cattails <strong>of</strong> which the hybrid TvDha x<br />

standlng 2 to 3 m high<br />

dominates. A t slightly higher<br />

elevations sedges are interspersed with<br />

cattafls. <strong>The</strong> high cattail produc-<br />

tivity and the low organic accumulation<br />

suggests that these habitats are<br />

flushed durJng high water periods. <strong>The</strong><br />

sedimentation process is relatively<br />

complete throughout the deltaic plain,<br />

though a few small lakes (e.g., Goose<br />

lake) stfll persjst.


interdfstrf butary bays. <strong>The</strong> landward<br />

slde is bordered by low, poorly<br />

developed transgressive beaches. <strong>The</strong>ir<br />

flanks are characterized by slightly<br />

higher, poor1 y developed natural levees<br />

which are permanently breached by<br />

crevasse channels. <strong>Lake</strong>ward these bays<br />

are exposed to wave action <strong>of</strong> the open<br />

lake. As clastic sediments are<br />

introduced into these bays, the basins<br />

w i l l evolve Into shallow interdistributary<br />

bays. Water depths range<br />

from 0.5 to 1.75 m and the bay bottom<br />

consists <strong>of</strong> fine sand. Occasionally<br />

in sheltered areas si 1 t deposits w i 11<br />

accumulate, but organic deposits are<br />

sparse through the habitat. Wave<br />

action prohibits the colonization <strong>of</strong><br />

dense emergents; however, n! spp.<br />

are prevalent. <strong>The</strong> deep water bays are<br />

confined to the Michigan portion <strong>of</strong> the<br />

de1 ta where the distributary channels<br />

are most active and where natural<br />

levees extend into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Common submersed aquatics include<br />

Characeae and f f s<br />

(Schloesser and Manny 1982). <strong>The</strong><br />

d istributjon <strong>of</strong> submersed macrophytes<br />

may be due to a lack <strong>of</strong> suitable<br />

substrate for attachment or excessive<br />

wave actlon.<br />

are best<br />

developed wlthin the <strong>St</strong>. <strong>Clair</strong> Delta<br />

but do sporadically occur along the<br />

Ontario shore1 ins. <strong>The</strong> beaches are<br />

wave deposited features standing<br />

approximately 30 cm above the lake. As<br />

noted in Figure 4, older beach deposits<br />

occur landward <strong>of</strong> the active<br />

transgressive beach in the shallow<br />

i nterdistributary bays. <strong>The</strong>y are<br />

composed <strong>of</strong> stratified coarse sand,<br />

gravel and organic debris. Although<br />

they vary in width, most beaches are<br />

about 5 m wfde. Borings through the<br />

beach deposits reveal alternating<br />

layers <strong>of</strong> coarse clastics and thin (1<br />

cm) wave deposited organic 1 enses.<br />

This clearly suggests that the features<br />

were deposited during periods <strong>of</strong><br />

erosion. During higher wave energy<br />

conditions the beaches are not<br />

breached; but rather overwash occurs,<br />

causing rafted debris to be deposited<br />

atop the shallow interdistributary<br />

marsh. Woody vegetation includfng<br />

eastern cottonwood (P-, 1,<br />

7. *<br />

willows (531 i x spp.) and staghorn sumac<br />

(m colonize the beaches.<br />

During higher water years sedges (Carex<br />

strictq) r reed-canary grass (<br />

udinace_a), and jewelweed (-<br />

spp.) commonly colonize the beaches.<br />

<strong>The</strong> latter three species are more<br />

common on the overwash deposits<br />

landward <strong>of</strong> the beaches.<br />

8. Shallow shelf enviroqg&i& border the<br />

nearshore zone <strong>of</strong> most <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>. <strong>The</strong> shelves are composed <strong>of</strong><br />

silt-sized clastic fragments. <strong>The</strong><br />

shelves are approximately 200 m wide,<br />

1.5 m fn depth and are turbid<br />

especially following storms. <strong>Wetlands</strong><br />

colonizing this habitat exist as a low<br />

dense fringe parallel to the shore and<br />

remaln exposed to lake effects such as<br />

wave action. This habitat is best<br />

displayed along the eastern shore <strong>of</strong><br />

the lake and is colonized wfth<br />

emergents and submergents. <strong>The</strong><br />

dominant vegetation is emergent<br />

macrophytes such as cattail. Other<br />

wetland species include lake sedge<br />

(Carex -1, large stands <strong>of</strong><br />

pickerel weed (Wderia EPCP-) ,<br />

water-mil foil ( k l v r l m<br />

hetero-<br />

-1, and white water lily<br />

(Nvmr>haea odorata) (Mudroch 19811,<br />

9, Barrier/l a-lexs~ occur south <strong>of</strong><br />

Mftchell Bay along the eastern<br />

shoreline <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong><br />

barriers are poorly developed and stand<br />

approximately 50 cm above the lake.<br />

However, they do provide some<br />

protection from wave action and<br />

seiches, especially during 1 ower water<br />

years. <strong>The</strong> lagoon is approximately<br />

1,000 m in width and i s characterized<br />

by a si 1ty bottom. Morphologically<br />

this habitat is similar to the shallow<br />

she1 f environment except for the<br />

occurrence <strong>of</strong> a low barrier, <strong>The</strong><br />

bottom sediments not only Include silts<br />

but 1 oose, partial 1 y decomposed organic<br />

fragments which may be j.~ or may<br />

have been transported into the lagoon<br />

by streams from the adjacent lake<br />

plain. Emergent macrophytes dominated<br />

by cattalls ( 1 occur in<br />

the area. <strong>The</strong> landward side <strong>of</strong> these<br />

wetlands has a gentle natural slope<br />

which in most fnstances along <strong>Lake</strong> <strong>St</strong>,<br />

<strong>Clair</strong> is abrupt1 y termfnated by


artificial dikes. It is important to<br />

note that the Ontario shoreline north<br />

<strong>of</strong> the Thames River has not experlenced<br />

signlf lcant coastal recession in recent<br />

years (Boulden 1975) whereas the south<br />

shore <strong>of</strong> the 1 ake has. This suggests<br />

that the barrier and adjacent wetlands<br />

retard wave action on this coastal<br />

reach.<br />

10. are broad zones<br />

i a1 wetl ands from<br />

well-drained upland surfaces. In some<br />

instances, such as on Harsens Island<br />

and along the eastern shoreline <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>r the transitional areas have<br />

been incorporated in diked farmland or<br />

water level-control led wetl ands and<br />

thus do not exist. However, good but<br />

rather iselated examples occur on<br />

Dickinson Island and east <strong>of</strong> <strong>St</strong>. John's<br />

Marsh in -bIichigan. <strong>The</strong> sediments<br />

consist <strong>of</strong> fibrous peat macr<strong>of</strong> ragments<br />

which are underlain by fine# mottled<br />

sand. <strong>The</strong> transition zone is normal 1 y<br />

not permanent1 y flooded and water<br />

tables are about 15 to 30 cm below the<br />

surface. <strong>The</strong> lakeward slde <strong>of</strong> this<br />

environment Is characterized by sedge<br />

sediments are derived from flood events<br />

and are usually fine sand and stlt wlth<br />

a sign1 f icant organic content. Natural<br />

levees adjacent to the channel are<br />

poorly developed and vegetation<br />

zonation difficult to detect. <strong>The</strong><br />

water level -in this wetland habitat is<br />

vartable and depends In part on the<br />

rdver dfscharge. Durlng dry years the<br />

wetlands may be stressed because <strong>of</strong> a<br />

low water table. <strong>The</strong>refore the wetland<br />

may be seasonal in nature. A t the<br />

confluence <strong>of</strong> river and lake, water<br />

tables are most <strong>of</strong>ten at or very near<br />

the surface and the floodbasin w i l l<br />

sustain dense emergents in most years.<br />

At the mouth <strong>of</strong> the Clinton Rlver dense<br />

stands <strong>of</strong> cattails have been observed<br />

for many years. Upstream the marshes<br />

grade into bottom1 and swamps.<br />

-.<br />

12. are a cultural modification<br />

<strong>of</strong> a wetland rather than a<br />

physical entity. Included in this<br />

category are transitional zones which<br />

have been dfked and drained for farming<br />

or recreational purposes. Since these<br />

tussocks (u spp.) interspersed with<br />

sparse cattails. Where water tables<br />

wetlands transcend physical boundaries<br />

such as soil/sediment types and slopest<br />

their morphologfcal framework is less<br />

are deeper, the transition zone is meaningful. On Harsens Island the<br />

colonized by a meadow which includes Michigan Department <strong>of</strong> Natural<br />

but is not limited to red osier dogwood<br />

(Carnus &lonifera)r gray dogwood (C.<br />

Resources has diked and drained much <strong>of</strong><br />

the shallow distributary bay area and<br />

1, panic grass (j2gmiam spp.1. the transition zone. <strong>The</strong> land cover <strong>of</strong><br />

and bluejoint grass. On the landward the central portion <strong>of</strong> the island is<br />

embl ing aspen (m in corn and grain crops which are utilis<br />

abundant. Because <strong>of</strong> ized as part <strong>of</strong> a waterfowl hunting<br />

the variable elevation and depth to<br />

groundwater thJs zone exhibits a<br />

management scheme. Wal pole Is1 and is<br />

the largest island in the delta. A<br />

diversity <strong>of</strong> wetl and plants. Similar large part <strong>of</strong> the island, which is now<br />

wetland species have been observed on<br />

the premodern surface <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

cultivated mainly for corn and soybean,<br />

was a1 so a shallow interdisciplinary<br />

Delta. Here local depressfons are bay. <strong>The</strong> diked marshes along the east<br />

underlain by clay pan which results in<br />

a perched water table which commonly<br />

shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> have long been<br />

used for private duck hunting. Wasupports<br />

dogwood/meadow vegetation,<br />

ter level in these marshes is regulated<br />

by pumps, dikes, and canals. <strong>The</strong><br />

fl, habltats are<br />

characterized by marsh vegetation<br />

marshes are landlocked and water depths<br />

range from 10 to 50 cm (Mudroch 1981).<br />

bordering a rfver which debouches into<br />

the lake, <strong>The</strong> Clinton River and<br />

Emergents colonize these marshes. Dominant<br />

macrophytes are the cattail and<br />

several small streams tributary to <strong>Lake</strong> sedge. Also large stands <strong>of</strong> pickerel<br />

<strong>St</strong>. Clafr support such wetlands, <strong>The</strong><br />

extent <strong>of</strong> the vegetated wetland is<br />

weed, water mil foil (M rio hyllum<br />

heteropbyll um) , and white*<br />

<strong>of</strong>ten restricted by a steep backslope occur in the channels within these<br />

paralleling the flood plain. <strong>The</strong> diked wetl ands.


Table 26 summarizes the identified<br />

morphological wetland settings Sn <strong>Lake</strong> <strong>St</strong>.<br />

Clalr in terms <strong>of</strong> selected physical<br />

characteristics, For thf s investigation<br />

the coastal and deltaic wetlands are<br />

classified into four major vegetative<br />

zones:<br />

1. Submergent (e.g., pondweeds,<br />

coontai 1 1<br />

2, Emergent (e.g.$ cattail, bulrush)<br />

3. Sedge/meadow (e.g. sedge* canary<br />

grass)<br />

4. Shrub (e.g., dogwood, willow, sweet<br />

gal el.<br />

Wetland diversity exhibited by submergent,<br />

sedgep and shrub vegetation is most<br />

evident in the abandoned channels (2) and<br />

the transitional (10) zones. Here,<br />

topography is variable, wave or river<br />

current action is low or not applicable<br />

and turbidity is low. Conversely the<br />

wetl and community is less diverse along<br />

river shoulders (l), crevasse (4), and<br />

estuarine/flood basfn (11) environments.<br />

<strong>The</strong>se rather monotypfc wetlands appear to<br />

be assocfated with hiqher wave/current<br />

action and higher turbidity levels. <strong>The</strong><br />

exception is the shallow interdistributary<br />

bay (5) environment which is composed <strong>of</strong><br />

dense and robust stands <strong>of</strong> cattails.<br />

Perhaps the combined perfodic exposure and<br />

a hlgher organic content <strong>of</strong> the substrate<br />

account for the dominance <strong>of</strong> the<br />

emergents.<br />

4.2 BlOLOGlCAL PRODUCTION<br />

Examination <strong>of</strong> energy flow in coastal<br />

wetlands shows that the entire<br />

heterotrophic component <strong>of</strong> the wetland Is<br />

dependent on organic matter produced<br />

during photosynthesis. In many cases,<br />

utilization <strong>of</strong> the material is through<br />

grazing <strong>of</strong> lfving tissues. Many species<br />

<strong>of</strong> waterfowl feed on various parts <strong>of</strong><br />

aquatfc hydrophytes. <strong>The</strong> seeds <strong>of</strong><br />

muskrat, mice, and deer browse heavily on<br />

several species <strong>of</strong> aquatic hydrophytes.<br />

Phytophagous fnsects are the food supply<br />

for several species <strong>of</strong> birds, fishes,<br />

reptiles, and mammals (Tilton et ale<br />

1978).<br />

A more significant form <strong>of</strong><br />

uttlization is the direct consumption <strong>of</strong><br />

detritus or alternatively, the microbial<br />

populations associ ated with organic<br />

particulate matter. Numerous inverte-<br />

brates re1 y completely on these food<br />

sources. <strong>The</strong>se organisms in turn form the<br />

food supply for fish, amphibians,<br />

reptiles, birds, and mammals.<br />

In addition to plants being a direct<br />

or indirect food source for many species<br />

<strong>of</strong> animals, aquatic plants provSde cover<br />

and nesting areas for waterfowl. Several<br />

species <strong>of</strong> fish9 such as the northern pike<br />

(b 3 uc i spawn in vegetated wetl and<br />

areas and muskrats (w &ethiu)<br />

prefer emergent vegetation for construction<br />

<strong>of</strong> feeding platforms and houses.<br />

Nutrient cycl ing is important to the<br />

continuation <strong>of</strong> primary and secondary<br />

production in wetlands (Figure 45).<br />

Coastal wetland vegetation immobilizes<br />

certain amounts <strong>of</strong> nutrients, a portlon <strong>of</strong><br />

which are released upon senescence and<br />

decay <strong>of</strong> the plants. Depending on the<br />

sedimentation characteristlcs <strong>of</strong> the<br />

wetlandsr nutrients are stored in the<br />

wetland as organic sediment or peat. In<br />

the marsh sol 1 sr microb 1 a1 processes<br />

transform some <strong>of</strong> the nutrients from<br />

organfc to inorganic forms. <strong>The</strong> net<br />

effect <strong>of</strong> these processes is generally a<br />

reduction In the concentration <strong>of</strong><br />

nutrients in water flowing through the<br />

wetl and. <strong>The</strong>refore, the coastal marshes<br />

are important in control 1 i ng nutrient<br />

loading to nearshore waters <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

Clai r (Tilton et a?. 1978).<br />

<strong>St</strong>udies <strong>of</strong> metals uptake by aquatic<br />

plants provide some information on<br />

nutrient cycling. Mudroch and Capobianco<br />

(1978) studied the uptake <strong>of</strong> metals from<br />

the water and sedtrnent by emergent,<br />

submersedl and floatfng-leaved plants fn<br />

marshes along the Ontario shore <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. Cl a i r. <strong>The</strong>y found that submersed


Table 26. Morphology and relative physical relationships <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong><br />

wetlands.<br />

Wet1 and Water Dominant Current/<br />

envl ronment Substrate deptha vegetatf on wave actionb Turbidfty<br />

(m)<br />

1. River shoulders sand -6.0 submergent high/l ow moderate<br />

2. Abandoned river silt/ -1.0 emergent/ 1 ow/NA 1 ow<br />

channel s organics subme rgent/<br />

sedge/scrub<br />

3. Recent1 y aban- silt/ -0.5 emergent/sedge med/NA high<br />

dwned channel s organics<br />

4. Crevasse deposlts sand -1.5 emergent high/med high<br />

5. Shallow intar- fine sand/ -0.3 emergent 1 ow/low 1 ow<br />

distributary bays arganics<br />

6, Deep water inter- f Ins sand -1.5 emergent/ high/hfgh high<br />

distributary bays 5u bme rgent<br />

7. Transgrasslve coarse i-0.5 sedge/scrub low/high N A<br />

beach deposlts sand/<br />

organics<br />

8. Shal low shelves silt/cl ay -1.0 emergent/ low/high htgh<br />

submergent<br />

9. Barrler/lagoon si lt/cl ay -1.0 emergent/ low/Jow hlgh<br />

compl ex submergent<br />

10. Transi tlonal fine rand +0.3 emergent/ NA/NA N A<br />

areas submergent<br />

11, Estuarfne<br />

floodbasin<br />

sand/s i 7 t +O. 2 emergent high/NA high<br />

12. Diked areas variable -0.1 emergent/ low/l ow moderate<br />

submergent<br />

" Below level <strong>of</strong> lake<br />

+ Feature usually above lakt3 level.<br />

NA Not appl fcable.


plants accumulated the highest concen-<br />

trations, considerably higher than the<br />

source water and sediment, and that these<br />

elements were retained in the plant<br />

tfssues until decay processes incorporated<br />

them into surface sediments.<br />

1. ENERQV 3. WATER 6. SEDIMENTS<br />

2. GASES 4. NUTRIENT$ 8. ANIMALS<br />

INVERTEBRATE<br />

Figure 45. Cycling <strong>of</strong> energy and material<br />

through the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal wetland<br />

ecosystem (Ti1 ton et a1 . 1978).<br />

Mudroch (1980) also studied the<br />

geochemical composition <strong>of</strong> sediment,<br />

uptake <strong>of</strong> nutrients and metals by<br />

macrophytes, and nutrient and metal<br />

composition in the water at Big Creek<br />

Marsh on the northeast shore <strong>of</strong> <strong>Lake</strong> Erie<br />

in Long Point Bay. <strong>The</strong> maximum<br />

concentrations <strong>of</strong> most metals (Pb, Nip Car<br />

Crr and Zn) in marsh sediments were loner<br />

than concentrations found in <strong>Lake</strong> Erje<br />

surf ici a1 muds, presumably due to uptake<br />

by the aquatic plants In the mud,<br />

Submerged plants (Ghim Spet Mvr1clgMlhm<br />

sp., and sp.) accumulated larger<br />

quantities <strong>of</strong> Cat Pbr Cur Nf, Cr, and Cd<br />

than emergent plants (m spp.1, but<br />

nutrfent concentrations were relatively<br />

uniform for all species (Table 27). She<br />

found that the biomass production <strong>of</strong> the<br />

macrophytes In the marsh was related to<br />

the subhydric soil fertility as we11 as<br />

the nutrient content in the marshwater.<br />

She concluded that short-term supplies <strong>of</strong><br />

nutrient-rich sewage to the surface<br />

subhydrlc soil layer can have a prolonged<br />

effect on macrophyte production by<br />

enrichment <strong>of</strong> the nutrient pool maintained<br />

In the perennial plant system.<br />

<strong>The</strong> flow <strong>of</strong> energy through a food<br />

chain is <strong>of</strong>ten represented by a pyramid<br />

which illustrates the quantitative<br />

relatfonshfps among the varfous trophic<br />

levels (Figure 46). Juday (1943) was one<br />

<strong>of</strong> the earliest investigators to Introduce<br />

trophic levels concepts, developed the<br />

Table 27. Concentrations <strong>of</strong> nutrients and metals in wetland plarlts at the<br />

stage <strong>of</strong> maximum development, Big Creek Marsh, Long Point, Ontario.<br />

Species<br />

a Data source: Mudroch (19801,<br />

-<br />

K N P C a<br />

-- - - .-. - -----


'-\<br />

--._--_------SOLAR ENERGY<br />

Figure 46. Energy pyramid in <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> coastal marshes (Upper Great <strong>Lake</strong>s<br />

Regional Commission).<br />

concept from studies <strong>of</strong> freshwater<br />

wetlands. He determined the vardous<br />

components <strong>of</strong> the aquatic population in<br />

Weber <strong>Lake</strong>, Wisconsinr as they existed in<br />

midsummer. <strong>The</strong> dissolved organic matter<br />

composed about 60% o f the total pyramid,<br />

the fish9 only 0.5Xr and the other animals<br />

slightly less than 5% <strong>of</strong> the total.<br />

In coastal marshes there are <strong>of</strong>ten<br />

four major sources <strong>of</strong> energy for aquatic<br />

consumers: 1) marsh detritus, 2) phyto-<br />

plankton production, 3) detritus from<br />

terrestrial sources brought in by drainage<br />

and, 4) planktonic material carried into<br />

the marsh from the open lake, Although<br />

much research remains to be done on food<br />

chain production and ecosystem energy<br />

re1 ationshi ps9 particular1 y <strong>of</strong> freshwater<br />

wetlandsr there are a few general<br />

principles which appear to have validity:<br />

PI food cycles rarely have more than five<br />

trophic levels, 2) the greater the<br />

separation <strong>of</strong> an organism from the basic<br />

source <strong>of</strong> energy (solar radiation), the<br />

less the chance that it w i l l depend solely<br />

upon the precedjng trophic 1 eve1 for<br />

energy* 3) at successively higher levels<br />

in the food cycle, consumers seem to be<br />

progressive1 y more efficient 4n the<br />

utilization <strong>of</strong> food supply, and 4) In<br />

wetl and successl on productivity and<br />

photosynthetic efficiency increase Prom<br />

01 igotrophy to eutrophy and then decl ins<br />

as the marsh undergoes senescence<br />

(Jaworski et ale 19779.<br />

A1 though no specf fjc works have been<br />

published on the fntricats structure <strong>of</strong><br />

energy flow in Great <strong>Lake</strong>s coastal<br />

wet1 ands* Ti %ton 6% a1 , (1978) have<br />

general ized the important processes based<br />

on studjes in other wetlands'. <strong>The</strong><br />

conversion <strong>of</strong> solar energy -into bjomass by<br />

autotsophs is perhaps the most important<br />

process. Conversion into a form avail able<br />

to heterotrophic organisms serves as the<br />

foundation for several complex and dynamic<br />

food webs.<br />

<strong>The</strong> coastal wetland community<br />

possesses two basic complexes <strong>of</strong><br />

interrelationships: 1) invertebrates ( primarily<br />

insects)o fishes, bf rdsr and<br />

mammals which util ize 1 ivi ng p7 ant tissues<br />

and 2) organisms which utilize detrftus or<br />

dead plant tissues, Living plant tissue<br />

(7 .e., algae, 'leaves, and<br />

rhizomes) serve as food for phytophagous<br />

animals such as stem boring and leaf<br />

mining insects as well as certain aphids<br />

and beetles. Many specfes <strong>of</strong> waterfowl<br />

graze extensively on plant material r and<br />

muskrats are important pl ant consumers.<br />

<strong>The</strong> next higher trophic level in the ftrst<br />

complex consists <strong>of</strong> animals whlch prey<br />

upon the phytophagous organisms. Spiders,<br />

predatory beetles, dragonfllesr certsln<br />

fishes, frogs, birdsr and small<br />

insectivorous mammals are important<br />

organisms <strong>of</strong> this upper trophic level.<br />

<strong>The</strong> second complex consists <strong>of</strong> a vast<br />

number <strong>of</strong> insect larvae which rely on<br />

organic detritus as a dlrect energy source<br />

or by stripping microbial populations from<br />

the surface <strong>of</strong> organic particles,<br />

Gastropods and annelids are also important<br />

organ isms in the detritophagous complex.<br />

Whatever residual that is not util fzed by<br />

these animals is subjected to further<br />

decomposition by bacterf a1 and fungal<br />

populations, As with the phytophagous<br />

complexr there exists in the detritophagous<br />

complex a wide spectrum <strong>of</strong> animals<br />

which prey on the detritus-feeding<br />

organisms. Several species <strong>of</strong> insects,<br />

amphibians# waterfowlr and mammals compose<br />

this level, and many <strong>of</strong> these species are<br />

not selective in thelr prey* utilizing<br />

organ f sms from both compl exes.<br />

Teal (19629 found that a smaller<br />

percentage <strong>of</strong> the total energy represented<br />

f n the primary product 1 on <strong>of</strong> wetl ands


passes through the phytophagous complex<br />

than the detritophagous complex.<br />

Submergent vegetation tends to be<br />

inhabited and grazed more heavily than<br />

emergent forms (Krecker 1939) because the<br />

submergent aquatic macrophytes lack the<br />

more impenetrable structural tissues<br />

prevalent in the emergent type. Estimates<br />

<strong>of</strong> the proportlon <strong>of</strong> material processed<br />

through each complex (Ti lton et a]. 19781,<br />

favors the detritus web (80% t o 95%) over<br />

the grazing web (5% to 20% 1. Ti 1 ton et<br />

a1 . (1978) a1 so point out that the<br />

Importance <strong>of</strong> an organism to an ecosystem<br />

may exceed its role in energy flow,<br />

Muskrats utilize only a fraction <strong>of</strong> the<br />

available energy stored in the 1 lve plants<br />

they cut and harvest In the wetlands* but<br />

they may be <strong>of</strong> signiffcant value to the<br />

detritophagous complex. Similarlyo the<br />

teeming populations <strong>of</strong> phytophagous<br />

insects may consume on1 y a small fraction<br />

<strong>of</strong> plant tissuet but through this activity<br />

may reduce the growth <strong>of</strong> host plants and<br />

the primary production <strong>of</strong> the wetland<br />

ecosystem.<br />

<strong>The</strong> energy budget for cattai 1 (Iyehg<br />

sp. marshes in Minnesota has been<br />

investigated by Bray (1962) and may<br />

provide insight for <strong>Lake</strong> <strong>St</strong>. Clafr<br />

wetlands. During the growing season the<br />

distribution <strong>of</strong> the various components <strong>of</strong><br />

solar radiation energy was as follows:<br />

a1 bedo (reflection1 22.0<br />

evapotranspiration 38.4<br />

conduction-convectlon 38.5<br />

primary production 1.1<br />

Thf s apparently 1 ow ut 11 f zat i on <strong>of</strong> sol ar<br />

radiation by sp. is cons'lstent wlth<br />

other wetland studfes and supports the<br />

general view that most plant communltfes<br />

utilize only 1% to 2% <strong>of</strong> the total solar<br />

energy for primary production (Ttlton el<br />

al, 1978). sp. In Austrian<br />

wetlands ranges from 1-24 t o 2.0% for Ma<br />

through July (Sieghardt 1973) and<br />

sp. in Georgfan estuarfes utilizes 1,4% <strong>of</strong><br />

total solar radiation (Tea? 19621, <strong>The</strong><br />

uni formity <strong>of</strong> these results suggests that<br />

emergent wetlands in the take <strong>St</strong>. Claf r<br />

system utfl ize approximately 1.2% <strong>of</strong> solar<br />

radiation for prdmary production.<br />

Emergent wet1 and communities are<br />

among the most productive areas on earth.<br />

West1 ake (1963) estimated that freshwater<br />

emergent macrophytes have a net primary<br />

productivity <strong>of</strong> 3,000 to 8,500 g/rnZ/yr<br />

comparable in productivity to salt marshes<br />

and tropical rajn forests. Productivity<br />

<strong>of</strong> freshwater submerged macrophytes ranges<br />

from 400 to 2,000 g/m2/yrr less than ha1 f<br />

<strong>of</strong> thefr marine counterpart. Site<br />

specif ic primary productivfty studies <strong>of</strong><br />

coastal emergent wetlands in <strong>Lake</strong> <strong>St</strong>,<br />

<strong>Clair</strong> do not exist. Table 28 lists the<br />

maximum blomass and productivity <strong>of</strong><br />

several emergent species (common in <strong>Lake</strong><br />

<strong>St</strong>. Clalr) for other areas <strong>of</strong> the Great<br />

<strong>Lake</strong>s region. Extrapolation <strong>of</strong> these data<br />

to the cattail marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Cl ai r<br />

can provide an approximate and reasonable<br />

estimate <strong>of</strong> product lvi ty, Jv~ha marshes<br />

appear to be one <strong>of</strong> the most productive<br />

communities, yielding vaf ues (approxfmately<br />

2,500 g/mz/yr) near the lower end<br />

<strong>of</strong> the range expected for freshwater<br />

emergent macrophytes.<br />

<strong>Lake</strong> <strong>St</strong>, Cl ai r, particularly Anchor<br />

Bay and Mitchell Bay, are known for thetr<br />

we1 1 developed submerged aquatic<br />

macrophyte beds. Dawson (19751 found that<br />

considerable spatial varfation in standing<br />

crop exists in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> bays (Table<br />

29). He est'imated mean dry weights <strong>of</strong><br />

varlous submerged communities to range<br />

from 4 g/m2 in sha71ow areas characterized<br />

by sparse cover <strong>of</strong> stonewo<br />

to 316 g/m2 in areas domfna<br />

<strong>of</strong> water-milfoll (<br />

1. Schloesser ( 1982)<br />

monthly abundance <strong>of</strong> submersed macrophytes<br />

in the <strong>St</strong>. <strong>Clair</strong> River and <strong>Lake</strong> <strong>St</strong>. elair<br />

during the 1978 growing season (Table 30).<br />

He found that the amount <strong>of</strong> submersed<br />

vegetation was low in early sprfng, No<br />

vegetatfon was Sound at the mouth <strong>of</strong> the<br />

Clinton Rtver or the head <strong>of</strong> the Detrolt<br />

River (Belle Isle) in April or May, and<br />

the only vegetation collected at <strong>St</strong>ag<br />

Islands Algonac and Anchor Bay during<br />

these months consfsted <strong>of</strong> decabSng or<br />

dormant material f ram the previous growf ng


Table 28. Biomass ,arid productivity <strong>of</strong> emergent wetland plants <strong>of</strong> the<br />

Great <strong>Lake</strong>s region.<br />

Species<br />

'~ata source: Tilton et a1. (1978).<br />

Maximum Net<br />

Common name b i omass product 1 on<br />

broad-1 eaved cattai l<br />

hybrfd cattail<br />

giant bur reed<br />

common arrowhead<br />

manna grass<br />

reed grass<br />

wild rice<br />

sp i ke-rush<br />

lake sedge<br />

beaked sedge<br />

river bulrush<br />

great bul rush<br />

s<strong>of</strong>t rush<br />

Mean<br />

(g/m2) (g/m2/yr)<br />

Tabla 29. Estimates <strong>of</strong> standing crop biomass for submergeni vegeta-<br />

tion in Anchor Bay, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (September, October 1974).<br />

Dominant Weedbed Range in Percent <strong>of</strong><br />

taxa in community area (km2) dry welght (g/m2) submergents<br />

TOTAL 108.8 4-3 16 100.0<br />

"~ata source: Dawson ( 1975).


