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(2009): Lichens in Scandinavia known mainly from Norwegian type ...

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GRAPHIS SCRIPTA 21 (<strong>2009</strong>) <strong>Lichens</strong> ma<strong>in</strong>ly <strong>known</strong> <strong>from</strong> <strong>Norwegian</strong> <strong>type</strong>s 10<br />

Lecidea hardangeriana H.Magn.<br />

Type: Norway, F<strong>in</strong>se, Hardangerjøkelen, alt.<br />

1250–1300 m, 1925, E. Frey 13606 (UPS!<br />

holo<strong>type</strong>). TLC: miriquidic acid.<br />

The poorly developed specimen shows close<br />

resemblance to Lecidea subplumbea Anzi<br />

accord<strong>in</strong>g to H. Hertel (pers. comm.), a name<br />

which most probably is a synonym of<br />

Miriquidica griseoatra, a rather variabel<br />

species <strong>in</strong> the poorly understood arctic-alp<strong>in</strong>e<br />

M. leucophaea complex (see Hertel &<br />

Rambold 1987). We can confirm that the<br />

holo<strong>type</strong> belongs <strong>in</strong> Miriquidica as it exhibits<br />

all the characters of that genus as def<strong>in</strong>ed by<br />

Hertel & Rambold (1987). More material is<br />

necessary to evaluate the taxonomy at species<br />

level, and we suspect this will prove to fall<br />

with<strong>in</strong> the variation of a more widespread<br />

taxon.<br />

Lecidea ileiformis Fr.<br />

Type: Norway, Dovre, M. N. Blytt (UPS!,<br />

holo<strong>type</strong>). TLC: Atranor<strong>in</strong>, psoromic and<br />

stictic acids, zeor<strong>in</strong>.<br />

This is a lecidioid member of the Lecanoraceae,<br />

and although it has a terricolous habitat, we<br />

regard it as a Calvitimela because of the<br />

<strong>in</strong>ternal characters of the apothecia. The spores<br />

(9–12 × 4–7 µm) falls with<strong>in</strong> the variation<br />

given by Haugan & Timdal (1994) for C.<br />

aglaea, but the chemistry differs. At the<br />

moment it is best regarded as a further<br />

chemo<strong>type</strong> of that species, but this variation is<br />

<strong>in</strong> need of further studies<br />

Lecidea <strong>in</strong>venusta H.Magn.<br />

Type: Norway, Granv<strong>in</strong>, near the top of<br />

Smøreggen, 1947, J. J. Havaas (UPS!,<br />

holo<strong>type</strong>). TLC: miriquidic acid.<br />

This <strong>type</strong> is <strong>in</strong> a rather poor condition and<br />

much eaten by snails. The rema<strong>in</strong><strong>in</strong>g thallus<br />

conta<strong>in</strong>s, however, miriqidic acid and the<br />

specimen certa<strong>in</strong>ly belongs <strong>in</strong> the Miriquidica<br />

leucophaea complex, just as Lecidea<br />

hardangeriana.<br />

Lecidea ludificans H.Magn.<br />

Type: Norway, Oppland, Jotunheimen, at the<br />

top of Galdhøpiggen, alt. 2468 m, 1947, G.<br />

Degelius (UPS!, holo<strong>type</strong>, Fig. 8). TLC:<br />

norstictic acid.<br />

This is a characteristic species with sh<strong>in</strong><strong>in</strong>g<br />

brown thallus and dist<strong>in</strong>ct, thick exciple (to<br />

about 100 µm) on the large, flat apothecia. It is<br />

certa<strong>in</strong>ly a member of the Lecidea praenubila<br />

complex, but stands out as most dist<strong>in</strong>ct among<br />

the many closely related, variable species (cf.<br />

Hertel 1977).<br />

It has not been recollected recently,<br />

certa<strong>in</strong>ly because of the rather <strong>in</strong>accessable<br />

locality on the top of Scand<strong>in</strong>avia’s highest<br />

mounta<strong>in</strong>. It is remarkable that two such rare<br />

species with arctic-alp<strong>in</strong>e relationships are<br />

found there (see also L. altissima above), a fact<br />

which strongly <strong>in</strong>dicates that this high<br />

mounta<strong>in</strong> was an ice-free nunatak, at least for<br />

parts of the quarternary glaciation. Curiously<br />

this has not been one of the peaks <strong>in</strong>cluded <strong>in</strong><br />

the heated debate among botanists and<br />

Figure 8. Lecidea ludificans, part of holo<strong>type</strong>.<br />

Bar = 2 mm.

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