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Mekelle University<br />

CDANR<br />

College of Dryland Agriculture and Natural Resources<br />

Community Management and Status of an Afro-alpine Ecosystem:<br />

The case of the <strong>Ethiopian</strong> <strong>Wolf</strong> at Mt. Abune Yoseph<br />

By<br />

<strong>Girma</strong> <strong>Eshete</strong><br />

A Thesis Submitted in Partial Fulfillment of the Requirement for the<br />

Master of Science Degree in Tropical Land Resource Management and<br />

Environmental Protection<br />

Advisors: <strong>Girma</strong>y Tesfay (PhD)<br />

Hans Bauer (PhD)<br />

Zelealem Tefera (PhD)<br />

Jorgelina Marino (PhD)<br />

June, 2010<br />

Mekelle<br />

i


Declaration<br />

This is to certify that this <strong>thesis</strong> entitled “Community Management and Status of an Afro-alpine<br />

Ecosystem: The Case of the <strong>Ethiopian</strong> <strong>Wolf</strong> at Mt. Abune Yoseph” submitted in partial<br />

fulfillment of the requirements for the award of the degree of Master of Science, in Tropical<br />

Land Resource Management, to the College of Dryland Agriculture and Natural Resources,<br />

Mekelle University, through the Department of Land Resource Management and Environmental<br />

Protection, done by Mr. <strong>Girma</strong> <strong>Eshete</strong> Genbere, (Id.No. CDANR/PR0004/01) is an authentic<br />

work carried out by him under our guidance. The matter embodied in this project work has not<br />

been submitted earlier for award of any degree or diploma to the best of our knowledge and<br />

belief.<br />

Name of the student <strong>Girma</strong> <strong>Eshete</strong> Genbere Signature & date ______________________<br />

Name of the Supervisors: <strong>Girma</strong>y Tesfay (PhD) Signature & date___________________<br />

Hans Bauer (PhD) Signature & date ____________________<br />

Zelealem Tefera (PhD) Signature & date _________________<br />

Jorgelina Marino (PhD) Signiture & date __________________<br />

ii


Abstract<br />

Mt. Abune Yoseph has high biodiversity and endemism which is representative of the Afro-alpine<br />

ecosystems. Our study employed socio-economic and ecological survey as a major<br />

methodological approach. For socio-economic data 120 randomly selected households from<br />

eight villages were interviewed and ecological data were collected from 110 circular sample<br />

plots of 5m radius marked along eight habitat assessment transects laid at 1km apart with 200m<br />

interval using a 1: 50,000 scale map and Global positioning system (GPS). The result of<br />

descriptive statistics showed that 80% households exploit economically important natural<br />

resources to support their livelihoods from Mt. Abune Yoseph and chi-square test indicated the<br />

presence of significant difference (P


Acknowledgements<br />

Above all, let endless thanks be to the Almighty God and his mother for their innumerable gifts,<br />

hold up and allowing me to reach another gesture in my career.<br />

I highly appreciate my supervisors Dr <strong>Girma</strong>y Tesfay and Dr Hans Bauer from Mekelle<br />

University Dr Zelealem Tefera from FZS and Dr Jorgelina Marino from Oxford University for<br />

their unreserved assistance and encouragement. I thank Dr. <strong>Girma</strong>y Tesfay and Dr. Hans Bauer<br />

for their generous and valuable comments on the content and depth of the paper continuously<br />

throughout the study. My special thanks go to Dr. Zelealem Tefera for giving me the opportunity<br />

to work at Mt. Abune Yoseph and organizing the fund for my project. He provided me with all<br />

necessary field materials and facilities. Generally, it was not possible to complete this work<br />

without his valuable support in every aspect. Many thanks go to Dr. Jorgelina Marino; she took a<br />

week time to assist me while I was in the field and trained me on basic ecological data collection<br />

methods. Her guidance and inspiration made this study a reality.<br />

I am grateful to N/Wollo EPLAUD, main sponsor; EWCP and FZS two sister NGOs working for<br />

the conservation of the <strong>Ethiopian</strong> wolf and Afro-alpine ecosystem sponsored my field study. I<br />

thank Chris Gorden and Anne-Marie Stewart heads of EWCP; Fekadu Lema, Zebiba Ahmed,<br />

Getachew Asefa, Bilemen Melkamu, Desiew Gelaw and Desale Melakie staff of FZS. I<br />

appreciate the material and moral contribution of my friends Getachew Shiferaw, Gessiu<br />

Teshome and all my staff members especially Ato Fentaw Awayehu, Desta Gessesse and Roman<br />

