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(2009): Lichens in Scandinavia known mainly from Norwegian type ...

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GRAPHIS SCRIPTA 21 (<strong>2009</strong>) <strong>Lichens</strong> ma<strong>in</strong>ly <strong>known</strong> <strong>from</strong> <strong>Norwegian</strong> <strong>type</strong>s 2<br />

theca hygrophila, Erioderma pedicellatum and<br />

Szcaw<strong>in</strong>skia leucopoda.<br />

Norway surpris<strong>in</strong>gly also has the driest,<br />

most cont<strong>in</strong>ental regions <strong>in</strong> Fennoscandia,<br />

those of the <strong>in</strong>ner, dry valleys of Opland and<br />

adjacent counties, <strong>in</strong> the ra<strong>in</strong> shadow of<br />

Jotunheimen (Ahlner 1949, Kleiven 1959),<br />

which conta<strong>in</strong> xerophilous steppe-elements, not<br />

found elsewhere <strong>in</strong> our region, e.g. Buellia<br />

asterella, Caloplaca tom<strong>in</strong>ii, Fulgensia<br />

desertorum, Gyalidea asteriscus, Phaeorrhiza<br />

sareptana, R<strong>in</strong>od<strong>in</strong>a terrestris, Squamar<strong>in</strong>a<br />

magnussonii, Ton<strong>in</strong>ia sculpturata and T.<br />

taurica.<br />

Norway reaches further north than any other<br />

of the Fennoscandian countries and has the<br />

highest mounta<strong>in</strong>s. This is reflected <strong>in</strong> arcticalp<strong>in</strong>e<br />

species, such as Arctocetraria andrejevii,<br />

Asah<strong>in</strong>ea chrysantha, Buellia adjuncta, Lobothallia<br />

alphoplaca and Rhizocarpon d<strong>in</strong>othetes.<br />

F<strong>in</strong>ally there rema<strong>in</strong>s a residue of rare<br />

species which does not necessarily fall <strong>in</strong>to<br />

these categories. They are all described <strong>from</strong><br />

Norway and have rema<strong>in</strong>ed <strong>known</strong> only, or<br />

ma<strong>in</strong>ly, <strong>from</strong> the <strong>type</strong>-collections. They<br />

therefore need particular attention, requir<strong>in</strong>g<br />

reexam<strong>in</strong>ation, preferably on freshly collected<br />

material to ascerta<strong>in</strong> their taxonomic and<br />

conservation status. There are two possible<br />

ma<strong>in</strong> reasons for their rarity:<br />

1. They are parts of poorly understood species<br />

complexes <strong>in</strong> difficult, not fully understood<br />

genera.<br />

2. They are genu<strong>in</strong>ely rare or overlooked.<br />

From a conservation po<strong>in</strong>t of view it is<br />

important to s<strong>in</strong>gle out the truly rare species<br />

with their only Scand<strong>in</strong>avian occurrence <strong>in</strong><br />

Norway, and to see if they represent phytogeographical<br />

elements to which also other rare<br />

lichens belong.<br />

We are not <strong>in</strong> a position to resolve the<br />

matter fully as we have been unable to secure<br />

the necessary new material for study, but we<br />

have prepared a commented list on them,<br />

hopefully <strong>in</strong>spir<strong>in</strong>g other colleagues to make an<br />

attempt to resolve the rema<strong>in</strong><strong>in</strong>g problems,<br />

which are long overdue.<br />

We have as far as possible studied the <strong>type</strong><br />

specimens microscopically and chemically<br />

(TLC). However, some <strong>type</strong>s are so m<strong>in</strong>ute that<br />

we have not dared to take more bits,<br />

particularly if studies of spores or chemistry<br />

would not be taxonomically important.<br />

List of species<br />

Acarospora impressula Th.Fr.<br />

Type: Norway, Akershus, L<strong>in</strong>døya, 1869, N.G.<br />

Moe (UPS!, lecto<strong>type</strong>, here selected, Fig. 1).<br />

TLC: no substancees detected.<br />

This is a rather nondescript species <strong>in</strong> the<br />

Acarospora badiofusca group. Magnusson<br />

(1929) po<strong>in</strong>ts out the dark, cont<strong>in</strong>uous, areolate<br />

crust with a negative C-reaction, the immersed<br />

apothecia, and the broadly ellipsoid spores. The<br />

collector of the orig<strong>in</strong>al material has noted that<br />

it grew only on dry rocks warmed by the<br />

afternoon sun, which is an <strong>in</strong>dication that it is<br />

one of the southern thermophilous elements,<br />

restricted to the Oslofjord-region (e.g. Anema)<br />

or also present <strong>in</strong> SE Sweden. This might<br />

account for its rarity, but the world distribution<br />

of this species does not confirm this. In Central<br />

Europe it is recorded as a high montane species<br />

Figure 1. Acarospora impressula, part of<br />

lecto<strong>type</strong>. Bar = 1 mm.

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