Table 30. Seasonal estimates <strong>of</strong> standing crop biomass for submersed<br />

rnacrophytes at several locations in the <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> sys~em (1978).<br />

Dominant<br />

Locatl on June July Rug Sept kt taxa<br />

<strong>St</strong>. <strong>Clair</strong> Rfver (<strong>St</strong>ag Is.) 72 267 103 38 15 SPP<br />

<strong>St</strong>. Clafr Delta (Algonacl 54 97 158 174 284<br />

Anchor Bay (Sand Is,) 85 113 135 133 146<br />

Clfnton River (mouth) 24 63 118 33 16 V a l l i w<br />

Detroit River (Belle Isle) 0 38 52 38 17 Yalli-<br />

iaowkaB<br />

a~ata source: Schloesser (1982).<br />

season. By mid-June dominant plants<br />

sprouted new growth at stem and leaf<br />

nodes, and by mid-July new growth which<br />

did not orfginate from overwintering<br />

dormant material was found at a17<br />

locations (Table 30). Higher maximum<br />

bfomass values were generally observed at<br />

locations where dormant overwinterfng<br />

vegetatjon was observed in early spring.<br />

Flgure 47 shows that the maximum biomass<br />

was reached In July in the <strong>St</strong>. <strong>Clair</strong> River<br />

(267g/m21; in August In the Gl inton River<br />

mouth (118 g/m2f and Detroit River head<br />

(53 g/m2); and in October in the <strong>St</strong>. <strong>Clair</strong><br />

Delta (284 g/m21 and Anchor Bay f 146<br />

g/m2). <strong>The</strong> submersed macrophytes became<br />

senescent and the biomass values decreased<br />

by late fall at all locations. <strong>The</strong><br />

results <strong>of</strong> thfs study indfcated that<br />

little or no submersed macrophyte growth<br />

takes place before June or after October<br />

in the rfver or lake. <strong>The</strong> dominant taxa<br />

found at each location are given in Table<br />

31,<br />

4.3 CQMMUNiTY PROCESSES<br />

Given the transittonal or ecotone<br />

character <strong>of</strong> wetlandsr the <strong>Lake</strong> <strong>St</strong>. Clafr<br />

wetlands are even more dynamic due to the<br />

pulsing nature <strong>of</strong> the lake level changes<br />

and the connectivlty due to the numerous<br />

distributary channels and embaymentst<br />

particularly wlthin the <strong>St</strong>. <strong>Clair</strong> Delta.<br />

By outlfning the community processes in<br />

these dynamlc and interrelated envi ron-<br />

ments, the high prlmary and secondary<br />

producti vi ty can be more full y understood.<br />

<strong>The</strong> <strong>Lake</strong> <strong>St</strong>. Clalr wetlands are<br />

actually composed <strong>of</strong> a variety <strong>of</strong> habitats<br />

f ncl ud 1 ng open ponds, cattai?/reed<br />

marshes, earthen dikes, barrler beaches,<br />

delta flats, and wooded swamps.<br />

Col7ectfvely these habitats are known as<br />

the coastal marsh community, Each habitat<br />

attracts its own species <strong>of</strong> plants* blrdsr<br />

mamnalsp reptiles, amphibianst and fn some<br />

cases, fish. <strong>The</strong> result Is more varfety<br />

in plant and animal life than in any other<br />

area <strong>of</strong> equal size fn the intarior <strong>of</strong> the<br />

bordering states and province. <strong>The</strong><br />

overall conditions <strong>of</strong> the uncontrolled<br />

coastal marshes are stf 17 very primitive,<br />

some having been visited by no more than a<br />

handful <strong>of</strong> people in the last several<br />

decades. W i thln those marshes where<br />

natural processss are allowed to take<br />

place, zonatfon and succession in response<br />

to changing envf ronmsntal condftions are


ANCHOR BAY<br />

MICHIGAN<br />

Sand Island<br />

\ LAKE<br />

Figure 47. Seasonal growth (dry weight) <strong>of</strong> dominant submersed macrophytes in the <strong>St</strong>.<br />

<strong>Clair</strong> River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-Detroi t River ecosystem (Schloesser 1982).<br />

Table 35 Dominant submersed aquatic macrophyte taxa <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

sys tern.<br />

Taxa<br />

-<br />

<strong>St</strong>. <strong>Clair</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Lake</strong> <strong>St</strong>. Clinton Detroit<br />

R'iver Delta Cl ai r River River<br />

(<strong>St</strong>ag Island) (Algonac) (Anchor Bay) (mouth) (Belle Isle)<br />

%ata sources: Dawson ( 19751 r Schloesser (1982).


among the important commun l t y processes.<br />

Water level fluctuationr and the resultant<br />

plant and anfmal responser is <strong>of</strong>ten the<br />

most sfgnificant drivjng force.<br />

Because the water level <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> is not stable, but has oscillated<br />

over 2 m during the period 1900 to 1977<br />

(Jaworski et a1. 19793, the wetland<br />

vegetation a1 ong the 1 ake margi ns exhibits<br />

pulse stability. <strong>The</strong> concept <strong>of</strong> pulse<br />

stability imp1 ies that it is these water<br />

level fluctuations which result tn the<br />

maintenance <strong>of</strong> the wetl and communities.<br />

Thus* even though wetland plant<br />

communfties retrogress during hlgh lake<br />

level periods and shift lakeward during<br />

low water level conditionsr these<br />

successional changes prevent the wet1 ands<br />

from being dominated by either aquatic or<br />

up1 and communities. Moreovers coastal<br />

wetlands do not become senescent or<br />

exhibit conversion t o terrestrial<br />

environments as compared to inlands<br />

pal ustrine wetl ands.<br />

It is important to distinguish<br />

between annual (or seasonal 1 water level<br />

fluctuations and longer term oscillations,<br />

With regard to annual water level changes<br />

fn <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, the average magnitude<br />

is approximate1 y 0.5 m, with highest<br />

levels occurrtng in June-July and lowest<br />

levels during the winter months. This<br />

seasonal water level pattern is opposite<br />

that in shallowr freshwater swamps which<br />

may dry up during the middle <strong>of</strong> the<br />

growing season. Cattails appear to be<br />

favored by low water levels in winter<br />

because oxygen diffuses through exposed<br />

stalks down into the roots and rhizomes<br />

(Jaworski et al. 1979).<br />

However, it is the longer term lake<br />

level osclllatlons which have the greatest<br />

effect on vegetation succession. In the<br />

Great <strong>Lake</strong>s the hydroperiod for these<br />

longer term oscillations is 8 to 20 years<br />

(U.S. Dept. <strong>of</strong> Commerce 1976). Near<br />

record low water levels - at about 174 m<br />

above datum (IGLD) - occurred in 1964-<br />

1965, whereas record highwater levels - at approximately 176 m above datum (IGLD) -<br />

took place during the 1972 to 1974<br />

<strong>The</strong> latepal shifts <strong>of</strong> the plant<br />

communities in response to water level<br />

fluctuations are illustrated by Figures 32<br />

and 33 <strong>of</strong> Section 3-2. DickYlnson Island<br />

was utiljzed for this dtscussion because<br />

it has an intact envi ronmental gradient<br />

over which vegetation shifts can occur.<br />

El sewhere on Warsens Island as we1 1 as on<br />

the Canadian side <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta9<br />

the cattail marsh ends abruptly against<br />

dikes or canals, and much <strong>of</strong> the sedge end<br />

other drier habftat communities have been<br />

devel oped.<br />

<strong>The</strong>se successional changes can be<br />

documented by a series <strong>of</strong> transects taken<br />

over time across the same survey 1 ine. As<br />

indicated by Figure 48, water level<br />

changes in the coastal wetlands <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. Clai r caused dramatic vegetation<br />

changes. Beginning in July 1964s one can<br />

observe extensive comunjties <strong>of</strong> cattail r<br />

sedge, and dogwood meadow, Howevers<br />

during the high water period <strong>of</strong> the 1970~~<br />

widespread ddeback occurred as far inland<br />

as the shrub/swamp frfnge which borders on<br />

the oak-ash hardwoods. During such<br />

condltlonsr much <strong>of</strong> the cattall marsh<br />

consisted <strong>of</strong> open water as well as<br />

communities <strong>of</strong> bulrushes and submersed<br />

aquatics. Notice that by the summer <strong>of</strong><br />

1977 r considerable reestablishment began<br />

to take place as water levels dropped<br />

somew hat.<br />

A plant commundty displacement model<br />

can be used t o predict community<br />

succession associated with water level<br />

fluctuati ons. As f 11 ustrated in Figure<br />

49s each wetland plant community has been<br />

placed at its proper elevation in relatlon<br />

to the long-term mean water level. <strong>The</strong><br />

length <strong>of</strong> the l ine under the plant<br />

community's name lndjcates the amount sf<br />

vertical shifting which that community<br />

exhibits. To use PhSs model* note that<br />

the communities shOft lakeward (to the<br />

left) durlng low water levels and displace<br />

communities along thef r lakeward margfn.<br />

To simulate S U C C ~ S S during ~ ~ ~ rfsing lake<br />

levels, shift the comunfttes to the right<br />

a1 ong the1 r respective dlspl acement 1 f nes,<br />

As one moves upward along the<br />

envi ronmenisl gradtent, the extent <strong>of</strong><br />

cornunity displacement is less and core<br />

areas persfst durfng both low and hlghinterim.<br />

Because <strong>of</strong> thess CYC~IC<br />

water water pert ads, f herefore* the shrub/swamp<br />

level oscillations, succession is a1 ways fringe zone shifts least and Os displaced<br />

taking place in the plant communities. less by the adjacent $1 ant comunf iy.


WEST<br />

Feet Drowned<br />

Tree-Shrub<br />

678<br />

670<br />

<strong>Lake</strong> Dead<br />

Cottonwood<br />

Cottonwood<br />

June 1975 EAST<br />

Meters<br />

A<br />

July - Qct. 1977<br />

July 1964<br />

Mixed<br />

- - . - - - - - - - - - - - - - - - -<br />

T-"I ---1-7- - T-*---T--- 7-----r--- --T- -7- -- --r<br />

+ 1x , _1T_11_1_T1_1r11_7-~T_I_ - -<br />

0 500 1000 1500 2000 2500 3000 3500 4000 4500 5000 5500 6000 6500 Ft.<br />

r-- --7""- ,<br />

0 200 400 600 800 1000 1200 1400 1800 1800 Meters<br />

Figure 48. Succes~iorial changes on Dickinson Island in response to water level fluctu-<br />

ations,<br />

flll UCTCHS<br />

hi." 1<br />

Shri~b/Swa~lip I<br />

I - - -<br />

Meadow -- ,,<br />

Sedge Mdrstl i -<br />

"" . . - . - - - - .- .- Llf6.0<br />

. . - . - - - - .- .- Llf6.0<br />

Mcan iakc Level 1<br />

Figtars 43. Wetland plant community dis-<br />

placement rnode'l for Dickinson island<br />

showing predicted response to water level<br />

changes,<br />

176<br />

175<br />

174<br />

As shown in Table 32, the wetland<br />

pl ant communities not on1 y change location<br />

but alter their areal extent in response<br />

t o these long-term water level<br />

oscillations. <strong>The</strong> dominance <strong>of</strong> the<br />

emergent cattail and sedge marsh during<br />

low-water levels is clear. In contrast,<br />

during high water periodsr the wetlands<br />

exhibit extensive floating leaved and<br />

submersed communities along with the<br />

cattails. <strong>The</strong> cattail marsh actual 1 y<br />

appears more widespread during high water<br />

conditions than it is because the dead<br />

cattail stalks <strong>of</strong>ten remain in place for<br />

several years. Notice, toor that the area<br />

<strong>of</strong> the shrub/swamp changes very 1 ittle as<br />

the water levels change. This situation<br />

prevails because the shrub/swamp community


Table 32. Vegetation cianges on Dickinson Island in response to<br />

lake level fluctuations.<br />

Percent <strong>of</strong> total areab<br />

Vegetation class Low water Mean level Hiah water<br />

1964 1975<br />

Open water<br />

Floating leaved/<br />

Submersed<br />

Emergents (cattails)<br />

Sedge marsh<br />

Dogwood meadow<br />

Shrub/Swamp<br />

Developed areas<br />

a Data source: Jaworski et a1 . (1979).<br />

b~ote: Total area lr130 hectares (2,800 acres).<br />

is slightly higher in elevation than the<br />

high water mark and it is difficult to<br />

separate dieback shrub from live brushy<br />

vegetation. Also? cattail is an ag-<br />

gressive invader compared to sedges and<br />

dogwood.<br />

<strong>The</strong> zonation exhibited by the wet1 and<br />

plant communities has previously been<br />

illustrated by Figures 26 and 33 and<br />

described in Section 3.2. <strong>The</strong>se plant<br />

zones, in order from upland to aquatic<br />

sites, are: shrub/swamp? dogwood meadow?<br />

sedge marsh? cattail marshr and open water<br />

floati ng/submersed commun f t i es . In many<br />

wetlands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> urban<br />

development and other land uses have<br />

progressed from the adjacent upland areas<br />

lakeward to the emergent marsh. On1 y on<br />

Dickinson Island, and to some extent in<br />

Bouvier Bay (i.e., <strong>St</strong>. John's Marsh)? does<br />

a complete environmental gradient? or full<br />

zonation <strong>of</strong> wetlands* exist today.<br />

Species diversity <strong>of</strong> invertebrates<br />

tends to be hlgher fn <strong>St</strong>. <strong>Clair</strong> Delta<br />

wetland ecosystems than f n nearshore<br />

benthic and pr<strong>of</strong>undal benthic zones <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Invertebrate productivtty<br />

also tends to be higher in wetlands<br />

compared to open water areas and ft<br />

appears that invertebrate productivity<br />

tends to. be higher in emergent wetlands<br />

compared to submergent wet1 ands (Ti 1 ton et<br />

al, 1978). Invertebrate populations in<br />

coastal wetlands are a food source for<br />

higher trophic levels in the food webt<br />

comprising a major fraction <strong>of</strong> the diet <strong>of</strong><br />

numerous species <strong>of</strong> fish* reptiles?<br />

amphibians, blrdsr and mammals.<br />

Several species <strong>of</strong> forage fish<br />

(spottail? black nose^ and emerald shiners9<br />

feed or spawn In wetland ecoysystems,<br />

<strong>The</strong>se forage fish are consumed by<br />

piscivorous fish as well as birds and<br />

mammals. toss <strong>of</strong> wetlands whjch produce<br />

forage fish can cause a reduction in the<br />

numbers <strong>of</strong> piscivorous fish as we11 as<br />

birds such as the kingfisher and common<br />

mergansers (Ti 1 ton et a1 . 1978).<br />

Zonation has also been demonstrated<br />

for larval fish populations in western<br />

<strong>Lake</strong> Erie (Cooper et al. 1981ar 1981b.<br />

1984; Mizera et ale 3.9811. A series <strong>of</strong><br />

studies in the estuaries and open waters<br />

<strong>of</strong> western <strong>Lake</strong> Erie has shown that the<br />

estuaries <strong>of</strong> the Maumee and Sandusky<br />

Rivers contained the hfghest densities <strong>of</strong>


larval fish when compared with other<br />

nearshore and <strong>of</strong>fshore areas. Glzzard<br />

shad, white bass, and freshwater drum<br />

dominated the estuarfne populatfons. <strong>The</strong><br />

highest densjty <strong>of</strong> yellow perch was found<br />

in nearshore areas assocfated wfth sandy<br />

bottoms, particularly north <strong>of</strong> Woodtick<br />

Peninsuia and in the vicinity <strong>of</strong> Locust<br />

P<strong>of</strong>nt. <strong>The</strong> followtng depth/denslty re-<br />

lationship was observed for thls species:<br />

<strong>The</strong> same general relationship was found<br />

for most species, fndfcatlng a greater<br />

preference for spawning and nursery<br />

grounds fn the coastal areas. A sfmilar<br />

pattern is suspected for <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Surveys conducted by Hatcher and Nester<br />

E1983)~ although not in the lake proper,<br />

show high densities Jn the <strong>St</strong>. <strong>Clair</strong> Delta<br />

and at the head <strong>of</strong> the Detroit River.<br />

Johnson (1984) belleved that the most<br />

important factor influencing production <strong>of</strong><br />

flsh larvae in #%stern <strong>Lake</strong> Erie marshes<br />

is predatfon. Although no information Is<br />

avaflable on predator-prey tnteractlons<br />

involving larval fJsh In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

wetlands* related research has Indicated<br />

habftat structure can be Smportant in<br />

medtatlng the outcome <strong>of</strong> aquatic predator-<br />

prey interactions. Glass (1971) found<br />

argemouth bass ( a4.L<br />

predat f on succes s the<br />

Ity <strong>of</strong> habitat structure increased.<br />

Crowder and Cooper (19791 concluded that<br />

areas <strong>of</strong> hSgh structural complexity should<br />

create an effective refuge for prey and<br />

onhance their survfval, However,<br />

(19791 found prey-sized bluegill (<br />

1 were strongly attracted to<br />

shade produced by I m diameter floats in<br />

wh~ch no submerged structure was present,<br />

He ~oncluded prey fish in shade can detect<br />

a predator (in sun) long before the<br />

predator can detect the prey.<br />

<strong>The</strong>refore prey specles and prey-<br />

sized fSsh may act1 vely escape predators<br />

by dfsappearfng Snto areas <strong>of</strong> high<br />

struct~ral complexity or by seeklng shade<br />

so as to gajn the visual advantage.<br />

Consequentlys fish larvae in wetlands may<br />

use floating habitat types to gain the<br />

vjsual advantage while other larvae may<br />

use submergent or emergent habitat types<br />

to hide from potential predators, Open<br />

water areas in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> coastal<br />

marshes may be expected to have low larval<br />

use during the day because <strong>of</strong> the<br />

increased success <strong>of</strong> visual predators in<br />

this habitat type, but these areass<br />

particular1 y the uncontrolled wetlands,<br />

may be Important to fish larvae at night,<br />

<strong>The</strong> coastal marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

attract large numbers <strong>of</strong> migratory<br />

waterfowl. Located at a crossing point on<br />

two major flyways, these marshes attract<br />

ducks from eastern Canada headlng for<br />

wintering grounds on the Mississippi River<br />

bottoms and ducks from the prairie<br />

provinces <strong>of</strong> Canada which winter along the<br />

Atlantfc coast. Beds <strong>of</strong> wfld celery are<br />

partjcul arl y attractive to great numbers<br />

<strong>of</strong> canvasbacks (m 1, red-<br />

heads ( ), and scaups<br />

(w<br />

spp. migrating southeastward to<br />

thef r wintering grounds on Chesapeake Bay<br />

(Andrews 1952).<br />

Lincoln (1935) introduced the concept<br />

that all populatlons <strong>of</strong> migratory birds<br />

adhere to their respectfve flyways as they<br />

make thef r semt annual fl lghts between<br />

breeding grounds and wintering grounds.<br />

Four distinctive flyways have been<br />

Identified for North America, two <strong>of</strong> whqch<br />

cross western <strong>Lake</strong> Erie and <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>: the Atlantic Flyway and the<br />

Mississippi Flyway, <strong>The</strong>se routes are used<br />

by all <strong>of</strong> the migratory waterfowl and<br />

other waterbirds which frequent the<br />

coastal wetlands. Each flyway has its own<br />

individual population <strong>of</strong> birdsp even for<br />

those specfes which have a broad<br />

contJnenta1 distribution. <strong>The</strong> breeding<br />

grounds <strong>of</strong> two or more flyways may, and<br />

<strong>of</strong>ten dor overlap broadly so that during<br />

the nesting season extensive areas may be<br />

occupfed by bjrds <strong>of</strong> the same species, but<br />

which belong to dffferent flyways.<br />

In the fall, the Atlantlc Flyway<br />

recsivas accretion <strong>of</strong> waterfowl from<br />

several fnterlor migration paths starting


at the breeding grounds on the Arctic<br />

tundra. Canada goose )V<br />

and df vfng ducks8 incl SI<br />

redheads and scaup come from their<br />

breeding grounds on the great northern<br />

plains <strong>of</strong> central Canaaa, follow the<br />

general southeasterly trend <strong>of</strong> the Great<br />

<strong>Lake</strong>s? rest in the <strong>St</strong>. <strong>Clair</strong> Delta, cross<br />

<strong>Lake</strong> Erje in the islands region, and<br />

continue over the mountains <strong>of</strong><br />

Pennsylvania to w i nter along the Atlantic<br />

coast in Chesapeake and Delaware bays.<br />

Concurrent1 y, dabbl ing ducks such as<br />

1, black ducks<br />

1, and blue-winged teals<br />

1 that have gathered in<br />

staging areas <strong>of</strong> southern Ontario<br />

(fncludfng the <strong>St</strong>. <strong>Clair</strong> Delta) also leave<br />

these feeding grounds, cross western <strong>Lake</strong><br />

Erie and proceed southwest over a course<br />

that leads down the Ohio and Mississfppi<br />

valleys (Mississippi Flyway), However,<br />

part <strong>of</strong> this duck population, upon<br />

reaching the <strong>St</strong>. <strong>Clair</strong> Flats, swings<br />

abruptly to the southeast, crosses the<br />

Appalachian mountains and winters along<br />

the Atlantic coast (Lincoln 1950).<br />

<strong>The</strong> Mississippi Flyway is easily the<br />

longest migration route <strong>of</strong> any in the<br />

Western Hemisphere. Its northern terminus<br />

is the Arctic coast <strong>of</strong> Alaska, whlle its<br />

southern end 1 ies In the Patagonia region<br />

<strong>of</strong> Argentina. Although the main path <strong>of</strong><br />

the flyway lies to the west <strong>of</strong> the Great<br />

<strong>Lake</strong>s major branches follow the southern<br />

trend <strong>of</strong> <strong>Lake</strong> Michigan and the<br />

southwestern trend <strong>of</strong> <strong>Lake</strong> Erle and the<br />

Maumee Rf ver valley. Some <strong>of</strong> the black<br />

ducks ma1 1 ardsr and teals that cross the<br />

Great <strong>Lake</strong>s in the viclnfty <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> do not turn abruptly to the<br />

southeast, but continue on to the<br />

southwest as members <strong>of</strong> the Mississippi<br />

Flyway bound for the Gulf <strong>of</strong> Mexico coast<br />

rather than the Atlantic seaboard. Fall<br />

migration is at Its peak in September and<br />

October, but the main shorebird passage is<br />

underway in August.<br />

Sprlng migration begins in late<br />

February with the appearance <strong>of</strong> rlng-<br />

billed gulls ( 1 In <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. Ma 11 bring<br />

h waterfowl mov-ntr ducks and loons<br />

( sp.1 appearing 5n open leads as<br />

blackbf rds (<br />

large numbers starting in late March. <strong>The</strong><br />

huge Canada goose movement normally takes<br />

place In early April. With April and May<br />

comes the major push <strong>of</strong> spring migration,<br />

especizlly the landbirds. Migrants are<br />

less selective than breeding birds In<br />

their choice <strong>of</strong> habitat, nevertheless,<br />

waterb i rds prefer shore1 lnes wf th pockets<br />

<strong>of</strong> vegetation. Coastal marshes and stream<br />

mouths comnly attract migrating dabbl f ng<br />

ducks. <strong>The</strong> open water shorelines<br />

concentrate the diving duck migrants and<br />

other waterbirds including loons, grebes,<br />

cormorants (<br />

For over three decades after Lincoln<br />

defined and mapped his four waterfowl<br />

flyways <strong>of</strong> North America, little new work<br />

was pub1 ished on the subject. <strong>The</strong>n,<br />

Bell rose (1968) pub1 ished his outstanding<br />

treatise on waterfowl migratlon corrldors<br />

east <strong>of</strong> the Rocky Mountains. In addition<br />

to these data and ground observatlons~<br />

Be1 1 rose used such advanced techn I ques as<br />

visual sightings from aircraft and radar<br />

su rvei l 1 ance. One impetus for undertaking<br />

this revision was the Increasing hazard to<br />

aircraft from migrating birds.<br />

Be1 l rose found that Llncol n s f 1 yway<br />

concept, although still valid on a grand<br />

scale, tended to oversfmpllfy the<br />

migratjon process. He observed that<br />

flyways were In reality e complex <strong>of</strong><br />

corridors (Figures 50-52). Each corridor,<br />

in turn? Is a web <strong>of</strong> routes as opposed to<br />

a single, narrow band rigidly followed by<br />

the waterfowl. <strong>The</strong> migration corridors<br />

represent passageways sac h connecting a<br />

serles <strong>of</strong> waterfowl habitats. Migration<br />

corridors differ from flyways in being<br />

smaller and more precisely deflnsd as to<br />

specles. Bell rose cons lders the f l yways<br />

proposed by Lincoln to be prfmarily<br />

geographical and secondarily biological,<br />

while his corridors are primarily<br />

biological and secondarif y geographical.<br />

As waterfowl migrate between breeding<br />

grounds and wfntering areas, they stop to<br />

rest and feed in wet1 ands, <strong>The</strong>se wetl ands<br />

2re referred to as nconcentratlon areas."<br />

<strong>The</strong> coastal wetl ands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Claf rr<br />

the lower Detroit Riverr and western <strong>Lake</strong>


DABBLING DUCKS<br />

ESTIMATED NUMBER<br />

USING A CORRIDOR<br />

PENNSYLVANIA<br />

Figure 50. Fa1 1 migration corridors for dabbling ducks (Tribe Anatini ) across<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Be1 l rose 1968).<br />

Erie provide some <strong>of</strong> the finest<br />

concentration areas <strong>of</strong> this type along the<br />

flyways. <strong>The</strong>se areas are characterized by<br />

an abundance <strong>of</strong> waterfowl foods, as well<br />

as by low wave energy and low human<br />

disturbance. Canvasbacks, redheads*<br />

animal foods, Canada geese and mallards<br />

also feed heavf ly on waste grains fn<br />

agricultural fields. Food availability<br />

may be more important than food<br />

preference, especially during the sprf ng<br />

migration when food supplies are less<br />

abundant. Food availabil ity f n wetlands<br />

is reduced by extreme high and low water<br />

levels, heavy siltationr turbidity, heavy<br />

hunting pressure* and other disturbances.<br />

Jones (1982) examined the food<br />

preference <strong>of</strong> wintering go1 deneye, greater<br />

scaup, and lesser scaup at the mouth <strong>of</strong><br />

the Detroit River. He determined that 30<br />

food taxa were utilized by the waterfowl<br />

including 10 plant and 20 animal<br />

references. Wild csler<br />

1, pondweeds (<br />

and other plant speci es made up over 60%<br />

<strong>of</strong> the food budget <strong>of</strong> most <strong>of</strong> the sampled


- -<br />

DIVING DUCKS<br />

ESTIMATED NUMBER<br />

USING A CORRIDOR<br />

Figure 51. Fa1 1 migration corridors for diving ducks (Tribe Aythyini) across<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Be1 lrose 1968).<br />

waterfowl. Miller (1943) and Bell rose<br />

(1976) have found that other diving ducks*<br />

particularly canvasbacks and redheads*<br />

feed on submersed aquatics such as those<br />

noted above as well as waterweed Elc&&<br />

s2amku&)<br />

In western <strong>Lake</strong> Erie marshes,<br />

Bednar f k (19751 found that preferred<br />

natural foods <strong>of</strong> diving ducks* such as<br />

wild celery* appeared to be more adversely<br />

affected by turbidity and siltation than<br />

foods <strong>of</strong> dabbling ducks and geese. In<br />

general, Great <strong>Lake</strong>s conservation agencies<br />

do not encourage the creation <strong>of</strong> resident<br />

wintering flocks# particularly in the <strong>Lake</strong><br />

<strong>St</strong>. Clafr region* because <strong>of</strong> the problem<br />

<strong>of</strong> waterfowl starvation during severe<br />

winters. Waterfowl that reach the spring<br />

breeding grounds in good condition tend to<br />

exhibit greater nesting success than those<br />

which are undernourished (Bell rose 1976).<br />

Q&l&lng duck <strong>The</strong>se ducks are so<br />

ca1 led because they normal 1 y do not dive<br />

below the water for food, but merely<br />

dabble on the bottom In shallow water.<br />

Annually about 17r500r000 ma1 1 ards and<br />

pintail s migrate down fl ight corrldors<br />

from Canada to the United <strong>St</strong>ates east <strong>of</strong><br />

the Rocky Mountains (Bellrose 19681, <strong>The</strong><br />

l argest portion, about 12r275r000s enter<br />

the geographfcal conffnes <strong>of</strong> the<br />

Mississippi Flyway from the northern Great<br />

Plains, About 20% <strong>of</strong> these birds continue<br />

across the Mississfppf Flyway and move<br />

down the Atlantic Flyway. In addition,<br />

anotRer 650,000 black ducks move south<br />

from Ontario and Quebec.<br />

_


CANADA GEESE<br />

ESTIMATED NUMBER<br />

USING A CORRIDOR<br />

75,100 - 150,000<br />

5,100 - 25,000<br />

Figure 52. Fall migration corridors for Canada geese (Branta canadensis) across<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Bellrose 1968).<br />

Several corridors carrying dabbllng As the name suggests,<br />

ducks cross the Great <strong>Lake</strong>s region (Flgure these ducks normally dive below the water<br />

50). An estimated 65,000 mallards, 35,000 for food. About 40200r000 diving ducks<br />

wigeons, and 25r000 pintalls move eastward annually migrate south into the United<br />

along the Chesapeake Bay Corridor. <strong>The</strong> <strong>St</strong>ates east <strong>of</strong> the Rockies (Bell rose<br />

corrldor starts In the upper Mfss'lssfppi 113681, Slightly over 609% <strong>of</strong> these are<br />

Rf ver Val ley and Progresses eastward scaups. Redheads are second fn abundance<br />

through Wlsconsin, Mlchiganr and Ohlo. It at ZO%5 while canvasbacks and ring-necked<br />

encompasses the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr ducks each form about 7% <strong>of</strong> the<br />

and <strong>Lake</strong> Erie f ram Algonacr Mlchfganr to populaticn. As with the dabbling ducksr<br />

Sandusky Bay. From these marshes it is a nume;.ous diving duck mfgration corridors<br />

645 km* non-stoP flight to Chesapeake Bay cross the Great <strong>Lake</strong>s region (Figure 51).<br />

where most <strong>of</strong> these ducks winter, <strong>The</strong><br />

Bl ack Duck Corridor extends southwest~ard <strong>The</strong> Southern Michigan Corrfdor takes<br />

from eastern Ontario5 across <strong>Lake</strong> <strong>St</strong>. the maln flow <strong>of</strong> diving duck passage from<br />

CIair to the confluence <strong>of</strong> the Wabash and eastern WJSCO~SSR~ across southern Mich-<br />

Qhfe, riversr and on south to Arkansas. lgan to Saginaw Bay and the <strong>Lake</strong> <strong>St</strong>.<br />

Approximately 35#QOO black ducks use this Cl a i r-Detroi t River-<strong>Lake</strong> Erte wet1 ands<br />

path, areas. Diving ducks congregate on SagJnaw<br />

100


Bay to the extent that peak numbers<br />

include 22,000 lesser scaupsr 22r000<br />

redheads, and 7,000 canvasbacks.<br />

Approximately 160 km to the south, peak<br />

populations <strong>of</strong> 380,000 lesser scaupsr<br />

260r 000 canvasbacks, and 42r000 redheads<br />

have been observed fram <strong>Lake</strong> <strong>St</strong>. Cl ai r to<br />

western <strong>Lake</strong> Erie. Although as many as<br />

15,000 diving ducks may winter on the<br />

Detroit River, at least 700,000 fly on<br />

Prom <strong>Lake</strong> <strong>St</strong>. Clalr to wintering grounds<br />

in the Atlantfc Flyway. This route is<br />

known as the Chesapeake Bay Corridor and<br />

is a similar route to the one taken by<br />

mallards and pintails.<br />

More than any other species<br />

<strong>of</strong> waterfowl, Canada geese have radical1 y<br />

altered their migration routes In the past<br />

few decades. Bell rose (1968) attrf buted<br />

this great change tn their migration<br />

habits to their rapid adoption <strong>of</strong> newly<br />

created waterfowl refuges, <strong>The</strong>y are still<br />

in the process <strong>of</strong> evolvtng new migration<br />

corridors, Currently, about lr300s000<br />

Canada geese leave Canada in the fall for<br />

wintering grounds in the United <strong>St</strong>ates.<br />

<strong>The</strong> largest number, 600~000 use the<br />

Atlantic Flyway, while another 475,000<br />

take the Mississippi Flyway. Most <strong>of</strong> the<br />

Atlantic Flyway crossings <strong>of</strong> the Great<br />

<strong>Lake</strong>s take place over <strong>Lake</strong> Ontarfor but<br />

one corridor uses the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> (Figure 52). <strong>The</strong> main migration<br />

corridor for Canada geese i n the<br />

Mississippi Flyway extends down the west<br />

shore <strong>of</strong> <strong>Lake</strong> Michigan,<br />

Mississippi River valley.<br />

then down the<br />

Snow geese (m 1 use<br />

the Misstssippi Flyway. Each October<br />

about 450r000 birds 1 eave Canada for<br />

wfntering grounds on the coastal marshes<br />

<strong>of</strong> Louisiana. <strong>The</strong> main corridors follow<br />

the east and west shores <strong>of</strong> <strong>Lake</strong> Michiganr<br />

converging fn the Mississippi River valley<br />

north <strong>of</strong> Loulsi ana. <strong>The</strong> easternmost<br />

flfght corridor, used by about 15,000<br />

geese, runs from the south end <strong>of</strong> James<br />

Bay to the marshes <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />

western <strong>Lake</strong> Erie, then turns south-<br />

westward across Indiana, A somewhat<br />

Jarger number <strong>of</strong> birds uses a corridor<br />

that extends from James Bay through<br />

Saginaw Bayr and then merges with the<br />

flight path from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>,<br />

4.4 WETaNDS AS HABITAT TO FISH AND<br />

WlLDLlFE<br />

Because the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands<br />

are composed <strong>of</strong> a variety <strong>of</strong> habitats<br />

including open ponds, cattail/reed<br />

marshesr earthen dikes, barrier beachesr<br />

delta flats, and wooded swamps, many<br />

species <strong>of</strong> plants and animals are<br />

attracted to the coastal marsh community.<br />

Of particular importance to fish and<br />

nil dl i fe, both resident and transient<br />

species# are the following considerat~ons:<br />

1) food chain production and energy flow,<br />

2) ffsh productionr spawningr and nursery,<br />

31 waterfowl migration, wintering and<br />

nesting, 4) mammal forager and 5)<br />

invertebrate habftat. Dole (1975) er-<br />

timated the density <strong>of</strong> wildlife in the<br />

vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Table 33).<br />

Most animals showed a stable trendr but a<br />

few important species are decreasing in<br />

abundance.<br />

Although there are scattered patches<br />

<strong>of</strong> wetlands around the perimeter <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>, the wetlands in the <strong>St</strong>. <strong>Clair</strong><br />

Delta provide the most valuable habitat<br />

for fish and wlldllfe, includtng water-<br />

fowl. Mot only ara these deltaic wetlands<br />

productlver they are diverse and exhibit<br />

high connectivity to the <strong>St</strong>. <strong>Clair</strong> River<br />

channels as well as to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

With regard to fish spawning in these<br />

wetlands, important species include the<br />

following: lake sturgeon* northern piker<br />

rnuskell unget carp, golden shiner, emerald<br />

shiner, common shinerr spottai l shlner,<br />

spotffn shiner, channel catf ishr bull-<br />

heads# rock basst pumpkf nseed, bl ueglll ,<br />

sunfish, and walleye (Figure 39). It has<br />

been suggested that ice fishing$ as f n<br />

Little Muscamoot Bay <strong>of</strong> Harsens Islandp is<br />

particul arl y good because wet1 ands are an<br />

important source <strong>of</strong> fish foods in winter<br />

(W. C. Bryant, MDNRI Ffsheries Division,<br />

pers. corn.). Howevers except for gtzzard<br />

shad, most f I sh mlgrate to deeper waters<br />

in fall and return to the shallows<br />

nearshore envi ronrnents fs1 lowing ice<br />

breakup (Werner and Manny 19791, Because<br />

wetlands warm up more quickly than <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong> in the sprfnge these coastal<br />

environments may be a crftical source <strong>of</strong><br />

food for ftsh attracted to the sharelines,


Gable 33, <strong>St</strong>atus <strong>of</strong> wildlife in ttge vicinity <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> and connecting waterways (1970).<br />

