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What Age is This Eagle? - American Birding Association

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Lake County,<br />

Florida; February<br />

2007. © Matthew<br />

Studebaker.<br />

Aremote video camera in<br />

Goldstream Provincial Park on Vancouver<br />

Island, Brit<strong>is</strong>h Columbia,<br />

recorded an unusual eagle as it foraged<br />

with Bald <strong>Eagle</strong>s of multiple age classes on<br />

8 December 2004. The unusual eagle was<br />

clearly larger and more massive than the<br />

nearby Bald <strong>Eagle</strong>s (Figs. 1 & 2, p. 30). On<br />

at least two occasions within a month after<br />

th<strong>is</strong> video was made, birders observed th<strong>is</strong><br />

unusual eagle perched and flying nearby.<br />

Most of the people who viewed<br />

th<strong>is</strong> eagle in the video and others who later saw it in<br />

person weren’t certain what it was they were seeing.<br />

Some thought that it was an unusual Bald, others felt<br />

that it was a Steller’s Sea-<strong>Eagle</strong>, and a few opted for a<br />

hybrid between the two.<br />

After viewing the video many times, including many<br />

stop-frames, and reading what observers have<br />

written about it, I’ve concluded that it wasn’t<br />

either of the eagle species, but a hybrid<br />

between them. Figs. 1–4 (p. 30) are stillframes<br />

taken from the video made by Darren<br />

Copley and first sent to me by David Allinson. Allinson<br />

had alerted me to the presence of the eagle. Comments<br />

on and links to pictures of th<strong>is</strong> unusual eagle were<br />

posted on the Frontiers of Bird ID and <strong>Birding</strong> Vancouver<br />

Island <br />

websites in December 2004. The material on these<br />

websites elicited comments from various persons, including<br />

those who saw the bird in person.<br />

<strong>What</strong> <strong>Age</strong> <strong>is</strong> Th<strong>is</strong> <strong>Eagle</strong><br />

Knowledge of th<strong>is</strong> eagle’s age <strong>is</strong> helpful in understanding<br />

what it <strong>is</strong>. Using a shorter video showing only the<br />

eagle in question, I was able to stop the frame often<br />

and look at the state of molt of the flight feathers. The<br />

resulting JPEG images were rather blurred but still usable.<br />

The eagle’s primaries show two waves of molt<br />

(the molt progression known as Stauffelmaüser), with<br />

P10 being a retained juvenal remex (Fig. 1). The bird’s<br />

secondaries show at least two ages of feathers. There-<br />

28<br />

B i r d i n g • J u l y / A u g u s t 2 0 0 8


fore, th<strong>is</strong> eagle <strong>is</strong> in its third annual plumage (Clark 2004); th<strong>is</strong><br />

plumage <strong>is</strong> synonymous with the Basic III plumage of Howell et<br />

al. (2003). See Clark (2004) and Pyle (2005) for d<strong>is</strong>cussions of<br />

primary wave molt and its use in aging raptors.<br />

Why Not a Bald <strong>Eagle</strong><br />

Throughout the video, th<strong>is</strong> odd eagle, compared to the various<br />

Bald <strong>Eagle</strong>s next to it, shows a much larger and squarer head, a<br />

P.O. Box 531467<br />

Harlingen, Texas 78550<br />

raptours@earthlink.net<br />

larger and deeper beak, a<br />

more massive body, and<br />

much longer nape feathers<br />

(Figs. 2 & 3). Its upperparts<br />

are much like that of a juvenile Bald <strong>Eagle</strong>, but it has a<br />

larger, orangey beak (Figs. 1–4). It shows a pale circle around the<br />

eye, a character of some Steller’s Sea-<strong>Eagle</strong>s (see Fig. 3, p. 69 in:<br />

Morioka et al. 1995) but not of Bald <strong>Eagle</strong>s. In addition, its underparts<br />

are uniformly colored (Figs. 3 & 4); those of immature<br />

Bald <strong>Eagle</strong>s usually show the breast contrastingly darker<br />

than the belly (Clark and Wheeler 2001). The eagle’s<br />

uppertail <strong>is</strong> uniformly gray<br />

(Fig. 4), a trait shown only by<br />

some juvenile Bald <strong>Eagle</strong>s<br />

(Clark and Wheeler<br />

2001); all older<br />

Hokkaido,<br />

Japan; February<br />

1992. © William S.<br />

Clark / VIREO.<br />

the Bald <strong>Eagle</strong> (opposite page) <strong>is</strong> widespread and fairly common in Alaska and elsewhere in northwestern<br />

north America, and the Steller’s Sea-<strong>Eagle</strong> (th<strong>is</strong> page) <strong>is</strong> of casual occurrence in Alaska. At least one mixed<br />

pair <strong>is</strong> known from Alaska, and it <strong>is</strong> possible that there are others. th<strong>is</strong> article describes an immature eagle<br />

in Brit<strong>is</strong>h Columbia in late 2004 that appears to be a hybrid between Bald <strong>Eagle</strong> and steller’s sea-<strong>Eagle</strong>.<br />

w w w . A B A . o r g 29


H Y B R I D<br />

E A G L E<br />

Fig. 1 Fig. 2 Fig. 3 Fig. 4<br />

Fig. 1. Bald <strong>Eagle</strong> × Steller’s Sea-<strong>Eagle</strong> hybrid. note the molt of the primaries, with P6 and P7 paler than P8, P9,<br />

and P1–P5. Videograb. Gulfstream Park, Brit<strong>is</strong>h Columbia; 8 December 2004. © Darren Copley.<br />