Sisay.<br />

I am thankful to my wife Agerie Debeb for her everlasting help and encouragement. She took<br />

alone the total load of house matters, including taking care of our children during my study.<br />

Without her it was not possible for me to reach this level.<br />

iv


List of Abbreviations and Acronyms<br />

ANRS<br />

ARDD<br />

a.s.l.<br />

BMNP<br />

BoFAED<br />

CBFM<br />

CSG<br />

EMA<br />

EPLAUD<br />

ESP<br />

EWCO<br />

EWCP<br />

EWNHS<br />

FZS<br />

GO<br />

HHs<br />

HHHs<br />

IIED<br />

IUCN<br />

Mt.<br />

NGO<br />

PAs<br />

SMNP<br />

TESFA<br />

U.S.A.<br />

WEPLAUO<br />

Amhara National Regional State<br />

Agriculture and Rural Development Department<br />

above sea level<br />

Bale Mountains National Park<br />

Bureau of Finance and Economic Development<br />

Community Based Forest Management<br />

Canid Specialist Group<br />

<strong>Ethiopian</strong> Mapping Agency<br />

Environmental Protection Land Administration and Use Department<br />

Environment Support Project<br />

<strong>Ethiopian</strong> <strong>Wolf</strong> <strong>Conservation</strong> Organization<br />

<strong>Ethiopian</strong> <strong>Wolf</strong> <strong>Conservation</strong> Program<br />

<strong>Ethiopian</strong> Wildlife and Natural History Society<br />

Frankfurt Zoological Society international NGO<br />

Governmental Organization<br />

Households<br />

Household heads<br />

International Institute for Environment and Development<br />

The World <strong>Conservation</strong> Union<br />

Mountain<br />

Non Governmental Organization<br />

Peasant Associations<br />

Simen Mountains National Park<br />

Tourism in Ethiopia for Sustainable Future Alternative<br />

United States of America<br />

Woreda Environmental Protection Land Administration and Use Office<br />

v


This piece of work is dedicated to the memories of my beloved father, <strong>Eshete</strong> <strong>Jenbere</strong> who took<br />

me out from the darkness of illiteracy to the light of education. Dad may the almighty God rest<br />

your soul in haven.<br />

vi


Table of contents<br />

page<br />

ABSTRACT ................................................................................................................................. III<br />

ACKNOWLEDGEMENTS ....................................................................................................... IV<br />

LIST OF ABBREVIATIONS AND ACRONYMS ................................................................... V<br />

LIST OF TABLES ....................................................................................................................... X<br />

LIST OF FIGURES .................................................................................................................. XII<br />

CHAPTER I: INTRODUCTION ................................................................................................ 1<br />

1.1. BACKGROUND ....................................................................................................................... 1<br />

1.2. STATEMENT OF THE PROBLEM .............................................................................................. 2<br />

1.3. HYPOTHESIS .......................................................................................................................... 3<br />

1.4. OBJECTIVES .......................................................................................................................... 4<br />

1.4.1. General Objective ......................................................................................................... 4<br />

1.4.2. Specific objectives ......................................................................................................... 4<br />

CHAPTER II: LITERATURE REVIEW .................................................................................. 5<br />

2.1. AFRO-ALPINE ECOSYSTEM IN ETHIOPIA ................................................................................ 5<br />

2.2. ETHIOPIAN AFRO-ALPINE ECOSYSTEM AND ITS ENDEMIC SPECIES ......................................... 7<br />

2.2.1. Afro-alpine fauna .......................................................................................................... 7<br />

2.2.1.1. The <strong>Ethiopian</strong> wolf ................................................................................................... 10<br />

2.2.2. Afro-alpine flora ......................................................................................................... 12<br />

2.3. NORTH WOLLO HIGHLANDS AFRO-ALPINE ECOSYSTEM ...................................................... 15<br />

2.4. COMMUNITY PERCEPTION AND ATTITUDE TOWARDS AFRO-ALPINE ECOSYSTEM ................. 17<br />

vii


2.5. IMPACT OF THE COMMUNITY ON THE AFRO-ALPINE ECOSYSTEM AND ITS WILDLIFE ............ 18<br />

2.6. SOCIO-ECONOMIC SIGNIFICANCE OF AFRO-ALPINE ECOSYSTEM .......................................... 20<br />

2.7. SUSTAINABLE COMMUNITY BASED AFRO-ALPINE ECOSYSTEM MANAGEMENT .................... 23<br />

CHAPTER III: MATERIALS AND METHODS .................................................................... 25<br />

3.1. STUDY AREA ....................................................................................................................... 25<br />