Wild1 lfe group<br />

and species Density Trend<br />

Big game<br />

White-tailed deer<br />

Waterfowl<br />

Ducks<br />

Geese<br />

Small game<br />

Cottontail rabbit<br />

Ring-necked pheasant<br />

Ruffed grouse<br />

Gray squ i rrel<br />

Fox squirrel<br />

Woodcock<br />

Mourning dove<br />

Bobwhite qua11<br />

Furbearers<br />

Muskrat<br />

M'lnk<br />

Beaver<br />

Weasel<br />

Raccoon<br />

Skunk<br />

Opossum<br />

Badger<br />

Non-game<br />

Woodchuck<br />

Red fox<br />

Gray fox<br />

Crow<br />

Rod squirrel<br />

Coyote<br />

Raptars<br />

Rare and endangered<br />

Bald eagle<br />

American osprey<br />

Unusual or unique<br />

Sandhi1 1 cranes<br />

Golden sag1 e<br />

a Data source: Dole (1975).<br />

1 ow i ncreasi ng<br />

high<br />

rned f um<br />

rned i urn<br />

high<br />

1 ow<br />

1 ow<br />

medi um<br />

1 ow<br />

high<br />

1 ow<br />

high<br />

rned i um<br />

1 ow<br />

rned i urn<br />

rned i um<br />

high<br />

high<br />

1 ow<br />

rned i urn<br />

med l um<br />

rned i um<br />

high<br />

1 ow<br />

1 ow<br />

rned f urn<br />

rned i urn<br />

rare<br />

stabl e<br />

increasing<br />

stable<br />

stab1 e<br />

stabl e<br />

decreasing<br />

stable<br />

stabl e<br />

stable<br />

stab1 e<br />

stabl e<br />

stabl e<br />

decreasl ng<br />

stable<br />

'lncreas'lng<br />

increasing<br />

stable<br />

stab1 e<br />

stable<br />

stabl e<br />

stabl e<br />

stable<br />

stabl e<br />

stabl e<br />

stabl e<br />

decreasing<br />

decreasing<br />

stabl e<br />

translent


With regard to wildlife habitats the<br />

<strong>St</strong>. <strong>Clair</strong> Delta wetlands are also<br />

exceptional, <strong>The</strong> main reasons for this<br />

outstandfng habitat qua1 l ty are: 1) large<br />

areas <strong>of</strong> relatively pristine wetlandsJ 2)<br />

habitat diversity (several wetlands<br />

exhibit cornpl ete envl ronmental con-<br />

tlnuums), and 3) relatively few barriers<br />

to movements. Dickinson Island and the<br />

Bassett Is1 and-Goose <strong>Lake</strong> areas represent<br />

relatively pristine wetl ands with direct<br />

hydrologic connection to <strong>Lake</strong> <strong>St</strong>. Clafr<br />

and few barriers to movements or<br />

migration. In Figure 31, one can readily<br />

see the habitat diversity, juxtaposition<br />

<strong>of</strong> environments, and mix <strong>of</strong> open water and<br />

vegetation which contributes to high<br />

wild1 ife values (We1 l er and Spatcher<br />

1965).<br />

<strong>The</strong> importance <strong>of</strong> the coastal<br />

wetlands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> with<br />

reference to waterfowl has a1 ready been<br />

discussed in Section 3.6. A1 though some<br />

waterfowl breeding does occur in the delta<br />

wetlands, the primary importance <strong>of</strong> the<br />

<strong>St</strong>, <strong>Clair</strong> River wetlands is in regard to<br />

resting and feeding habitat during<br />

migration. Because <strong>of</strong> availability <strong>of</strong><br />

preferred plant and invertebrate foods<br />

(Dawson 19751, as well as fts location<br />

along the Atlantic and Missfssippi Flyways<br />

(Be1 1 rose 1968, 19761, these wetlands<br />

serve as concentration areas and stopover<br />

points for both dfving and dabbling ducks.<br />

Habitat degradation along western <strong>Lake</strong><br />

Erie and Saginaw Bay, due to excessive<br />

turbidity, siltation and water quality<br />

decline (Martz and Ostyn 1977; U.S.<br />

Department <strong>of</strong> Interior 19771, i s<br />

associated with greater re1 iance on the<br />

<strong>St</strong>. <strong>Clair</strong> Delta wetlands. In this regards<br />

the waterfowl habltat deficien~les<br />

predicted for southeast Michigan as <strong>of</strong> the<br />

year 2000 are most significant (U.S.<br />

Department <strong>of</strong> Interior 1971).<br />

Large numbers <strong>of</strong> furbearers as we11<br />

as birds, reptiles, and amphibians have<br />

been identified in the <strong>St</strong>. <strong>Clair</strong> Delta<br />

wetlands (Herdendorf et al, 1981~).<br />

Muskrats abound in the cattafl marshes and<br />

raccoons are particul arly numerous where<br />

large trees occur. In comparison to the<br />

Canadian wetl ands, whfch consist largely<br />

<strong>of</strong> cattafl marshes and open-water areass<br />

the Dickinson Island wetlands on the<br />

American side <strong>of</strong> the delta exhibit much<br />

greater habitat diversity. Given access<br />

by boat via the abandoned channel which<br />

winds across Dickinson Island, both<br />

consumptive and nonconsumptive users can<br />

easfly enjoy this rich wildlife resource.<br />

To evaluate fish, wildlife and other<br />

values <strong>of</strong> wetlands, the Michigan Depart-<br />

ment <strong>of</strong> Natural Resources has developed a<br />

field evaluation manual (Wolverton 1981).<br />

However, the <strong>St</strong>. <strong>Clair</strong> Delta wetlands have<br />

not been rated by thfs agency.<br />

Neverthel essJ because <strong>of</strong> its exceptional<br />

fish, waterfowl, and other wildlife<br />

values, along with its accessibility to<br />

users and re1 at i vel y undeveloped nature,<br />

particularly Dickinson Is1 and# these<br />

wetl ands may we11 be among the mast<br />

valuable in the Great <strong>Lake</strong>s region.


5.4 WETLAND DISTURBANCE<br />

CHAPTER 5. HUMAM 81MPA67S AND APPLiED ECOLOGY<br />

A new awareness has arisen in the<br />

last decade over the alarming loss <strong>of</strong><br />

wetlands. In Michigan, 71% <strong>of</strong> the coastal<br />

wetlands have been lost (Jaworski and<br />

Raphael 1978) and much <strong>of</strong> the remaining<br />

42r420 ha have been altered because <strong>of</strong><br />

degraded water quality, changes in<br />

hydrology, and other factors, However,<br />

losses and degradation are not solely due<br />

to human intervention. Western <strong>Lake</strong> Erie<br />

has suffered a significant loss <strong>of</strong> aquatic<br />

vegatatlon, possfbl y because <strong>of</strong> changing<br />

1 ake 1 eve7 s, water movements, and carp<br />

feeding habits (Core 1948; <strong>St</strong>uckey 1971).<br />

Saiches and flooding may temporarily<br />

reduce emergent wetlands; however,<br />

regeneration usually occurs. Converse1 y,<br />

human impacts such as dredging, filling,<br />

diking* and draining have a permanent<br />

adverse impact on the resource base.<br />

Adverse impacts in the <strong>St</strong>. <strong>Clair</strong><br />

wetlands began wlth the exploitation <strong>of</strong><br />

fish and wildlife prior to European<br />

contact and continued Into the 1850s.<br />

Howeverr these impacts were modest and the<br />

losses somewhat replaceable through<br />

natural reproducllon. Impacts which fol-<br />

lowed were more permanent. We w i l l<br />

discuss the adverse impacts within a<br />

historical framework spanning about 125<br />

years (Figure 53). This is how our<br />

present problems emerged and this is the<br />

way they have been addressed.<br />

Historicallyr modlficatfon <strong>of</strong> the<br />

wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> began shortly<br />

after European contact. Historical<br />

accounts suggest that the wetlands <strong>of</strong> Lak?<br />

<strong>St</strong>, <strong>Clair</strong> were not excessively exploited<br />

by the fur traders as were other wetlands<br />

<strong>of</strong> the Great <strong>Lake</strong>s such as those along<br />

western <strong>Lake</strong> Erie. An abundance <strong>of</strong><br />

wildlife and fish attracted farmers from<br />

the settlement at Detroit. <strong>The</strong> desirable<br />

quallty and quantity <strong>of</strong> fish and wildlife<br />

led to the establlshment <strong>of</strong> fishing and<br />

huntlng clubs. In the mid-19th century*<br />

one such club, the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Fishing<br />

and Shooting Club, spent $80r000 to<br />

improve the property it occupied in the<br />

delta (Jenks 1912).<br />

Historic maps reveal that the<br />

wet1 ands were not impacted sfgnif icantly<br />

In the 1850s (Meade 18571, Although<br />

maritime commerce was carried on through<br />

the lake and its distributary channels,<br />

emergent aquatics were abundant and<br />

diverse throughout the <strong>St</strong>. <strong>Clair</strong> Delta and<br />

the mouth <strong>of</strong> the Clinton River. Jenks<br />

(1912) noted that wild rice (Zlrsnls<br />

aauatlca) was abundant as were at least<br />

116 specfes and 25 varieties <strong>of</strong> wetland<br />

macrophytes, including sedges. Fish and<br />

wildlife were exploited in a non-<br />

commercial capacity in the area. In 1857<br />

New Baltimore was the only community in<br />

the northern part <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Significant wetland disturbances are<br />

associated with two interrelated cultural<br />

processes: agricultural development, which<br />

was followed by urban growth. <strong>The</strong><br />

surveyor-general <strong>of</strong> the United <strong>St</strong>ates<br />

reported in 1815 that a large part <strong>of</strong> the<br />

southeastern regfon <strong>of</strong> the Michigan<br />

territory was swamp and practically<br />

worthless. As early as 1826 attempts were<br />

made for reclamation; but It was not until<br />

September 1850 that a general swampland<br />

law was enacted. Its purpose was to allow<br />

for draining and dikfng <strong>of</strong> nworthless<br />

public lands, lying as marshes or subject<br />

to periodic overflow by adjacent water-<br />

coursesw (Donaldson 19701,<br />

<strong>The</strong> 1850 swamp act stimulated<br />

significant wetland alteration. By 1873


Drec?ging, Filling, Rulkheadir~g<br />

- URBANIZATION -<br />

Figure 53. Historic trends in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetland disturbance,<br />

the land cover between the Detroit and establishment <strong>of</strong> the <strong>St</strong>, <strong>Clair</strong> Ship Canal,<br />

Clinton Rivers was in agriculture. <strong>The</strong> a 6 m channel dredged through South<br />

north shore <strong>of</strong> the lake a1 so supported Channel for commerclal navigation.<br />

agriculture. Establ ished villages such as<br />

New Baltimore grew, and new urban centers When the Grand Trunk Railway, the<br />

such as Anchorville were founded. thirdmode, wasconstructed, it dissected<br />

<strong>Wetlands</strong> at the head <strong>of</strong> the Detr<strong>of</strong> t Rlver the remaining wetlands in several places.<br />

were also being dredged. On the <strong>St</strong>, <strong>Clair</strong> However, on this shore1 ine most <strong>of</strong> the<br />

Delta, hardwoad forests <strong>of</strong> the premodern wetlands had a1 ready been drained so the<br />

deltawere cleared for farmland. About effect <strong>of</strong> this railroad was not<br />

half <strong>of</strong> the modern delta on Harsens Island particularly adverse.<br />

was diked.<br />

Several sportsmen's clubs were<br />

Three <strong>of</strong> trans~ortat'on which between 1850 and 1873 in<br />

were by the mid-1870s Ontarlo on portions <strong>of</strong> the delta which are<br />

provided improved access to the shoreline<br />

by boat, For example, in<br />

and An ra'lway was Ontario the <strong>St</strong>e, Anne Club and the Canada<br />

constructed the perimeter <strong>of</strong> the Cl ub were founded, while In Mlchigan the<br />

lake Detroit to A1gonacv <strong>The</strong> Grande Paint- and the Old Clubs were<br />

was On the h4gher ground' establ j shed. Hotels Lo accommodate summer<br />

generally av<strong>of</strong>dfng the remaining<br />

tourjsts from Detroit were constructed on<br />

between New BaltimOre and<br />

the South Channel levee. Since the levees<br />

*lgonacr the bed was constructed were law and subject to periodic flood'ing,<br />

through <strong>St</strong>. John's Marsh# a shall0w jnter- bul k-headf ng began to appear. Iso1 ated<br />

distributary bay col onired with emergent fill jng and bulkheadjng was most fntense<br />

aquatics, a1 ong South Channel f the ~aSn navigatfon<br />

A second mode <strong>of</strong> transport with channel) and In ii few Isolated localities.<br />

adverse impacts on the delta was the <strong>The</strong> first road on the <strong>St</strong>, ClaJr Delta was<br />

105


constructed on the west side <strong>of</strong> South<br />

Channel to allow access and settlement. In<br />

contrast, the shore1 ine from the ihames<br />

River northward remained unaltered except<br />

for 1 imited settlement in the northern<br />

port1 on <strong>of</strong> Wal pol e Is1 and.<br />

Channel ization for commercial nav-<br />

igatfon began in earnest In 1886. <strong>The</strong><br />

U. S. Congress authorized the deepening <strong>of</strong><br />

the ClSnton River (Figure 54) to 2.4 m,<br />

and <strong>Lake</strong> <strong>St</strong>. Claf r and the <strong>St</strong>. <strong>Clair</strong> Flats<br />

Channel to a depth <strong>of</strong> 8.4 m. By 1892 the<br />

<strong>St</strong>. <strong>Clair</strong> Rlver was dredged as we1 1. On<br />

the lower portlon <strong>of</strong> South Channel,<br />

channels were dredged through the natural<br />

levees into Big Muscamoot Bay and<br />

bulkheading and backfilling for summer<br />

houslng contlnued. By 1903 the modern<br />

delta wetlands (the shal low inter-<br />

distributary bay) <strong>of</strong> Harsens Island<br />

continued to be drained, diked, and used<br />

for agriculture.<br />

<strong>The</strong> period 1900-1930 was the<br />

beginning <strong>of</strong> urbanlzation. Channels and<br />

roads were constructed a1 ong Middle and<br />

South Channels. <strong>The</strong> electric rail road was<br />

removed and replaced by highway M-29,<br />

linking Algonac with communities to the<br />

west. Recreational boating facflfties<br />

began to appear. Urban areas and<br />

waterways expanded at the expense <strong>of</strong><br />

agriculture. Urbantzation more than<br />

doubled and agricultural land use declined<br />

to one-half in the <strong>St</strong>, John's Marsh area<br />

(Table 34 and Figures 55 and 56). Direct<br />

lmpacts on the wetlands contlnued.<br />

In 1945 Intense commercIa1 and urban<br />

growth began. Urban strip development<br />

extended from Detroit t o Algonac.<br />

Dredging, f i 11 f ng, and channel ization<br />

consumed and drained a sjgnificant portion<br />

af <strong>St</strong>, John's Marsh. Urban development<br />

also extended along the perimeter <strong>of</strong><br />

Harsens Is1 and. To improve waterfowl<br />

habitat, most <strong>of</strong> the remalnlng portion <strong>of</strong><br />

Marsens Island was diked and planted i n<br />

corn and other crops. Although most <strong>of</strong><br />

the Ontarto portfon <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Flats<br />

stfll remadns as the Walpole Island Indian<br />

Reserve, south <strong>of</strong> Wftchell Bay wetlands<br />

have been drained and channelized for<br />

agriculture. Dfkes were constructed to<br />

a1 1 e vi ate f 1 ood i ng and control water<br />

levels fn formerly unrestricted wet1 ands.<br />

<strong>The</strong> diking projects on Harsens Island and<br />

fn Ontario extended into the cattail<br />

marsh, Sedge and dogwood/meadow wet1 ands<br />

which are located at or just above the<br />

water table? were the areas initially<br />

drained for agrfculture. An analysis <strong>of</strong><br />

coastal land use, based on 1973<br />

topographfc maps during record high lake<br />

levels reveals that the only significant<br />

areas <strong>of</strong> intact wetlands in Michigan were<br />

within the interdistributary basins <strong>of</strong> the<br />

<strong>St</strong>. Cl ai r Flats and a small area south <strong>of</strong><br />

the Clinton River. In 1985 the Michigan<br />

wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> represented<br />

only some 29000 hectares <strong>of</strong> the land<br />

cover.<br />

Commerci a1 and recreational traffic<br />

has also had a considerable impact on <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. Bulk cargo and recreational<br />

boating have requ 1 red channel deepening.<br />

New channel and marine construction on the<br />

<strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> shoreline continues.<br />

Public maintenance dredging to accommodate<br />

1 ake carriers and private recreational<br />

craft is estimated to be 122,000 m3<br />

annual 1 y ( Rap hael et a1 . 1974) exclusive<br />

<strong>of</strong> the <strong>St</strong>. Clalr River. <strong>The</strong> construction<br />

<strong>of</strong> highway 1-94 from Detroit northward to<br />

Port Huron has improved access to the <strong>Lake</strong><br />

<strong>St</strong>, <strong>Clair</strong> shoreline. Reduced driving time<br />

from Anchor Bay and Algonac to Detrolt has<br />

encouraged suburban devel opment,<br />

partlcularly along the western shoreline<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. All <strong>of</strong> these factors<br />

have stimul ated development <strong>of</strong> the<br />

shoreline and have had adverse impacts on<br />

many <strong>of</strong> the wetlands.<br />

In part, because <strong>of</strong> the exceptional<br />

fish and wildlife productivity and<br />

Table a34. Land use change at <strong>St</strong>. John's<br />

Marsh.<br />

Land<br />

use -.LcuL- ..1974<br />

ha acres ha acres<br />

Urban 126 313 320 793<br />

Agriculture 822 2,035 431 1,066<br />

Waterways 21 52 109 269<br />

a Data source: Roller (1977 1.


Figure 54. Aerial photograph <strong>of</strong> the mouth <strong>of</strong> the Clinton River, Macomb County,<br />

Michigan, showing a coastal wetland (Verchev's Marsh) isolated from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> by<br />

channelization and residential development (June 1975).


Figure 551, Aerial photograph <strong>of</strong> <strong>St</strong>. John's Marsh on Bouvier Bay, and North Channel,<br />

<strong>St</strong>, <strong>Clair</strong> Delta (April 1949), Note wetlands established during low lake levels. Com-<br />

pare with Figure 56,


Figure 56. Aerial photograph <strong>of</strong> North Channel, <strong>St</strong>, <strong>Clair</strong> De'l ta, and <strong>St</strong>. John's Marsh<br />

on Bouvier Bay (May 1980). Compare with Figure 55, Note wetlands lost during high<br />

lake levels, residential development, dredge spoil sites, and highway (M-29) bi fur-<br />

eating marsh.


ecreational value <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalrr<br />

visitors from Mfchigan and Ontarfo<br />

continue to be attracted to the lake. <strong>The</strong><br />

lake is readfl y accessible to the urban<br />

areas <strong>of</strong> metropolitan Detroit and Windsor<br />

as well as numerous small urban areas such<br />

as Flint, Saginaw, Port Huron, and Sarnia.<br />

<strong>Lake</strong> <strong>St</strong>. Cl air is the most valuable Great<br />

<strong>Lake</strong>s area for non-salmonid sport fish in<br />

Mlchlgan. For example, in 1975* nearly<br />

half <strong>of</strong> the total Great <strong>Lake</strong>s fishing<br />

effort was expended on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

(Jaworski and Raphael 1978). Furthermorer<br />

fur (rwskrat and raccoon) production <strong>of</strong><br />

the area is among the highest in the<br />

<strong>St</strong>ate.<br />

Wlth increased commerce 4n the Great<br />

<strong>Lake</strong>s, a cut<strong>of</strong>f channel was dredged at<br />

South Channel where it debouches into <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. To accomodate Seaway traffic,<br />

a 8-m channel was dredged in 1952 through<br />

the interdistributary marsh for a dfstance<br />

<strong>of</strong> 10 km. Seaway Island, a large disposal<br />

site now occupies a portion <strong>of</strong> the<br />

Jntardistributary marsh. To improve<br />

recreational boating facillties and<br />

mitigate against further erosion,<br />

bulkheading and filling along all<br />

urbanized shore1 ines have occurred over<br />

the decades. Seawalls extendfng we1 1<br />

above hfyh water levels have replaced the<br />

natural levees which extended south from<br />

Harsens Island into <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, <strong>The</strong><br />

number <strong>of</strong> crevasse channels which<br />

transported water, sediment, and nutrients<br />

into Muscamoot Bay has decreased.<br />

<strong>The</strong> degeneratfon <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

wetlands follows a historical pattern<br />

similar to that <strong>of</strong> other large wetland<br />

complexes in the United <strong>St</strong>ates, Initially<br />

stlrnulated by the 1850 swamp acts,<br />

agricultural activity occurred. A second<br />

signfffcant stlmulant was access5billty<br />

dus t o ssveral transportatton<br />

devslapments. Inter-urban electric<br />

raf 1 roads, excursfon vessels, and<br />

eventually, roads and interstate highways<br />

made the wetlands and the shoreline<br />

readily accessible. Ef f l cient transport<br />

routes have encouraged urban and suburban<br />

expanston, changfng summer residences to<br />

year-round housing ,<br />

By contrast, access to the delta and<br />

the shoreline <strong>of</strong> Ontario is more dffficult<br />

because <strong>of</strong> the lack o f roads.<br />

Furthermore, southern Ontario is less<br />

populous and much <strong>of</strong> the eastern portion<br />

<strong>of</strong> the delta is an Indtan reserve, <strong>The</strong>se<br />

physical and social factors have not<br />

encouraged as much wetland loss as seen in<br />

Michigan* except at the northern end <strong>of</strong><br />

Walpole Island which has experienced<br />

wetl and loss.<br />

With the exception <strong>of</strong> the electric<br />

railroad through <strong>St</strong>. John's Marsh9 wetland<br />

losses occurred on the periphery <strong>of</strong> the<br />

resource; that is, modification began at<br />

the boundary between the marsh and the<br />

adjacent upland. In the <strong>St</strong>. <strong>Clair</strong> Delta<br />

agriculture was initiated on the premodern<br />

surface and gradual 1 y expanded southward<br />

to the transition (dogwood/meadow) zone<br />

and eventual f y into the shall ow inter-<br />

distributary bays. <strong>The</strong> natural levees<br />

were modifled by bulkheading in the 1870s.<br />

Filling, aided by bulkheads, extended into<br />

the deep interdistributary bays and onto<br />

the river shoulders. This pattern <strong>of</strong><br />

development has continued to the present<br />

because wetland boundaries are difficult<br />

to ascertain, especial 1 y i n the Great<br />

<strong>Lake</strong>s where temporary lower lake levels<br />

provide a false perception <strong>of</strong> the position<br />

<strong>of</strong> the shore1 ine.<br />

Adverse impacts to the artificial<br />

(diked wetrands1 and 11 natural<br />

envi ronments identif led in section 4.1 as<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands are shown in Table<br />

35. With the exception <strong>of</strong> the recently<br />

abandoned channels, which are in Ontar i or<br />

and the transgressive beaches which are<br />

generally Is01 ated, al? the wet1 ands have<br />

had adverse impacts. Bulkheads and fills<br />

have been constructed on the river<br />

shoul ders, crevasses, and margins <strong>of</strong> the<br />

deep-water Interdistributary bays for over<br />

a century. To construct roads, spoil was<br />

dredged from the landward side <strong>of</strong> the main<br />

channels which inadvertently led to the<br />

channel ization <strong>of</strong> the shallow bays.<br />

Bulkheads are a common structural feature<br />

along many shorelines <strong>of</strong> the lake and<br />

delta and appear to be detrimental to<br />

submersed aquatics ( Schl oesser and Manny<br />

1982). Bulkheads constructed on river<br />

shoulders occupy the shallowest portion <strong>of</strong><br />

the rlvsr. Consequently, as dredge and<br />

f i 11 occursr submersed wetl ands are<br />

df spf aced i akeward where bottom attachment<br />

fs more difffcult because <strong>of</strong> stronger<br />

currents and greater depths.


Table 35. Adverse impacts to the various morphology types <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> wetlands.<br />

Wet1 and<br />

morphology type<br />

Impact Exampl e<br />

1. River shoulders bulkheaded, South Channel<br />

dredged dl sposal<br />

2. Abandoned river drainedr agriculture Harsens Island<br />

channels<br />

3. Recently abandoned no change<br />

channels<br />

4. Crevasse deposits channel 1 zed,<br />

bulkheaded<br />

Bassett Channel<br />

South Channel<br />

5. Shallow inter- di ked/drained, Harsens Is1 and<br />

dlstrlbutary bays agrtculture~<br />

channel i zed<br />

6. Deep lnterdis- bulkheadedr<br />

tributary bays channel ized<br />

Harsens Is1 and<br />

7. Transgress1 ve minor urban Big Muscamoot Bay<br />

beaches settlement,<br />

bul kheaded<br />

* ,<br />

8. Shallow shelves dikedr filled, Mitchell Bay<br />

channel i zed<br />

9. Barrier/l agoon channel fzed<br />

*<br />

~orth <strong>of</strong> Thames River<br />

10. Transi tional areas diked, agriculturer<br />

filled for residenti<br />

a1 devel opment<br />

Harsens Is1 and<br />

11. Estuarl ne/<br />

floodbasf n<br />

12. Diked<br />

diked/drainedr <strong>St</strong>. John's Marsh<br />

agriculture/urban<br />

d i ked/drai ned<br />

East shore1 i ne<br />

(orfgf nally shelves) (O~tarlo)


To accommodate rec reattonal boating<br />

and commerci a1 shippl ng, channels to<br />

depths <strong>of</strong> 8 m have been dredged on the<br />

perimeter and In the basin <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />

<strong>Clair</strong>. Also there is considerable<br />

residential pressure on the shore1 fne to<br />

include the wetlands <strong>of</strong> the basin,<br />

Bu?kheadtngr ftlling, dtking9 and djsposal<br />

<strong>of</strong> dredged materials have occurred since<br />

the early 1900s.<br />

Because <strong>of</strong> the need for commercial<br />

navigation and recreational boating fn<br />

take <strong>St</strong>. <strong>Clair</strong>, a significant amount <strong>of</strong><br />

channel dredging has occurred. <strong>The</strong> U.S.<br />

Army Corps <strong>of</strong> Engineers, which Is<br />

responrj-i ble for harbor maintenance, has<br />

conducted dredging and disposal activities<br />

at the Cl inton and <strong>St</strong>. Claf r Rfvers and<br />

the 8 m navigation channel bisecting the<br />

lake, In Ontario the Department <strong>of</strong> Public<br />

Works mafntains the <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f<br />

Channel and the harbor at Wallaceburg,<br />

east <strong>of</strong> the <strong>St</strong>, Clalr Rlver Delta. To<br />

maintafn channels at the proscribed<br />

depths, the aceumul ated sedlment was<br />

normal 1 y hydraul Jcall y dredged and<br />

dfsposud <strong>of</strong> in convenient areas such as in<br />

designated portions <strong>of</strong> the open lake.<br />

<strong>The</strong> volume <strong>of</strong> materfals dredged for<br />

take <strong>St</strong>. ClaJr and the <strong>St</strong>, <strong>Clair</strong> River<br />

navjgatlon projects Is shown on Table 36.<br />

Over the 52-year perJodp about 782,800 m3<br />

were dredged from U.S, project sites in<br />

the two areas. More recent data indicate<br />

that from 1971 through 1984 dredging<br />

activity in <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> totaled 382r300<br />

m3, <strong>The</strong> average for the 14-year period Is<br />

27t300 m3 annually which is considerably<br />

less than the historlcal average <strong>of</strong><br />

313,200 m3 for <strong>Lake</strong> <strong>St</strong>. Clalr. On the<br />

Cl inton RIver the 1971-1984 total was<br />

45,000 d, which approximates the historic<br />

average.<br />

<strong>The</strong> Ontario maintenance dredging and<br />

new work dredging averaged about 195,800<br />

m3 annually. Most <strong>of</strong> the dredging in<br />

Ontario is confinad to the <strong>St</strong>, <strong>Clair</strong><br />

Cut<strong>of</strong>f Channel (182,400 d/year).<br />

In 1970 the U.S. Congress enacted<br />

Pub1 dc Law 91-611 (River and Harbor Flood<br />

Control Act), which authorized the Corps<br />

to construct, operate, and malntain<br />

confined disposal sites for containment <strong>of</strong><br />

polluted dredged spoil in the Great <strong>Lake</strong>s<br />

for a period not to exceed 10 years. At<br />

that timer the <strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel<br />

and <strong>Lake</strong> <strong>St</strong>, Clajr sediments were 100%<br />

polluted9 and the Clinton River was 85%<br />

polluted (Raphael et al. 19741. <strong>The</strong><br />

pollutant was primarily mercury, although<br />

the conventional pollutants were also<br />

beginning to reach limits set by the U.S.<br />

Table 36. Hiituricaa dredging total 5 <strong>of</strong> Corps <strong>of</strong> Engineers projects in <strong>Lake</strong><br />

<strong>St</strong>. Cldir 1320-1972.<br />

---<br />

Mafntenance New work Annual<br />

d redgi ng dredgl ng Total average<br />

Channel oF Laire <strong>St</strong>. <strong>Clair</strong> 2~894r400 13~634.600 1Sr529,000 317,900<br />

Cl inton Rlver 237 r 400 96,900 334,300 6,400<br />

<strong>St</strong>. Cl air River 3 ,125,300 20~879~200 24r004r500 461,600<br />

a Data source: Raphael et aS, (19741.<br />

i 12


Envl ronmental Frotactfon Agency. To<br />

comply wlth PL 91-631, two conflned<br />

disposal sites with a combtned cdpaclty <strong>of</strong><br />

1.5 mllfion m3 were constructed at the<br />

north end <strong>of</strong> Dfckinson Island. Both sltes<br />

were placed on tho premodern surface <strong>of</strong><br />

the <strong>St</strong>. <strong>Clair</strong> Delta. <strong>The</strong> west site<br />

enclosed an area <strong>of</strong> about 22 hectares and<br />

the east site about 48 hectares (U.S. Army<br />

Corps <strong>of</strong> Engineers 1974). In Ontarfo the<br />

<strong>St</strong>. <strong>Clair</strong> Cut<strong>of</strong>f Channel was constructed<br />

in the early 1950s to accommodate Seaway<br />

traffic along South Channel, <strong>The</strong> channel<br />

was dredged through the fnterdistrlbutary<br />

bay and was 1 inked to the main navigation<br />

channel in <strong>Lake</strong> <strong>St</strong>. Clalr. Dredge spoll<br />

was put Sn a disposal sltu (Soaway Island)<br />

along the east bank <strong>of</strong> South Channel. <strong>The</strong><br />

facflity covers an area <strong>of</strong> about I1 km2,<br />

which was prlmarlly colonized wfth<br />

emergent macrophytes typical <strong>of</strong> an Inter-<br />

dl stributary bay snvl ronment.<br />

Private or contract dredglng (under<br />

Sec.10 permit <strong>of</strong> the U. S. River and<br />

Harbor Act <strong>of</strong> 1899) is done by local<br />

cftizens or homeowners to fmprove access<br />

to <strong>Lake</strong> <strong>St</strong>. Clalr or to construct<br />

boatslfps along the lake or the<br />

dfstributaries (Figure 57). <strong>The</strong> data are<br />

not normally tabu1 ated; however, based on<br />

our estimates, private dredging volumes<br />

probably average about 7,600 to 11,400 m3<br />

annually. SSgnff tcant areas <strong>of</strong> dradglng<br />

for restdential development have occurred<br />

Figure 57. Commercial development tin Har-<br />

sens Is1 and wetland (August 1984).<br />

on the shoreline <strong>of</strong> Anchor Bay, <strong>St</strong>. John's<br />

Marsh, at the mouth <strong>of</strong> the Clinton RIver<br />

and along the distrfbutary channels <strong>of</strong> the<br />

dsl ta, parttcul arl y North and South<br />

channels. Prfvate dredging in btarfo is<br />

lim~ted to approach channels for<br />

recreational craft Jn Mitchell Bay and a<br />

marina complex near Belle River on the<br />

south shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr.<br />

Dfking fragments coastal wetlands,<br />

separating the managed parcels from the<br />

adjacent littoral and upland environments.<br />

General1 y, as the slre <strong>of</strong> the wetland is<br />

diminfshed, so Is the vegetative and<br />

wildlife diversity (Larsen 1973).<br />

Furthermore, dik 1 ng <strong>of</strong> wet1 ands reduces<br />

vegetation diversity because water level<br />

controls are established to maintain a<br />

particular vegstatfon type or envlron-<br />

mental condftlon. <strong>St</strong>abll $zing water<br />

levels tends to eliminate those plant<br />

species whfch raqul re low-water periods<br />

for regenerat 1 on, and promotes domfnance<br />

<strong>of</strong> a few tolerant spectes such as cattafl<br />

( spp. 1 and woody plants (Keddy and<br />

Reznfcek 1984).<br />

<strong>The</strong> hydrologic isolation <strong>of</strong> dlked<br />

systems 1s reflected In their greatly<br />

reduced flsh use and simpllfled<br />

1 nvertebrate commun5t~esr particularly<br />

where wlnter ice and summer temperatures<br />

reduce df ssol ved oxygen and otherwise<br />

reduce the water qua1 ity. <strong>The</strong> effect <strong>of</strong><br />

earthen dtkes on waterfowl mfaratlon,<br />

other wild1 ffe movementsr and prtrdatfon Is<br />

not well known. Howevert DennTs and North<br />

(1981) have shown a dramatlc decline in<br />

the use <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> marshes by<br />

true marsh-dwellfng specles such as the<br />

Amerlcan wigeon (<br />

), green-wfnged tea2 (<br />

ood duck (&.&<br />

1968-69 to 1976-77 the use by these<br />

dabbl lng ducks decl fned by 73% durf ng the<br />

spring pertod. Autumn marsh use also<br />

declfned by 40%. Howeverr the<br />

investigators noted that sprlng use<br />

generally provides a batter f ndicatlon <strong>of</strong><br />

the value <strong>of</strong> the wetlands to mfgrant<br />

waterfowl si nee the managemnt techn l quas<br />

whfch were used during the fall huntfng<br />

seasons are not Implemented during the<br />

sprf ng,


Discovery <strong>of</strong> mercury contaml natfon In<br />

tha <strong>St</strong>. Clafr Rfver-<strong>Lake</strong> <strong>St</strong>. Clafr-Detrolt<br />