Fig. 2. Bald <strong>Eagle</strong> × Steller’s Sea-<strong>Eagle</strong> hybrid. the hybrid has an orangey bill and <strong>is</strong> much larger overall than the<br />

juvenile Bald <strong>Eagle</strong> in the foreground. Videograb. Gulfstream Park, Brit<strong>is</strong>h Columbia; 8 December 2004. © Darren Copley.<br />

Fig. 3. Bald <strong>Eagle</strong> × Steller’s Sea-<strong>Eagle</strong> hybrid. not only <strong>is</strong> the hybrid larger in overall size than the juvenile Bald<br />

<strong>Eagle</strong> in the foreground, but it also has a relatively larger and squarer head. Videograb. Gulfstream Park, Brit<strong>is</strong>h<br />

Columbia; 8 December 2004. © Darren Copley.<br />

Fig. 4. Bald <strong>Eagle</strong> × Steller’s Sea-<strong>Eagle</strong> hybrid. note the gray upperside of the tail. Videograb. Gulfstream Park,<br />

Brit<strong>is</strong>h Columbia; 8 December 2004. © Darren Copley.<br />

Bald <strong>Eagle</strong>s have a white or pale gray uppertail, with a<br />

wide dusky subterminal band. Bald <strong>Eagle</strong>s in their<br />

third annual plumage show a white crown and cheeks,<br />

and usually white spots on the back forming a pale triangle<br />

(Fig. 5).<br />

Why Not a Steller’s Sea-<strong>Eagle</strong><br />

Steller’s Sea-<strong>Eagle</strong>s of all ages have deeply wedged<br />

white tails, with those of<br />

juveniles usually showing<br />

black tips (Wheeler and<br />

Clark 1995, Clark and<br />

Wheeler 2001; see also<br />

Fig. 3, p. 69 in: Morioka et<br />

al. 1995). Th<strong>is</strong> pattern <strong>is</strong><br />

shown by the Steller’s in<br />

its third annual plumage<br />

depicted in Fig. 6. The uppertail<br />

of the eagle in<br />

question was completely<br />

dark gray (Fig. 4), like<br />

that of some juvenile Bald<br />

<strong>Eagle</strong>s. As seen in the<br />

video and commented on<br />

Fig. 5. Basic III Bald <strong>Eagle</strong>.<br />

note the white crown and<br />

cheeks as well as the dusky<br />

beak. Boundary Bay, Brit<strong>is</strong>h<br />

Columbia; 6 February 2005.<br />

© William S. Clark.<br />

by people who saw th<strong>is</strong> eagle in flight, the tail wasn’t<br />

extremely wedge-shaped, like that of Steller’s (Fig. 6),<br />

but rather was somewhat wedge-shaped, although not<br />

strongly so (see Wheeler and Clark 1995, Clark and<br />

Wheeler 2001; see also Figs. 8 & 9, p. 72 in: Morioka<br />

et al. 1995).<br />

Steller’s Sea-<strong>Eagle</strong>s fly with their wings in a strong dihedral<br />

(Wheeler and Clark 1995, Clark and Wheeler<br />

2001). However, Allinson wrote<br />

on the <strong>Birding</strong> Vancouver Island<br />

website after seeing the eagle in<br />

flight, “I’m afraid th<strong>is</strong> bird has<br />

rather typical Baldie flat dihedral<br />

in flight. The tail, while more<br />

pointed than typical Baldies, <strong>is</strong><br />

still rounded when fanned out<br />

and nothing like what it should<br />

be for Steller’s.”<br />

Why a Hybrid Between<br />

These Two Species<br />

The eagle in question has characters<br />

of both species and, as d<strong>is</strong>cussed<br />

in the previous paragraphs,<br />

apparently <strong>is</strong>n’t either.<br />

For example, it doesn’t show the<br />

small crest of feathers on the forehead<br />

shown by some perched<br />

Steller’s (Figs. 1–6 & 17 in:<br />

Morioka et al. 1995). Moreover, it<br />

30<br />

B i r d i n g • J u l y / A u g u s t 2 0 0 8


has a uniformly gray upper tail, a trait never shown on<br />

Steller’s Sea-<strong>Eagle</strong>. Additionally, it shows an orangey<br />

beak, a character never shown by immature Bald <strong>Eagle</strong>.<br />

No other eagle looks like th<strong>is</strong> one.<br />

Mixed Pair<br />

A mixed pair of adult Bald <strong>Eagle</strong> and Steller’s Sea-<strong>Eagle</strong><br />