3.1.1. LOCATION ........................................................................................................................ 25<br />

3.1.2. Topography ................................................................................................................. 27<br />

3.1.3. Climate ........................................................................................................................ 27<br />

3.1.4. Soil .............................................................................................................................. 27<br />

3.1.5. Vegetations and wild animals ..................................................................................... 28<br />

3.1.6. Water resources .......................................................................................................... 28<br />

3.2. METHODS ............................................................................................................................ 29<br />

3.2.1. Interview ..................................................................................................................... 29<br />

3.2.2. Population demography .............................................................................................. 30<br />

3.2.3. Ecological survey ........................................................................................................ 30<br />

3.3. DATA ANALYSIS ................................................................................................................. 31<br />

CHAPTER IV: RESULTS AND DISCUSSION ...................................................................... 32<br />

4.1. SOCIO-ECONOMIC CHARACTERISTIC AND ECONOMIC POTENTIAL OF MT. ABUNE YOSEPH .. 32<br />

4.1.1. Household characteristics .......................................................................................... 32<br />

4.1.2. Socio-economic potential of Mt. Abune Yoseph Afro-alpine ecosystem ..................... 33<br />

4.2. COMMUNITY PERCEPTION TOWARDS MT. ABUNE YOSEPH AFRO-ALPINE ECOSYSTEM........ 38<br />

4.3. COMMUNITY PERCEPTION TOWARDS ETHIOPIAN WOLF ....................................................... 42<br />

viii


4.4. HUMAN-LIVESTOCK INTERFERENCE TO AFRO-ALPINE ECOSYSTEM AND THE RESULTING<br />

CONFLICT ................................................................................................................................... 46<br />

4.4.1. Human-livestock interference pressure and its impact ............................................... 46<br />

4.4.2. Human conflict with <strong>Ethiopian</strong> wolf at Mt. Abune Yoseph ......................................... 53<br />

4.5. ECOLOGICAL ASSESSMENT OF MT. ABUNE YOSEPH AFRO-ALPINE ECOSYSTEM .................. 55<br />

4.5.1. Households response on <strong>Ethiopian</strong> wolf presence/ absence....................................... 55<br />

4.5.2. Determination of wolf distribution by rodent burrow................................................. 58<br />

4.5.3. Determination of <strong>Ethiopian</strong> wolf abundance by sighting at Mt. Abune Yoseph ......... 62<br />

CHAPTER V: CONCLUSION AND RECOMMENDATIONS ............................................ 66<br />

5.1. CONCLUSION ....................................................................................................................... 66<br />

5.2. RECOMMENDATIONS ........................................................................................................... 67<br />

REFERENCES ............................................................................................................................ 70<br />

APPENDIX .................................................................................................................................. 75<br />

ix


List of Tables<br />

Table 1: Composition and diversity of diet from wolves scat analysis at Mt. Abune Yoseph ..... 19<br />

Table 2: Sample HHs characteristics of Mt. Abune Yoseph ....................................................... 32<br />

Table 3: Villages and sample HHs got economic benefit from Mt. Abune Yoseph ................... 33<br />

Table 4: HHs economic resource benefit from Mt. Abune Yoseph with respect to distance ....... 34<br />

Table 5: Type of economic benefits obtained from Mt. Abune Yoseph (n =120) ....................... 35<br />

Table 6: Price of major economic goods and services from Mt. Abune Yoseph in 1year (n =120)<br />

....................................................................................................................................................... 36<br />

Table 7: HHs average estimated birr obtained from Mt. Abune Yoseph in one year ................... 37<br />

Table 8: Distribution of Nominal variables employed in logistic regression ............................... 39<br />

Table 9: Descriptives of continuous variables employed in logistic regression (n=120)............. 39<br />

Table 10: Factors determining the perception of respondents towards Mt. Abune Yoseph Afroalpine<br />

ecosystem and its management .......................................................................................... 40<br />

Table 11: Factors determining the perception of respondents towards <strong>Ethiopian</strong> wolves ........... 44<br />

Table 12: HHs reason why they perceive wolves good or bad species at Mt. Abune Yoseph ..... 45<br />

Table 13: Human population in woredas that have Afro-alpine ecosystem in North Wollo ........ 46<br />

Table 14: Livestock grazing period at Mt. Abune Yoseph (n =120) ............................................ 47<br />

Table 15: Livestock population growth trend in woredas that have Afro-alpine ecosystem........ 48<br />

Table 16: Reasons for change in wolf number trend at Mt. Abune Yoseph (n=120)................... 49<br />

Table 17: HHs response on whether or not faced wild animal problem ....................................... 51<br />