RJver-<strong>Lake</strong> Erie system in 1970 generated<br />

great pub1 l c Interest In the mercury<br />

levels <strong>of</strong> water and flsh, Subsequent<br />

banntng <strong>of</strong> sport and commercial ffshlng<br />

led to testing for trace mercury by<br />

varjous <strong>St</strong>ate and Federal agencies to<br />

def f ne the extent <strong>of</strong> the pollution and to<br />

determf ne its major sources. Two chemfcal<br />

plants, Dow Chemical Corporatlon, Sarnia*<br />

Ontarfo, and BASF Wyandotte Chemical<br />

Corporatlon, Wyandotte, Mfchfgan, were<br />

fdentff fed as major contributors to the<br />

mercury pollution <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> and<br />

<strong>Lake</strong> Erie (Waiters et al. 19741, <strong>The</strong>se<br />

companfes manufactured chlorf ne gas and<br />

caustlc soda by the alactrolytlc process,<br />

whlch lnvolves the slectrolysis <strong>of</strong> brfne<br />

to produce chlorlne gas and a mercury-<br />

sodium metal ama'lgarn. <strong>The</strong> amalgam 1s<br />

contacted wlth water to produce sodium<br />

hydroxf de, the mercury metal belng<br />

recycled ta tho afectrotytfc coll, <strong>The</strong><br />

rate <strong>of</strong> accfdental mercury release from<br />

these two plants has been sstfmated to be<br />

25 kg per day from 1950 to 1970 for the<br />

Dow Chsmlcal plant and S to 10 kg per day<br />

from 5939 to 1970 for the OASF Wyandotte<br />

plant (Federal Water Oualfty<br />

Admfnfstratfsn l970).<br />

Waltars @t at, (19741 calculated that<br />

228 metrlc tons <strong>of</strong> mercury pol Jution had<br />

been laadad to the top 20 cm <strong>of</strong> western<br />

<strong>Lake</strong> Erie sediment durlng tho p~rlod 1939<br />

to 1971, Sodlment cores takan at the<br />

mouth <strong>of</strong> the Detr<strong>of</strong>t Rfver and wastern<br />

<strong>Lake</strong> Erica y folded surfaca mercury valutls<br />

up to 3.8 ppm whleh generally decreased in<br />

concentratton exyonentfally wfth depth<br />

f FIgurs $81. Several yoars after the<br />

chlor-alkalf plants diminished aperation<br />

ths area was agafn carad (Wflson and<br />

Wdlturs 19789 vfth analyses shovfng that<br />

recent deposits covered the high1 y<br />

contrrnlnated sedfnrcsnt wfQh a thrn layer <strong>of</strong><br />

new material vhlch had mercury<br />

concentrat l ons approaching backgraund<br />

lsvsls 6B,b ppm). As a result af these<br />

discharges, marcury fn flsh <strong>of</strong> take <strong>St</strong>.<br />

Clnlr and western <strong>Lake</strong> Erfe was a major<br />

contaminant problem In the early 1970s.<br />

Levels <strong>of</strong> total mercury in walleye<br />

C L 1 col lectsd from<br />

Figure 58. Mercury concentration in sedi-<br />

rrient core from western <strong>Lake</strong> Erie near<br />

mouth <strong>of</strong> Detroit River (Halters et al.<br />

1974). Note mercury-enriched surface zone<br />

over1 yi ng sediment havi ny natural back-<br />

ground mercury level s (dashed 1 ine).<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> have decl ined from over 2<br />

ug/g In 1970 to 0.5 ug/g In 1980. In<br />

western <strong>Lake</strong> Erie, 1968 levels <strong>of</strong> mercury<br />

were 0.BA3ug/g as compared to only 0.31<br />

ug/g in 1976. <strong>The</strong> rapid environmental<br />

response subsequent to the cessation <strong>of</strong><br />

the point source discharges at Sarnia,<br />

Ontarlo, and Wyandotte* Michigan, can be<br />

attributed to rapid flushing <strong>of</strong> the <strong>St</strong>.<br />

Cl a1 r-Detroit Rf ver system, the high load<br />

<strong>of</strong> suspended sediment delivered to western<br />

<strong>Lake</strong> Erier and t h e high rate <strong>of</strong><br />

productivity (International Jolnt Com-<br />

missjon 1981).<br />

Mudroch and Capob-ianco (1978) studied<br />

the relatlonshlp between the concentration<br />

<strong>of</strong> seven metals In sediment and<br />

marshwater, and uptake <strong>of</strong> these metals by<br />

emergent, submersed, and fl oat1 ng-l eaved<br />

plants growing In wetlands located on the<br />

east shore <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong>y found<br />

a hf gh correl atton between metals and<br />

organfc carbon contents In the sediments.<br />

<strong>The</strong> accumulation <strong>of</strong> metals In plants<br />

varied fram specfes to species (Table 37)<br />

and showed a complex relatfanshlp with<br />

metal concentratfons i n the sed i ment .<br />

Water-mf l foil ( spp.) and<br />

pondweed ( lated more<br />

metal than the ather aquatic plants.


Table 37. Metals concentratiorls in livirly plant tissues from Big Point<br />

Marsh, Ontario on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. a<br />

Metals (ggm)<br />

Specles Pb Zn C r N! Cd Go Cu<br />

a Data source: Mudroch and Capobianco (1978).<br />

Metal concentrations In roots <strong>of</strong> broad-<br />

leaved cattat purple<br />

loosestrjfe ( , and<br />

pickerel weed were<br />

found to be hlgher than in the aboveground<br />

biomass. <strong>The</strong> metals taken up by marsh<br />

flora are retafned In the tissues and<br />

after decay they become a part <strong>of</strong> the<br />

marsh surface sediment.<br />

Jaworskf and Raphael (1976) compiled<br />

comprehensfve data on wetland loss for the<br />

Mlchfgan sjde <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. In 1873<br />

the U.S. portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

supported 7,274 hectares <strong>of</strong> wetland<br />

vegetation (TabTe 381. By 1973 this<br />

habitat had dwlndled to 2,020 hectares.<br />

Significant losses not on1 y occurred on<br />

the <strong>St</strong>. <strong>Clair</strong> Delta and <strong>St</strong>. John" Marsh<br />

but on the entire margin <strong>of</strong> the lake as<br />

well. Gaukler Polnt at the head <strong>of</strong> the<br />

Detroit River was colonized with 187<br />

hectares <strong>of</strong> wetlands and the Clfnton River<br />

had over 1,295 hectares <strong>of</strong> habitat at its<br />

mouth and f t s flood basln. Some coastal<br />

areas* particular1 y north <strong>of</strong> the Cl i nton<br />

River appear to have been drained for<br />

agriculture in the 1860s suggestfng that<br />

the 1868-1873 data represent rninjrnum<br />

rather than maxlmum wetland acreage.<br />

In contrast to Michigan's wetlands<br />

being lost to the ong<strong>of</strong>ng urbanizationr<br />

Ontarfo wetlands are being lost to<br />

agriculture, <strong>The</strong> wetlands from the Thames<br />

River north to Chenal Ecarte dwindled Prom<br />

about 3,600 hectares in 1965 to 2,900<br />

hectares in 1970 (McCul lough 1982). About<br />

700 hectares or nearly 20% <strong>of</strong> the<br />

prlvately owned resource base was lost<br />

over the 5-year period. Figure 59 and<br />

Table 38 reveal specific areas <strong>of</strong> wetland<br />

loss <strong>of</strong> the coastal Ontarlo wetlands<br />

excluding the Walpole Island Indian<br />

Reserve. Drai nlng for agricultural use<br />

has accounted for 91% <strong>of</strong> the wetland loss<br />

whereas marina and cottage development has<br />

consumed 9% <strong>of</strong> the resource. Clearly,<br />

urban pressure Is presently a problem In<br />

Michigan and agriculture pressure a<br />

problem in coastal Ontario,<br />

Although the Ontario shore <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>, <strong>Clair</strong> is low lying and subject to<br />

flooding, erosion has on7 y been tdentified<br />

as a hazard from the Thames Rfver westward<br />

(Boul den 1975). This suggests that<br />

permanent loss <strong>of</strong> wetland habitat north <strong>of</strong><br />

the Thames River 1s due t o human<br />

interventton. During the record high lake<br />

l eve1 fn the early 1970~~ about lrOOO<br />

hectares <strong>of</strong> emergent shore1 f ne marsh f rom<br />

Mitchell Bay southward to the Thames River<br />

were temporarily lost (McCullough 19821.<br />

This loss was tempered in part by the<br />

flooding <strong>of</strong> transitjon vegetation which<br />

occurred on the upland (east) margfn <strong>of</strong><br />

the wetlands, Slmflarly the delta and<br />

Anchor Bay are subject to flooding, though<br />

erosion ds not identified as a serfous<br />

problem (Great <strong>Lake</strong>s Basfn Commission


Table 38. -<br />

a<br />

Michigan and Ontario wetland losses on <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong>.<br />

Mlchigan<br />

1 ocat i on<br />

1868-73<br />

(ha)<br />

1973<br />

(ha) Loss (ha)<br />

<strong>St</strong>. <strong>Clair</strong> Flats 5,473<br />

Swan Creek 75<br />

Marsac Point 6 1<br />

New Baltimore 2 1<br />

Salt River 162<br />

Cl i nton River 1,295<br />

Gaukler Polnt 187<br />

Total 7,274 2 r 022 5,252<br />

Ontar lo 1965 to 1973 Wet1 and<br />

1 ocat i on Area loss (ha) type Cause<br />

Thames Rfver 59<br />

Thames Rlver mouth 7 4<br />

Bradley Marsh 327<br />

Balmoral Marsh 11<br />

Snake Island Marsh 156<br />

<strong>St</strong>. Lukes Bay 22<br />

Patricks Cove 5 9<br />

Mitchell Bay 7<br />

Mud Creek Marsh 167<br />

Tota 1 882<br />

diked<br />

open<br />

diked<br />

diked<br />

diked<br />

diked<br />

diked<br />

d I ked<br />

d l ked<br />

agriculture<br />

mari ne/cottage<br />

construction<br />

agriculture<br />

agriculture<br />

agriculture<br />

agriculture<br />

agriculture<br />

agriculture<br />

agriculture<br />

a Data sources: Jaworski and Raphael (19761, McCullough (1982).<br />

1975). <strong>The</strong> recent losses here are due to<br />

dlktng and/or ffllfngr fn most Instances,<br />

Par urban growth.<br />

<strong>The</strong> wetlands <strong>of</strong> <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> hav~<br />

been mapped from older navfgatton charts<br />

and recent topagraphlc maps CFlgure 60).<br />

Thfs figure illustrates signiffcant losses<br />

between 1873 and 1968 on the periphery <strong>of</strong><br />

the lake, However, the most apparent<br />

1 mpacts are evident in the Clinton River,<br />

the <strong>St</strong>. Clai r Delta* and the eastern shore<br />

<strong>of</strong> the lake. It is ev4dent that the<br />

wetlands in all three areas have been<br />

modified along their margins. <strong>The</strong><br />

wetlands on the eastern shore1 ine were<br />

approximately 2.5 km wide. Since 1873<br />

they have been impacted from the landward<br />

side and are now about 0.8 km in width.<br />

<strong>The</strong> progression <strong>of</strong> losses Jn the <strong>St</strong>. <strong>Clair</strong><br />

Delta follows a similar geographic<br />

pattern. <strong>The</strong> losses in both areas were<br />

fnf tial ly stimulated by agriculture. In<br />

the Cl inton River area, wetl and losses<br />

occurred from both the landward as well as<br />

the 1 akeward boundary. Fundamental lyr an<br />

1 sol ated wetl and has been created.


Figure 59. Wetland losses along the Ontario shore1 ine <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. C'lair<br />

(McCul lough 1982).


Figure 60. Comparison <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clai r<br />

1968.<br />

Table 39 represents a summary <strong>of</strong><br />

wetland losses in <strong>Lake</strong> <strong>St</strong>. Clalr. Since<br />

the topographic quadrangles and the<br />

navigation charts only dlsplay emergent<br />

wetlands, the area values <strong>of</strong> submergent<br />

aquatics were estimated. <strong>The</strong> re1 atlve<br />

1 osses between the historical perf ods are<br />

evldent. <strong>The</strong> losses on Table 39 are<br />

greater than that noted by Jaworski and<br />

Raphael (1976) because thais data did not<br />

include what is now surm!sed to be<br />

submersed vegetation, <strong>The</strong> percentage <strong>of</strong><br />

wetland area lost fn Michfgan is greater<br />

than In Ontario (45% vs. 34X)r but the<br />

actual area lost in Ontario exceeds that<br />

lost in Michigan. In Michigan as well a5<br />

fn Ontario* more <strong>of</strong> the wetland base was<br />

l ost along the shore1 ines than in the<br />

delta. Slnce the coastal zone wetlands<br />

are parallel to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, their<br />

accesslbilfty plays an faportant role in<br />

thefr modification. <strong>The</strong> <strong>St</strong>. Clafr Delta<br />

conversely projects into <strong>Lake</strong> <strong>St</strong>. Clafrr<br />

limfting accessibi7ity and hence adverse<br />

LAKE<br />

38.968<br />

coastal wetlands distribution in 1873 and<br />

impacts. It should not be inferred from<br />

the data that the wetland quality and<br />

diversity have not changed over the 95-<br />

year period. In sum, 9r 136 hectares <strong>of</strong><br />

the resource base have been lost.<br />

Both natural disturbances, such as<br />

water-level fluctuations (as the Great<br />

<strong>Lake</strong>s water levels oscill ate) as we1 7 as<br />

cultural disturbances ( ~ . g . diking) ~<br />

affect the wetland ecosystems. A s<br />

reflected In Eugene Odurnts qlpulse<br />

stability concept," some <strong>of</strong> these<br />

disturbances can be beneficial. In this<br />

section, however, the lmpacts <strong>of</strong> adverse<br />

cultural effects have been reviewed. Of<br />

particular concern are the following: 1)<br />

wet1 and loss* 2) dikl ng, fragmentationr<br />

and loss <strong>of</strong> hydrologic connectivity and<br />

3) changes in the environmental gradient<br />

and plant communities.


Table 39. Comparison <strong>of</strong> 1873 and 1968 <strong>Lake</strong> <strong>St</strong>, <strong>Clair</strong> wetland areas.<br />

Location<br />

<strong>St</strong>. <strong>Clair</strong> Delta 5,414 3,077 9,641 7,234 15,055 10,311<br />

Cl 1 nton River 19192 248 -- -- 1,192 248<br />

Remaining Mich.<br />

shore1 I ne 1,900 806 -- -- 1,900 806<br />

Remai njng Ontario<br />

shore1 1 ne -- -- 4,219 1,862 4,219 1,862<br />

-- -- -_IL<br />

Total 7,506 4,131 13,860 9,096 22,366 13.227<br />

As discussed in this section, during<br />

the period 1873-1968? approximately 9,136<br />

hectares <strong>of</strong> coastal wetlands have been<br />

lost. This amounts to a 41% reductlon in<br />

the original coastal wetl and resource base<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> greatest<br />

percentage loss has been on the American<br />

side and the greatest area loss on the<br />

Canadian side. More wetlands have been<br />

destroyed along the shorel ines than in the<br />

delta. In additionr except for Dickinson<br />

Island and <strong>St</strong>. John's Marsh* the upper<br />

zones (i.e.# the sedge marsh-shrub-swamp<br />

fringe) were converted to other land uses.<br />

<strong>The</strong> juxtapositfon <strong>of</strong> corn fields and diked<br />

cattail marshes on <strong>St</strong>. Anne Island is a<br />

case in point. At present, nearly half <strong>of</strong><br />

these extant wetl ands along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

consist <strong>of</strong> cattail communities.<br />

Loss <strong>of</strong> coastal wetlands along the<br />

Michigan sfde <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr results in<br />

a loss <strong>of</strong> wetland function and value. For<br />

example, public drains installed to<br />

improve run<strong>of</strong>f now occupy former creek<br />

channels, which no longer benefit from the<br />

flood water storage, sedfment trapping,<br />

and nutrient uptake afforded by the<br />

natural wetlands. Nor do the remalnlng<br />

wetlands along the river mouths and<br />

shorel ines, which have been reduced f n<br />

size, partially developed (especi a1 1 y on<br />

the lakeward sfdeIo and otherwfse<br />

impacted, have the fish and wildlife<br />

valves they once had.<br />

hectares* fn the <strong>St</strong>. Clafr Delta are now<br />

diked. This includes <strong>St</strong>. John's Marsh,<br />

which is isolated from <strong>Lake</strong> <strong>St</strong>. Clafr by<br />

earthen berms, but is not subject to water<br />

level controls. With the exception <strong>of</strong> <strong>St</strong>.<br />

John's Marsh, the upper part <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> Flats <strong>St</strong>ate Game Area, and a few<br />

other small parcels, over 75% <strong>of</strong> the diked<br />

wetlands are colonlred by cattails and<br />

associated submersed aquatic comunlties.<br />

<strong>The</strong> phrase "diked cattail marshesn is<br />

indeed appropriate. What is important<br />

here is that it fs the remainfng open-<br />

system wetl ands which are contributing to<br />

flsh production, wild1 lfe habitat, and<br />

dlving duck feeding <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. In<br />

contrast, the diked wetlands provide more<br />

<strong>of</strong> a single purpose - that <strong>of</strong> attracting<br />

dabbling ducks, especially mallards for<br />

waterfowl hunters.<br />

<strong>The</strong> adverse impact <strong>of</strong> isolatfng and<br />

fragmenting wetlands by means <strong>of</strong> roadbeds,<br />

canals, earthen dikes, and other<br />

developments appears not to be fully<br />

recognfzed. For example, many conser-<br />

vat1 onists 1 n Michigan are advocating the<br />

preservation <strong>of</strong> <strong>St</strong>. John's Marsh, but feb<br />

call for an increase In Its hydrologfc<br />

connectfvfty to <strong>Lake</strong> <strong>St</strong>, Clafr. more over^<br />

unless a wetland fs physically destroyed,<br />

not mere1 y f ragmentedr most wet1 and<br />

researchers would not refer to an Is01 ated<br />

wetland as be'lng gtlost.gt<br />

Approxlmately one-half <strong>of</strong> the Current coastal development not on1 y<br />

remafnfng coastal wetlands, f .e., 5,280 results .fn fragmentation and loss <strong>of</strong><br />

119


hydrologic connectivity, but also<br />

frequently Is assocfated with the loss <strong>of</strong><br />

As indtcated in Appendix M and on<br />

Figure 61r most <strong>of</strong> the ownership <strong>of</strong> the<br />

upper p1 ant communities (Jaworski and larger* extant coastal wetlands 1 ies in<br />

Raphael 1976). <strong>The</strong>refore, most <strong>of</strong> these public hands. Exceptions are <strong>St</strong>, Johnts<br />

extant wetlands conslst <strong>of</strong> just cattail<br />

and submersed aquatfc communities, rather<br />

Marsh, whlch the <strong>St</strong>ate <strong>of</strong> Michigan is<br />

attempttng to purchase from the remal n l ng<br />

than a compl eta wet1 and communfty private ownersr and Walpole Is1 and,<br />

contfnuum. Changfng <strong>of</strong> water levels In Ontarior which forms an Indian Reserve.<br />

the Great <strong>Lake</strong>s w i l l t then, result in the Many <strong>of</strong> the undfked coastal wetlands on<br />

loss <strong>of</strong>the current functfon fn a given<br />

wetland as opposed to the shift <strong>of</strong> this<br />

theCanadian side <strong>of</strong> the <strong>St</strong>. CJaSr Delta<br />

are owned by the Province <strong>of</strong> Ontario, wlth<br />

function laterally in accord with the the exception <strong>of</strong> Walpole Island, Canada's<br />

vegetatfon movements. In contrast, as<br />

exemplified by Dickinson Island, wetlands<br />

Department <strong>of</strong> the Environment (DOE) owns<br />

the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> National Wildlife Areap<br />

which are connected directly to <strong>Lake</strong> <strong>St</strong>. whereasDucks Unlimited has ownership OF<br />

<strong>Clair</strong> and sxhlbit a full environmental<br />

gradient, tend to experience lateral<br />

diked marshes south <strong>of</strong> Mitchell Bay.<br />

shifts In function and values.<br />

Furthermore, they are mafntained at little<br />

<strong>The</strong> land ownership <strong>of</strong> the Michigan<br />

portion <strong>of</strong> the delta is shown in Figure<br />

or no cost to the pub1 ic. 62. Two open water parcels, f.e.* Units A<br />

and B <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Waterfowl Refuge,<br />

Small, isolated wetlands near<br />

developments such as suburban housing*<br />

marinas, etc., oxhibit proximity and <strong>of</strong>fsite<br />

Impacts. Proximity impacts include<br />

ambfent noise levels as well as human and<br />

pet intrusions. Off-site affects center<br />

on nutrfent and sediment loading resulting<br />

f rem wlnd and water transport mechanisms.<br />

Xf the extant parcel <strong>of</strong> wetland is zoned<br />

for resldentlal or some other intensive<br />

use, there are pressures for filling and<br />

development. Fl rep as a disturbance,<br />

seams to be limfted to the cattafl and<br />

sedge marshes <strong>of</strong> Dickinson Island<br />

(Jaworskl et a1, 1979).<br />

5.2 WETLAND OWNERSHIP AND<br />

MANAGEMENT<br />

Wetland ownarship Js in the hands <strong>of</strong><br />

individuals as we91 as <strong>St</strong>ate, provincial<br />

and Federal agencies. <strong>The</strong> management <strong>of</strong><br />

the wetlands by these groups follows<br />

df verse strategies. <strong>The</strong> current ownership<br />

<strong>of</strong> the coastal wetlands as well as the<br />

zoning and long-range plans wf 11 strong? y o 5 10 15 20<br />

affect the future use <strong>of</strong> these Slatule M~les<br />

envfronmenls. In thfs section, an attempt<br />

has been made to project the future land Figure 61. Location map <strong>of</strong> individual<br />

use <strong>of</strong> these areas, and by usinpa those coastal wetlands <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

p r ogect 1 ans address the issues <strong>of</strong> River-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ecosystem; character-<br />

managemnts restorationl and arti f i ci a1 istics and ownership <strong>of</strong> each wetland are<br />

construct don <strong>of</strong> wet1 ands, given in Appendix M.<br />

728


Figure 62. Ownership map <strong>of</strong> the Michigan portion <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta.<br />

are administered by the Michigan DNR.<br />

Approximately 40% <strong>of</strong> Dickinson Island is<br />

in private ownership, especially the<br />

northern and eastern shores as well as a<br />

block in the south-central area. In<br />

addition, the U,S. Army Corps <strong>of</strong> Engineers<br />

manages two disposal sites on Dickinson<br />

Is1 and along North Channel, and there are<br />

private bottom1 and patents along the<br />

distributary channels <strong>of</strong> both Dickinson<br />

and Harsens Islands. Although the<br />

Department <strong>of</strong> Natural Resources is slowing<br />

the residential and summer home<br />

development along these dfstributary<br />

channels, the land remains in private<br />

ownership. At present, the lower part <strong>of</strong><br />

Harsens Is1 and, and much <strong>of</strong> Dickinson<br />

Is1 and, and those unpatented wet1 ands and<br />

adjacent shall ow-water bottomlands would<br />

be considered part <strong>of</strong> Michigan's <strong>St</strong>. <strong>Clair</strong><br />

Flats <strong>St</strong>ate Game Area. Ownership <strong>of</strong> the<br />

small, undj ked wet1 and parcels scattered<br />

along Anchor Bay and along the westerns<br />

eastern, and southern shores sf <strong>Lake</strong> <strong>St</strong>.<br />

Cl air is generally private.<br />

At present, in the delta there are<br />

two basic management stradagjes: 1)<br />

natural and 2) managed for waterfowl<br />

hunting and perhaps breeding as well. <strong>The</strong>


pub1 icly owned* open wetland systems are<br />

natural responding to seasonal and 1 ong-<br />

term water level fluctuatlons. In<br />

comparf son, the diked wetl ands consf st<br />

largely <strong>of</strong> cattail marshes that are<br />

managed by means <strong>of</strong> electric pumps, flood<br />

gates and weirs. In several managed<br />

marshes, crops are grown, then flooded to<br />

facll itate waterfowl feedfng.<br />

No sf gnif fcant changes in the wetland<br />

management practices are antlclpated. It<br />

fs expected that the status quo w i l l<br />

generally continue. Further diking <strong>of</strong><br />

wetl ands is strong1 y d 1 scouraged because<br />

<strong>of</strong> the adverse impacts discussed earller,<br />

With regard to the management <strong>of</strong> the<br />

currently dlked areas, It i s recommended<br />

that waterfowl breedlng be encouraged In<br />

a1 1 <strong>of</strong> the contained marshlands by means<br />

<strong>of</strong> nesting platforms and areas. Moreover,<br />

to facilitate flsh use, perhaps the<br />

wetlands could be opened up during the<br />

spring spawning season or at least during<br />

those years when no water-level<br />

manipulat 1 on is necessary for waterfowl<br />

management. Creating artlf icial islands<br />

and allowing better-drained areas to<br />

revert to shrub or maadow communities may<br />

also add dlverslty.<br />

DFTRO/T RIVER<br />

---*.<br />

.,. - - ~.<br />

KILOMt lHf S<br />

Figure, 63. Location map <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

diked coastal wetlands (see text for<br />

explanation <strong>of</strong> numbers).<br />

Other practices, such as experimental<br />

marsh burning* have improved the waterfowl<br />

loafing population in Ontario's diked<br />

wetlands (Ball 1984). Fertflfration<br />

however, is not recomnended at this time.<br />

Wherever possible, hydrologic connectivity<br />

should be increased.<br />

<strong>St</strong>. Clalr Delta wetlands are diked: <strong>St</strong>.<br />

Johnrs Marsh (no. l)r lower Harsens Island<br />

(no. 2)s lower Walpole Island near Goose<br />

<strong>Lake</strong> (no. 31, lower <strong>St</strong>. Anne Island (no.<br />

4 ) ~ and Mitchell Bay-Thames Rtver mouth<br />

regfon (no. 5). Nearly one-ha1 f <strong>of</strong> the<br />

total delta wetlands, some 5,280 hectares,<br />

Dlcklnson xsland~ because <strong>of</strong> Its 'pen<br />

system nature and nt act ronment<br />

gradients. must be managed se~aratel~ as<br />

it may be the<br />

wetland in the Great <strong>Lake</strong>s region,<br />

Keepfng P t an open system, with only<br />

passive recreation permitted, w i l l Insure<br />

are separated from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> by<br />

earthen dfkes. Most <strong>of</strong> these djked<br />

environments occur .fn zones where cattai 1s<br />

natural 1 grow and el evat range<br />

between 175 and 176 m above Internatfonal<br />

<strong>Lake</strong>s datum*<br />

Its multiple functions for fish, Except for the wetlands in <strong>St</strong>, John's<br />

waterfowl, other wildlffe* and food chain Marsh and those along western <strong>Lake</strong> <strong>St</strong>.<br />

support, Once fil ledr the confined <strong>Clair</strong>, most <strong>of</strong> these dfked marshlands are<br />

disposal sitesrnay be used for wildlife, managed for waterfowl purposes. Dikjng<br />

but not for a theme park as suggested by and water-level manlpulatlon <strong>of</strong> wetlands<br />

some local residents, <strong>The</strong> provfslon <strong>of</strong> for the benefit <strong>of</strong> waterfowl hunters is a<br />

public utilfties, on either Dickinson or traditional practice in the Great <strong>Lake</strong>s<br />

Harsens fslands* would Jnitiate reglon among both private and public<br />

uncontrol 1 ed development pressures. shooting clubs. <strong>The</strong> earthen dfkes Impound<br />

As indfeated<br />

water insfde the marshlands during towwater<br />

perfods in the Great takes, but also<br />

in Figure 53# several large areas <strong>of</strong> the f~ncti~n to protect the marsh from erosion<br />

12.2<br />

A


and excessive high water during above-<br />

average lake level conditions, <strong>The</strong> <strong>St</strong>.<br />

<strong>Clair</strong> Flats <strong>St</strong>ate Game Area, Canada Club*<br />

and the <strong>St</strong>e, Anne Club marshes are among<br />

the venerable diked wetlands managed for<br />

waterfowl huntlng during the fall<br />

migration,<br />

<strong>St</strong>. Johnfs Marsh, whlch straddles<br />

Hjghway M-23 in Clay Township9 is included<br />

as a diked wetl and because the l i ne <strong>of</strong><br />

houses and access road along Bouvier Bay<br />

(<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. C1 air) hydrologically<br />

separate it from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> (Figure<br />

56). Approximately 217 hectares <strong>of</strong> dfked<br />

marshlands are sftuated west <strong>of</strong> hlghway M-<br />

299 and about 478 hectares lie east <strong>of</strong><br />

this roadway. Interior dikes and the<br />

roadbed with only one bridge (or culvert)<br />

further fragment this wetl and and preclude<br />

water exchange, Although there are five<br />

openfngs to Bouvier Bay, water exchange<br />

and fish access are limited.<br />

Much <strong>of</strong> lower Harsens Island is<br />

contafned within the <strong>St</strong>. <strong>Clair</strong> Flats <strong>St</strong>ate<br />

Game Area (1,085 hectares) and is managed<br />

by Michtgan DNR as a public waterfowl<br />

hunting facility. In the upper section,<br />

which consists <strong>of</strong> 515 hectares, crops such<br />

as corn are grown and flooded in fall to<br />

attract migratory waterfowl, <strong>The</strong> more<br />

lakeward portion <strong>of</strong> the game area is<br />

essential1 y a water-level control led<br />

cattail marsh which provides cover and<br />

some breeding habitat for the waterfowl.<br />

During the 1974-75 waterfowl hunt, 76% o f<br />

the harvest from the 60 blinds on the game<br />

area consisted <strong>of</strong> mallard along with<br />

lesser numbers <strong>of</strong> black duck, pintail,<br />

green-winged teal and American wigeon<br />

( Jaworski and Raphael 1978).<br />

In Ontario, diking i s commonly<br />

practiced (Table 38). As shown on Figure<br />

63, diked area no. 3 consists <strong>of</strong> three<br />

diked systems located on lower Walpole<br />

Island. <strong>The</strong> area <strong>of</strong> these dfked<br />

marsh1 ands totals 1,097 hectares.<br />

Cattails and submerged aquatfcs appear to<br />

dominate the vegetation. Some <strong>of</strong> the<br />

wetl ands <strong>of</strong> the northernmost d ?ked area<br />

are now being utilized for cultivated<br />

crops.<br />

Diked area no. 4 on Figure 63 refers<br />

to two dfked systems on lower <strong>St</strong>. Anne<br />

Island. <strong>The</strong> diked area located on the<br />

west side <strong>of</strong> Chenal Ecarte consists <strong>of</strong> 660<br />

hectares, whereas the dlked system on the<br />

east side is comprised <strong>of</strong> 415 hectares.<br />

Aerfal photographs indicate dense colonies<br />

<strong>of</strong> cattails in these managed wetlands,<br />

with few canals or other human<br />

disturbances. <strong>The</strong> most 1 akeward diked<br />

wetland on lower Walpole Island also has<br />

few canals across it. Pumping stations<br />

and flood gatas or welrs are employed to<br />

manage water levels here and at the other<br />

waterfowl shooting club marshes.<br />

Along eastern <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> many <strong>of</strong><br />

the coastal wetlands are diked. Diking is<br />

done by the local farmers who wish to<br />

cultivate as close to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> as<br />

possible. In addition, at Mitchell Point<br />

there is a managed wetland, and two more<br />

exist near the mouth <strong>of</strong> the Thames River.<br />

<strong>The</strong> areas <strong>of</strong> these three dfked systems<br />

are: 194 hectares, 393 hectares, and 500<br />

hectares. <strong>The</strong> latter two are designated<br />

as Area IV and Area 111, respective1 yr on<br />

the Tflbury (Ontario) Quadrangle.<br />

Moreoverr east <strong>of</strong> Area IV, there is a 156-<br />

hectare, square-shaped diked area. A<br />

total <strong>of</strong> 1,329 hectares were planimetered<br />

on current maps <strong>of</strong> this coast. Cattails<br />

appear to be the domfnant vegetation type,<br />

<strong>The</strong> adverse impacts <strong>of</strong> diking coastal<br />

wet1 ands are poor1 y understood. Never-<br />

theless, several adverse impacts include:<br />

1) loss <strong>of</strong> fishery function, 2) blockage<br />

<strong>of</strong> water exchange, 3) loss <strong>of</strong> food cha'ln<br />

support, 4) fragmentation and isolation <strong>of</strong><br />

diked wetlands, and 5) lower diversity and<br />

water quality <strong>of</strong> diked areas. Loss <strong>of</strong><br />

fish habitat and associated fishery<br />

functions is perhaps the principal adverse<br />

impact <strong>of</strong> coastal wetland diking. Diking<br />

has long been a source <strong>of</strong> conflict between<br />

waterfowl- and f ishery-resource managers.<br />

Clearly, most fish9 with the exceptton <strong>of</strong><br />

extremely tolerant species such as carp<br />

and bullheads, do not spawn in dfked<br />

wetl ands. Furthermore, l arval and juve-<br />

nile fish cannot easily disperse Into nar<br />

tolerate the f 1 uctuatlng water temper-<br />

atures and water qua1 Jty <strong>of</strong> these<br />

contained systems. <strong>The</strong> authors have<br />

noticed that predator fish tend to forage<br />

only fn those coastal wetlands where lake<br />

water masses f nvade the open wetlands,<br />

For example, on Dickfnson Island one may<br />

observe 'l argemouth bass and bl uegi 71 s on? y


Figure 6$1 i'ier ial jir~otoornijii <strong>of</strong> !)i~kin*<strong>of</strong>~ Xs'lana wctlancis, <strong>St</strong>, <strong>Clair</strong> DPI ta {May 19R0je<br />

