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Floodplain hay meadows along the river Tisza in Hungary

Floodplain hay meadows along the river Tisza in Hungary

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extensive cattle farm<strong>in</strong>g, fish<strong>in</strong>g (e.g. of <strong>the</strong> traditional species “csík”<br />

(Barbatula spp.)), catch<strong>in</strong>g crabs and also by trad<strong>in</strong>g crops and livestock<br />

(Andrásfalvy, 2007; Frisnyák, 1990).<br />

In addition to <strong>the</strong> local farm<strong>in</strong>g system, transhumance also existed.<br />

In <strong>the</strong> process of extensive livestock farm<strong>in</strong>g, <strong>the</strong> grey cattle were<br />

driven from <strong>the</strong> summer pastures to <strong>the</strong> w<strong>in</strong>ter pastures of <strong>the</strong><br />

floodpla<strong>in</strong>, which was covered by a more productive sward (Andrásfalvy,<br />

2007). Pigs, liv<strong>in</strong>g on <strong>the</strong> <strong>river</strong><strong>in</strong>e <strong>meadows</strong> and marshes dur<strong>in</strong>g<br />

summer like “semi-wild” boars, spent <strong>the</strong> w<strong>in</strong>ter <strong>in</strong> <strong>the</strong> vast oak<br />

woodlands of <strong>the</strong> upper reaches of <strong>the</strong> <strong>Tisza</strong>, where <strong>the</strong>y fattened<br />

on <strong>the</strong> acorns to reach <strong>the</strong>ir sell<strong>in</strong>g weight (Bellon, 2003). From <strong>the</strong><br />

neighbour<strong>in</strong>g mounta<strong>in</strong>s, large herds were also transferred to <strong>the</strong><br />

Great Pla<strong>in</strong> after <strong>the</strong> summer.<br />

At <strong>the</strong> beg<strong>in</strong>n<strong>in</strong>g of <strong>the</strong> 19 th century, <strong>the</strong> need for <strong>the</strong> expansion of<br />

arable land <strong>in</strong>creased as a result of <strong>the</strong> rise <strong>in</strong> crop prices and <strong>the</strong><br />

growth <strong>in</strong> population. This land was <strong>in</strong>itially obta<strong>in</strong>ed by <strong>the</strong> prevention<br />

of regular flood<strong>in</strong>g of <strong>the</strong> <strong>meadows</strong> and marshes. This led to<br />

<strong>the</strong> dry<strong>in</strong>g out and rapid sal<strong>in</strong>isation of <strong>meadows</strong> <strong>in</strong> many areas. The<br />

vast dra<strong>in</strong>ed marshes and fens were ploughed. A negative effect of<br />

keep<strong>in</strong>g <strong>the</strong> water out of <strong>the</strong>se fields was <strong>the</strong> <strong>in</strong>crease of <strong>the</strong> annual<br />

floods <strong>in</strong> <strong>the</strong> area close to <strong>the</strong> <strong>river</strong>.<br />

Build<strong>in</strong>g high levees<br />

Moreover, because of woodland clearance <strong>in</strong> <strong>the</strong> mounta<strong>in</strong>s catchments<br />

of <strong>the</strong> <strong>river</strong> bas<strong>in</strong>, <strong>the</strong> Great Pla<strong>in</strong> received a greater volume<br />

of water at a faster rate. To solve <strong>the</strong>se problems, high levees were<br />

built from 1884 onwards, with a f<strong>in</strong>al total length of 3000 km (Dunka<br />

et al., 1996). These embankments divided <strong>the</strong> floodpla<strong>in</strong> <strong>in</strong>to a “floodfree”<br />

area (<strong>in</strong>ner-dike area) and <strong>the</strong> “floodway”(outer-dike area),<br />

which is regularly <strong>in</strong>undated. Due to <strong>the</strong> levees be<strong>in</strong>g built close to<br />

<strong>the</strong> <strong>river</strong>bed and <strong>the</strong> isolation of <strong>the</strong> former <strong>meadows</strong> and marshes<br />

from <strong>river</strong> floods, numerous habitats became considerably degraded<br />

and, <strong>in</strong> addition, <strong>the</strong> biodiversity of <strong>the</strong>se regions also substantially<br />

decreased. Careless alteration of land use has led to <strong>the</strong> accumulation<br />

of seem<strong>in</strong>gly unsolvable economic and social problems (Mihók et al.,<br />