Table 18: Distribution of sample HHs that faced different problem by wildlife ......................... 52<br />

Table 19: Mean ±SE number of sheep preyed in the last 10 years at Mt. Abune Yoseph (n=120)<br />

....................................................................................................................................................... 53<br />

x


Table 20: HHs response on wolf observation, observation time and trend at Mt. Abune Yoseph 56<br />

Table 21: HHs wolf observation frequency, wolf seen at once and wolf estimation at Mt. Abune<br />

Yoseph (n =120) ........................................................................................................................... 56<br />

Table 22: Analysis of rodent burrows by slope and mean comparison using Tukey-Kramer HSD<br />

....................................................................................................................................................... 59<br />

Table 23: Bivariate fit analysis of rodent burrow by altitude ...................................................... 60<br />

Table 24: Analysis of rodent burrow by vegetation height and mean comparison using Tukey-<br />

Kramer HSD ................................................................................................................................. 61<br />

Table 25: Analysis of rodent burrow by vegetation cover and means of rodent burrow at each<br />

level of vegetation cover ............................................................................................................... 62<br />

Table 26: Bivariate analysis of wolf sighting by altitude ............................................................. 63<br />

Table 27: Analysis of wolf sighting by slope and mean comparisons using Tukey-Kramer HSD<br />

....................................................................................................................................................... 64<br />

Table 28: Analysis of wolf sighting by vegetation height and mean comparisons using Tukey-<br />

Kramer HSD ................................................................................................................................. 65<br />

xi


List of Figures<br />

Figure 1: From top to bottom: Subordinate female feeding pups and Adult male <strong>Ethiopian</strong> wolf<br />

....................................................................................................................................................... 11<br />

Figure 2: <strong>Ethiopian</strong> wolf sighting on flat slope, short and open grassland ................................... 14<br />

Figure 3: The endemic giant lobelia from Mt.AbuneYoseph Afro-alpine ecosystem .................. 15<br />

Figure 4: Location of the study Area ............................................................................................ 26<br />

Figure 5:.Land ownership status and economic benefit from Mt. Abune Yoseph ....................... 36<br />

Figure 6: People selling firewood in the market ........................................................................... 37<br />

Figure 7: Farming inside Afro-alpine ecosystem by clearing lobelia ........................................... 51<br />

Figure 8: HHs response on sheep predators at Mt. Abune Yoseph for the last 10 years ............. 54<br />

Figure 9: One way Analysis of Rodent burrows by Slope .......................................................... 59<br />

Figure 10: One way analysis of rodent burrow by vegetation height ........................................... 60<br />

Figure 11: Short Afro-alpine vegetation (E. pinifolius) with rodent burrow from Mt. Abune<br />

Yoseph .......................................................................................................................................... 61<br />

Figure 12: One way analysis of wolf sighting by slope ................................................................ 63<br />

Figure 13: One way analysis of wolf sighting by vegetation height ............................................ 64<br />

xii


xiii


Chapter I: Introduction<br />

1.1. Background<br />

The highlands of Ethiopia are a massif of volcanic origin formed between 70 and 5 million years<br />

ago. The Great Rift Valley divides the massif into a larger northern area and a smaller<br />

southeastern area (Malcom and Ashenafi, 1997). One of the characteristics that distinguish<br />

Ethiopia from all other African countries is the extent of its highland, with 50% of all the lands<br />

above 2,000 m and 80% of all the lands above 3,000 m in Africa (Yalden and Largen, 1992).<br />

These mountains enabled the evolution of an abundance of endemic animal and plant species<br />

confined to the Afro-alpine ecosystem (Yalden, 1983).<br />

In Ethiopia Afro-alpine grasslands and ericaceous heathlands extend above 3,400 m a.s.l,<br />

covering about 5000 km 2 . The entire area of Afro-alpine ecosystem in sub Saharan Africa is<br />

11,887 km 2 , of this 73% is found in Ethiopia. Sixty percent of this habitat occurs in three<br />

mountain ranges (Bale, Arsi and Simen) with the remainder scattered in smaller isolated areas<br />

(Marino, 2003). The Afro-alpine ecosystem and surrounding highland support an exceptionally<br />

high diversity of rare and endemic species, recognized as one of the 34 biodiversity hotspots in<br />

the world. Among the endemic mammals the endangered <strong>Ethiopian</strong> wolf (Canis simensis),<br />

Gelada baboon (Theropithecus gelada) and Walia Ibex (Capra walia) are the major.<br />