Molr, df~+jn,%rl~ fr-rrn top IC,;I'.;TA! f1t2r~hf"~ :n ~?F~E>'I' Ray (rrsff ~irie!, Vpv.fh ch;?nne'l ic +t<br />

t8?c?i!rwl Fq iti tilf2 tv~tt~)rli ~f LOP pirotagraph,<br />

the ?ti$, Il'lfj~ll?~<br />

324


in cl rcul ating marsh waters, particularly<br />

near culverts* or where lake water masses<br />

prevai 1,<br />

Di k i ng prevents water exchange<br />

between the wetland and the adjacent water<br />

body. Thusp the waters <strong>of</strong> the canals and<br />

open-water areas <strong>of</strong> the contafned sites<br />

stagnate, cover with f f lm, and exhibit<br />

l arge diurnal temperature ranges (Mudroch<br />

and Capobianco 1978). Although the<br />

dissolved oxygen levels In the dlked<br />

wet1 ands remain above 75% satu rat1 on<br />

(Figure 22)r in response to atmospheric<br />

dfffusion and photosynthetic activityr<br />

excessive daytime water temperatures in<br />

summer reduce the dissolved oxygen<br />

concentrations to critical levels. During<br />

the cold winter months with abundant snow<br />

cover, oxygen depletion may occur under<br />

the ice cover, resulting in fish kf 11s and<br />

die-<strong>of</strong>f <strong>of</strong> invertebrates.<br />

zooplanktonr and ye1 low perch caught over<br />

macrophyte-free areas contained hfgher<br />

numbers <strong>of</strong> these invertebrates than did<br />

fish caught over macrophyte areas. <strong>The</strong><br />

paradox in these data may be explafned by<br />

marsh drafnage which 4s associated with<br />

large numbers <strong>of</strong> f I? ter-feeding zooplankton<br />

and yellow perch regardless <strong>of</strong><br />

the occurrence <strong>of</strong> vegetated substrates.<br />

Only two areas<br />

<strong>of</strong> coastal wetland are jdentifjed for<br />

restoration. First, the function and<br />

value <strong>of</strong> <strong>St</strong>. John's Marsh could be<br />

increased with 1 mproved hydro1 og ic<br />

connectivity (Figure 56). It fs<br />

recommended that the culverts connecting<br />

this wetland to Bouvier Bay be en1 arged to<br />

f aci 11 tate greater water mass exchange,<br />

Also, water exchange across M-29 could be<br />

Improved with the addltion <strong>of</strong> another<br />

larqe culvert close to Pearl Beach.<br />

~dditionally, the water quality near the<br />

definitive research has Perch Isle Point subdivision should be<br />

been conducted which the monitored9 and some <strong>of</strong> the fnterfor dikes<br />

exceptional fish production in <strong>Lake</strong> <strong>St</strong>. west <strong>of</strong> N-29 should be breached or<br />

<strong>Clair</strong> (Hatcher 1982) r it is be1 ieved that<br />

the wetlands are important in the food<br />

webs. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta wetlands are<br />

important sources <strong>of</strong> nutrients,<br />

particul ate organic matter, phytop? ankton,<br />

small zooplankton, and drift organisms,<br />

Zoopl ankters probably f $1 ter the preferred<br />

food items out <strong>of</strong> the marsh drainage.<br />

Figure 64 illustrates drainage from the<br />

delta wetlands. Leach (1980) found that<br />

the waters from the main channels <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> River are lower in nutrients<br />

than the waters derived from the<br />

tributaries on the Canadian side <strong>of</strong> the<br />

delta. Because Chenal Ecarte and the<br />

other Ontario tributaries Incorporate<br />

considerable flow from the adjacent<br />

wetlands, it is felt that these waters<br />

contain important quantities <strong>of</strong> nutrients,<br />

detrftus, and living matter for the food<br />

webs <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>.<br />

Poe ( 1983 attempted to demonstrate<br />

the relationship <strong>of</strong> the aquatic<br />

macrophytes and associated perlphyton to<br />

the fish production <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, In<br />

spite <strong>of</strong> some positive relatfonships<br />

establ ished i n some sampl l ng stationsp<br />

Poets data for Muscanoat Bay seemed<br />

contradictory. Sampling <strong>of</strong> Muscamoot Bay<br />

revealed that macrophyte-free areas had<br />

higher numbers <strong>of</strong> macrophytoplankton and<br />

removed.<br />

Second1 y the fragmented wetland<br />

complex south <strong>of</strong> the Cllnton Rfver Estuary<br />

should be restored (Figure 543, Perhaps<br />

this area should be comprehensively<br />

planned since a private developer has<br />

already attempted t o construct<br />

subdivisions in that area. <strong>The</strong> Detro<strong>St</strong><br />

District Corps <strong>of</strong> Engineers refers to this<br />

threatened wetland as Verchev9s Marsh. A<br />

comprehensivs plan which fncludes the<br />

funct-ion <strong>of</strong> the Black Creek* the impact <strong>of</strong><br />

the Metropol i tan Beach development# and<br />

connectivtty to <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> is<br />

suggested. Restoratfon <strong>of</strong> a wetland on<br />

this western side <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clajr could<br />

partially mitigate the historic wetland<br />

losses in this coastal stretch, Moreover*<br />

certaln fdsh such as northern pike and<br />

1 argemouth bass require spawning habitat<br />

ddstributed every 8 to 10 km along a coast<br />

because they are not Isng-dqstance<br />

migrants (Jaworski and Raphael 1978).<br />

No al-ti-fi-<br />

cjaf construction <strong>of</strong> wetlands is<br />

recommended at this time because the<br />

current wet1 ands appear adequate and <strong>Lake</strong><br />

<strong>St</strong>, <strong>Clair</strong> Js very productfve, If the<br />

current wetland resource base can be


conserved and/or Improved, then the important ecological cornmunlt~es, <strong>The</strong>se<br />

present ecology <strong>of</strong>: the lake may well be coastal wotlandst par-tfcularly the <strong>St</strong>.<br />

assured. ClaJr Delta wetlands, have had and should<br />

continue to have signSPIcant functions 3n<br />

perform<br />

va'uable functions for<br />

munfcf~alft'es* <strong>The</strong>y ~rovlde habjtats for<br />

desfred plants and animals, they filter<br />

pollutants and sedfments from storm<br />

the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> ecosystem. Conclusions<br />

fn thls sectlon are based an physical.<br />

blologfcal econamfc, and poi lt-jcal<br />

factors.<br />

drainage, and they protect coastal <strong>The</strong> rocent history <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>,<br />

structures from wave attack, On the other ~1 a and fs cha ratter zed by<br />

handr wetlands are <strong>of</strong>ten barriers to lake rel stabf 1 as wel 1 as h<br />

access and are susce~tibfe to function and value. Gjven the general<br />

filng* and which<br />

in turn lmpaf rs thef r natural functfons.<br />

degradatfon whlch has taken place the<br />

lower Great <strong>Lake</strong>s durfng the past several<br />

<strong>The</strong> fmportant functfons <strong>of</strong> wetlands the future prospscts for the<br />

are <strong>of</strong>ten dfsregarded by those wishing to<br />

are<br />

develop an area, <strong>St</strong>ate <strong>of</strong> Michfgan reasonably brfght* As coastal wetlands fn<br />

has recogn'lzed this and has forrnul ated<br />

adjacent lakes the re' st've<br />

pol cy to wetlands.<br />

~h~ 'Importance <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. Clalr wetlands<br />

Shore1 ands Protection and Management Act is increasfng* there has a<br />

<strong>of</strong> 1970. provfdss programs whlch include 4X' loss In the <strong>Lake</strong> <strong>St</strong>* wet'ands8<br />

the protection <strong>of</strong> wetlands in the Great<br />

and Of the extant wet'ands are dfkedr<br />

<strong>Lake</strong>s shorelands. At the local level* a<br />

special problsm facfng declslon-makers is<br />

the delta wetlands have not suffered<br />

degradation comparable to that <strong>of</strong> Green<br />

Saginaw Bay, and westerr1 <strong>Lake</strong> Erie.<br />

the tdentf f fcatjon and regulation <strong>of</strong><br />

adequate buffer areas surrounding the,<br />

desf gnated wetlands,<br />

Tha physfcal framework <strong>of</strong> <strong>Lake</strong> <strong>St</strong>,<br />

Cl alr wstl ands partla1 ly explafns the high<br />

In Mlchlgan, once "areas <strong>of</strong> concernw productf vl ty <strong>of</strong> these wet1 ands. Firstr<br />

have bean establ lshed~ the level <strong>of</strong> the total area <strong>of</strong> coastal wetlands Is<br />

control must be determined. Speclal use large In comparfson to the relatively<br />

psrmlts and 1 lsts <strong>of</strong> acceptable<br />

csnstruetton near wetlands provfde a<br />

small sfze <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr. Second,<br />

these deltafc wetlands are dlstrfbuted<br />

method <strong>of</strong> mlnlmizfng wetland dlsturbance over a large area <strong>of</strong> the northern part <strong>of</strong><br />

by permitting only those %ctlvltfes which the 1 ake, Thf rd, physical dlvorsity<br />

contrfbute least to change. Wet1 ands are fact 1 ltatss mu1 tfple subsystems. This<br />

unsuitable for most klndg <strong>of</strong> development. creates numerous aquatic habttat and<br />

D'isruptfve dredging and fdlf fng are <strong>of</strong>ten coastal wet1 and interfaces which are<br />

necessary to create a usable area for the located in remote areas and thus buffered<br />

constrirctfon af bulld.fngsr parkfng areas, fr~n cultural 4mpacts. In addition, the<br />

and access foads. On-sfte sanitary sewage <strong>St</strong>. Clalr Delta Is geologlcal1y stable.<br />

treatment is dffficu'ft because <strong>of</strong> poor Subsldenca Is not occurring and delta<br />

percoIat4on and the high water table.<br />

Recregtfori usas can be developed around<br />

extensi on or erosion have not been<br />

slgnificant over the last century.<br />

wetlands in a manner compatfble wfth<br />

natural systems, Camping# canoeing, and Qn the ecosystem level, as well as an<br />

nature tratls make use <strong>of</strong> the water a local habltat scalet the hydrology and<br />

recrezrtfonel value <strong>of</strong> wet1 ands. with such hydrolog ic connectlvl ty <strong>of</strong> the <strong>St</strong>, Clafr<br />

precautionsr construc9lan need not. cause Delta wetlands are most favorable.<br />

excessive damage, Re? atlvel y cool, hfghly-oxygenated, and<br />

5.3. PROSPECTS FOR THE FUTURE<br />

non-turbld waters enter from <strong>Lake</strong> Huron<br />

and pass through the deltaic wetlands,<br />

<strong>The</strong> df stributaries and crevasse channels<br />

(1 ocally celled "hfghwaysn) dl stribute<br />

this high-qua1 f ty tnput water throughout<br />

<strong>The</strong> freshwater wetlands along <strong>Lake</strong> the wetlands, For example, an abandoned<br />

<strong>St</strong>, CT a1 r w $1 1 canttnue to corratltuts dlstrftsutery channel permits water flow<br />

7%


across the entire length <strong>of</strong> Djckinson<br />

Island. Given a residence time <strong>of</strong> on1 y 7<br />

to 9 days in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r circulatlon<br />

is well established and little stagnation<br />

occurs. Water exchange between the lake<br />

and coastal wetlands is a1 so f aci 1 itated<br />

by seiches.<br />

<strong>The</strong> we? 1-known turbidity problems <strong>of</strong><br />

other coastal wetland systems? such as<br />

Green Bay and Saginaw Bay, are not a major<br />

adverse impact in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

wetlands. SiltatJon and turbidity are low<br />

in the deltaic wetl ands because the <strong>St</strong>.<br />

Cl ai r River has a very low suspended load<br />

and the bedload consists <strong>of</strong> fine sand.<br />

Some local turbidity occurs at the mouth<br />

<strong>of</strong> the Cl inton and Thames Rivers durlng<br />

spring run<strong>of</strong>f and flooding. Also, within<br />

<strong>Lake</strong> <strong>St</strong>. Clai r, resuspension <strong>of</strong> bottom<br />

sediments, as In Anchor Bay during storms,<br />

can discolor the water over 1 arge areas.<br />

Wetland erosion is not a problem<br />

along <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> due to the relatively<br />

short fetch and shallowness <strong>of</strong> the lake<br />

basin. Coastal erosion resulting from<br />

eustatically rising water levels and f ram<br />

long-term lake level fluctuations is not a<br />

significant problem as it is in <strong>Lake</strong> Erie,<br />

Because the deltaic wetlands are largely<br />

open systems that slope lakeward, lake<br />

level oscillations do not result in<br />

wetl and die-back or senescence ' as is true<br />

in many other coastal wet1 and systems.<br />

Rather, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> wetlands shift up<br />

or down the gradient, maintaining their<br />

productivity at various 1 ake levels.<br />

Dickinson Is1 and is an excellent example.<br />

Biological 1 y, these coastal wet1 ands<br />

exhibit relatively high habitat and<br />

species diversity, Productivity in <strong>St</strong>.<br />

<strong>Clair</strong> Delta wetlands is also high. Such<br />

conditfons manifest themselves in surplus<br />

waterfowl foods and Impressive flsh<br />

catches. Open wetlands systems, with<br />

complete envi ronmental gradients and<br />

numerous ecotones, exhibit "pulse"<br />

stability and high productivity. Although<br />

further research is needed to define the<br />

importance <strong>of</strong> the flux <strong>of</strong> particulate<br />

organic matterr drift organisms, and other<br />

organic foods from the wetlands into <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>r there i s an apparent<br />

association <strong>of</strong> waterfowl r game fish, and<br />

other wildlife with these wetland<br />

discharges. <strong>The</strong> high dissolved oxygen<br />

levels and high flushing rates,<br />

accompanied by relatively low contaminant<br />

and turbid1 ty levels* are positively<br />

correlated with the high productlvfty.<br />

From an economic vjewpoi nt these<br />

coastal wetlands exhibit high fish and<br />

wildlife values. Because these functions<br />

and values are linked to the ecology <strong>of</strong><br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, they cannot easil y be<br />

replaced or mitigated. Wfth regard to<br />

human uses, passive recreation and open<br />

space designation appear to be more<br />

suitable than intensf ve activl ties. Uses<br />

involving hydro1 ogic modification <strong>of</strong> the<br />

wetlands and permanent intrusions <strong>of</strong>ten<br />

create large and irreparable<br />

disturbances. It i s expected that urban<br />

development w i l l result in the filling or<br />

destruction <strong>of</strong> most coastal wetlands along<br />

western <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, northward to <strong>St</strong>.<br />

John's Marsh. Moreover, south <strong>of</strong> Mitchell<br />

Bay, agricultural encroachment may is01 ate<br />

all but the cattail marsh fringe. It is<br />

unlikely that the Clinton River mouth and<br />

the <strong>St</strong>. John's Marsh wetlands w i l l be<br />

restored. Neverthelessr because <strong>of</strong> the<br />

recognized value <strong>of</strong> the remaining<br />

wetlands, we believe that most <strong>of</strong> the<br />

extant wetlands w i l l be preserved in their<br />

present state.<br />

<strong>The</strong> future impacts on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

wetlands are dependent upon leg is1 ative<br />

efforts and public awareness in the Great<br />

<strong>Lake</strong>s basin <strong>of</strong> the value <strong>of</strong> the resource<br />

base. Legislatively, the <strong>St</strong>ate <strong>of</strong><br />

Michigan has been a leader i n<br />

environmental protection <strong>of</strong> wet1 ands. For<br />

example, the followfng laws have been<br />

enacted :<br />

--<br />

Public Act 247 <strong>of</strong> 1955, as amended*<br />

prohibits constructing or dredgi ng any<br />

artificial body <strong>of</strong> water that would<br />

ultimately connect with a Great <strong>Lake</strong>.<br />

This act also requires a permit from<br />

the Department <strong>of</strong> Natural Resources to<br />

f i 11 any submerged 1 ands in a Great<br />

<strong>Lake</strong>, including <strong>Lake</strong> <strong>St</strong>, Clalr.<br />

ment Act -- PublJc Act 245 <strong>of</strong> 1970t as<br />

amended* designates wetlands adjacent<br />

to a Great <strong>Lake</strong> or a connecting<br />

waterway such as the <strong>St</strong>. <strong>Clair</strong> River as<br />

envf ronnental areas necessary to<br />

preserve flsh and wildl $fee


connectivity Is good and the wetland<br />

diversity Is well displayed, Converse1 yr<br />

regulates wet1 ands through several many wetlandsp especially In Ontarfor have<br />

important laws relating to shorelands<br />

and submerged 1 ands.<br />

been diked and are less diverse s'lnce<br />

there Ss a dominance <strong>of</strong> cattail. It %s<br />

<strong>of</strong>ten pointed out that <strong>Lake</strong> <strong>St</strong>.,Cla%r is<br />

In Ontarfo, wetlands are viewed as located tn waterfowl flyways and is aw<br />

recreational land and are thus taxed as important staging area for migration, a<br />

such. Taxation <strong>of</strong> agricultural land is fact which gustlffdes dfking to some<br />

less than recreational 1 and. This degree, Howevers as noted earlier, the<br />

unbalanced tax structure ls an incent-fve economlc value <strong>of</strong> sport fishing greatly<br />

to drain and dlke wetlands In Ontario for<br />

agrlcul tural purposes (G. B. McCulloughr<br />

exceeds waterfowl hunting and trapping<br />

combined, More specifically, the economic<br />

Canadian Wildlife Service; pers. comm.1, value <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> to sport flsh'ing*<br />

Several wetlands north <strong>of</strong> the Thames River including coldwater species, has a value<br />

including huntlng clubs lands have <strong>of</strong> $59641 per wetland hectare/year<br />

succumbed to the tax pressure and have (Jaworski and Raphael 197131. To detach<br />

been converted to agriculture. the sportfish habltat from the wetland<br />

ecosystem may well decrease wetland<br />

It is most fortunate that large dlversity and the economic value <strong>of</strong> the<br />

tracts <strong>of</strong> wetlands are owned by <strong>St</strong>ate or <strong>St</strong>. Clalr Delta. Also the cost for dike<br />

Federal agencies. Assuming that the construction and maintenance in view <strong>of</strong><br />

governling agencies wf 11 preserve the decreasing diversity ought to be<br />

resource9 most <strong>of</strong> the Michigan portion <strong>of</strong><br />

the <strong>St</strong>, <strong>Clair</strong> Delta w j l l be protected from<br />

considered.<br />

future development. In Ontario the Coastai erosion is modest on the<br />

Wal pole Island Indian Reserve encompasses shore1 lnes <strong>of</strong> the laker and selches and<br />

Seaway, Squirrel, Wal pole and <strong>St</strong>. Anne flooding are beneficial flushing agents<br />

Islands. <strong>The</strong> preservation prospects for and nutrient transport mechanisms to the<br />

this portion <strong>of</strong> the modern delta are open-water bodies <strong>of</strong> the bays and lake.<br />

reasonably bright. D i k 1 ng preserves a rather monoiyp fc<br />

wetland9 prohibits adequate hydrologfc<br />

<strong>The</strong> MichZgan DNR has made a concerted connectlvityr and decreases wetland<br />

and sustained effort to inform the publlc utlllty In a low wave energy environment.<br />

<strong>of</strong> the value, in terms <strong>of</strong> economic Dikfng--to preserve wetlands in higher<br />

returns, <strong>of</strong> the <strong>St</strong>ate's wetlands. Through wave energy environments or in coastal<br />

funded wetland research and literatures areas dominated by fine clastic sediments<br />

the assets<strong>of</strong> the wetland resources have or where geological subsidence fs<br />

been dissemfnated to the public, occurring such as western <strong>Lake</strong> Erie-appears<br />

to have some justiflcation and<br />

In spite <strong>of</strong> legislative an3 public trade-<strong>of</strong>f value. In <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> the<br />

awareness efforts# there are still areas envf ronmental (i .eeP hydrological and<br />

sf concern. Both in Michigan and Ontario, geological 1 considerations ought to<br />

bul kheading and f i 11 i ng for small encourage wetlands preservatlon to be<br />

residential development continuer particu- dfrected t o optlmurn hydrologic<br />

larly along the natural levees <strong>of</strong> South connectivity between the resource and the<br />

and M-lddle channels and the shelf areas, open lake,<br />

In spite <strong>of</strong> Michfgan PA 247 and PA 24Sr<br />

dfked disposal sftes for polluted dredge In November 1984, a "Great <strong>Lake</strong>s<br />

spoil were const rlscted on the premodern Coastal Wet1 ands Col loqui um, " co-sponsored<br />

def tr? <strong>of</strong> Dickinson Island in the mid- by the Sea Grant Program and Envfronment<br />

f97Ds, Although Michigan@s efforts to Canada* was held on the campus <strong>of</strong> Michigan<br />

preserve these coas;tal wetlands have been <strong>St</strong>ate University to explore natural and<br />

posltfve, alteratton is continuing. manipulated water levels in Great <strong>Lake</strong>s<br />

wetlands, Resides pr~vidjng a forum for<br />

Dick;nson Island pr~vides an example the exchange <strong>of</strong> current research<br />

<strong>of</strong> a coastal wetland which has not been linformationr the rneetjng resulted In the<br />

signfficantly altered, <strong>The</strong> hydrological establ lshment <strong>of</strong> a network <strong>of</strong> wet1 and<br />

1 28


ecolog~sts and managers In the Great <strong>Lake</strong>s<br />

region and plans for the pub1 fcation <strong>of</strong> a<br />

quarterly newsletter for such workers,<br />

Although the conference p<strong>of</strong>nted out the<br />

need for more research djrected toward<br />

understanding processes and wild1 ife<br />

utllizatjon <strong>of</strong> the coastal marshes* an<br />

equally important consSderatlon was the<br />

alarming loss <strong>of</strong> wetlands. If there was a<br />

central theme <strong>of</strong> the conference* it would<br />

have to be that the conferees were in<br />

agreement that the existing coastal<br />

wetlands must be vigorously preserved if<br />

these valuable habitats are to survjve<br />

Intense development pressures.<br />

Gary B. McCullough <strong>of</strong> the Canadian<br />

Wild1 i fe Service documented the extensive<br />

conversion <strong>of</strong> coastal marshes to<br />

agricultural 1 ands along the Ontario shore<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> which has taken place in<br />

the past several decades. He estimated<br />

that 36% <strong>of</strong> the original wetlands have<br />

been lost. Much <strong>of</strong> the remaining wetland<br />

area south <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong> Delta is<br />

protected by dikes and water level control<br />

structures. John Ba7 1 <strong>of</strong> the University<br />

<strong>of</strong> Guelph reported that in these marshes,<br />

such as the ones owned by Ducks Unlimited<br />

or within the <strong>St</strong>. Clalr National Wildlife<br />

Area (Canada)# waterfowl utilization can<br />

be enhanced by clearing open areas in the<br />

cattail marshes (Ball 1984). To attract<br />

migrating ducks the optimal size <strong>of</strong> the<br />

opening appears to be approximately 50 m.<br />

<strong>The</strong> col 1 oqu ium concluded with the remarks<br />

<strong>of</strong> Harold H. Prince, underscoring the<br />

"signaturew which coastal wetlands impart<br />

to the fish and wildlife resources <strong>of</strong> the<br />

region. He pointed out that the<br />

significance <strong>of</strong> these marshes can be<br />

appreciated by recognizing: 1) that so<br />

many animals have embraced wet1 and<br />

habitats as part <strong>of</strong> their 1 ife strategy<br />

and 2) that wetlands are indicators <strong>of</strong><br />

llttoral and upland environmental quality.<br />

<strong>The</strong> ideas put forth at this conference are<br />

1 ikely to shape the direction <strong>of</strong> wetlands<br />

research in the Great <strong>Lake</strong>s region for the<br />

next decade.


Allen, H.L. 1971. Primary productivity,<br />

chemo-organotrophy, and nutritional<br />

interactions <strong>of</strong> epiphytic algae and<br />

bacteria on macrophytes in the<br />

1 ittoral <strong>of</strong> a 1 ake. Ecol . Monogr.<br />

41(2): 97-127.<br />

Andrews, R. 1952. A study <strong>of</strong> waterfowl<br />

nesting on a <strong>Lake</strong> Erie marsh. M.S.<br />

<strong>The</strong>sis. Ohio <strong>St</strong>ate Univ., Columbus.<br />

85 PP.<br />

Assel, R.A., F.H. Quinn, G.A. Leshkevich,<br />

and S. J. Bolsenga. 1983. Great <strong>Lake</strong>s<br />

ice atlas. U.S. Dept. Commerce, Great<br />

<strong>Lake</strong>s Envir. Res. Lab. Ann Arbor, MI.<br />

115 pp,<br />

Ayres, J.C. 1964. Currents and related<br />

problems at Metropol itan Beach, <strong>Lake</strong><br />

<strong>St</strong>. <strong>Clair</strong>. Great <strong>Lake</strong>s Res. Div.<br />

Spec. Rep. No. 20, Univ. Michigan,<br />

Ann Arbor. ,55 pp.<br />

Bajorunasr S.J. 1968. River ice jams; a<br />

literature revlew. U.S. Army Corps <strong>of</strong><br />

Engineers. Res. Rep. 5-5, Detroit*<br />

MI. 570 pp.<br />

Baumgartner, L . L. 1942, Ecological survey<br />

<strong>of</strong> muskrat populations and habitats.<br />

Mich. Dep. Nat. Resour., Proj. Rep.<br />

14-R. 128 pp.<br />

Bayly, I. 1975. Preliminary study <strong>of</strong> the<br />

nutrient regime <strong>of</strong> marshland <strong>of</strong> the<br />

<strong>St</strong>. <strong>Clair</strong> National Wildlife Area -<br />

Dover Township, Ontario. Rep. to<br />

Can. Wildl. Serv., London, Ontario,<br />

27 PP*<br />

Bednarik, K.E. 1956. <strong>The</strong> muskrat in Ohio<br />

<strong>Lake</strong> Erie marshes. Ohio Dep. Nat.<br />

Resour., Ohio Div. Wildl. 67 pp.<br />

Bednar i kr K. E. 1975. Environmental impact<br />

assessment report, Magee Marsh<br />

Wildlife Area dike construction. Ohio<br />

Dep. Nat. Resour,, Crane Creek Wildl.<br />

Exp. <strong>St</strong>a., Oak Harbor. 13 pp.<br />

Behler, J.L. and F.W. King. 1979. <strong>The</strong><br />

Audubon Society field gufde to North<br />

American reptiles and amphibians.<br />

Alfred A. Knopf, New York. 743 pp.<br />

Bellrose, F.C. 1968. Waterfowl migration<br />

Baker, DeG.r and J.G. <strong>St</strong>rub, Jr. 1965. corridors east <strong>of</strong> the Rocky Mountains<br />

Climate <strong>of</strong> Minnesota. Part 111:<br />

In the United <strong>St</strong>ates. Illinois Nat.<br />

temperature and its appl ication. History Survey Biol, Notes. No. 61.<br />

Unfv. Minnesota, Agr. Expr. <strong>St</strong>a.<br />

Tech. Bull. 254. 43 pp.<br />

24 PP-<br />

Ball, J.P. 1984. Habitat selection and Bellrose, F.C* 1976- Ducks* geese* and<br />

optimal foraging by mallards; a field swans <strong>of</strong> North America* 2nd ed.<br />

experfment. M.S. <strong>The</strong>sfs. Unfv. <strong>St</strong>ockpol e Books* Harrf sburg, PA. 544<br />

Guelphr Guelph, Ontario. 44 pp.<br />

PP.<br />

Bates, R.L., and J.A. Jackson, eds. 1980.<br />

Glossary <strong>of</strong> geology, 2nd ed, Am.<br />

Berry* E.G. 1943, <strong>The</strong> Amnicalfdae <strong>of</strong><br />

Mfchigan: d4stributfonr wology and<br />

Geol. Instit.# Falls Church,<br />

PP*<br />

VA. 749 taxonomy. Unfv. Michigan, Mus. Zool.<br />

Misc. Publ. 57. 68 pp.