2006; Rakonczay, 2002). As a result of water management, common<br />

pastures had been parcelled by <strong>the</strong> end of <strong>the</strong> 19 th century and <strong>the</strong><br />

traditional transhumance <strong>in</strong> <strong>the</strong> floodpla<strong>in</strong> <strong>meadows</strong> also ceased. The<br />

extensive farm<strong>in</strong>g us<strong>in</strong>g <strong>the</strong> ancient grey cattle decl<strong>in</strong>ed significantly,<br />

and <strong>the</strong> “semi-wild” pig breeds also disappeared (Andrásfalvy, 2007).<br />

Fur<strong>the</strong>r dra<strong>in</strong><strong>in</strong>g and <strong>in</strong>tensification<br />

By <strong>the</strong> 1960’s, even <strong>the</strong> rema<strong>in</strong><strong>in</strong>g <strong>in</strong>land waters on previous floodpla<strong>in</strong>s<br />

beyond <strong>the</strong> levees had been dra<strong>in</strong>ed. The arid grasslands,<br />

hav<strong>in</strong>g replaced <strong>the</strong> dried-out <strong>meadows</strong>, had been mostly ploughed<br />

by that time. This situation was worsened with <strong>the</strong> <strong>in</strong>troduction of<br />

<strong>the</strong> s<strong>in</strong>gle-crop system established on vast fields, toge<strong>the</strong>r with <strong>the</strong><br />

<strong>in</strong>creased use of chemicals and fertilizers. The replacement of <strong>the</strong><br />

floodpla<strong>in</strong> <strong>meadows</strong> and native woodlands with plantations of alien<br />

species (hybrids of Populus canadensis, Frax<strong>in</strong>us pennsylvanica and<br />

Acer segundo) and <strong>the</strong> establishment of large-scale cultivation on<br />

<strong>the</strong> floodway also commenced at this time. However, an <strong>in</strong>terest<strong>in</strong>g<br />

dualism can be observed concern<strong>in</strong>g <strong>the</strong> utilisation of <strong>the</strong> floodway:<br />

on floodpla<strong>in</strong> <strong>meadows</strong> and pastures extensive, small-scale peasant<br />

farm<strong>in</strong>g (graz<strong>in</strong>g, mow<strong>in</strong>g, etc.) survived until <strong>the</strong> 1980’s, simultaneously<br />

with <strong>the</strong> socialist large-scale farm<strong>in</strong>g and forestry system. The<br />

reason was that woodlands and wooded <strong>hay</strong>fields smaller than 50<br />

ha were placed under <strong>the</strong> ownership of <strong>the</strong> farmers’ co-operatives<br />

<strong>in</strong>stead of forestry. Thus, <strong>the</strong>ir traditional, extensive utilisation may<br />

have played a significant role <strong>in</strong> <strong>the</strong> preservation of <strong>river</strong><strong>in</strong>e species<br />

and biodiversity.<br />

Although <strong>in</strong> <strong>the</strong> second half of <strong>the</strong> 20th century most floodpla<strong>in</strong><br />

<strong>meadows</strong> ly<strong>in</strong>g <strong>in</strong> <strong>the</strong> floodway were converted <strong>in</strong>to <strong>hay</strong>fields, grazed<br />

pastures also rema<strong>in</strong>ed until <strong>the</strong> 1980’s. The treeless grasslands, <strong>the</strong><br />

wooded pastures and <strong>hay</strong>fields and even <strong>the</strong> grass layer of <strong>the</strong> sparse<br />

woodlands, were grazed and mown. The graz<strong>in</strong>g and mow<strong>in</strong>g pattern<br />

of <strong>the</strong> grasslands was determ<strong>in</strong>ed by <strong>the</strong> diverse topography, <strong>the</strong><br />

water regime and <strong>the</strong> climatic conditions.<br />

Recent abandonment of <strong>the</strong> <strong>river</strong> valley<br />

In <strong>the</strong> early 1990’s <strong>the</strong> socio-economic changes (re-privatisation,<br />

decreased support for animal husbandry) considerably altered <strong>the</strong><br />

use of <strong>the</strong> floodway grasslands. Subsidisation of animal husbandry<br />

reached its m<strong>in</strong>imum level and as a consequence less milk, meat and<br />

wool was produced. S<strong>in</strong>ce <strong>the</strong> local market system had not been developed<br />

by that time due to <strong>the</strong> previous central control, <strong>the</strong> number<br />

of livestock dramatically and rapidly decreased <strong>in</strong> <strong>the</strong> settlements<br />