close to Juneau in Tagu Inlet has been together for<br />

more than 10 years during the breeding season<br />

.<br />

Many birders during th<strong>is</strong> time have chartered a helicopter<br />

and flown to see the adult Steller’s. Recently, another<br />

adult has been seen repeatedly on the Nuskagak River<br />

near Dillingham in southwestern Alaska . It<br />

<strong>is</strong> certainly possible, given the remoteness and vastness<br />

of the Bald <strong>Eagle</strong>’s range in Alaska, that other such<br />

mixed pairs occur. An immature hybrid from such a<br />

mixed pair might be expected to travel to the coast and<br />

then south to Vancouver Island during the winter.<br />

There are no publ<strong>is</strong>hed records of hybridization<br />

within the genus Haliaeetus. However, there are many<br />

records of hybrid raptors within other genera, for example,<br />

in the genus Buteo (Clark and Witt 2006), in<br />

Milvus kites (Sylvén 1977), and in Aquila eagles (Helbig<br />

et al. 2005).<br />

Th<strong>is</strong> most unusual eagle shown in the video taken in<br />

December 2004 at Goldstream Provincial Park <strong>is</strong> evidently<br />

neither a Bald <strong>Eagle</strong> nor a Steller’s Sea-<strong>Eagle</strong>, but<br />

rather has characters of both species. It <strong>is</strong> almost certain<br />

from its shared characters that th<strong>is</strong> eagle <strong>is</strong> a juvenile<br />

hybrid between the two, perhaps from the mixed<br />

pair of adults at Tagu Inlet, Alaska.<br />

Acknowledgments<br />

I thank David Allinson for telling me about th<strong>is</strong> unusual<br />

eagle and David Copley for providing me copies<br />

of the CD with images of the bird. Todd Katzner and<br />

the National Aviary in Pittsburgh provided the photo of<br />

the Steller’s Sea-<strong>Eagle</strong>, and I. Szabo made constructive<br />

comments on previous drafts.<br />

Literature Cited<br />

Clark, w.s. 2004. wave molt of the primaries of Accipitrid raptors, and its<br />

Fig. 6. Basic III Steller’s Sea-<strong>Eagle</strong>. note the massive beak<br />

and white tail on th<strong>is</strong> captive individual. National Aviary,<br />

Pittsburgh, Pennsylvania; 17 May 2007. © Erin Estell.<br />

use in ageing, pp. 705–804 in: r.d. Chancellor and B.-u. Meyburgh,<br />

eds. Raptors Worldwide: Proceedings of the Vth World Conference on<br />

Birds of Prey, Budapest. world working group on Birds of Prey, Berlin.<br />

Clark, w.s., and B.K. wheeler. 2001. A Field Guide to Hawks of North America.<br />

Houghton Mifflin, Boston.<br />

Clark, w.s., and C.C. witt. 2006. First known specimen of a hybrid Buteo:<br />

swainson’s Hawk (Buteo swainsoni) × rough-legged Hawk (B. lagopus).<br />

Wilson Journal of Ornithology 118:42–52.<br />

Helbig, A.J., i. seibold, A. Kocum, d. liebers, J. irwin, u. Bergman<strong>is</strong>, B.u.<br />

Meyburg, w. scheller, M. stubbe, and s. Bensch. 2005. genetic differentiation<br />

and hybridization between greater and lesser spotted <strong>Eagle</strong>s<br />

(Accipitriformes: Aquila clanga, A. pomarina). Journal of Ornithology<br />

146:226–234.<br />

Howell, s.n.g., C. Corbin, P. Pyle, and d.i. rogers. 2003 the first basic<br />

problem: A review of molt and plumage homologies. Condor<br />

105:635–653.<br />

Morioka, t., n. yamagata, t. Kouchi, and t. Kawata. 1995. The Birds of Prey<br />

of Japan. Bun-ichi sōgō shuppan, tokyo.<br />

Pyle, P. 2005. remigial molt patterns in north <strong>American</strong> Falconiformes as<br />

related to age, sex, breeding status, and life-h<strong>is</strong>tory strategies. Condor<br />

107:825–836.<br />

sylvén, M. 1977. Hybridization between red Kite Milvus milvus and Black<br />

Kite M. migrans in sweden in 1976. Vår Fågelvärld 36:38–44.<br />

wheeler, B.K., and w.s. Clark. 1995. A Photographic Guide to North <strong>American</strong><br />

Raptors. Princeton university Press, Princeton.<br />

w w w . A B A . o r g 31

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