The <strong>Ethiopian</strong> wolf is listed as Critically Endangered by the IUCN (2008) Red List. This<br />

specialized carnivore lives only in Afro-alpine habitats, generally above 3,200 m a.s.l., and feed<br />

mainly on rodents, many of which endemic to the <strong>Ethiopian</strong> highlands.<br />

In the Northern highlands of the Amhara region, <strong>Ethiopian</strong> wolf populations are smaller and<br />

more fragmented than those South east of the Rift Valley (Marino, 2003). These habitats are also<br />

1


subjected to greater land use pressures. Mt. Abune Yoseph range is one of the main habitats for<br />

wolf populations in Northern Ethiopia, also home to Gelada baboons and many bird species, the<br />

giant Lobelia (Lobelia rhyncopetalum) and short Afro-alpine vegetation. Unless measures are<br />

taken, these species are all vulnerable to extinction.<br />

1.2. Statement of the Problem<br />

According to the U.S. Census Bureau 2006 report, the world‟s population is expected to reach 7<br />

billion in 2012. Providing a long-term food supply for the growing global population without<br />

damaging ecosystems is one of the greatest challenges facing mankind. This is also an acute<br />

problem in the <strong>Ethiopian</strong> highlands where 80% of the country‟s 70 million people live in the<br />

highlands, with significant pressure for agricultural development. Ethiopia also has the largest<br />

national herd of domestic livestock in Africa, increasingly using Afro-alpine areas to graze<br />

leading to overgrazing and erosion. This heightened competition between livestock and wildlife<br />

cause fragmentation and loss of habitat with its endemic species (Hurni, 1986).<br />

The Abune Yoseph massif of North Wollo suffers from diverse levels of human intervention<br />

mainly from agricultural activities and livestock grazing which dramatically modified the natural<br />

landscape. Natural Afro-alpine habitat has contracted over the last decades and currently remains<br />

only above 3,700 m a.s.l. The future survival of wolves in Northern Ethiopia especially in Mt.<br />

Abune Yoseph is in question (Ash, 2000; Marino, 2003). In view of the persistent human impact<br />

on the massif immediate action is required. To design conservation actions research is needed to<br />

investigate the current socio-economic and ecological potential of Mt. Abune Yoseph including<br />

its flagship species, the <strong>Ethiopian</strong> wolf.<br />

2


<strong>Ethiopian</strong> wolves depend on Afro-alpine habitats with enough rodents to survive. According to<br />

Delany (1972), the main factors accounting for the distribution and abundance of rodents are the<br />

nature of vegetation through the food and cover it provides and the extent to which these habitat<br />

components are modified by large mammals and human activity. Previous studies of Mt. Abune<br />

Yoseph‟s rodents (Tekilu, 2008) and field survey carried out by EWCP from 1997 to 2000,<br />

identified rodent species and attempted to measure biomass but no research looked in detail at<br />

the vegetation and environmental factors that determine the distribution and abundance of<br />

rodents and <strong>Ethiopian</strong> wolf in this area.<br />

A number of studies Kellert (1985), indicated that people who perceived economic benefits and<br />

enjoy unrestricted access to natural resources usually support ecosystem conservation efforts, but<br />

these did not consider how local communities perceives the Afro-alpine ecosystem and its<br />

endemic wildlife at Mt. Abune Yoseph.<br />

Therefore, this research was conducted to fill the above mentioned gaps of the area by testing the<br />

following leading hypo<strong>thesis</strong>.<br />

1.3. Hypo<strong>thesis</strong><br />

<br />

Community perception towards common natural resources (Afro-alpine ecosystem and<br />

<strong>Ethiopian</strong> wolf) determines their sustainable utilization.<br />

<br />

Ecologic and anthropogenic factors determine the distribution and abundance of<br />

<strong>Ethiopian</strong> wolf.<br />

3


1.4. Objectives<br />

1.4.1. General Objective<br />

The general objective of the study was to investigate the socio-economic potential and natural<br />

resources assessment of Mt. Abune Yoseph Afro-alpine ecosystem for the improvement of local<br />

community livelihoods through sustainable management.<br />

1.4.2. Specific objectives<br />

<br />

To evaluate the socio-economic potential of the Mt. Abune Yoseph Afro-alpine<br />

ecosystem for the improvement of surrounding community livelihoods.<br />

<br />

To assess perception of the local community towards the Afro-alpine ecosystem and<br />