<strong>along</strong> <strong>the</strong> <strong>river</strong>. The management of <strong>the</strong> <strong>meadows</strong> and wooded pastures<br />

of <strong>the</strong> floodway ceased and this has resulted <strong>in</strong> <strong>the</strong> slow <strong>in</strong>vasion<br />

by shrubs, which is consistent with <strong>the</strong>ir orig<strong>in</strong> from woodland<br />

clearance. At <strong>the</strong> same time, toge<strong>the</strong>r with <strong>the</strong> abandonment of <strong>the</strong><br />

pastures, <strong>the</strong> cultivation of several arable fields also ceased, and<br />

<strong>the</strong>se areas were <strong>in</strong>vaded <strong>in</strong> a few years by <strong>the</strong> previously <strong>in</strong>troduced<br />

Amorpha fruticosa. The sudden <strong>in</strong>crease <strong>in</strong> <strong>the</strong> amount of propagules,<br />

toge<strong>the</strong>r with <strong>the</strong> high-level floods of <strong>the</strong> late 1990’s, led to <strong>the</strong> serious<br />

<strong>in</strong>festation of all <strong>the</strong> grasslands, wooded <strong>hay</strong>fields and woodlands.<br />

The swiftly spread<strong>in</strong>g Amorpha fruticosa destroys <strong>the</strong> natural<br />

vegetation by <strong>the</strong> deep shad<strong>in</strong>g of its dense population as well as by<br />

means of allelopathy (Szigetvári and Tóth, 2002). When <strong>the</strong> <strong>in</strong>dividuals<br />

<strong>in</strong>vad<strong>in</strong>g a certa<strong>in</strong> area reach <strong>the</strong> age of 4-5 years, <strong>the</strong>y form a<br />

closed canopy, and <strong>the</strong> grassland becomes featureless, <strong>the</strong> character<br />

species disappear, and only species resistant to shad<strong>in</strong>g can survive.<br />

The floodpla<strong>in</strong> <strong>hay</strong> <strong>meadows</strong> of <strong>the</strong> region<br />

Due to flood control, ancient floodpla<strong>in</strong> <strong>hay</strong> <strong>meadows</strong>, with a nearnatural<br />

water regime and water supply can only be found with<strong>in</strong> <strong>the</strong><br />

floodway of <strong>the</strong> <strong>river</strong> <strong>Tisza</strong>. <strong>Floodpla<strong>in</strong></strong> <strong>meadows</strong> ly<strong>in</strong>g beyond <strong>the</strong><br />

levees have been ei<strong>the</strong>r cultivated or have dried out, thus <strong>the</strong>y have<br />

become sal<strong>in</strong>ified and have changed <strong>in</strong>to Achillea steppes. Accord<strong>in</strong>g<br />

to recent studies, floodpla<strong>in</strong> <strong>meadows</strong> isolated from <strong>the</strong> <strong>river</strong> totally<br />

dry out and become sal<strong>in</strong>ised with<strong>in</strong> 50-60 years. They are replaced<br />

by dry, short grass-dom<strong>in</strong>ated steppes typically with Achilleo-Festucetum<br />

pseudov<strong>in</strong>ae vegetation (or Lythro-Alopecuraetum <strong>in</strong> regions of<br />

lower elevation), <strong>the</strong> soil of which accumulates alkali salts at depth<br />

(Molnár and Borhidi, 2003).<br />

In <strong>the</strong> Pleistocene and lower Holocene, <strong>the</strong> meander<strong>in</strong>g <strong>river</strong>s of <strong>the</strong><br />

Great Pla<strong>in</strong> deposited semi-circular po<strong>in</strong>t bars. These curved ridges<br />

can be observed on <strong>hay</strong> <strong>meadows</strong> anywhere <strong>in</strong> <strong>the</strong> floodpla<strong>in</strong> <strong>along</strong><br />

<strong>the</strong> <strong>Tisza</strong>. The narrow depressions between <strong>the</strong>se ridges, which support<br />

marshes and sedge-dom<strong>in</strong>ated communities, usually had a direct<br />

connection with <strong>the</strong> <strong>river</strong> <strong>in</strong> past centuries. Today, <strong>the</strong>se are covered<br />

by smaller or wider patches of marsh vegetation dom<strong>in</strong>ated by<br />

Glyceria maxima or Phalaris arund<strong>in</strong>acea large sedges such as Carex<br />

gracilis, and beds of Phragmites australis or Typha latifolia. The top of<br />

<strong>the</strong> curved ridges are covered by Peucedanum offic<strong>in</strong>ale dom<strong>in</strong>ated<br />

meadow-steppes, while on <strong>the</strong>ir slopes and at lower elevations, tall<br />

herb communities and different types of marshes have established.<br />

The <strong>meadows</strong> have properly adapted to <strong>the</strong> rapidly chang<strong>in</strong>g water<br />

supply and, with <strong>the</strong>ir dynamically chang<strong>in</strong>g species composition,<br />

