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IDY TE1>t' I- - American Museum of Natural History

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1970<br />

controlled by the nature <strong>of</strong> the underlying<br />

object, and about equal numbers <strong>of</strong> specimens<br />

represent concave, flat, and convex external<br />

pr<strong>of</strong>iles. The umbonal region <strong>of</strong> the left valve<br />

varies from slightly convex to moderately convex,<br />

and the beak area rises only slightly above<br />

the cardinal area in a hinge view. The principal<br />

growth gradient <strong>of</strong> the left valve umbonal area<br />

is commonly downward (infracrescent), rarely<br />

forward (procrescent). Commonly, the beak is<br />

situated medially along the hinge, and in numerous<br />

specimens it is slightly anterior <strong>of</strong> the midpoint.<br />

The subequal auricles are set apart from the<br />

main body <strong>of</strong> the valve by flattening and by<br />

subauricular sinuses <strong>of</strong> the valve margin; these<br />

indentations are essentially equal in a single<br />

valve. The vertical pr<strong>of</strong>ile beyond the attachment<br />

area tends to be slightly convex to flat in<br />

both left and right valves. The curvature <strong>of</strong> the<br />

NEWELL AND BOYD: PERMIAN BIVALVIA<br />

surface as viewed along the plane <strong>of</strong> commissure<br />

is constant, but some individuals exhibit a<br />

steeper posterior or anterior slope. Although the<br />

main body <strong>of</strong> the valve is approximately equilateral,<br />

some specimens are slightly extended<br />

forward or backward.<br />

Early growth <strong>of</strong> the right valve produced an<br />

infracrescent, unattached pectiniform stage<br />

(fig. 25H, I). This stage persisted to a height <strong>of</strong><br />

2 to 4.5 mm., beyond which the valve became<br />

cemented to the substrate (fig. 26C). The few<br />

specimens with a well-preserved umbonal area<br />

show a byssal notch in the pectiniform stage.<br />

The trace <strong>of</strong> the notch, delineated by growth<br />

lines, varies from 1 to 2 mm. long. In most specimens<br />

the pectiniform stage is not sharply differentiated<br />

from the adjacent part <strong>of</strong> the valve. In<br />

one specimen the hinge line <strong>of</strong> the pectiniform<br />

stage is along the edge <strong>of</strong> the triangular cardinal<br />

area and anterior to the apex.<br />

FIG. 26. Pegmavalvula gloveri Newell and Boyd, new species, Permian, Road Canyon Formation, Glass<br />

Mountains, western Texas. A, B. Left valve, U.S.N.M. No. 155006; U.S.N.M. 721j. C, D. Right valve,<br />

U.S.N.M. No. 154987, cemented at umbo to shell fragment; U.S.N.M. 721s. E. Shell fragment, U.S.N.M.<br />

No. 154988, showing cylindrical spines; U.S.N.M. 721j. F, G. Holotype, U.S.N.M. No. 154989, left valve;<br />

U.S.N.M. 726d. A-D, F, G, x 1; E, x 2.<br />

265

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