<strong>Ethiopian</strong> wolf at Mt. Abune Yoseph.<br />

<br />

To evaluate the human-livestock interference and the resulting conflict with wolf in Afroalpine<br />

ecosystem at Mt. Abune Yoseph.<br />

<br />

To determine wolf presence/absence and relative abundance in the ecosystem in relation<br />

to vegetation parameters (height, cover) and environmental factors (slope, altitude and<br />

rodent burrow) at Mt. Abune Yoseph.<br />

4


Chapter II: Literature Review<br />

2.1. Afro-alpine ecosystem in Ethiopia<br />

Ethiopia‟s unique geology sets it apart from other African countries. In the high mountains of<br />

East Africa the climate determines three broad altitudinal belts: an Afro-alpine zone, subafroalpine<br />

and montane forests or grassland belt (Kingdon, 1990).<br />

The <strong>Ethiopian</strong> massif is characterized by marked altitudinal variations that produce a range of<br />

climates affecting plant and animal distribution and the concentration of people (Yalden and<br />

Largen, 1992). Most people live in the fertile highlands of the Woina Dega (warm temperate)<br />

and Dega (temperate) zones. The Wurch zone (alpine, >3,000m) is colder and less favorable for<br />

agriculture and broadly corresponds with the distribution of habitats suitable for the <strong>Ethiopian</strong><br />

wolf. Within the Wurch, ericaceous heathlands dominated by Phillipia and Erica spp. (Miehe<br />

and Miehe, 1993) are replaced at higher elevations by the Afro-alpine grasslands, scrubs and<br />

meadows above about 3,200m.<br />

The Afro-alpine areas experiences regular night frosts, diurnal temperature fluctuations that far<br />

exceed seasonal ones, and intense irradiation. Marked altitudinal variations characterize the<br />

climate, fauna and flora of the <strong>Ethiopian</strong> highlands (Yalden and Largen, 1992) temperature<br />

decreases with altitude and rainfall begins to decrease above 3,850 m (Hillman, 1986; Sillero-<br />

Zubiri, 1994) determining an altitudinal component in the diversity of Afro-alpine plants and<br />

rodents.<br />

The most recent glaciers retreated within the last 10,000 years and as the climate warmed the<br />

Afro-alpine ecosystem became restricted to the highest mountains. The end of the Pleistocene<br />

5


ought a change in the climate, and the extensive <strong>Ethiopian</strong> Afro-alpine steppes shrunk to their<br />

present state, reducing the habitat available to <strong>Ethiopian</strong> wolves by an order of magnitude<br />

(Gottelli and Sillero-Zubiri, 1992). Global warming during the last 10,000 years progressively<br />

confined the Afro-alpine ecosystem to the highest mountains, and 60% of all <strong>Ethiopian</strong> land<br />

above 3,000m has been converted to farmland.<br />

The central Sanetti Plateau located within the Bale Mountains National Park (BMNP) in<br />

Southwestern Ethiopia is one of the most distinctive of all <strong>Ethiopian</strong> habitats and represents the<br />

type of characteristic Afro-alpine vegetation that would have spread over a wide area of highland<br />

during the peak of the glacial. Apart from Bale and Arsi in Southern highlands, Afro-alpine<br />

ecosystems are located in the Simien Mountain National Park (SMNP) the largest Afro-alpine<br />

range North of the Rift Valley and the third most important in wolf population in Ethiopia. The<br />

other Afro-alpine units are North Wollo (1150 km 2 above 3200 m), South Wollo (1220km 2 ),<br />

Mt.Guna (960 km 2 ), Mt.Choke (500km 2 ) Guassa-Menz (124km 2 ) and Gosh Meda (90km 2 )<br />

(Marino, 2003). The major three primary mountain ranges: Bale, Arsi and Simen receive some<br />

recognition as protected areas under the EWCO. In Northern highlands 680 km 2 of Afro-alpine<br />

and sub-afroalpine habitat in the Simen Mountains (ANRS) only 180 km 2 fall within the SMNP<br />

and the remainder the buffer zone around the park. One area of Afro-alpine habitat in Menz<br />

(ANRS) is managed by a committee of the community (Ashenafi et al., 2005), and all other<br />

including the Mt. Abune Yoseph have no formal protection.<br />

6


2.2. <strong>Ethiopian</strong> Afro-alpine ecosystem and its endemic species<br />

In Ethiopia the altitudinal variation produces varied macro and micro climatic conditions that<br />

affect nature and humans and contribute to the formation of diverse ecosystems. The flora of<br />

Ethiopia is estimated to comprise between 6,500 and 7,000 species, of which about 10% are<br />

considered to be endemic. The fauna is also highly diverse: 277 species of mammals, 861 species<br />

of birds, 201 species of reptiles, 63 species of amphibians, 150 species of fish and 324 species of<br />

butterflies are known from Ethiopia. Among these 31 mammals, 16 birds, 24 amphibians, 9<br />

reptiles and 40 fish are believed to be endemic (Hillman, 1993).<br />

Comparing provinces in Ethiopia Wollo has the lowest Afro-alpine endemic richness, despite<br />

having more than 4000 km 2 of land above 3000 m and three peaks over 4000 m, representing<br />