<strong>the</strong>y tolerate both <strong>the</strong> longer humid and dry periods. Nowadays,<br />

<strong>the</strong>y are partially under nature conservation management, and <strong>the</strong><br />

authorities try to susta<strong>in</strong> <strong>the</strong>ir current state by mechanized mow<strong>in</strong>g.<br />

Under natural conditions, with <strong>the</strong> cessation of mow<strong>in</strong>g or graz<strong>in</strong>g,<br />

<strong>the</strong> floodpla<strong>in</strong> <strong>meadows</strong> are slowly encroached by <strong>the</strong> native <strong>river</strong><strong>in</strong>e<br />

trees (Salix alba, S. fragilis, S. purpurea, Populus alba, P. nigra, Quercus<br />

robur, Frax<strong>in</strong>us angustifolia ssp. pannonica, Ulmus campestris and U.<br />

laevis) or shrubs (Viburnum opulus, Euonymus europaeus, Sambucus<br />

nigra and Crataegus monogyna), as <strong>the</strong>y orig<strong>in</strong>ated from <strong>the</strong> cleared<br />

woodlands. This tendency was utilised by humank<strong>in</strong>d with <strong>the</strong> development<br />

of wooded pastures and <strong>hay</strong>fields, where some young trees<br />

were left to grow, establish<strong>in</strong>g a special landscape characteristic of<br />

<strong>the</strong> <strong>Tisza</strong> floodpla<strong>in</strong>. Due to <strong>the</strong> cessation of management, <strong>the</strong> condition<br />

of <strong>the</strong> wooded pastures and <strong>hay</strong>fields gradually decl<strong>in</strong>ed and<br />

<strong>the</strong>y are replaced by thickets of <strong>in</strong>troduced species over a relatively<br />

short period of time. Though <strong>the</strong>y have great value as examples of<br />

traditional land-use support<strong>in</strong>g high biodiversity, <strong>the</strong>ir preservation<br />

on <strong>the</strong> non-protected areas has not as yet been resolved.<br />

<strong>Floodpla<strong>in</strong></strong> mesotrophic <strong>meadows</strong><br />

<strong>Floodpla<strong>in</strong></strong> mesotrophic <strong>meadows</strong> were once <strong>the</strong> most frequent<br />

meadow type <strong>along</strong> <strong>the</strong> <strong>Tisza</strong>. Even today <strong>the</strong>y serve as a matrix for<br />

<strong>the</strong> floodpla<strong>in</strong> <strong>hay</strong> <strong>meadows</strong>. Under natural conditions, <strong>the</strong>y are wet<br />

dur<strong>in</strong>g <strong>the</strong> greater part of <strong>the</strong> year, and often <strong>in</strong>undated. They are<br />

characterised by meadow and marsh species and dom<strong>in</strong>ated by Alopecurus<br />

pratensis, Poa pratensis, Festuca pratensis, Dactylis glomerata<br />

and Carex melanostachya. The average height of <strong>the</strong> sward is about<br />

0,5-1 metre and species composition is determ<strong>in</strong>ed by <strong>the</strong> duration<br />

of water cover. The characteristic species are Clematis <strong>in</strong>tegrifolia,<br />

Galium rubioides, Cardam<strong>in</strong>e pratensis, Galega offic<strong>in</strong>alis, Gratiola offic<strong>in</strong>alis,<br />

Viola elatior, Rumex confertus, Thalictrum flavum, Thalictrum<br />

lucidum, Cirsium canum, Cichorium <strong>in</strong>tibus, Symphytum offic<strong>in</strong>ale, Iris<br />

pseudacorus, Galium palustre, Lysimachia vulgaris, Lythrum virgatum,<br />

L. salicaria, Ranunculus repens, Past<strong>in</strong>aca sativa and Lychnis flos-cucculi.<br />

Sometimes <strong>the</strong>y may have fen characteristics due to water logg<strong>in</strong>g.<br />

This is <strong>in</strong>dicated by <strong>the</strong> presence of species such as Serratula t<strong>in</strong>ctoria,<br />

Ophioglossum vulgatum and Gentiana pneumonan<strong>the</strong>. In <strong>the</strong> <strong>meadows</strong>,<br />

<strong>the</strong> characteristic colourful species <strong>in</strong>clude those that are often also<br />

typical of <strong>the</strong> willow-poplar woodlands, such as Leucojum aestivum.<br />

240<br />

241

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