Notes on the acanthaceae of Java - DWC

44 NOTES ON THE ACANTHACEAE OF JAVA
it was afterwards united by MOORE. It diHers. however. conspicuously
from that genus in the structure of the inflorescenee. the pollen grains and
the ,testa. In ENGL. u. PRANTL. Nat. Pflanzenfam. IV 3 b. p. 344. 1895.
LINDAU transferred two more species to this new genus. namely ]usticia
dichotoma BI. (syn.: Raphidospora dichotoma Nees) and Dianthera term inalis
Fawcet.
S. MOORE (in Joum. of Bot. LX. p. 355. 1922) was certainly not
ignorant of LINDAU's remarks on the taxonomie position of these species
when he referred ,them' to the section Euryantheae of the amplified genus
Ptyssiglottis. but he was apparently not sufficiently acquainted with the
structural peculiarities of the pollengrains to appreciate their taxonomie
value. The section Euryantheae comprised. apar,t from the type species of
the genus. Pt. radicosa (Nees) T. And .• the three species which LINDAU
had referred to Strophacanthus. further Leda subcordata Clarke and a
presumably new species. which he called Pt. Zollingeri. In basing the
latter on ZOLLINGER n. 2210. he apparently overlooked that NEEs had
founded his Rhaphidospora ten elia on a specimen of ZOLLINGER bearing
the same number and evidently identical with the specimen in the herbarium
of the British Museum on which he himself based his new species.
No example of this plant was available to me. but HOCHREUTINER (in
Candollea V. p. 242. 1935) aker a re-examination of the type came to
the conclusion that it must be regarded as a variety of Strophacanthus
dichotomus (BI.) Lindau (syn.: Raphidospora dichotoma Nees). differing
from the type of the latter in a somewhat greater length of the .ïnflorescence
and in the absence of the glandular pubescence on the rachis and
branchlets of the inflorescenee and on the calyx.
As the reduction of the genus Strophacanthus to Ptyssiglottis is in no
way justified. the genus is to be restored. The following species belong
to it:
*collinus (T. And.) Lindau in Engl.. Bot. Jahrb. XVIII. p. 58. 1893 et in
EngI. u. Prantl. Nat. Pflanzenfam. IV 3 b. p. 334. 1895 (syn.:
]usticia collina T. And. in Journ. Linn. Soc. IX. p. 515. 1867;
Dianthera collina (T. And.) Cl arke in Hook.f .• Fl. Brit. Ind. IV.
p. 543. 1885)
membranifolius (Miq.) Brem. n. combo (syn.: Rhaphidospora membrani­
[olia Miq .• FI. Ind. Bat. II. p. 850. 1857; - ]usticia dichotoma BI..
Bijdr. FI. Ned. Ind .• p. 783. 1826. nom. illeg. nam non RottI. in
Nov. Act. Acad. Nat. Cur. IV. p. 221. 1803 quae est Rhinacanthus
nasutus (L.) Lindau; Peristrophe dichotoma (BI.) Hassk. (comb.
illeg.). Cat. Hort. Bog. ed. 2. p. 152. 1844 et in Tijdschr. Nat.
Gesch. V. p. 129; Leptostachya dichotoma (BI.) Nees (comb.
illeg.) in DC .• Prodr. XI. p. 379. 1847; Rhaphidospora dichotoma
(BI.) Nees (comb. illeg.) op. cito p. 500. specimine filippino excl.
quod. ad Rh. luzonensem (Clarke) Brem. ,no combo (Justicia Clarke)
pertinet; Dianthera dichotoma (BI.) Clarke (comb. illeg.) in
Hook.f.. FI. Brit. Ind. IV. p. 543. 1885; Strophacanthus dichotomus