Mirge, E.A. 2894, A report on a collectfun Churchlllr ESP., Jr. and S.1. l-ow"is, 1924,<br />

<strong>of</strong> Cladocera, mostly from take <strong>St</strong>.<br />

Gfafrr Mtchfgan, Pagss 43-47 fn J. E.<br />

Food and feeding In fresh-water<br />

mussels. Bull, U.S. Bur. FSsh.<br />

Welghard, ed., A bfologScal 39:439-471.<br />

examfnatlan a% take <strong>St</strong>. Clalr. Bull,<br />

Mfch. flsh Corn. No, 4. Clarke, A.M. 1901. Tho freshwater molluscs<br />

Boreman# J., ed. 1976, Gruat <strong>Lake</strong>s ffsh<br />

egg and 1 arv&a tdantiffcat'lon: Proc,<br />

<strong>of</strong> a workshap, U.S. Ffsh Wlldl. Serv.<br />

FWS/W5-76/B. 220 pp.<br />

<strong>of</strong> Canada. tdatfonal Museums <strong>of</strong><br />

Canada, Ottawa. 446 pp.<br />

Cola. G.A. 1983. Textbook <strong>of</strong> 1 imnologyt<br />

3rd ad. @. V. Mosby Co., <strong>St</strong>. Louis.<br />

B~uidsn~ R.S. r ed. 1975, CanadaiOntarlo<br />

Great <strong>Lake</strong>s shore damage survey,<br />

Tech. Rap. Envlr, Canada and Ontarto<br />

MOn, Nat. Resaur, Toronta, 97 pp,<br />

Brant, R,A, and C,E, Herdandorf. 2971.<br />

Delfnlatfan <strong>of</strong> Great takas sstuartss.<br />

Pagss 710-718 & Proc. 15th Conf.<br />

Graat <strong>Lake</strong>s Wos,r Xnt. A$soc. Great<br />

<strong>Lake</strong>s Ros,<br />

Bray, 3.8, l962. EstimsLori <strong>of</strong> energy<br />

budgets for a (sattnlll marsh,<br />

Scfence 1368 1119-1120*<br />

Brlcksri K.S.* F.4, Glrfcker, and J.E.<br />

Gannon, 1976, Distrlbutlon and<br />

abundance aF zwplanktan dn the U.S.<br />

watars <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>r 1973. J,<br />

Great lakes Bas, 2( 2 1 :256-271,<br />

Brock, 1",C). 1970, Phetasynthesfs by elgal<br />

spfphytes af in Ever-<br />

gladas Netlonal Perk. Bull. Mar, 5ci.<br />

26~952-956.<br />

Brawn, C,t, X9B3, Producttvlty <strong>of</strong> iiquatlc<br />

mircrophyter and ask~oclated par'fphyton<br />

In Loke <strong>St</strong>, Clalrt a data report.<br />

U,S, Ftsh Wtldl . Sorv,, Grsat Lakas<br />

Ffsh. Lab,* knn Arbor, MI 41 po.<br />

Butzsr* K,W, 1972. Recent hfstory sf an<br />

Ethlopfan dalta. Unfvsrslty <strong>of</strong><br />

CRl~ago Press, Chicago. 184 pp,<br />

401 pp.<br />

Colie, L,J. 1903, <strong>The</strong> delta <strong>of</strong> thu <strong>St</strong>.<br />

Clais Rivsr. Gaologfcal Surv. Mlch.<br />

19 : 1-28<br />

Coleman* J .M, 1976, Deltas: processes <strong>of</strong><br />

dapositlon and models for axplora-<br />

tfon. Contlnulng Education Publ . Co.<br />

1nc.r Champatgn, It. 102 pp.<br />

Comntt R. 1975. A Fleld gulda to reptfles<br />

and amphfbfans <strong>of</strong> aastsrn and central<br />

North Amurjca. Noughton MSFflln Co.#<br />

Boston. 429 pg.<br />

Cooper, C,L. I W, C, Partholomewr C. E.<br />

Eiordendorf, J. N. Reuttar, and F. C.<br />

Snyder. 19Bla. Limnetlc larval ffsh<br />

<strong>of</strong> the Maumas and Sandusky River<br />

estuaries. J. Great <strong>Lake</strong>s Rcs.<br />

7(11:51-54.<br />

Cooper, C.1 ,, M,R, Henfkonr and C,F,<br />

Werdsndorf. 1981b. Llmnatlc larval<br />

fish In the Ohlo portlan <strong>of</strong> the<br />

western basln <strong>of</strong> taka Erie, 1995-<br />

1976. J. Gruat Lakas Rss. 7(3) :326-<br />

329,<br />

Cooper, Get. J .3, Mfzora, and C.E.<br />

H~rdendorf. 1984. Distribut +on,<br />

abundance and antra J nmtrnt <strong>of</strong> larval<br />

ffrhes In the western and csntral<br />

basins <strong>of</strong> <strong>Lake</strong> Erfe. U.S. Envfron.<br />

Prot. Agency, Project Summary EPA-<br />

690/53-84-017. 3 pp.<br />

Marlan, 1975, Bistfibutlern <strong>of</strong> statas<br />

and count fes af waterfowl spe~cles Corer F.C. 1948, <strong>The</strong> flora <strong>of</strong> the Erls<br />

harvested during 1961-1978 huntfng islands, an annotated llst <strong>of</strong><br />

sealon, U,S, Ffsh Wild3. S~arv,, Spec, vdsrular plants. Ohio <strong>St</strong>tte Univ,*<br />

ScF, Rep, 187, Wash-Sogtonr D, C, 132 Franz <strong>The</strong>odore <strong>St</strong>one Lab. Gontrib. 9.<br />

PP 106 PP*


Cowardln* L.M., V. Carter, F.C. Golet, and<br />

E.T. La Roe. 1979. Classification <strong>of</strong><br />

wetlands and deepwater habitats <strong>of</strong><br />

the United <strong>St</strong>ates. U.S. Fish Wildl.<br />

Serv. FWS/OBS-79/31, Washington, D.C.<br />

103 pp.<br />

Crowderr L.B., and W.E. Cooper, 1979.<br />

<strong>St</strong>ructural complexity and f i sh-prey<br />

interactions in ponds: a point <strong>of</strong><br />

view. Pages 2-10 D.L. Johnson and<br />

R,A. <strong>St</strong>ein, eds. Response <strong>of</strong> fish to<br />

habitat structure in standing water.<br />

N. Central Div. Am. Fish, Soc., Spec.<br />

Pulb. No. 6, Bethesda, Md.<br />

Dawson? S.A. 1975. Waterfowl food<br />

production and util ization i n Anchor<br />

Bay9 <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Michigan. M.S.<br />

<strong>The</strong>sis* Univ. Michiganr Ann Arbor.<br />

124 pp.<br />

Dennis, D.G., and R.E. Chandler. 1974.<br />

Waterfowl use <strong>of</strong> the Ontario<br />

shoreline <strong>of</strong> the southern Great <strong>Lake</strong>s<br />

during migration. Pages 55-65 H.<br />

Boyd* ed.? Canadian Wildlife Service<br />

Waterfowl <strong>St</strong>udies in Eastern Canada.<br />

Can. Wildl. Serv. Rep. Ser.29,<br />

Ottawa, Ontario.<br />

Dennis, D.G., G.B. McCullought N.R. North,<br />

and R.K. Ross. 1981. An update<br />

assessment <strong>of</strong> migrant waterfowl use<br />

<strong>of</strong> the Ontario shorelines <strong>of</strong> the<br />

southern Great <strong>Lake</strong>s. Can. W i 1 d 1 .<br />

Serv. Rep. Ser. (manuscript rep. 1<br />

Dennis, D.G.? and N.R. North. 1981.<br />

Waterfowl use <strong>of</strong> the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

marshes during migration in 1976-77<br />

versus 1968-69. Can. Wildl. Serv.<br />

Rep. Ser. (manuscript rep.)<br />

Derecki, J.A. 1982. Effect <strong>of</strong> channel<br />

changes in the <strong>St</strong>. <strong>Clair</strong> River since<br />

1900. U. S. Dep. Commerce Great <strong>Lake</strong>s<br />

Envir, Res. Lab Rep. ERL GLERL-40#<br />

Ann Arbor, MI 20 pp.<br />

Derecki, J.A. 1983. Travel times in the<br />

Great <strong>Lake</strong>s connecting channels. U.S.<br />

Dep. Commerce Great <strong>Lake</strong>s Envir, Res.<br />

Lab Rep, No. 370, Ann Arbor, MI 12<br />

PP*<br />

Dolep S,A., ed. 1975. Great <strong>Lake</strong>s basin<br />

framework study? Appendix 17:<br />

Wildlife. Great <strong>Lake</strong>s Basin C~rnrn.~<br />

Ann Arbor? MI 140 pp.<br />

Doneldson, T. 1970. <strong>The</strong> public domain: its<br />

history with statistics. Johnson<br />

Reprint Corp., New York. 1,343 pp.<br />

Donohoe, R. 1966. Muskrat reproducti on in<br />

areas <strong>of</strong> control led and uncontrol 1 ed<br />

water level units. J. Wildl. Manage.<br />

30(2) :320-326.<br />

Dorrr J.A., and D.F. Eschman. 1970.<br />

Geology <strong>of</strong> Michigan. University <strong>of</strong><br />

Mich. Press, Ann Arbor. 476 pp.<br />

Duane, D.B. 1967. Characteristics <strong>of</strong> the<br />

sediment load in the <strong>St</strong>. C:air River.<br />

Pages 115-132 In Proc. 10th Conf.<br />

Great <strong>Lake</strong>s Res., Int. Assoc. Great<br />

<strong>Lake</strong>s Res.<br />

Edmondson, W .T. ed, 1959, Ward and<br />

Whipple's fresh-water bio10gyr 2nd<br />

ed. John Wiley and Sons, Inc.,<br />

York. 1~248 pp.<br />

New<br />

Eichenlaub, V,L. 1979. Weather and climate<br />

<strong>of</strong> the Great <strong>Lake</strong>s regfon. Unfversity<br />

<strong>of</strong> Notre Dame Press# South Bend, IN.<br />

335 pp.<br />

Esrhemeyer, P.H. 1950. Life history <strong>of</strong> the<br />

walleye, m n vltreum Yltreum<br />

(Mitchell), in Michigan. Mich. Dep.<br />

Conserv. Bull. Inst. Fish Res. No. 3.<br />

99 PP*<br />

Federal Water Qua1 i ty Administration.<br />

1970. Investfgation <strong>of</strong> mercury in the<br />

<strong>St</strong>. <strong>Clair</strong> River-<strong>Lake</strong> Erie system,<br />

U.S. Dsp. Interior, FWQA, Chlcago.<br />

108 pp.<br />

Ferguson, R.G., and A.J. Derksen. 1971,<br />

Migrations <strong>of</strong> adult and juvenile<br />

walleyes ( Yif;r<br />

-1 In southern <strong>Lake</strong> Huron, Laue<br />

<strong>St</strong>. <strong>Clair</strong>r <strong>Lake</strong> Erie and connecting<br />

waters. J, Fish. Res. Bd. Canada.<br />

28: 1133-1142,


Fuller, S.L. 1974. Clams and mussels<br />

(Mol1usca:Bivalvla). Pages 215-273<br />

C.W. Hart, Jr. and S.L. Fuller* eds.<br />

Pollution ecology <strong>of</strong> freshwater<br />

invertebrates. Academic Press# New<br />

York. 389 pp.<br />

Gilbert, G.K. 1890. <strong>Lake</strong> Bonnevllle. U.S.<br />

Geol. Surv. Monog. No. 1, Washington,<br />

D.C. 438 pp.<br />

Great <strong>Lake</strong>s Basin Commiss$on 1975, Shore<br />

use and erosfon. Appendix 12 Great<br />

<strong>Lake</strong>s Basan Framework <strong>St</strong>udy. Great<br />

<strong>Lake</strong>s Basin Corn., Ann Arbor, MI.<br />

111 pp. + Appendix.<br />

Great <strong>Lake</strong>s Commission. 1984. Ice<br />

conditions hamper start <strong>of</strong> the 1984<br />

inter1 ake commerce. Great <strong>Lake</strong>s<br />

Newsletter, Vol . 24(6) 23-4.<br />

Greeson, P.E.r J.R. Clark, and J.E. Clark,<br />

Glass, N.R. 1971. Computer analysis <strong>of</strong> eds. 1979. Wetland functions and<br />

predation energetics in the values the state <strong>of</strong> understanding.<br />

largemouth bass. Pages 325-363 JJ Proc. National <strong>Wetlands</strong> Symposium.<br />

B.C. Patten, ed. Systems analysis and Tech, Publ. No. TPS79-2. Am. Water<br />

simul ati on ecology, Vol . 1. Academic Resour. Assoc. Minneapol is, MN. 674<br />

Press, New York. PP e<br />

Goodnight, C.J., and L.S. Whitley. 1960. Haas, R.C. 1978. <strong>The</strong> muskellunge in <strong>Lake</strong><br />

Oligochaetes as indicators <strong>of</strong> <strong>St</strong>. <strong>Clair</strong>. Am, Fish, Soc. Spec. Publ.<br />

pollution. Pages 139-142 In Proc. 11. pp 334-339.<br />

15th Industrial Waste Conf.. Purdue<br />

Univ. , West Lafayette, IN Harman, W.N. 1974. S n a i l s<br />

(Moll usca:Gastropoda) . Pages 275-312<br />

Goodrich, C.r and H. Van der Schalie. a C.W. Hart, Jr., and S.L. Fuller,<br />

1932. <strong>The</strong> naiad species <strong>of</strong> the Great<br />

eds. Pollution ecology <strong>of</strong> freshwater<br />

<strong>Lake</strong>s. Univ. Michigan, Mus. Zool. invertebrates. Academic Press, New<br />

Occas. Pap. 238. 14 pp.<br />

York. 389 pp.<br />

Goodwin, C.E. 1982. A bird-flnding guide<br />

to Ontario. University <strong>of</strong> Toronto<br />

Press, Toronto, Ontario. 248 pp.<br />

Goodyear* C.D.r T.A. Edsallr DIM. Dempsey,<br />

G.D. Moss, and P.E. Polanskf. 1982a.<br />

Atlas <strong>of</strong> the spawning and nursery<br />

areas <strong>of</strong> Great <strong>Lake</strong>s fishes. Vol. VI:<br />

<strong>St</strong>. <strong>Clair</strong> River. U.S. Fish Wildl.<br />

Serv,, FWS/O%S-82/52rAnn Arbor, MI.<br />

18 PP*<br />

Goodyear, C.D., T.A. Edsallr D.M. Dempsey,<br />

G.D. Moss, and P.E. Polanskl. 1982b,<br />

Atlas <strong>of</strong> the spawning and nursery<br />

areas <strong>of</strong> Great <strong>Lake</strong>s fishes. Vol VII:<br />

<strong>St</strong>. <strong>Clair</strong> <strong>Lake</strong>. U.S. Fish Wildl,<br />

Serv., F#S/OBS-82/52. Ann Arbor, MI.<br />

35 PP.<br />

Hatcher, C.O. 1982. <strong>The</strong> structure and<br />

productivity <strong>of</strong> fish communities in<br />

the <strong>St</strong>. <strong>Clair</strong> - Detroit River<br />

Ecosystem, as effected by shore1 i ne<br />

and lakebed modifications. U.S. Fish<br />

Wild. Serv.8 Great <strong>Lake</strong>s Fish. Lab.<br />

Res. Comp. Rep. 51 pp.<br />

Hatcher, C.O., and R.T. Nester. 1983.<br />

Distribution and abundance <strong>of</strong> fish<br />

larvae in the <strong>St</strong>. <strong>Clair</strong> and Detroit<br />

Rlvers. U.S. Fish Wildl, Serv.8 Great<br />

<strong>Lake</strong>s Fishery Lab., Ann Arbor, MI,<br />

Admln. Rep. 83-5. 41 pp.<br />

Hayes, B.N. 1964. An ecological study <strong>of</strong> a<br />

wet prai rle on Harsens Island,<br />

Michigan. Mlch. Bot. 3(3) :71-83.<br />

Goodyear, C.D.* T.A. Edsall, D.M. Dempsey, Helfman, G.S. 1979. Fish attraction to<br />

G,D. Moss, and P.E. Polanski. 1982~.<br />

Atlas <strong>of</strong> the spawning and nursery<br />

floating objects in lakes. Pages 49-<br />

57 In D.L. Johnson and R.A. <strong>St</strong>ein,<br />

areas sf Great <strong>Lake</strong>s fishes, Vo1. eds. Response <strong>of</strong> fish to habftat<br />

VIII: Detroit River. U.S. Fish Wildl. structure in standing water. N.<br />

Serv., FWS/BS-82/52,<br />

33 PP*<br />

Ann Arbor, MI. Central Div. Am. Fish. SOC. Spec.<br />

Publ. No. 6. Bethesda. MD.


Herdendorf, C.E. 1970. take Erle physlcal Wildl. Serv.8 Great <strong>Lake</strong>s Fish. Lab.,<br />

1 4mnol ogy cruise, midsummer 1967. Ann Arbor, Mich., Admin. Rep. 80-2.<br />

Ohfo Dep. Nat, Resour., Div. Geol. 87 PP=<br />

Surv. Rep. Invest. 79. 77 pp.<br />

Herdendorf, C.E. 1982. Large lakes <strong>of</strong> the<br />

world. J. Great <strong>Lake</strong>s Res. 8(3):379-<br />

412.<br />

Herdendorf, C.E.8 S.M. Hartley, and M.D.<br />

Barnes, eds. 1981a. Fish and wildlife<br />

resources <strong>of</strong> the Great <strong>Lake</strong>s coastal<br />

wetlands within the United <strong>St</strong>ates.<br />

Vol. I: Overview. U,S. Fish Wildl.<br />

Serv., Washington, D. C. FWS/OBS-<br />

81/02-vl. 469 pp,<br />

Herdendorf, CdEer and S.E. Herdendorf.<br />

1983. Flora and fauna <strong>of</strong> the islands<br />

region <strong>of</strong> western <strong>Lake</strong> Erie. Oh1 o<br />

<strong>St</strong>ate Unlv.. Cent, <strong>Lake</strong> Erie Area<br />

Res. Tech. Rep, 285. 190 pp.<br />

Hiltunen, 3.K. 1971. Limnological data<br />

from <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, 1963 and 1965.<br />

Natl. Mar, Fish, Serv.8 Data Rep. 54.<br />

45 PP*<br />

Hunt, G.S. 1962. Water pollution and the<br />

ecology <strong>of</strong> some aquatic invertebrates<br />

in the lower Detroit River. Pages 29-<br />

49 In Proc. 5th Conf. Great <strong>Lake</strong>s<br />

Res. Div., Univ. Michigan, Ann Arbor.<br />

International 301 n t Commf sslon ( IJC) ,<br />

1981. Report on Great <strong>Lake</strong>s water<br />

quality for 1981. IJC, Water Qua1 i ty<br />

Board* Windsor, Ontario. 74 pp.<br />

Herdendorf, C.E.8 S.M. Hartley* and M.D.<br />

Barnes* eds. 1981b. Fish and wild1 ife<br />

resources <strong>of</strong> the Great <strong>Lake</strong>s coastal<br />

wetlands within the United <strong>St</strong>ates.<br />

Vol. 3: Overview. U.S. Fish Wildl.<br />

Jaworsk f , E. 1981. <strong>The</strong> economics <strong>of</strong><br />

wetland protection. Pages 1-10 In A.<br />

Champagne* ed. Proc. Ontario Wet1 ands<br />

Conf. Ryerson Polytech. Institute,<br />

Toronto, Ontario. Sept. 1981.<br />

Serv.? Washington* D.C.<br />

81/02-v3. 505 pp.<br />

FWS/OBS-<br />

Jaworski* E. * J .R. McDonald* S. McDona1 dr<br />

and C.N. Raphael. 1977. General<br />

Herdendorf, C.E., S.M. Hartley* and M.D. functions and values <strong>of</strong> freshwater<br />

Barnes? eds. 1981c. Fish and wildlife wetlands in the glaciated midwest.<br />

resources <strong>of</strong> the Great <strong>Lake</strong>s coastal Michigan <strong>St</strong>ate Univ., Kellogg<br />

wetlands within the United <strong>St</strong>ates, Biological <strong>St</strong>ation. 69 pp.<br />

Vol. 4: <strong>Lake</strong> Huron. U.S. Fish Wildl.<br />

Serv.<br />

811'02-v4.<br />

Washington* D.C.<br />

792 pp,<br />

FWS/OBS-<br />

Jaworski, E., and C.N. Raphael. 1973. A<br />

morphological comparison <strong>of</strong> the <strong>St</strong>.<br />

<strong>Clair</strong> and Mississippi deltas. Proc.<br />

Assoc. Am. Geograp. 5(1):121-126.<br />

Jaworski, E., and C.N. Raphael. 1976.<br />

Modification <strong>of</strong> coastal wetlands <strong>of</strong><br />

southeastern Michigan and management<br />

a1 ternatives. Mich. Acad, 8(3) :303-<br />

317.<br />

Jaworski* E.8 and C,N. Raphael. 1978,<br />

Fish, wildlife, and recreational<br />

Hiltunen, J.K. 1980. Compositlonr<br />

values <strong>of</strong> Michigan's coastal<br />

distribution* and denslty <strong>of</strong> benthos<br />

wetlands. U.S. Fish Wildl. Serv..<br />

in the lower <strong>St</strong>. <strong>Clair</strong> River, 1976-<br />

Twin Cities* MN. 209 pp.<br />

. .<br />

1977. U.S. Fish Wildl. Serv., Great<br />

<strong>Lake</strong>s Fish. Lab., Ann Arbor. Mich.~ Jaworski, E,p eN ~ ~ ~ p. h J . ~ ~ 1<br />

Admin. Rep. 80-4. 27 pp.<br />

Mansfield, and B.B. Williamson, 1979,<br />

Hiltunen* J.K., and B.A. Manny, 1982.<br />

Impact <strong>of</strong> Great <strong>Lake</strong>s water level<br />

fluctuations on coastal weti ands.<br />

Distribution and abundance <strong>of</strong> ProJect No. 5-045-MICH. Inst. Water<br />

macrozoobenthos in the Detroit River Res.8 Michigan <strong>St</strong>ate Univ., East<br />

and <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>* 1977. U.S. FOsh Lansing. 351 pp.


Jaworsk i E., G.N.<br />

Mansf l%ldr and B.B.<br />

Raphael r P,J.<br />

W l l l iamson, 1981,<br />

Korkfgianl I.I.9. 1963, Channel changes fn<br />

the <strong>St</strong>. Cfafr RIver slncs 1933. J.<br />

Impact <strong>of</strong> Great <strong>Lake</strong>s water level<br />

f 1 uctuat tons on coastal wstl ands.<br />

Pages 106-300 In C.E, Herdendorf,<br />

S.M. Hartley, and M,D. Barnes, eds.<br />

Waterways and Harbor Dlvi sf onr Proc,<br />

her. Soc, Cfvii Eng. 89tMay)rl-14.<br />

Krecker, F.W. 1939. A comparative study <strong>of</strong><br />

Ffsh and wdldlife resources <strong>of</strong> the the anfmal populatfon <strong>of</strong> certa'ln<br />

Great <strong>Lake</strong>s coastal wetlands w4thfn submerged aquatic plants. <strong>Ecology</strong><br />

the Unlted <strong>St</strong>ates. Volume 1:<br />

Ovsrvfew. U.S. Flsh Wlldl. Ssrv.,<br />

20(4) :553-562,<br />

Washington* D.C. FWS/08S-B1/02-~1. La R~cqusr A. 1966. Pleistocene Mollusca<br />

Jenks, W.L. 1912. History <strong>of</strong> <strong>St</strong>, Clafr<br />

<strong>of</strong> Ohlo. Ohio Dtv. Geol. Surv. Bull,<br />

62, Pt. 1. 111 pp.<br />

County, Lewis Publfshing Co.*<br />

Chlc&ga. 2 vols. 904 pp. <strong>Lake</strong> <strong>St</strong>. Clai r Advisory Committee, 1975,<br />

Jennlngs, H.S. 1894. A llst <strong>of</strong> the<br />

Fishlng chart and gulde, <strong>Lake</strong> <strong>St</strong>,<br />

Claf rr <strong>St</strong>, <strong>Clair</strong> RIver, and upper<br />

Rotatorla <strong>of</strong> the Eraat <strong>Lake</strong>s and some Detr<strong>of</strong>t River. <strong>St</strong>. <strong>Clair</strong> Consultants,<br />

<strong>of</strong> the Inland lakes <strong>of</strong> Mlchlgan.<br />

Bull. Mlch. Flsh Corn. NO. 3. 34 pp.<br />

Royal Oak, MI. 12 pp.<br />

Johnsonr 0.1, 1984, Preference <strong>of</strong> fish<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> Advfsory Committee, 1983,<br />

Ftshing chart and guide, <strong>Lake</strong> <strong>St</strong>,<br />

larvae for habftat types in a C1airr <strong>St</strong>, Clafr River" and upper<br />

contrallsd Laks Erfs wetland. Ohfo<br />

<strong>St</strong>ate Univ,, Water Res. Center,<br />

Detro4t Rfver. Revfsed ad. Mt.<br />

Clamens. MI. II pp.<br />

Columbus. 22 pp.<br />

Larsen, J.S,, ed. 1973. A gulda to<br />

Johnston, D,A, 1977. Popul atfon dynaml~s important characteristfcs and values<br />

<strong>of</strong> walleye [ XI32 <strong>of</strong> freshwater wetlands 3n the<br />