18% of the total for Ethiopia and Eritrea (Yalden, 1983). According to Saavedra (2009) the<br />

explanation for apparent lower diversity may reside in that the areas has been overlooked by<br />

botanists, including the mountains of Abune Yoseph, Abohoy Gara, Gubalafto, Delanta and<br />

Amba Ferit.<br />

2.2.1. Afro-alpine fauna<br />

The demands posed by an extreme climate and terrain influenced speciation, particularly for the<br />

smaller organisms such as rodents. Rodents are inherently well suited as the dominant herbivores<br />

of the Afro-alpine ecosystem (Marino et al., 1999). In cold conditions at high altitude, small<br />

homoeothermic species benefit from being larger, and indeed the community of Afro-alpine<br />

rodents is characterized by a high proportion of large species (Sillero-Zubiri et al., 1995).<br />

7


The present community of high altitude rodents include four endemic rodents that are confined to<br />

areas of 3,000 m a.s.l (Tachyoryctes macrocephalus, Stenocephalemys albocaudata, Lophuromys<br />

melanonyx and Megadendromus nikolausi); and six other non endemic extend into areas of 3000<br />

m a.s.l (Dendromus lovati, Arvicanthis blicki, Stenocephalemys griseicauda, Praomy salbipes,<br />

Praomys ruppi and Mus mahomet). There are at least a further 16 species of non endemic<br />

rodents, although four of endemic rodents (Tachyoryctes splendens, Otomys typus, Lophuromys<br />

flavopunctatus and Arvicanthis abyssinicus) are found together higher than 3000 m (Hillman<br />

1986; Yalden 1983; Yalden and Largen 1992; Sillero-Zubiri et al., 1995).<br />

L. melanonyx, S. albocaudata and A. blicki are characteristic of the Afro-alpine belt (Sillero-<br />

Zubiri et al. 1995). Yalden (1988) suggested that O. typus might be an Afro-alpine moorland<br />

specialist. L. flavopunctatus and S. griseicauda were characteristic of the montane grasslands. O.<br />

typus, decreased in abundance in trap returns as altitude increased through the ericaceous and<br />

Afro-alpine belts (Sillero-Zubiri et al., 1995).<br />

Within Afro-alpine areas, fine-scale associations were identified between rodents and vegetation<br />

types, largely determined by drainage and soil characteristics in relation to local topography.<br />

Stratified counts and rodent trapping demonstrated some clear habitat association between<br />

species and microhabitat conditions (Sillero-Zubiri et al., 1995). This habitat selection behavior<br />

helped to identify predictors of rodent abundance and distribution, and to interpret species‟<br />

adaptations to survive the harsh Afro-alpine environment.<br />

The giant molerat (T. macrocephalus) is currently confined to the Bale Mountains only (Yalden<br />

and Largen 1992) where it shows clear cut habitat selection between vegetation types, and<br />

between sub-habitats. Giant mole rats feed mainly on grasses with some dicotyledons, but<br />

8


apparently do not favor Helichrysum (Yalden 1983). Lack of suitable food may therefore limit<br />

their abundance in the ericaceous belt, where major plant types are woody Philippia and Erica<br />

species. Indices of giant mole rat biomass for Helichrysum dwarf-scrub and ericaceous belt were<br />

only 1/5 and 1/150, respectively of those in Afro-alpine grasslands (Marino, 2003).<br />

The role of the Afro-alpine rodent community in limiting the distribution of <strong>Ethiopian</strong> wolves<br />

was studied in Bale by looking at the relationship between wolf abundance and the species<br />

composition, relative abundance and activity pattern of the rodent community in various Afroalpine<br />

habitats (Sillero-Zubiri et al., 1995). In Bale mole rats were a vital determinant of wolf<br />

presence but in other Afro-alpine ecosystems small rodents also determine the presence and<br />

distribution of <strong>Ethiopian</strong> wolves (Ashenafi et al., 2005).<br />

The burrowing habits of giant mole rats have a modifying effect on the landscape and the<br />

composition of vegetation communities (Miehe and Miehe, 1994). Mima mounds are the soil<br />

accumulated from burrowing. A correlation between the presence of mima mounds and the<br />

abundance of mole rat signs supports the hypo<strong>thesis</strong> that mole rat abundance could determine<br />

wolf population densities (Marino, 2003).<br />

Birds of prey rely heavily on rodents (in Bale), but their main predator is another species unique<br />

to Ethiopia, the <strong>Ethiopian</strong> wolf (C. simensis) endemic to the Afro-alpine ecosystem of the<br />

