Biodiversity, carbon storage and dynamics of old northern ... - BPAN.fi
Biodiversity, carbon storage and dynamics of old northern ... - BPAN.fi
Biodiversity, carbon storage and dynamics of old northern ... - BPAN.fi
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Figure 3.1. Disturbance factors at various spatial <strong>and</strong> temporal scales in boreal<br />
forests. The spatial scale will run from single trees to several st<strong>and</strong>s or parts <strong>of</strong><br />
catchment areas <strong>and</strong> the temporal from sub-annual to centuries<br />
Time scale<br />
Often Rare<br />
Rot + wind,<br />
competition,<br />
scenesence<br />
Flooding,<br />
erosion<br />
Wind-throws,<br />
droughts<br />
Insect attacks,<br />
large storms<br />
Severe <strong>fi</strong>res<br />
Small<br />
Spatial scale<br />
Large<br />
Modi<strong>fi</strong>ed from Jonsson & Siitonen 2012.<br />
Shorohova et al. (2011) summarized current underst<strong>and</strong>ing <strong>of</strong> natural<br />
disturbance regimes <strong>and</strong> the related <strong>dynamics</strong> determining the natural<br />
prevalence <strong>and</strong> development <strong>of</strong> <strong>old</strong>-growth boreal forests (Table 3.1).<br />
They concluded that in contrast to forests in western North America <strong>and</strong><br />
western Siberia, boreal forests <strong>of</strong> northwest Europe tend to be dominated<br />
by less severe, non-st<strong>and</strong>-replacing <strong>fi</strong>res <strong>and</strong> other disturbances, letting<br />
more varied <strong>and</strong> smaller scale disturbances shape forest development <strong>and</strong><br />
structure, between the rather infrequent severe, st<strong>and</strong>-replacing <strong>fi</strong>res.<br />
Forests are then mainly dominated by small-scale gap <strong>dynamics</strong>, with<br />
cohort <strong>dynamics</strong> where <strong>fi</strong>res <strong>of</strong> medium severity occur, giving a complex<br />
<strong>and</strong> <strong>fi</strong>ne-grained forest structure. As the proportion <strong>of</strong> <strong>old</strong>-growth forests<br />
in a l<strong>and</strong>scape is determined by the frequency <strong>and</strong> severity <strong>of</strong> recurrent<br />
st<strong>and</strong>-replacing disturbances (<strong>fi</strong>res, storms, insect outbreaks), the low<br />
frequency between severe disturbances in boreal forests in Fennosc<strong>and</strong>ia<br />
should imply that <strong>old</strong>-growth, multi-aged forests may originally have covered<br />
a considerable area <strong>and</strong> lasted for rather a long time. In western <strong>and</strong><br />
<strong>northern</strong> Norway, the broken topography <strong>and</strong> moister climate should<br />
have made large, st<strong>and</strong>-replacing <strong>fi</strong>res very rare, but local storm damage<br />
<strong>and</strong> avalanches would probably have been quite common.<br />
Studies <strong>of</strong> natural forest <strong>dynamics</strong> in the Nordic countries have been<br />
concentrated to boreal forests (mainly <strong>of</strong> the <strong>northern</strong> <strong>and</strong> middle boreal<br />
zones), with little speci<strong>fi</strong>c information on the natural <strong>dynamics</strong> <strong>of</strong><br />
temperate forests (Peterken 1996). This is mainly due to the long-lasting<br />
<strong>and</strong> extensive human influence in temperate forests (cf below), leaving<br />
virtually no suitable cases for studies <strong>of</strong> such <strong>dynamics</strong>, although some<br />
selected forest tracts with long forest continuity can be found, e.g., the<br />
<strong>Biodiversity</strong>, <strong>carbon</strong> <strong>storage</strong> <strong>and</strong> <strong>dynamics</strong> <strong>old</strong> <strong>northern</strong> forests 29