CBU~) and yellow perch ( f 2 ~ ~ ~ 3 northeast. Water Resour. Ras. Center<br />

~~~YBSEBBZ) in <strong>Lake</strong> <strong>St</strong>, Clalr, Pub. 31., Unfv. <strong>of</strong> Massachusettst<br />

especf a1 1y durfng 1970-1976. J, Fish hh~rst. 35 pp.<br />

Res. Board Can, 34~1868-1877.<br />

Leach# J.H, 1972. Distribution <strong>of</strong><br />

Jones, J,J, 1982, Potential effects <strong>of</strong> chlorophyll g and related variables<br />

ulntsr shiplng on divlng ducks<br />

wintarlng on the Dstrolt Rlver. M.S.<br />

<strong>The</strong>sisr Unfv, Mlchfganr Ann Arbor. 76<br />

PP*<br />

In Ontarlo waters <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clalr,<br />

Pages 80-86 Proc. 15th Conf. Great<br />

<strong>Lake</strong>s Res,, Znt. Assoc. Great Lakas<br />

Ras .<br />

Judayl 6. 1943, <strong>The</strong> summer standfng crop Leach. J.H, 1373, Seasonal dfstrfbutfon,<br />

<strong>of</strong> plants and an2mals In four<br />

Wfsconsln lakes, Trans. Wfs. Acad,<br />

Scf . Arts Lett. 342103-135,<br />

comporltlon and abundance <strong>of</strong><br />

rooplankton In Ontarfo wafers <strong>of</strong> <strong>Lake</strong><br />

<strong>St</strong>. Cl at r. Pages 54-64 $n Proc, 16th<br />

Conf. Great <strong>Lake</strong>s Res., Int, Assoc.<br />

Great <strong>Lake</strong>s Res.<br />

Keddyr P.A., and ,%,A, Reznlcek, 1984.<br />

Vsgetailon dynamS cs and water-level<br />

fluctuatfons on the shore7 fnss <strong>of</strong> the Leach, 1980= Lfmnologfcal sampl lng<br />

Great <strong>Lake</strong>s. Discussion Paper, fntsnslty in <strong>Lake</strong> <strong>St</strong>. C?alr fn<br />

<strong>Wetlands</strong> Conf, Nove&er 1984# Kal Jogg relation to distribution <strong>of</strong> water<br />

C%nlarp Mlcklgan <strong>St</strong>ate UnJv. 21 pp, massss, J. Grsat <strong>Lake</strong>s Res. 6(21:141-<br />

145.<br />

Kall@yr A.H,# D,S, middle ton^ and W,P.<br />

~fckell, 1963, Bfrcfs <strong>of</strong> the Detro<strong>St</strong>- Lincoln, F.C, X935. <strong>The</strong> waterfowl flyways<br />

Windsor area, Cranbr~ok Inst. <strong>of</strong> <strong>of</strong> North Amerlca. U.S. Dep. Agr.<br />

Sciencs, Detr<strong>of</strong>t, I19 pp*<br />

Clrc. 342, 12 pp.<br />

1136


Lincolnp F,C. 1950. Mjgration <strong>of</strong> bfrds.<br />

U.S, Fish Wildl. Serv. Circ. 16. 102<br />

PP 0<br />

Lyon, J.G. 1979, Analyses <strong>of</strong> coastal<br />

wetland characteristics: the <strong>St</strong>,Clalr<br />

Flats, Michigan. M.S, <strong>The</strong>sis, Un'iv.<br />

Michigan9 Ann Arbor. 65 pp.<br />

Mandelbaum, H. 1966. Sedimentatl on in the<br />

<strong>St</strong>. <strong>Clair</strong> River delta. Page 192-202<br />

j~) Proc. 9th Conf. Great <strong>Lake</strong>s Res.,<br />

Int. Assoc. Great <strong>Lake</strong>s Res.<br />

Mandel baumo H. 1969. Analysis <strong>of</strong> sedlment<br />

cores from Big Muscamoot Bay and<br />

Goose Bay in the <strong>St</strong>. <strong>Clair</strong> River<br />

delta. Pages 271-299 111 Proc. 12th<br />

Conf. Great <strong>Lake</strong>s Res., Int. Assoc.<br />

Great <strong>Lake</strong>s Res,<br />

Marsh, C.D. 1895. Cyclopidae and Calanidae<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr, <strong>Lake</strong> Michigan and<br />

certain lnland lakes <strong>of</strong> Michigan,<br />

Bull. Mich. Fish Comm. No. 5. 24 pp.<br />

Michigan Water Resources Commfssion. 1975.<br />

Limnological survey <strong>of</strong> the Michigan<br />

portion <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. Clafr~ 1973.<br />

Mich. Dep. Nat. Resour., Lansing. 48<br />

PP *<br />

Miller, 3.S. 1943. Waterfowl survey,<br />

Saginaw Bay-<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>-<strong>Lake</strong> Erie.<br />

Mich. Dep. Nat. Resour. Proj. No. 13-<br />

R., Lansing. 132 pp.<br />

Millfc, D.F. 1979. <strong>The</strong> epiphytic diatom<br />

flora <strong>of</strong> three species <strong>of</strong> aquatic<br />

vascular plants common to three <strong>Lake</strong><br />

Erie marshes. M.S, <strong>The</strong>sis, Bowling<br />

Green <strong>St</strong>ate Univ.* Bowling Green, OH.<br />

205 pp.<br />

Mizerar J.J., C.L. Cooper, and C.E.<br />

Herdendorf. 1981. Limnetfc larval<br />

fish in the nearshore zone <strong>of</strong> the<br />

western basin <strong>of</strong> <strong>Lake</strong> Erie. J. Great<br />

<strong>Lake</strong>s Res. 7(1):62-64.<br />

Modlin, R.F., and J. Gannon. 1973. A<br />

contribution to the ecology and<br />

dfstributfon <strong>of</strong> aquatic acari n the<br />

Martz, G.F. and D. Ostyn. 1977. Recovery <strong>St</strong>. Lawrence Great <strong>Lake</strong>s. Trans. Am.<br />

patterns <strong>of</strong> banded scaup harvested in Microsc. Soc. 92(2):217-224.<br />

Michigan, 1964-73. Mich. Dep. Nat.<br />

Res. , W i ldl . Div- Lansing, Rep. 2753- Morgan, J .P. 1970, Depositional processes<br />

17 PP. and products i n the deltaic<br />

environment In J.P. Morganr ed.<br />

Del talc sedfmentati on: modern and<br />

McCullough, G.B. 1982. Wetland losses in ancient. Soc. Econ. Paleontol.<br />

<strong>Lake</strong> <strong>St</strong>. Claf r and <strong>Lake</strong> Ontario. Mineral, 312 pp.<br />

Pages 81-89 In A. Champagner ed.<br />

roc. Ontarlo <strong>Wetlands</strong> ~onf. Ryerson Mudroc h A. 1 g 80. B i ogeoc hem q<br />

Polytech. Institute. Toronto, investigation <strong>of</strong> Bfg Creek Marshr<br />

Ontario. Sept. 1981. <strong>Lake</strong> Erie, Ontario. J. Great <strong>Lake</strong>s<br />

Res. 6(4) :338-347.<br />

Meade* G.G. 1857. Chart <strong>of</strong> the <strong>St</strong>. <strong>Clair</strong><br />

flats. Office <strong>of</strong> <strong>Lake</strong> Surveys,<br />

Washingtonr D.C. One map.<br />

MUdroch, A, 1981. A study <strong>of</strong> selected<br />

Great <strong>Lake</strong>s coastal marshes. Environ.<br />

Can., Natl. Water Res. Inst., Sci.<br />

Ser. No.<br />

PP<br />

122. Burlington, Ontario. 44<br />

Meeks, R.L.# and R.D. H<strong>of</strong>fman. 1980. Bird<br />

populations common to the Sister<br />

Is1 ands--the role <strong>of</strong> Navarre Marsh.<br />

Toledo Ed1 son Companyr Toledor OH. 60<br />

PP*<br />

Messman, L., R. Reppert, and E. <strong>St</strong>okhiv.<br />

1977. Wetland values, interfin<br />

assessment and evaluation method-<br />

ology, U.S. Army Inst. for Water<br />

Resour., Fort Belvolr, VA. 5-61 pp.<br />

Mudroch, A., and J. CapobSanco, 1978,<br />

<strong>St</strong>udy <strong>of</strong> selected metals in marshes<br />

on <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Ontario. Arch,<br />

Hydrobfol. 84(1):87-108,<br />

Murtyr T.S., and 8.3. Palavarapu. 1975.<br />

Reconstruction <strong>of</strong> some <strong>of</strong> the early<br />

storm surges on the Great <strong>Lake</strong>s. J.<br />

Great <strong>Lake</strong>s Res. f(ll:116-129.


Pennak, R.W. 1.978. Fresh-water inverte-<br />

brates <strong>of</strong> the United <strong>St</strong>ates9 2nd ed.<br />

John Wiley 8 Sons, New York. 803 pp.<br />

Peterson, R.T. 1980. A field gulde to the<br />

birds east <strong>of</strong> the Rockies. Houghton<br />

Mifflin Co., Boston. 384 pp.<br />

Pezzetta, J.M. 1968. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> River<br />

delta. Ph.D. Dissertation, Univ.<br />

Michigan, Ann Arbor. 168 pp,<br />

Phillips, D.W., and J.A. McCulloch. 1972.<br />

<strong>The</strong> c1 imate <strong>of</strong> the Great <strong>Lake</strong>s Basin.<br />

Envlron. Can. Clfmatol. <strong>St</strong>ud. No. 20.<br />

Toronto, Ontario. 79 pp.<br />

Pieters, A.J. 1893, <strong>The</strong> plants <strong>of</strong> <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong>. Bull. Mich. Fish Comm. No, 2.<br />

11 PP.<br />

Poe, T.P. 1983.- Food habits <strong>of</strong> larval<br />

yellow perch as a potential indicator<br />

<strong>of</strong> water and habitat quality. U.S.<br />

Fish Wlldl. Serv.? Great <strong>Lake</strong>s Fish.<br />

Lab., Ann Arbor, Mich. Admin. Rep.<br />

83-2. 21 pp.<br />

Pospichal, L.B. 1977. <strong>St</strong>. John's Marsh<br />

management plan, <strong>St</strong>. <strong>Clair</strong> County?<br />

Michigan, Mich. Dep. Nat. Resour.,<br />

Lansing. 17 pp.<br />

Price, 3 .W. 1963. A study <strong>of</strong> the food<br />

hablts <strong>of</strong> some <strong>Lake</strong> Erie fish. Ohio<br />

Biol, Surv. Bull. 2( 1) : 1-89.<br />

Qulnn, F.H. 1976. <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

hydrologic transfer factors. U. S.<br />

Dep. Commer. Great <strong>Lake</strong>s Envir. Res.<br />

Lab. ERL GLERL-10, Ann Arbor, MI. 16<br />

PP.<br />

Qufnn, F,H., and R,N. Kelley. 1983. Great<br />

<strong>Lake</strong>s monthly hydrologic data. U.S.<br />

Dep. Commer. Great <strong>Lake</strong>s Envir. Res.<br />

Lab Report ERL GLERL-26, Ann Arbor,<br />

MI. 79 pp.<br />

Raphael* C.N.* E. Jaworski, C.F. Ojala,<br />

and D,S. Turner. 1974. Future<br />

dredging quantitfes fn the Great<br />

<strong>Lake</strong>s. U.S, Envfron. Prot. Agency<br />

Ecol. Res. Ser. 660/3-74-029,<br />

Corvall is, OR. 219 pp.<br />

Raphael, C.N., and E. Jaworskf. 1979.<br />

Economic vaf ues <strong>of</strong> fish, wild1 lfe and<br />

recreation fn Michigants coastal<br />

wet1 ands. J. Coastal Zone Manage.<br />

5(3) :181-194.<br />

Raphael, C.N., and E. Jaworski. 1982. <strong>The</strong><br />

<strong>St</strong>. <strong>Clair</strong> River Delta, a unique lake<br />

delta. Geograph. Bull. 21(2) :7-28.<br />

Reighard, J.E. 1894. A biological<br />

examination <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Bull.<br />

Mich. Fish Corn. No. 4. 61 pp.<br />

Reppert, R.T., and W. Sigleo. 1979.<br />

Wet1 and values: concepts and methods<br />

for wetland evaluation. U.S. Army<br />

Inst. for Water Resour., Fort<br />

Belvoir, VA. 30 pp.<br />

Roller* N,E. 1976. Changes in the wetlands<br />

and land-use patterns <strong>of</strong> <strong>St</strong>. John's<br />

Marsh, <strong>St</strong>. <strong>Clair</strong> County, Michigan.<br />

Environ. Res. Inst. Mich.r Ann Arbor.<br />

8 PP.<br />

Roller, N.E. 1977. Remote sensing <strong>of</strong><br />

wetlands. Office <strong>of</strong> University<br />

Affairs. Natl. Aero. Space Admin.*<br />

Washington, D.C. 156 pp.<br />

Ruthven, A.G., C. Thompson, and H.T.<br />

Gaige. 1928. <strong>The</strong> herpetology <strong>of</strong><br />

Michigan. Univ. Mus. Handbook Ser.<br />

No. 3. Univ. Michigan, Ann Arbor. 50<br />

PP<br />

Sanderson, M. 1980. <strong>The</strong> climate <strong>of</strong> the<br />

Essex Region - Canada's southland.<br />

Dep. <strong>of</strong> Geography, Univ. Wlndsor.<br />

Windsor, Ontario. 105 pp.<br />

Sawyer, C.N. 1954. Factors involved in<br />

disposal <strong>of</strong> sewage effluents to<br />

lakes. Sewage Ind. Wastes. 26(3 1 :317-<br />

325.<br />

Schelske, C.L., and J.C. Roth. 1973.<br />

Llmnological survey <strong>of</strong> <strong>Lake</strong>s<br />

Michigan, Superior, Huron, and Erfe.<br />

Great <strong>Lake</strong>s Res. Div., Univ.<br />

Michigan, Ann Arbor Publ. 17. 178<br />

PP.


Schloesser, D.W. 1982. Seasonal growth <strong>of</strong><br />

submersed aquatic macrophytes in the<br />

<strong>St</strong>. Cl air-Detroit River ecosystem.<br />

U.S. Fish Wildl. Serv., Great <strong>Lake</strong>s<br />

Fish. Lab., Ann Arborp Mich., Admin.<br />

Rep. 82-5. 23 pp.<br />

Schloesserp D.W., and P.A. Manny. 1982.<br />

Distribution and re1 ative abundance<br />

<strong>of</strong> submersed aquatic macrophytes in<br />

the S t . <strong>Clair</strong>-Detroit River<br />

<strong>St</strong>uckey, R.L. 1968. Distributional history<br />

<strong>of</strong> Butornu umbel 1 a t u (flowering<br />

rush) in the western <strong>Lake</strong> Erie and<br />

<strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> region. Mich. Bot.<br />

7: 134-142.<br />

<strong>St</strong>uckey, R.L. 1971. Changes <strong>of</strong> vascular<br />

aquatic flowering plants during 70<br />

years in Put-in-Bay Harbor, <strong>Lake</strong><br />

Erie, Ohio. Ohio J. Sci. 71(6):321-<br />

342.<br />

U*S* Fish<br />

Great <strong>Lake</strong>s Fish. Lab.,<br />

Mich., Admin. Rep. 82-7.<br />

Ann Arbor*<br />

49 pp.<br />

stuckey, R. L, 1972. Taxononly and<br />

distribution <strong>of</strong> the genus<br />

(Cruciferae) in North America. Sida<br />

Contrih. Bot. 279-430.<br />

Schloesser, D.W., and B.A. Manny. 1984.<br />

Distribution <strong>of</strong> Eurasian water-<br />

mil foil (Mvrioohvllum ~oic-1 in<br />

the <strong>St</strong>. <strong>Clair</strong>-Detroit River system in<br />

1978. J. Great <strong>Lake</strong>s Res. 10(3):322-<br />

326.<br />

Scott, S.L., ed. 1983. Field guide to the<br />

birds <strong>of</strong> North America. National Geo-<br />

graphic Soc. Washington, D.C. 464 pp.<br />

Shelemon, P.J. 1971. <strong>The</strong> Quaternary<br />

deltalc and channel system in the<br />

central great valley, Cal i forn ia.<br />

Ann, Pssoc. Am. Geogr. 61(3):427-440.<br />

Sieghardt, H. 1973. Utilization <strong>of</strong> solar<br />

energy and energy content <strong>of</strong><br />

di f ferent organs <strong>of</strong> Phra~mi tes<br />

somrnu Trin. Pol, P,rch. I-cydrobiol .<br />

20:151-156.<br />

Smith, F. 1894, List <strong>of</strong> protozoa and<br />

mollusca observed in <strong>Lake</strong> <strong>St</strong>. Clai r<br />

in the summer <strong>of</strong> 1893. Pages 42-44<br />

3. E. Heighard, ed., A biological<br />

examination <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. Bull.<br />

Mich. Fish Comm. No. 4.<br />

<strong>St</strong>ansbery, I?. H. 1960. <strong>The</strong> Unionf nae<br />

(Molluscar Pelecypoda, Naiadacea) <strong>of</strong><br />

Fishery Bay, South Bass Island, <strong>Lake</strong><br />

Erie. Ph.D. Diss., Ohio <strong>St</strong>ate Univ.,<br />

Columbus. 216 pp.<br />

<strong>St</strong>ockner, J .G., and F.A. Armstrong. 1971.<br />

Periphyton <strong>of</strong> the experimental 1 akes<br />

area, northwestern Ontario. J. Fish.<br />

Res. Board Can. 28(2):215-229.<br />

<strong>St</strong>uckey, R.L. 1975. Aquatic vascular<br />

plants <strong>of</strong> the Sandusky River Basin.<br />

Pages 295-333 in D.B Baker, hl.P.<br />

Jackson, and P.L. Prater, eds. Proc,<br />

Sandusky River Basin Symposium.<br />

PLUARG/ I nt. Joint Comm., Great <strong>Lake</strong>s<br />

Regional Office, Windsorr Ontario.<br />

475 pp.<br />

<strong>St</strong>uckey, R.L. 1979. <strong>The</strong> declfne <strong>of</strong><br />

flowering lake plants. Pages 21-31 In<br />

J. Rousmaniere, ed. <strong>The</strong> enduring<br />

Great <strong>Lake</strong>s. W. W. Norton and Co.,<br />

New York.<br />

Taftp C.E., and C. W. Taft. 1971. <strong>The</strong><br />

algae <strong>of</strong> western <strong>Lake</strong> Erie. Ohio<br />

Biol. Surv. Bull. 4(1):1-189.<br />

Teal, J .M. 1962. Energy flow in the salt<br />

marsh ecosystem <strong>of</strong> Georgia. <strong>Ecology</strong><br />

43 : 614-624.<br />

Tilton, D.L., P.H. Kadlec, and P.R.<br />

Schwegler. 1978, <strong>The</strong> ecology and<br />

values <strong>of</strong> Michigan s coastal<br />

wetlands. U.T. Fish Wildl. Serv.,<br />

Twin Cities, MN. 98 pp.<br />

Tiner, R.W., Jr. 1984. <strong>Wetlands</strong> <strong>of</strong> the<br />

United <strong>St</strong>ates: current status and<br />

recent trends. Fish Wildl. Serv.,<br />

U.S. Dep, <strong>of</strong> Interior 11. S. Govt.<br />

Print. Officer Washington, P.C. 59<br />

PP<br />

U.S. Army Corps <strong>of</strong> Engineers. 1974,<br />

Proposed diked disposal area on<br />

Dickinson Island, <strong>St</strong>. Glafr County,<br />

Michigan. Final Environniental Impact<br />

<strong>St</strong>atementar U.S. Army District*<br />

Detroit* MI. 235 pp.


U.S. Department <strong>of</strong> Agriculture, 1974. Soil<br />

survey <strong>of</strong> <strong>St</strong>. <strong>Clair</strong> County, Vichfgan.<br />

U.S. Govt. Print. Office, Washington,<br />

D.C. 113 pp.<br />

U.S. Department <strong>of</strong> Commerce. 1976. Great<br />

<strong>Lake</strong>s water levels9 1860-1975.<br />

National Cceanic and Atmospheric<br />

Administrat ion, Washington, D.C. 187<br />

PP-<br />

U.S. Department <strong>of</strong> Interior. 1971. Flyway<br />

habitat management unit project. Dl v.<br />

Wildl. Refuges, Bur, Sport Fish.<br />

Wildl. FHMUP Rep. 8. Washington, D.C.<br />

5 PP*<br />

U.S. Department <strong>of</strong> Interior. 1977. <strong>The</strong><br />

Nation's renewable resources-an<br />

assessment* 1975. For. Res. Rep, 21.<br />

For. Serv. Washfngton, D.C. 243 pp.<br />

Urban, D. 1968. <strong>The</strong> ecology <strong>of</strong> raccoons on<br />

a managed waterfowl marsh in<br />

southwestern <strong>Lake</strong> Erie. M.S. <strong>The</strong>si sr<br />

Ohio <strong>St</strong>ate Univ., Columbus. 90 pp.<br />

Urban, D. 1970. Raccoon populations,<br />

movement patterns, and predation on a<br />

managed waterfowl marsh. J. W i ldl .<br />

Manage. 34(2) :372-382.<br />

Wagner, hi. J . , ed. 1977. Endangered,<br />

threatened and rare vascular plants<br />

in Michigan. Mich. Dep. <strong>of</strong> Nat.<br />

Resour., Lansing. 24 pp.<br />

Walters, L.J., T.L.. Kovack, and C.E.<br />

Herdendorf . 1974. Mercury occurrence<br />

in sediment cores from western <strong>Lake</strong><br />

Erfe. Ohio J. Scf. 74(1):1-19.<br />

Weller, M.W., and C.E. Spatcher, 1965.<br />

Role <strong>of</strong> habitat in the distribution<br />

and abundance <strong>of</strong> marsh birds. Agr.<br />

Home Econ. Exp. <strong>St</strong>a. Iowa <strong>St</strong>ate<br />

Univ,, Spec. Rep. No. 43. Ames. 31<br />

PP *<br />

Wernert M,T., and B.A. Manny. 1979. Fish<br />

distribution and limnological<br />

conditions under ice cover in Anchor<br />

Bay, <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, 1979. U.S. Fish<br />

Wildl. Serv., Great <strong>Lake</strong>s Fish. Lab.,<br />

Ann Arbor, Mich,, Admin. Rep. 80-1.<br />

58 PP*<br />

Westlake, D.F. 1963. Comparison <strong>of</strong> plant<br />

productivity. Biol . Rev. 38:385-425.<br />

Wetzel, R.G. 1975. Limnology. W. B.<br />

Saunders c0.9 Philadelphia. 743 pp.<br />

Wightman, W.R. 1961. <strong>The</strong> <strong>St</strong>. <strong>Clair</strong> Delta,<br />

M.S. <strong>The</strong>sis, Univ. Western Ontario,<br />

London. 142 pp.<br />

Wilson, J., and L.J. Walters. 1978.<br />

Sediment-water-biomass interaction <strong>of</strong><br />

toxfc metals fn the western basin,<br />

<strong>Lake</strong> Erie. Ohio <strong>St</strong>ate Univ., Cent.<br />

<strong>Lake</strong> Erie Area Res. Tech. Rep. 96.<br />

113 pp.<br />

Winner, J.M,, A.J. Oud, and R.G. Ferguson.<br />

1970. Plankton productivity studies<br />

in <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>, Pages 640-650 In<br />

Proc. 13th Conf. Great <strong>Lake</strong>s Res.,<br />

Int. Assoc. Great <strong>Lake</strong>s Res.<br />

Wolcott, R.M. 1903. <strong>The</strong> North American<br />

species <strong>of</strong> Limnesja. Trans Am.<br />

Microsc. Soc. 24:85-107.<br />

Wolfertr D.R,, W.N. Busch, and C.T. Baker.<br />

1975. Predation by fish on walleye<br />

eggs on a spawning reef in western<br />

<strong>Lake</strong> Erie. 1969-71. Ohio J. Sci.<br />

75(3) :118-125.<br />

Wolverton, C. 1981. Manual for wetland<br />

evaluation techniques. Operational<br />

draft. Div. Land Resour. Prog.# Mich.<br />

Dep. Nat. Resour., Lansing. 30 pp.<br />

Wright, L. D., and J .M. Coleman. 1972.<br />

River delta morphology: wave cl imate<br />

and the role <strong>of</strong> the subaqueous<br />

pr<strong>of</strong>ile. Science 176(4032):282-284.


DIMENSIONS OF LAKE ST. CLAIR AND ST. CLAIR RIVER WETLANDS BY UNITED STATES<br />

AND CANADA QUADRANGLE MAPSa<br />

Jurisdiction and<br />

quadrangle map<br />

MICHIGAN :<br />

Wayne County<br />

Be1 1 e Is1 e Quadrangle<br />

Detroit Quadrangle<br />

Subtotal<br />

Macomb County<br />

Grosse Point Quadrangle<br />

Mt. Cl emens Quad rangl e<br />

New Haven Quad rangl e<br />

Subtotal<br />

<strong>St</strong>. C1 air County<br />

A1 gonac Quad rangl e<br />

Marine City Quadrangle<br />

New Baltimore Quadrangle<br />

Port Huron Quadrang1 e<br />

<strong>St</strong>. Cl ai r Quadrangle<br />

<strong>St</strong>, <strong>Clair</strong> Flats Quadrangle<br />

Subtotal<br />

MICHIGAN TOTAL<br />

ONTARIO:<br />

Essex County<br />

Be1 le River Quadrangle<br />

Riverside Quadrangle<br />

<strong>St</strong>oney Point Quadrangle<br />

Ti1 bury Quadrangle<br />

Subtotal<br />

Coastal length Area <strong>of</strong><br />

af wet1 ands wet1 ands<br />

(km) (km2)


Jurisdiction and<br />

quad rangl e map<br />

Kent County<br />

Mi tche11 Bay Quadrangle<br />

Ti 1 bury Quadrangle<br />

Wall aceburg Quadrangle<br />

APPENDIX A (CONTINUED)<br />

Subtotal<br />

Lambton County<br />

Courtright Quadrangle<br />

Johnston Channel Quadrangle<br />

Mi tchel 1 Bay Quadrangle<br />

Sarn i a Quad rangl e<br />

Seaway Is1 and Quadrangle<br />

Sombra Quadrangle<br />

Wall aceburg Quadrangle<br />

Wal pol e Is1 and Quad rangl e<br />

Coastal 1 ength Area <strong>of</strong><br />

<strong>of</strong> wet1 ands wet1 ands<br />

( km? ( km2 I<br />

Subtotal 64.4 161.69<br />

ONTARIO TOTAL 102.4 231.33<br />

--<br />

LAKE ST, CLAIR/ST. CLAIR RIVER TOTAL 188.2 382.91<br />

%ata sources: United <strong>St</strong>ates Geological Survey, Dept. Interior, 7.5-minute<br />

Quadrangle Maps (1:24,000 scale?; Canada Department <strong>of</strong> Energy, Mines? and<br />

Resourcest 7.5-minute Quadrangle Maps (1:25,000 scale),


APPENDIX A (CONCLUDED)


MEAN MONTHLY ST. CLAIR RIVER FLOWSp 1900 TO 1980a<br />

Flaw In cubic meters per second<br />

WGTfl<br />

YEAR JAN FED MAR &PR M&Y JUN JUL AUO SEP OCT NOV DEC MERN<br />

1900 4190 3960 3960 5130 5240 5300 5440 5470 5530 5530 5640 5440 5077<br />

1901 4700 3570 4360 3600 3499 5720 5780 5780 5660 5610 5550 5300 5105<br />

1902 4130 4300 5180 5270 5320 5440 5410 5490 5380 5270 5300 5150 9141<br />

1903 3990 3880 4000 5240 5240 5320 5440 5440 5490 9610 5440 5150 3102<br />

1904 4330 4190 4360 5300 5520 3720 5780 5800 5780 5800 5490 5350 5304<br />

190% 3570 3940 4560 5550 566.0 5780 5930 5860 5360 5330 5720 5610 5321<br />

1306 5470 4250 4810 5460 5780 5780 5830 5800 5720 5610 5550 5150 5458<br />

1907 4420 4110 4900 5580 5640 5720 5830 5800 5330 5720 5810 5490 5395<br />

1908 3360 3740 4730 5440 5610 57YO 5330 5830 5630 '5550 5440 5380 3258<br />

1909 4810 3450 4360 25210 5380 5520 5L;SC) 5520 3490 f;3E:0 5210 5(:)40 5E1'3(1


APPENDIX B (CONCLUDED)<br />

Flow in cubfc meters per second<br />

WGTD<br />

YEW AN FEB I"ICIR APR MAY JUN JUL CSUG SEB OCT NOV DEC MEAN<br />

a Data source: U,S, Army Carps <strong>of</strong> Engineers.


ALGAE OCCURRING IN COASTAL WETLANDS AND NEARSHORE WATERS OF LAKE ST. CLAIRa<br />

Taxa Form and habitat<br />

Phylum Cyanophyta (blue-greens)<br />

Class Myxophyceae<br />

Order Chroococcal es<br />

Family Chroococcaceae<br />

Order Harmogonal es<br />

Famlly Osclllatoriacaas<br />

Family Nostocacaae<br />

SP *<br />

Fa<br />

Phylum Euglsnaphyta Cauglenoids)<br />

C7 ass Eugl enophyceas<br />

Order Eugl snal es<br />

Family Euglenaceae<br />

246<br />

Colonial; planktonic<br />

Colonial; planktonlc<br />

Colonial; planktonic<br />

Colonial; planktonic<br />

Colonial ; pl anktonfc<br />

Colonial ; p1 anktonic<br />

Colonial; gl anktanic<br />

Colonfal; planktonic<br />

Colonfal; pl anktonic<br />

Fi l amentous; pl ankton ic<br />

Fil amentous; sessile 8<br />

pl anktonfc mats<br />

Filamentous; planktonlc<br />

Colonial; among marsh p1 ants8<br />

sessile & planktonic<br />

Sol itary; pl anktonic<br />

Solftary; planktanic<br />

Sol i tary; pl ankton jc<br />

Sol ftary; p1 anktonic


APPEND SX c ( CrMlf NUED 1<br />

f axa Form and Rabftat<br />

Phylum Chrysophyta (golden-browns)<br />

Class Xanthophyceae (yellow-greens)<br />

Ordor Hoterococeales<br />

Family Chlorothaclacsae<br />

Qrdar kleterotrichal es<br />

Famlly Trlbonematacsaa<br />

SP *<br />

Order Hatoroslphonalas<br />

Farnfly Vauchorlacaae<br />

Class Chrysophyceas (goldan algae1<br />

Qrdcs Chrysomonadalas<br />

Famfly khromnadacsao<br />

Class Bacl llarlophycaa Idlatms3<br />

Order Centrales (centrfc dlatomsl<br />

Famlly Corcfnadfscacsas<br />

w<br />

Order Psnoal es ( psnnate d 1 atoms)<br />

Fanlly Frayllarlaceae<br />

Faml l y Achnanthacaae<br />

sP<br />

Family Navicul aceas<br />

Sol<strong>St</strong>ary; among marsh plants<br />

8 planktanfc<br />

Filamentous; planktonlc<br />

Filamentous; among marsh plants<br />

Filamentous; among marsh plants<br />

Sol ltary~ planktonlc<br />

So1 ftary; pl anktanic<br />

Sat ftary; planktontc<br />

$01 ftary; planktonfc<br />

Sot i tary; plankton f c<br />

$01 1 tary; planktanf c<br />

Sol 1 taryt planktonfc<br />

sol ltary; pl anktonds<br />

Sol I tary; p1 anktonfc<br />

Sol ftary; pf anktanfcr &<br />

attached to aquatic plants<br />

Sol ltary; ptanktonlc


f axa<br />

Famfly Gomphonsmaceas<br />

Family Cymbellaceas<br />

Phylum Pyrrhsphyta (fire algae)<br />

Class Dfnaphycsas (din<strong>of</strong>lagellates)<br />

Qrdar Psridiniales<br />

FnmZ f y Per 4 dl n 1 aceat3<br />

Phylum Chlarophyta (greens)<br />

Class Chlorophycsaa<br />

Ordar Volvoealas<br />

Famfly Ghlamydmanadacea~<br />

SP *<br />

BrQas Tetrasporal @s<br />

Fa<br />

Order Mf crssporal ss<br />

Fanfly Mlcrosporacsao<br />

spa<br />

Order Chaetopharaf ss<br />

5%<br />

Farm and habitat<br />

Sol itary; yl anktanfc<br />

So1 1 tasy; p7 anktonic<br />

Sol Ztary; planktonlc<br />

Sol l tary; planktonlc<br />

$01 1 tery; planktonic<br />

Calonlal; planktonfc<br />

Colonlal; planktonlc<br />

Colonial; planktonlc<br />

Colanfal; planktonic<br />

Colonlal; glanktanfc<br />

F$lamsntous; planktonic<br />

Filamentous; attached to<br />

aquatJc plants & debrfs


APPENDIX C (CONTINUED)<br />

Taxa Form and habitat<br />

Fami l y Protococcaceae<br />

Family Coleochaetaceae<br />

Order Cl adophoral es<br />

Fami 1 y Cl adophoraceae<br />

SP<br />

--<br />

SP 0<br />

--<br />

m b o r v a n u m<br />

Ped.i_arj.truml2ahsmanhtusum<br />

Order Oedogonial es<br />

Fami 1 y Oedogon i aceae<br />

Order Chl orococcal es<br />

Family Hydrodictyaceae<br />

ped i astrm -taturn<br />

-m-<br />

-mtetras<br />

Sorastrums<strong>of</strong>nulasuol<br />

Fami 1 y Coel astraceae<br />

Family Botryococcaceae<br />

,l2uulu<br />

Solitary; planktonic attached<br />

to organic debris<br />

Filamentous; attached to<br />

organic debris<br />

Fil amentous; attached to<br />

aquatic p1 ants<br />

Fi1 amentous; p1 anktonic<br />

Filamentous; attached to<br />

aquatic plants<br />

Colonial ; pl anktonic<br />

Colonial; planktonic 8<br />

among marsh plants<br />

Colonial; planktonic 8<br />

among marsh plants<br />

Colonial ; p1 anktonic<br />

Colonial ; pl anktonic<br />

Colonial; planktonic<br />

Colon i a1 ; pl ankton i c<br />

Col on i a1 ; pl ankton ic<br />

Colonial ; planktonic<br />

Colonial ; planktonic<br />

Colonial ; p1 anktonic 8<br />

among marsh plants<br />

Colon i a1 ; pl ankton i c<br />

Colon i a1 ; among marsh pl ants<br />

Colonial ; planktonic<br />

Colonial; planktonic


Taxa Form and habitat<br />

Famt 7 y Scenadesmaceae<br />

Order Zygnsmatales<br />

Family Zygnematacsae<br />

Family Ossmfdiateas Cdasmlds)<br />

So1 ltary; pl anktonfc<br />

Sol l tary; p1 anklon lc<br />

Colon4al; pl anktonfc<br />

Colonial; amang marsh plants<br />

Salltary; planktonlc<br />

Colonf a1 ; pl anktonlc<br />

Sol ltary; among marsh pl ants<br />

Colonial; planktonjc<br />

Colonlal; planktonlc<br />

Colonlal; planktonlc<br />

Colonial ; among marsh plants<br />

Colonial; amng marsh plants<br />

Colonf a1 ; pl anktsnic<br />

Colonfal; planktonlc<br />

Filamentous; planktonic<br />

Filamentous; marsh pools<br />

Filamantous; plantonfc<br />

Solitary; among marsh plants<br />

Sol i tary; amang marsh plants<br />

Solitary; among marsh plants<br />

Solltnry; among marsh plants<br />

Sulftary; among marsh plants<br />

Sol ftary; amang marsh plants<br />

Saldtary; among marsh plants<br />

Solitary; attached Lo debrfs<br />

Salltary; among marsh plants<br />

Solitary; among marsh plants<br />

5cZltaryy among marsh plants<br />

Sol l tary; amang marsh plants<br />

Sclftary; among marsh plants<br />

Solttary; among marsh plants<br />

So? f tary; among marsh pl ants<br />

Salltary; planktanfc & sessile<br />

Sol 4 tary; among marsh plants<br />

Solitary; among marsh plants<br />

Solltary; among marsh plants<br />

Sot ttargr; air*orzy marsh pl atits<br />

Sol f fary; among marsh plants


WPENDIX C (CONCLUDED)<br />

Taxa Form and habitat<br />

Sol i tary; among marsh p1 ants<br />

Sol i tary; among marsh plants<br />

Sol i tary; among marsh plants<br />

So1 i tary; among marsh pl ants<br />

Sol i tary; among marsh pl ants<br />

Fil amentous; among marsh pl ants<br />

Fil amentous; among marsh pl ants<br />

Filamentous; among marsh plants<br />

Sol i tary; among marsh plants<br />

Sol 1 tary; among marsh pl ants<br />

Sol i tary; among marsh pl ants<br />

So1 itary; among marsh pl ants<br />

Fi 1 amentous; among marsh pl ants<br />

Filamentous; among marsh plants<br />

Sol 1 tary; among marsh plants<br />

Solitary; among marsh plants<br />

So1 i tary; among marsh p1 ants<br />

Sol i tary; among marsh pl ants<br />

Sol i tary; among marsh pl ants<br />

Filamentous; among marsh plants<br />

Solitary; among marsh plants<br />

Sol i tary ; among marsh pl ants<br />

Sol i tary; among marsh plants<br />

Sol itary; p1 anktonic &<br />

among marsh plants<br />

So1 i tary; p1 anktonic &<br />

among marsh plants<br />

Sol itary; among marsh plants<br />

Sol itary; planktonic<br />

Sol itary; among marsh pl ants<br />

So1 dtary; p1 anktonic &<br />

among marsh plants<br />

Sol itary; among marsh pl ants<br />

Sol itary; among marsh pl ants<br />

So1 itary; among marsh pl ants<br />

So1 i tary; among marsh plants<br />

Sol i tary; among marsh pl ants<br />

Solitary; among marsh plants<br />

Sol itary; among marsh plants<br />

Sol itary; among marsh pl ants<br />

Sol i tary; among marsh plants<br />

a Data sources: Pfeters (189311 Winner et al. (19701, Taft and Taft (19713s<br />

Michigan Water Resources Commdssion (1975).


Phylum Protozoa fpsotoroansl<br />

C1 ass Sarcad l na<br />

Qrdar<br />

Class @Illopksre leflfalosf<br />

Order HaleP~lckEda<br />

FamSly Tracheldnldaa<br />

Order 925g(st,rlchlda<br />

Famfly Tlnttnfitda6<br />

ZWPLANKVQN OCGURRING Xt\B THE COASTAL WETLARDS<br />

AW NEARSHORE WATERS QF LAKE ST, CtAXWa<br />

1 52<br />

Abundance status<br />

Rare<br />

Cornon<br />

Camn<br />

Cornon<br />

Abundant<br />

Cornon


APPENDIX 0 (~oNTIMUED)<br />

Spec 1 es Abundance status<br />

Order Psri trichida<br />

Family Vorticallldae<br />

Family Ophrydiidae<br />

Phylum Rotatoria (rotifsrs)<br />

