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PLIENSBACHIAN (LOWER JURASSIC) BIOSTRATIGRAPHY AND ...

PLIENSBACHIAN (LOWER JURASSIC) BIOSTRATIGRAPHY AND ...

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Type species. Leptaleoceras leptum Buckman, 1918<br />

(p. 284, PI. 26, fig. la, b) by original designation.<br />

Rem arks. Shells evolute; whorl section compressed,<br />

elliptical. Flanks convex, ventrolateral shoulder<br />

rounded. Venter narrow, carinate; keel flanked by<br />

narrow flat zones that become sulcate on the body<br />

chamber of some species.<br />

Ornamentation consists of dense, slightly sinuous<br />

ribs that arise at or just above the umbilical shoulder,<br />

trend more or less rectiradiately on the flanks, then<br />

fade out at the ventrolateral shoulder where they<br />

project slightly. Innermost whorls, up to diameters of<br />

10 mm, are smooth.<br />

Leptaleoceras is distinguished from Arieticeras by<br />

its greater rib density and the nonsulcate venter<br />

observed on all but the body chamber of some species<br />

of Leptaleoceras [e.g., L. accuratum and L. insigne<br />

(Braga, 1983)].<br />

A g e and distribution. Leptaleoceras is primarily a<br />

Tethyan form but is occasionally found in the Boreal<br />

Realm. In Europe it is found in the middle Upper<br />

Pliensbachian, in association with A rie tic e ra s<br />

algovianum . In western North America, Leptaleoceras<br />

is found associated with Arieticeras and Fanninoceras,<br />

indicating a Late Pliensbachian age (Smith et al.,<br />

1988).<br />

Leptaleoceras sp. aff. L. accuratum (Fucini, 1931)<br />

Plate 13, figures 1-6<br />

aff. *1931 Arieticeras(7) accuratum Fucini, p. 107,<br />

PI. 8, fig. 10.<br />

aff. 1934 A rie tic e ra s p se u d o ra d ia n s (Reynes).<br />

Monestier, p. 63, PI. 8, fig. 62; non<br />

figs. 61, 68.<br />

1964a Leptaleoceras pseudoradians (Reynes).<br />

Frebold, p. 15, PI. 4, non PI. 5, figs. 4, 5.<br />

1970 Leptaleoceras pseudoradians (Reynes).<br />

Frebold, p. 443, PI. 2, fig. 2 (1964<br />

material partly refigured)<br />

aff. 1980 A r i e t i c e r a s a c c u r a t u m (F u c in i).<br />

Wiedenmayer, p. 109, PI. 17, figs. 15-18.<br />

1981 L e p ta le o c e ra s cf. L . p s e u d o r a d ia n s<br />

(Reynes). Imlay, p. 40, PI. 11, figs. 12, 13.<br />

71981 Arieticeras cf. A . domarense (Meneghini).<br />

Imlay, p. 39, PI. 10, fig. 15; non figs. 1, 2,<br />

6-14, 22.<br />

aff. 1983 Leptaleoceras accuratum (Fucini). Braga,<br />

p. 256, PI. 11, figs. 27-29; PI. 12,<br />

figs. 1-10.<br />

1988 Leptaleoceras aff. accuratum (Fucini).<br />

Smith et al., PI. 4, fig. 9.<br />

Material. Eight specimens, poorly to moderately well<br />

preserved as external and internal moulds in calcareous<br />

concretion matrix.<br />

Measurements.<br />

Specim en D UD U W H W H D WW W W D W W W H P R H W<br />

G SC 83106 38.5 17 44 a 13 34 __ _ — 26<br />

G SC 83107 39 19 49 a 13 33 - — - 25<br />

G SC 83109 a25 alO 40 a9 36 - — — a25<br />

G SC 83110 31.5 14 44 11 35 — — - 27<br />

G SC 83111 a70 33 47 a 18 26 a8 — — 22<br />

G SC 84967 a45.5 a l9 42 a l 5 33 a5 - — 27<br />

M ean — _ 44.3 — 32.8 _ _ ___ ___<br />

Si. Dev. — 3.3 — 3.5 — — — —<br />

D escription. Evolute, whorl section compressed,<br />

ogival. Umbilical wall low, shallow; umbilical shoulder<br />

rounded. Flanks slightly convex; ventrolateral shoulder<br />

rounded. Venter bears a distinct keel flanked by two,<br />

narrow, flat zones or slight sulci in some larger<br />

specimens.<br />

Ornamentation varies with growth. The innermost<br />

whorls are smooth to a diameter of 3 to 4 mm. The<br />

middle whorls bear fine, dense, sinuous ribs that have<br />

a slightly rursiradiate overall trend. On the upper part<br />

of the flanks the ribs project strongly adorally. At<br />

diameters greater than about 35 mm (or UD =<br />

13-15 mm) ribbing becomes less dense, more sinuous,<br />

and more rursiradiate.<br />

Discussion. In his discussion of L. accuratum, Braga<br />

(1983) defined a new morphotype, L. accuratum<br />

" p re a ccu ra tu m ". This morphotype, which has the<br />

same stratigraphic range, differs from typical forms of<br />

L. accuratum by its slightly more densely ribbed inner<br />

whorls and more sinuous ribs that maintain their<br />

sinuosity with growth, rather than becoming straighter,<br />

as seen in L. accuratum proper.<br />

The Spatsizi specimens of L. sp. aff. L. accuratum,<br />

together with the other North American material in the<br />

synonomy, compare well with the llpreaccuratum "<br />

material illustrated by Braga (1983), the only difference<br />

being the slightly greater rib density of the North<br />

American specimens (Fig. 16). Previous assignment of<br />

the North American material to L. pseudoradians<br />

(Frebold, 1964a, 1970; Imlay, 1981) is questioned here<br />

due to the significantly greater rib density, maintained<br />

throughout growth, of L. pseudoradians (Fig. 15).<br />

Braga (1983) does note, however, that there is a

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