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Piping Plover - U.S. Fish and Wildlife Service

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North Carolina, in 2005-2006, <strong>and</strong> piping plover activity was concentrated in 12 areas<br />

totaling 2.2 km 2 (Cohen et al. 2008). Noel <strong>and</strong> Ch<strong>and</strong>ler (2008) observed high fidelity of<br />

b<strong>and</strong>ed piping plovers to 1 km - 4.5 km sections of beach on Little St. Simons Isl<strong>and</strong>,<br />

Georgia.<br />

WM 2.2.1.2 Survival:<br />

The most consistent finding in the various population viability analyses (PVAs)<br />

conducted for piping plovers (Ryan et al. 1993, Melvin <strong>and</strong> Gibbs 1996, Plissner <strong>and</strong><br />

Haig 2000, Wemmer et al. 2001, Larson et al. 2002, Calvert et al. 2006, Brault 2007) is<br />

the sensitivity of extinction risk to even small declines in adult <strong>and</strong>/or juvenile survival<br />

rates.<br />

Efforts to partition survival within the annual cycle are beginning to receive more<br />

attention, but current information remains limited. Drake et al. (2001) observed no<br />

mortality among 49 radio-marked piping plovers (total of 2,704 transmitter days) in<br />

Texas in 2007-2008. Cohen et al. (2008) documented no mortality of radio-tracked<br />

wintering piping plovers at Oregon Inlet in 2005-2006, but this was based on only seven<br />

individuals. Analysis of South Carolina resighting data for 87 b<strong>and</strong>ed piping plovers<br />

(78% Great Lakes breeders) in 2006-2007 <strong>and</strong> 2007-2008 found 100% survival from<br />

December to April 1 (J. Cohen, Virginia Polytechnic Institute <strong>and</strong> State University, pers.<br />

comm. 2009 ). Noel et al. (2007) inferred two winter (November to February)<br />

mortalities 2 among 21 b<strong>and</strong>ed (but not radio-tagged) overwintering piping plovers in<br />

2003-2004 <strong>and</strong> nine mortalities among 19 overwintering birds during the winter of 2004-<br />

2005 at Little St. Simons Isl<strong>and</strong>, Georgia. LeDee (2008) found higher apparent survival 3<br />

rates during breeding <strong>and</strong> southward migration than during winter <strong>and</strong> northward<br />

migration for 150 adult (i.e., after-hatch year) Great Lakes piping plovers.<br />

Mark-recapture analysis of resightings of uniquely b<strong>and</strong>ed piping plovers from seven<br />

breeding areas by Roche et al. (2009) found that apparent adult survival declined in four<br />

populations <strong>and</strong> increased in none over the life of the studies 4 . Some evidence of<br />

correlation in year-to-year fluctuations in annual survival of Great Lakes <strong>and</strong> eastern<br />

Canada populations, both of which winter primarily along the southeastern U.S. Atlantic<br />

1 However, of those birds, one unique <strong>and</strong> one non-uniquely b<strong>and</strong>ed piping plover were seen in the first<br />

winter <strong>and</strong> were resighted multiple times in the second fall at the same location but were not seen during<br />

the second winter; whether these two birds died in the fall or shifted their wintering location is unknown<br />

(Maddock et al. 2009).<br />

2 Noel et al. (2007) inferred mortality if a uniquely b<strong>and</strong>ed piping plover with multiple November to<br />

February sightings on the survey site disappeared during that time <strong>and</strong> was never observed again in either<br />

its nonbreeding or breeding range. Note that most of these birds were from the Great Lakes breeding<br />

population, where detectability during the breeding season is very high.<br />

3 “Apparent survival” does not account for permanent emigration. If marked individuals leave a survey<br />

site, apparent survival rates will be lower than true survival. If a survey area is sufficiently large, such<br />

that emigration out of the site is unlikely, apparent survival will approach true survival.<br />

4 Data were analyzed for 3-11 years per breeding area, all between 1998 <strong>and</strong> 2008.<br />

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