Class Monogononata (sfngle ovary rotifers)<br />

Order Flosculariacea<br />

Famf 1y Conochil idae<br />

unicorniq<br />

Family Tostudinellidae<br />

Order Co? 1 othecacea<br />

Famfly Collothscidae<br />

Order Ploimacea<br />

Family Notommatldae<br />

Family Synchaetidae<br />

Family Gastropodidae<br />

Occasional<br />

Common<br />

Rare<br />

Uncomnon<br />

Common<br />

Abundant<br />

Common<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Occasional<br />

Occasional<br />

Abundant<br />

Abundant<br />

Rare<br />

Rare<br />

Occasional<br />

Common


Spec 1 os Abundance status<br />

Rare<br />

Bcca5fonal<br />

ff are<br />

Rare<br />

Rare<br />

Occasional<br />

Rare<br />

Rara<br />

Rare<br />

Rare<br />

Abundant<br />

Rare<br />

Csmn<br />

kcas tonal<br />

kcas f anal<br />

Rare<br />

Ris re<br />

Rare<br />

Raro<br />

Rare<br />

Abundant<br />

Abundant<br />

Cornan<br />

Cmon<br />

Raro<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Ram<br />

Ra ra<br />

Rare<br />

Rare


APPENDIX D (CONTINUED)<br />

Species Abundance status<br />

Family Tylotrochidae<br />

--<br />

llxxEaG<br />

eriorlslghniq auadranu<br />

Phylum Arthropoda (joint-legged animals)<br />

Cl ass Crustacea<br />

Order Cl adocera (water fleas)<br />

Family Daphnidae<br />

aaPhnfa-<br />

DnDhn_aw-<br />

DaD.hnia-<br />

DaPbn.la-<br />

DaDhnLsaul_ex<br />

BaBhnlsret;racurvn<br />

--<br />

Fami 1 y Leptodoridae<br />

Le~todoril hindtii<br />

Family Sididae<br />

Uahmxam -era1 anum<br />

smd-<br />

Fami 1 y Hol oped i dae<br />

LlmQCm<br />

Family Bosminidae<br />

Bosmlna-<br />

EubPsminm.coreaon5<br />

Fami 1 y Chydoridae<br />

Alona-<br />

Alona-<br />

Blneanuadranaularis<br />

81snella SPP*<br />

-a<br />

Gdmmaas-<br />

Family Macrothrici dae<br />

Rare<br />

Uncommon<br />

Fare<br />

Rare<br />

Common<br />

Common<br />

Occasional<br />

Uncommon<br />

Common<br />

Occasional<br />

Uncommon<br />

Uncornmon<br />

Rare<br />

Occasional<br />

Abundant<br />

Abundant<br />

Uncommon<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Rare<br />

Occasional<br />

Rare<br />

Rare<br />

Rare


Spec 3 es<br />

Abundance status<br />

Unc mion<br />

hhc oarnor?<br />

SuBardtrr. bZ&rpaet Jc~Zda 4 hat-past lc<strong>of</strong>d cogspr~ds)<br />

e drrrf 1 y blasrpacticldao<br />

rn Rare


APPEMDlX E<br />

VASCULAR AQUATIC PLANTS KCURRING IN LAKE ST, CLAIR WETLANDSa<br />

Ecologicaf niche and species Abundance statusb<br />

Class Angiospermae (flowering p1 ants)<br />

Subclass Monocotyledonae (monocots)<br />

Qrdsr Pandanal es<br />

Family Typhaceae (cattails)<br />

(narrow-leaved cattail)<br />

road-1 saved cattail<br />

(hybrid cattail)<br />

Family Sparganium (bur reeds)<br />

(giant bur reed)<br />

Order A1 ismales<br />

Famlly Alismataceae (water plantains)<br />

(water plantain)<br />

mmon arrowhead)<br />

Family Butomaceae (flowering rushes)<br />

(flowering rush)<br />

Order Grami nal es<br />

Fami 1 y Grami neae (grasses) [bluejoint grass1<br />

tlesnake grass)


Ecological ndche and specles Abundance status<br />

Famf 1 y Cyperaceae (sedges)<br />

(tussock sedge)<br />

(redrooted cyperusl<br />

(rusty cyperus)<br />

f redfooted spiks-rush)<br />

unt spike-rush)<br />

1 (American bulrush,<br />

(dark green bulrush)<br />

(river bulrush)<br />

sat bul rush)<br />

Order Xyrldalas<br />

Fsmif y Pontedsriaceao (ptckeral weads)<br />

(water stargrass, mud plantain)<br />

(p fckarel wwd)<br />

Qrder ifllalaa<br />

Famlly Juncaceae (rushes)<br />

(s<strong>of</strong>t rush)<br />

(knotted rush)<br />

CTorreyrs rush1<br />

Ordnr arch Idalss<br />

Family Orchidaceae (orchf ds)<br />

(prairfe fringed orchfdl<br />

5ubcIass Olcatyladanas tdfeotsl<br />

Order Sal fcales<br />

Famfly Sallcacaae (willaws)<br />

C eastern ca.t;tonwood)<br />

(quaking aspen)<br />

y wlSSaw9


WPENDXX E (CONTINUED)<br />

Ec<strong>of</strong> ogical niche and species Abundance status<br />

Order Myrical es<br />

Family Jug1 andaceae (walnuts)<br />

(shagbark hickory)<br />

Order Fagal es<br />

Family Fagaceae (beeches)<br />

bicolar (swamp white oak)<br />

(burr oak)<br />

(pin oak)<br />

Order Urtical es<br />

Fami 1 y Ul maceae (elms 1<br />

Y.lnrus mericana (American elm)<br />

Famlly Urticaceae (nettles)<br />

Ilrtica dioica (stinging nettle)<br />

Order Polygonal es<br />

Family Polygonaceae (smartweeds)<br />

Pol va- am~hi b ium (water smartweed)<br />

Po1 vgfau~~ ~occine~ (swamp persicaria)<br />

Po1 v m ~onvol- (black bindweed)<br />

Pol vaow Mi f o l u (nodding smartweed)<br />

& d y g p~nsvl ~ ~ ~ vanica (Pennsylvania smartweed,<br />

pi nkweed)<br />

pol vgonum punctat~ (water smartweed)<br />

R j 3 vertici,ll atus (swamp dock)<br />

Order Ranales<br />

Family Nymphaeaceae (water lilies)<br />

Jutes (American water-lotus)<br />

Nu~hu & v e ~ ( spatterdock, ye1 1 ow water 1 $1 y 1<br />

Order Papaverales<br />

Family Cruciferae (mustards)<br />

Wamine pensvlvania (bitter cress)<br />

IWiQJB i2-1 (marsh cress)<br />

Order Rosa1 es<br />

Family Crassulaceae (orpines1<br />

Pent- (ditch stonecrop)<br />

Fami 1 y Rosaceae ( roses)<br />

(sf 1 verweed)<br />

mp rose)


Order SapSnd&lea<br />

F emf 3 y Anacardf aceao I sum&< b b<br />

(btaghorn sumac)<br />

family Acoraceact Imaplus)<br />

MS % tsf3ver maple)<br />

Ut-4c.r hbal alas<br />

FamfXy eorna~sae (woyruodsl<br />

gray dqiwsrjda<br />

# r ran a5 4er d~xjhwjd<br />

1


AQPEHDIX E ( CONTINUED 1<br />

Ecological niche and species Abundance status<br />

Family Boragfnaceae (borages)<br />

(common comfrey)<br />

Family Solanaceae (nightshades)<br />

(nightshade)<br />

Order P<strong>of</strong>emoniales<br />

Family Labiatae (mints)<br />

(water horehound)<br />

Family Acanthaceae (acanthids)<br />

(water wfllow)<br />

mad-dog sku1 lcap)<br />

Order Rub i a1 es<br />

Family Rubiaceae (madders)<br />

~ccidentali~ (buttonbush)<br />

Order Campanul a1 es<br />

Family Lobelliaceae (lobellas)<br />

(blue lobelia)<br />

Order Asteral es<br />

FamSly Composftae (composites)<br />

(nodding beggar tick)<br />

(purple-stemmed swamp beggar tick)<br />

(Hi11@s thistle)<br />

(bonset)<br />

Cl ass Ang i ospermae ( f 1 ower i ng pl ants 1<br />

Subclass Monocotyledonae (monocots9<br />

Order Na j adal es<br />

Fami 1 y Potamogatonaceae ( pondweeds)<br />

(knotty pondweed)<br />

Subclass Dicotyledonae Cdicots)<br />

Order Po1 ygonal es<br />

Family Polygonaceae (buckwheats)<br />

(water smartweed)<br />

(swamp smartweed)


APPENDIX E (CONCLUDED)<br />

Ecological niche and species Abundance status<br />

Order Xyridales<br />

Family Pontederiaceae (pickerel weeds)<br />

(mud plantain, water stargrass) C<br />

Subclass Dicotyledonae (dicots)<br />

Order<br />

-<br />

Ranal ss<br />

Family Ranunculaceae (buttercups)<br />

Jonairastris (water crowfoot)<br />

Order Myrtal es<br />

Family Ha1 oragidaceae (water mi 1 foils)<br />

Mvrf0~h~l1t,@ a1 terniflor_urn (1 Jttle water mil foil<br />

(water mil foil 1<br />

~~riophvllwm s~icatw (water milfoil)<br />

Family Onagraceae (evening primroses)<br />

(water primrose)<br />

Class Angiospermae (flowering plants)<br />

Subclass Monocotyledonae (monocots)<br />

Order AraZ es<br />

Family Lemnaceae (duckweeds)<br />

Lemnq $rI scu1 q (star duckweed)<br />

Subclass Dicotyledonae (dicots)<br />

Order Ranales<br />

Family Ceratophyllaceae (hornworts)<br />

Cerato~hvllace_ae demersw (hornwort, coontail) A<br />

Order Polemon fa1 es<br />

Family Lentibulariaceae (bladderworts)<br />

(bl adderwort)<br />

"0ata sources: <strong>St</strong>uckey (1968, 1972, 1975, 19791, J aworski and Raphael<br />

( 1978) Herdendorf et a1 . (1981~).<br />

b~bundance<br />

status: A = abundant, c = common, 0 = cccasional, R = rare,<br />

U = uncormnon.


BENTHIC MACROINVERTEBRATES OF NEARSWORE LAKE ST, CLAIR AND CONNECTING<br />

WATERWAYS (EXCLUDING MOLLUSKSIa<br />

Classfflcatfun and spscfss<br />

Phylum Porffera fsgongas)<br />

Class 0mosponglae (horny spangas)<br />

Famll y Spang11 l ldae ( fresh-water spanyes)<br />

SPP *<br />

Phylum Costante& ata (61 ass Cnf darfal<br />

Class Wydrozea (kydrozoans)<br />

Famtly Wydrldas<br />

SPP<br />

Phylum Platyhslmlnthes (flatworms)<br />

Class Turbel? aria t p1 anarl ansl<br />

Ordar Trfcladlda (triclads?<br />

Frtmlly Plannrlldae (planasfans1<br />

Phylum Namatoda (roundworms)<br />

Gl asa Adsnaphorao<br />

Family Trlpylfdae<br />

SP *<br />

Phylum Annelids (sagmented worms?<br />

GI ass Pal ychaeta<br />

FamE 1 y Sabal l idae ( Fan warmsr feather-duster worms1<br />

Class Bllgochaata laquatic aarthwarms)<br />

f emf l y Lun&r?cul idae


WPENDZX F (CONTINUED)<br />

Cl assi f ication and species<br />

Famdly Tubificidae (sludge worms)<br />

Famf ly Naididae<br />

sj2ik.h-<br />

Unc;.f- -<br />

Fami 1 y Gl ossoscol eci dae<br />

i2zIEE&<br />

Class Hirudinea<br />

--<br />

(leeches)<br />

Family Glossiphoniidae<br />

He1 &dell a $riserial is<br />

Order Pharyngobdellida<br />

Family Erpobdellidae


Classffjcatlon and spscfes<br />

Phylum Arthrapada (joint-lsggod anlmalsl<br />

C1 ass Arachndda<br />

Order Acarina (mites and tIcks1<br />

Superfamily Mydracarfna dwatgr rnftss)<br />

Famfly Limnssifdao<br />

Fami l y Wygrabatidaa<br />

sp,<br />

Family Plonidae<br />

Cl ass Crustacea<br />

Subclass Bstracoda (ssod shrimps)<br />

Famfly Cypridae<br />

SPP *<br />

Subcl ass Ma1 acast raca<br />

Qrder Isopoda (aquatlc sowbugs1<br />

Family Assllldas<br />

Ordsr Amphfpoda fstdeswfmrs, scuds)<br />

Family Gamaridae<br />

Order Becapoda tcrayff shesr shrimps)<br />

Subardsr Raptantfa (crayfJskss1<br />

Famlly Gambarldas (crayffshosb<br />

SP *<br />

Class Xnsacta (a Msxapodal /Insects)<br />

Order Ephemroptera (may fl 3 @r,!<br />

Fsmlly Baetfscfdas<br />

5P *


APPENDIX F (CONTINUED)<br />

Classiffcat4on and species<br />

Family Ephemeridae (burrowing mayflies)<br />

Fami 1 y Pol ymi tarci dae<br />

Order Odonata (dragonflies and damselflies)<br />

Suborder Anisoptera (dragonfl ies)<br />

Family Gomphidae<br />

SP<br />

Suborder Zygoptera (damselflies)<br />

Family Coenagrionidae (narrow-winged damselflies)<br />

SP *<br />

Order Hemiptera (bugs)<br />

Family Corixidae (water boatmen)<br />

SP<br />

-<br />

Family Hydropsychidae (net-spinning caddfsflles)<br />

SP.<br />

SP.<br />

Order Trjchoptera (caddi sf1 ies)<br />

Family Hydroptilidae (micro-caddisflies)<br />

HyGm@m SP.<br />

- SP *<br />

Family Polycentropodidae (tube-making caddisflies)<br />

SP.<br />

Family Psychomylldae (trumpet-net caddisflies)<br />

l2admuh SP*<br />

Family Mol annidae<br />

SP *<br />

Family leptocerldae (long-horned caddisfl les)<br />

!&xiz&a SP.<br />

SP *<br />

SP *


Cl asst f icaP3on and speclas<br />

Order Coleaptera (bsotles)<br />

Farnlly Hallplldaa (crawling water bsotles)<br />

Famtly Elmfdae (rlffla beetles)<br />

sp.<br />

Order DIptera ltrus flles)<br />

Faml l y Chaobor ldaa (phanton midges)<br />

SP *<br />

Family Culicidas (mosquftaesf<br />

SP *<br />

'8ata saurcos: Wo3cotf ( 1903 1 @ Edmondsan ( 1959), Hunt ( 19621, Hf 1 tunen<br />

tE971r 1BBQ)r Modlin and Gwnnon 6197311 Pennak (1S78Is Mrdsndarf et al.<br />

il$8lcIt Mfltunen and Manny (1982)r Hardsndorf and Hardsndorf (1983).


TAXONOMIC LISTING OF FRESHWATER MOLLUSCA OF LAKE ST. CLAIR<br />

COASTAL MARSHES, NEARSHORE WATERS, AND TRIBUTARY MOUTHSa<br />

Specf es<br />

Cl ass Gastropoda (snai l s)<br />

Subclass Prosobranchia (gill-breathing snails)<br />

Order Mesogastropoda<br />

Superfamily Viviparidae (mystery snails)<br />

Famlly Vivfparidae (mystery snails)<br />

816) brown mystery sna?l<br />

r 1834) banded mys<br />

(Gray, 1834) (= Y,<br />

oriental mystery snail or Japanese snail<br />

Superfamily Valvatacea<br />

Family Valvatidae (valve snails)<br />

Valvat3 w e s s q Walker, 1906 flat valve snail<br />

Valvata piscf nal is (Mu1 ler, 1774) European valve snail<br />

slncer~ sincera Say, 1824 ribbed valve snail<br />

trlcarinata (Say, 1817) three-keeled valve snail<br />

Superfamily Rissoacea<br />

Family Hydrobiidae (spire snails)<br />

clncinnatiensis (Anthony, 1840) campeloma spire<br />

snail<br />

(Baker, 1928) fl at-ended spi re snail<br />

r, 1928) Pilsbryps spire snail<br />

Say, 1817) ordinary splre snail<br />

Pilsbry, 1898 small spire snail<br />

(Say, 1825) deep water spire snail<br />

Family Truncatellidae (looplng snails><br />

(Say, 1817) river bank looping snail<br />

Family Bfthynjidae (faucet snails)<br />

l2mULb (Lfnnaeus, 1767) faucet snafl<br />

Superfamily Cersithiacea<br />

Family Pleuroceridae (horn snai 1s)<br />

Rafinesque* 1831 fl at-sided horn snaS9<br />

(Menke, 1830) Great <strong>Lake</strong>s horn snaS3<br />

--


Spsc .i tss<br />

APPENDIX C; (CONTINUED)<br />

Subclass Pulmansla Qfuny-breathing snafls)<br />

Ordsr Basomatophosa<br />

Superfamlly Lymnaeacea<br />

Famlly Lymnaefdea (pond snaf 1s)<br />

(<strong>St</strong>reng, 18961 shouldered northern fossarfa<br />

sa, laall graceful fossarja<br />

(Say, 18251 modest fassaria<br />

(Lea, 1841) amphlblaus fossarfa<br />

er, 1907) small pond snall<br />

{Say, 1817) AmerScan ear snail<br />

nney, 18671 slsnder pond snall<br />

y pond snall<br />

1817) great pond snafl<br />

(Say, 1817) lake stagnjcola<br />

sr, 19069 mini aturg<br />

(Sayt 18211 comon stagnfcola<br />

(Say, 1821) striped staynlsola<br />

Superfamlly Physacea<br />

Family Physfdaa (tadpals snaffs)<br />

Say, 1821 tadpols snafl<br />

eman, 1841 solid lake physa<br />

(Llnnaeu~~ 1758) polished tadpole snafl<br />

Superf amll y Pl anorbacoa<br />

Family Pl anarbldae (ramshorn snaj 1 s)<br />

ETyron, 1866) flatly colled gyraulus<br />

Baker, 1932 great carinate<br />

(Sayc 1816) larger eastern ramshorn<br />

Fa e f reshwatsr l impets9<br />

.G. Adams, 18411 dusky lily-pad lfmpet<br />

(Tyron, 1863) oval lake lfmpet<br />


Cl ass Pel ecypoda<br />

Order Eul amel l ibranchia<br />

Superf am1 1 y Un i onacea<br />

Family Unionidae (pearly mussels)<br />

Subfamily Ambleminae (button shells and relatives)<br />

(Say, 1817) three-ridge<br />

Rafinesque, 1820) plg-toe<br />

(Rafi nesgue, 1820) maple-leaf<br />

Lea, 1831) warty-back<br />

(Rafinesque, 1820) purple pimple-back<br />

inesque, 1820) spike, or lady-finger<br />

(Conrad, 1836) false pig-toe<br />

Subfamily Anodonti nae ( f l oater mussels)<br />

Raf inesque, 1820) brook wedge mussel<br />

Say, 1819 ridged wedge-mussel<br />

(Barnes, 1823) white heel-splitter<br />

29) brook l asmigona<br />

er fluted shell<br />

1 (Say, 1825) mudpuppy<br />

(Lea, 1834) cyl lndrical floater<br />

ay, 1829 common floater<br />

paper pond-she1 l<br />

SugAxku ur,ululaUs (Say, 1817) squaw-foot<br />

Subfamily Lampsi1 inae (1 amp-mussels)<br />

(Rafinesque, 1820) kidney shell<br />

gue, 1820 three-horned warty-back<br />

Jruncilla maciformis (Lea, 1828) fawn%-foot<br />

Truncilla fsunc- Rafinesque, 1820 deer-toe<br />

U (Say, 1817) pink heel-spl itter<br />

(Barnes, 1823) lilliput mussel<br />

(Raf inesque, 1820) round hickory-nut<br />

Raffnesque, 1820) olive hickory-nut<br />

Rafinesque* 1820) fragile paper-shell<br />

(Barnes, 1823) mucket<br />

, 1817) polnted sand-shell<br />

(Lamarck, 1819) bl ack sand-she1 1<br />

r 1820 wavy-rayed lamp-mussel<br />

(Barnes, 1823) fat mucket<br />

f Barnes, 1823 > pocket-book<br />

, 1831) bean villosa<br />

(Leal 1838) northern rfffle shell<br />

sque, 1820) trieorn pearly mussel


Spec l sf;<br />

APPENDIX &; 6.<br />

SuperfamDly Sphaarlacsa<br />

Famlly Corblculfdae (71ttle basket clams)<br />

lMullor, 2774) Aslatls clam<br />

Famlly Sphaerifdae (ffngornafl clams and poa clams)<br />

Subfamily Sphasrilnaa (fingarnail clams)<br />

nnaous, 17583 European flngernafl clam<br />

ssfn, 1876 Arctic-alpine Flagernall clam<br />

(Say, 18221 rho&<strong>of</strong>d ffngernall clam<br />

y, 18161 grooved ffngernatl clam<br />

f lannarck, liWl8) strfated flngernaf I clam<br />

Mullsr, 27743 faks fingarnail clam<br />

(Say, 18221 swamp flngernafl clam<br />

fme, 18511 pond ffngernadl clan<br />

(Say, 1829) long ffngsrnafl clam<br />

Subfamtly Pi%ldl%nao Epsa clams or pill clams)<br />

(Muller, 17741 greater European pea clam<br />

Roper, 1890 giant northern pea clam<br />

me, 1852 Adamt% pea clam<br />

(Poli, 139%) ubdqultous poa clam<br />

Prfme, 1852 rfdged-beak pea clam<br />

Primer 1852 rsufid pea clam<br />

d, 1898 rfver pea clam<br />

Prlmsr 1852 rusty poa clam<br />

(Sheppardr 1895) Hsnslowts paa clam<br />

Clersint 1886 Lilljeborgqs pea clam<br />

1836 quadrangul ar plll cl am<br />

fisr L832 shlny pea clam<br />

rlme, 1852 fat pea clam<br />

Malm, 1855 shart-sndsd pea clam<br />

Primer 1852 triangular poa clam<br />

Primat 1851 globular pea clam<br />

rkll 1895 Walkorqs pea clam<br />

@lasainl 1877 Arctic-alplns poa clam<br />

SLerkf r 1895 perforatad pea clam<br />

--<br />

& OaLa sources: Gosdrlch and Van dor fehalla (&93211 Berry (19431, <strong>St</strong>ansbary<br />

I196OT* Clarke (X98l1,


Species<br />

APPENDIX H<br />

HABITAT PREFERENCE OF FRESHWATER MOLLUSCA OF LAKE ST. CLAIR<br />

COASTAL MARSHES AND NEARSHORE WATERS<br />

Shallow Deep Gravel<br />

((2 m) (>2 m) Mud Sand & rock Plants Comments<br />

Eutrophf c<br />

Eutrophic<br />

Rare<br />

Rare


Spec t as<br />

APPENDIX W ICQNIINUED)<br />

Shall sw Deep Gravel<br />

((2 m) f >2 m) Mud Sand 8 rock PI ants Cornants<br />

x Eutrophic<br />

x Eutraphlc<br />

x Rare<br />

X<br />

x EutrophSc<br />

X<br />

x Eutrophic<br />

X<br />

X<br />

X<br />

X<br />

X<br />

X<br />

X<br />

X<br />

X<br />

Rare


Species<br />

Shall ow Deep Gravel<br />

((2 m) (>2 n) Mud Sand d rock Plants Comments<br />

X X X<br />

x x Rare<br />

X X<br />

X X<br />

X X X<br />

X X X X<br />

X X X<br />

X<br />

Rare<br />

a~ata<br />

sources: Goodrich and Van der Schal ie (1932) Berry (19431, Clarke<br />

( 1981 1.


Famlly and species<br />

Fsmfly Aclpenserldas<br />

FISH KNOWN OF4 F'Rt SlSMED TO U"I"TI_IZE THE COASTAL WETiADuUS<br />

OF THE ST, CLAXR RXVEM AND LAKE ST, CtAIRa<br />

( sturgeon 1<br />

Famfly kspfsostsldee (gars)<br />

l angnase gas) Coprm~n Gomon<br />

(spotted gar) Unc<strong>of</strong>nnrun Dncorman<br />

Family fso~ldas (pfkesl<br />

Cnorthsrn plksl Abundant Camn<br />

(musks1 1 ungo)<br />

(grass pfckarsl 1<br />

Camon<br />

Camon<br />

Uncomon<br />

Comn<br />

Famfl y Umbrf dae (mudm9 nnawsl<br />

(central mudmf nnow?<br />

Fa I rigs)<br />

(gfttard shad)<br />

Cyalden ~hjnerf<br />

blackchin shiner)<br />

spattat 1 shf narl<br />

femrald shingar)<br />

sand shiner)<br />

g fathead mS nhov I<br />

Camon<br />

Rare<br />

Ra rs<br />

Comn<br />

Camn<br />

Abundant<br />

Ataerndant<br />

Abtnradant<br />

Abundant<br />

Csmn<br />

Itncomn<br />

Comn<br />

Rare<br />

Rare<br />

Camn<br />

Cornon<br />

Abundart<br />

Abundant<br />

Camon<br />

C O ~ R<br />

Abundant<br />

Abundant


Famlly and species<br />

Family Catostomidea (suckers)<br />

(white sucker)<br />

e chubsucker)<br />

(spotted sucker)<br />

Family Ictaluridae (catfishes)<br />

.li&&uE (channel catfish)<br />

ma1 urus me1 as ( bl ack bull head 1<br />

Jctalurus ~ u l o s (brown u ~ bull head)<br />

Lctalurug natalis (yellow bullhead)<br />

Ngturus gvrinu (tadpole madtom)<br />

Family Cyprinodontidae (killifishes)<br />

Fundulu dlaohanur (banded killifish)<br />

Family Centrarchidae (sunfishes)<br />

Bmblo~l ites ~ e s t r i s ( rock bass)<br />

je~omi3 c~anellus (green sunfish)<br />

) I s albbosus (pumpkinseed)<br />

~ o c h i r u s (bluegill<br />

Micro~teru dolomiea (smal lmouth bass)<br />

Micro~teru salmoide~ (largemouth bass)<br />

Pomoxi~ gaaulari~ (white crappie)<br />

~1iaromacu1 atu (black crappie)<br />

Family Percidae (perches)<br />

fiheostoma exile (Iowa darter)<br />

Etheosto~ D I (johnny darter)<br />

perca f 1 avescens (ye1 1 ow perch<br />

Percina wades (1 ogperchl<br />

<strong>St</strong>izostedion vitreu~ (walleye)<br />

Family Perichthyidae (basses)<br />

Nrone &- (white bass)<br />

Family Sciaenidae (drums)<br />

A~lodinotus grunniens (freshwater drum)<br />

Family Gasterosteidae (sticklebacks)<br />

Culea inconstans (brook stickleback)<br />

Family Cottidae (sculpins)<br />

(mottled sculpin)<br />

-- --<br />

aorta source: Herdendorf et al. (1981~).<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Uncommon<br />

Common<br />

Common<br />

Common<br />

Common<br />

Common<br />

Common<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Uncommon<br />

Common<br />

Common<br />

Abundant<br />

Abundant<br />

Common<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Common<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Uncommon<br />

Common<br />

Common<br />

Common<br />

Common<br />

Common<br />

Common<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Uncommon<br />

Common<br />

Common<br />

Abundant<br />

Common<br />

Common<br />

Common<br />

Common<br />

Common<br />

Conimon<br />

Common


APPENDIX J<br />

WPWXRIANS AND REPTILES OGCURRING IN THE COASTAL WETLANDS<br />

OF LAKE ST. CLAIR AND THE ST. CLAlR RIVERa<br />

Spec 1 er Abundance status<br />

Class Amphfbfa (amphfbians)<br />

Order Gaudata (salamanders)<br />

PPY<br />

Crsd-spotted newt)<br />

Caastsrn tfger salamander)<br />

(blue-spotted sal amander)<br />

( spattad sal amanderf<br />

sd-backed salamander)<br />

(four-toed salamander)<br />

Order Saltantfa Cfroys and toads)<br />

tern chorus frog)<br />

% crfckot frog)<br />

Class Regtffla Creptflasl<br />

Ordar Squaaata (snakes and llzards)<br />

(northern water snake1<br />

(northern rlbbon snake)<br />

(northern red-ballfsd snake)<br />

brawn snake)<br />

mf dl and brown snake)<br />

f~astern hagnose snake)<br />

(northern rtngnsclk snake)<br />

rn $moth green snake)<br />

Comon<br />

Comon<br />

Common<br />

Comn<br />

Coman<br />

Abundant<br />

Rare<br />

Abundant<br />

Cam0<br />

Comon<br />

Comon<br />

Gomon<br />

Comon<br />

Comn<br />

Abundant<br />

Camnon<br />

Comn<br />

Abundant<br />

Abundant<br />

Uncomnon<br />

Abundant<br />

Uncomn<br />

Comon<br />

Comon<br />

Comn<br />

Coman<br />

Comn


Spec S es Abundance status<br />

feLd. (blue racer)<br />

eastern fox snake)<br />

rat snake)<br />

(eastern ml1 k snake)<br />

(eastern massasauga)<br />

Order Testudf nes (turtles)<br />

Z, ser~entlnq (snapping turtle)<br />

Common<br />

Rare<br />

Rare<br />

C0mm0~<br />

Uncomnon<br />

Comnon<br />

Rare<br />

Common<br />

Common<br />

Abundant<br />

Comon<br />

Rare<br />

a~ata sources: Ruthven et al. (1928) * Conant (1975). Behler and King (1979).<br />

Herdendorf et a1 . (1981~).


BIRDS KCURRING IN THE VICINITY OF LAKE ST. CLAIR<br />

AND ST. CLAZR RIVER COASTAL WETLANDS3<br />

Spec f es Resf dance statush<br />

Class Avas (Bfrds)<br />

Qrdar Anserifarms<br />

Famlly Anatfdae (swans# go@$@, and ducks1<br />

SubfamJly Anserinae (swans and ge@sa)<br />

Trfbe Cygnlni<br />

(tundra swan)<br />

Subfamily Andtinaa Iducksl<br />

Trlbe CafrZninl<br />

(wood Buck)<br />

(dabbl f ng ducksf<br />

whfte-fronted goose)<br />

(canvasback)


Tribe Oxyurini<br />

APPENDIX K (CONTINUED)<br />

fflehead) W<br />

(hooded merganser) M<br />

common merganser) M<br />

( red-breasted merganser) M<br />

(ruddy duck)<br />

Order Podicipedi formes<br />

Family Podicipedidae (grebes)<br />

orned grebe) M<br />

(pied-billed grebe) BR<br />

Order Pelecaniformes<br />

Family Pelecanidae (pelicans)<br />

(white pelican) M<br />

Residence status<br />

Order Gruiformes<br />

Famlly Ra11 idae (rails, moorhensp and coots)<br />

BR<br />

(Virginia rail BR<br />

BR<br />

(common moorhen) BR<br />

merican coot) BR<br />

Order Ciconi i formes<br />

Family Ardeidae (herons)<br />

(American bittern) BR<br />

BR<br />

YR<br />

BR<br />

BR<br />

(green-backed heron) BR<br />

f bl ack-crowned BR<br />

n-ight-heron)<br />

Order Charadrllformes<br />

Family Charadri idae (plovers)<br />

(semi palmated plover) M


- OIIJL<br />

h<br />

e-. L<br />

UI a3 CI<br />

wl Q. I<br />

a, n-<br />

F Ira -g<br />

%--QIW L.r<br />

o m a c m a<br />

c m-- m am<br />

~ o a m c c<br />

a?= a a<br />

X3rCh mtn<br />

C<br />

vlL"-rn@a at-<br />

L@P- .P m a<br />

@i).t-)@-W--- L<br />

116s XQIF m*, Q<br />

+totmwa~<br />

t +rn-mu~=<br />

a Cno 0-r 0 @F-<br />

E-=--uiQ. LP-ar-<br />

s<br />

x -<br />

u ."-<br />

at-<br />

3 3<br />

1 Cn<br />

x<br />

u w<br />

FO. c<br />

i r - F3


WPEMDXX K (CONTINUED)<br />

Spec f es Residence status<br />

Family Accipitridae


APPENDIX K (CONCLUDED)<br />

Spec l es Residence status<br />

Subfamily Emberizinae (sparrows)<br />

(swamp sparrow) Y R<br />

(savannah sparrow) BR<br />

a~ata sources: Peterson ( 19801, Herdendorf et a1 . ( 1981~1, Scott (1983).<br />

b~esi<br />

dence key :<br />

YR - Year-round resident BR - Breeding range (spring and summer)<br />

M - Migrant W - Winter range


APPENDIX L<br />

MAMMALS OCCURRING IN THE COASTAL WETLANDS OF LAKE ST. CLAIR<br />

AND THE ST. CLAIR DELTAa<br />

Species Abundance status<br />

Order Marsupialla (marsupials)<br />

Family Didelphidae<br />

(Vi rgfnia opossum)<br />

Order Lagomorpha<br />

Family Leporidae<br />

(eastern cottontail)<br />

(European hare)<br />

Order Rodentia<br />

Famlly Sciuridaa<br />

(woodchuck)<br />

ray squf rral<br />

rrel )<br />

( red squirrel 1<br />

Family Cricetl dae<br />

Order Carnivora<br />

Family Canidae<br />

Yu1pcs. Lu.lxa (red fox)<br />

Famil y Procyon idae<br />

( raccoon 1<br />

(muskrat 1<br />

Family Mustel idae<br />

frenat~ (long-tailed weasel)<br />

Order Art i odactyl a<br />

Family Cervf dae<br />

(striped skunk)<br />

(white-talled deer)<br />

'~ata sources: Hayes ( 1964) r Herdendorf et a1 . ( 1981~).<br />

Common<br />

Rare<br />

Occasional<br />

Uncommon<br />

Uncommon<br />

Occasional<br />

Abundant<br />

Qccas ional<br />

Occasional<br />

Occasfonal<br />

Occasional<br />

Camon<br />

Rare


APPENDIX M<br />

OWNERSHIP OF MAJOR LAKE ST, CLAIR WETLANDS<br />

Wetland Name Re1 a t i ve Water-1 eval Veg .<br />

Pub1 i c Private areab controlC typed<br />

-- ---<br />

WAYNE COUNTY, MICHIGAN<br />

I. Belle Isle M/ F<br />

MACOMB COUNTY* MICHIGAN<br />

2, Black Creek -<br />

3. Clinton Rtver Estuary M<br />

4. Betvidere Bay -<br />

5. Salt River Estuary -<br />

ST. CLAJR COUNTY, MICHIGAN<br />

6. Marsac Paint .<br />

7, Swan Cresk Estuary -<br />

8. <strong>St</strong>. Johnts Marsh S<br />

9. Dickinson Island S/F<br />

10. Harsens Is1 and S<br />

11. Bslle Rfvar Estuary -<br />

12. Pine River Estuary M<br />

13, Marysvi 11 e -<br />

14, Port Huron -<br />

ESSEX COUNN, ONTARIO<br />

25, Peach Island<br />

16. Ruscom River<br />

17, Thames River Estuary<br />

KENT COUNTY p ONTARIO<br />

18, Bradley Marsh<br />

19. BJg P<strong>of</strong>nt Marsh<br />

20, Tacky Marsh<br />

21. MJtchell Pafnt Marsh<br />

(Moon Island Marsh)<br />

22. Mitchell Bay<br />

CPlntal1 and Mud Creek<br />

marshes9<br />

23, Val 1 aceburg<br />

(Bear Creek and<br />

Pigeon marshes9


APPENDIX M (CONCLUDED)<br />

Wetland Name Re1 at i ve Water-1 evel Veg .<br />

Public Private area control type<br />

LAMBTON COUNTY, ONTARIO<br />

24. <strong>St</strong>. Anne Is1 and<br />

25, Wal pol e Is1 and<br />

26. Squirrel Island<br />

27. Bassett Island<br />

28. Seaway Island<br />

29. Port Lambton<br />

30. Fawn Island<br />

31. <strong>St</strong>ag Island<br />

-<br />

-<br />

32, Sarnia M<br />

a<br />

Ownership Key: b~rea Key: dVeg. Type Key:<br />

F = Federal S = Small (>500 ha) S = Submergent<br />

S = <strong>St</strong>ate or Provincial M = Medium (500-lrOOO ha) E = Emergent<br />

M = Municipal<br />

SC = Shooting Club<br />

L = Large (>l,000 ha)<br />

PM = Private, multiple owners CWater-1 eve1 Control Key:<br />

PS = Private, single owner D = Diked, managed marsh<br />

R = Indian Reservation U = Uncontroll ed wet1 and<br />

'~ote: Between Bradley Marsh (18) and Mitchell Bay (22) several shooting<br />

clubs operate small marshes, including Recess Club, Balmoral Club, <strong>St</strong>. Lukes<br />

Clubr Rex Club, Bay Loge Club* Rex/Cadotte Marsh, and Rankin/Sloan Marsh.


<strong>The</strong> <strong>Ecology</strong> <strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong> <strong>Wetlands</strong>:<br />

A Communi ty Pr<strong>of</strong> i 1 e<br />

I R eW Dote<br />

September 1986<br />

b Ibr(.)<br />

Charles E. Herdendorf ,I C. Nicholas Raphael, Eugene ~aworski<br />

6. P<strong>of</strong>formln(l O~anlzotlon Rem. No<br />

Authors' Affiliations: la CmimctfTesk/Work Unit NO.<br />

lCenter for <strong>Lake</strong> Erie Area Research<br />

<strong>The</strong> Ohio <strong>St</strong>ate University<br />

Columbus, OH 43210<br />

U w m O~llaniatlon Nome and Address<br />

National <strong>Wetlands</strong> Research Center<br />

Fish and Wildlife Service<br />

U.S. Department <strong>of</strong> the Interior<br />

Washington, DC 20240<br />

Geography<br />

and Geology 11. Controct(C) w Grnnt(O) No.<br />

Eastern Michigan University ,<br />

Ypsilanti, MI 48197<br />

la Abstnct (Limit: loo words) I<br />

This publication reviews the ecological data and information on the wetlands<br />

<strong>of</strong> <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong>. <strong>The</strong> lake, whicn is a part <strong>of</strong> the Great <strong>Lake</strong>s ecosystem, is<br />

situated between southeastern Michigan and southwestern Ontario. <strong>Lake</strong> <strong>St</strong>.<br />

<strong>Clair</strong> contains a substantial portion <strong>of</strong> the remaining Great <strong>Lake</strong>s coastal<br />

wetlands, with extensive wetlands occurring in the vicinity af the <strong>St</strong>. <strong>Clair</strong><br />

River Delta. <strong>The</strong>se wetlands are among the most productive areas in the Great<br />

<strong>Lake</strong>s ecosystem. This publication summarizes the geologic history <strong>of</strong> the<br />

region leading to the development <strong>of</strong> wetlands and the present environment;<br />

biological production and community organization in the <strong>Lake</strong> <strong>St</strong>. <strong>Clair</strong><br />

wetlands are also examined. <strong>The</strong> publication closes with a chapter on applied<br />

ecology which addresses issues such as wetland disturbance, management, and<br />

future prospects for these wetlands.<br />

LI. Mllltlfian/O$wt.Ended Terms<br />

Nutrient dynamics<br />

lrophic interactions<br />

Invertebrates<br />

Birds<br />

Great <strong>Lake</strong>s<br />

Navigation<br />

Rnthropogenic i n f 1 uences<br />

nt j 9<br />

- ---<br />

. Security Class (Thts Rrwm 21. No <strong>of</strong> Psges<br />

Unclassified<br />

Unlimited distribution -- -<br />

I Swd ANS1-339.11))<br />

1 Unclassified<br />

22. Pncc<br />

I<br />

ORIOMAL FORM 272 (4-77)<br />

(Forwrly NJIS-35)<br />

Dapartment <strong>of</strong> ComWrC4

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!