<strong>Ethiopian</strong> highlands, its diurnal habits and distinctive coat render this species conspicuous.<br />

Moreover, the rodents‟ distribution and diurnal activity coincide with <strong>Ethiopian</strong> wolves‟ diurnal<br />

and solitary foraging habits, and their confinement to Afro-alpine habitats over 3,000 m (Marino<br />

et al., 1999).<br />

9


In addition, Afro-alpine ecosystem of Ethiopia is home for several unique populations of<br />

threatened endemic large mammals such as <strong>Ethiopian</strong> wolf, the walia ibex and the gelada<br />

baboon.<br />

2.2.1.1. The <strong>Ethiopian</strong> wolf<br />

The <strong>Ethiopian</strong> wolf is an Afro-alpine specialist endemic to the highlands of Ethiopia (Marino,<br />

2003). It survives where there is an Afro-alpine vegetation community and an associated rodent<br />

community. Unlike many carnivores, pack members of <strong>Ethiopian</strong> wolves forage and feed alone<br />

on small diurnal rodent prey, which they commonly and actively dig out from their burrows<br />

during the day time. In the Bale Mountains, the <strong>Ethiopian</strong> wolf feeds primarily on the giant mole<br />

rat and other species of Murinae rodents (Sillero-Zubiri et al. 1995). Other food items include<br />

rock hyrax (Procavia capensis), young antelopes and lambs (Sillero-Zuberi and Gottelli, 1995;<br />

Ashenafi, 2001).<br />

<strong>Ethiopian</strong> wolf density, measured both from observation and road counts, correlated positively<br />

with the total biomass index and the biomass index for diurnal species, but not for nocturnal<br />

species. Also a positive correlation was detected between rodent burrows and wolf sign<br />

(droppings or diggings) along habitat assessment transects (Sillero-Zubiri et al., 1995).<br />

The <strong>Ethiopian</strong> wolf is a medium-sized Canid with a reddish coat, distinctive white markings,<br />

long legs and elongated muzzle. Males are significantly larger than females in terms of body<br />

mass. The face, ears and upper parts of the muzzle are red. Ears broad, pointed, and directed<br />

forward. Characteristic facial markings include a white ascending crescent below the eyes, and a<br />

small white spot on the cheeks. The throat, neck, chest and under (ventral) parts is white.<br />

Boundary between the red coat and the white markings is sharp and well defined. The contrast of<br />

10


white markings against the red coat increases with age and social rank in both sexes; the female's<br />

coat is generally paler than the male's. The long, slender legs are reddish outside, with inner<br />

aspect white. Front feet has five toes, hind feet with four. The area around the anus is white. Both<br />

sexes have short tail, becoming a black stripe leading to a thick brush of black-tipped guard hairs<br />

(Sillero-Zubiri et al., 2004).<br />

Figure 1: From top to bottom: Subordinate female feeding pups and Adult male <strong>Ethiopian</strong> wolf<br />

11


The continued decline of the <strong>Ethiopian</strong> wolf's has prompted IUCN Canid Specialist Group<br />

(CSG) to list as critically endangered, in acknowledgment of its extremely high risk of extinction<br />

in the wild.<br />

2.2.2. Afro-alpine flora<br />

The <strong>Ethiopian</strong> Afro-alpine vegetation is dominated by grass and low growing shrubs from which<br />

the peculiar giant lobelias (Lobelia rhyncopetalum) emerge. Hedberg (1964) working on<br />

mountains in East Africa identified five major Afro-alpine vegetation communities,<br />

Dendrosenecio wood lands, Helichrysum scrub, Alchemilla scrub, Tussock grass land, Bogs and<br />

related communities. The last four, but not the first are found on <strong>Ethiopian</strong> mountains. Variation<br />

in moisture and topography determines which species predominates.<br />

Helichrysum scrub (usually the small gray shrub Helichrysum splendidum) covers large areas of<br />

drier (


melanantha, Dianthoseris schimperi, Erigeron alpinum, Haplocarpha rueppellii, Haplosciadium<br />

abyssinicum, Ranunculus oreophytus, and Swertia spp.), others sometimes form colonies of<br />

cushion shape (e.g. Myosotis keniensis, Sagina abyssinica, Sagina afroalpina, and Wahlenbergia<br />

pusilla) or have creeping habits (e.g. Alchemilla microbetula; Cerastium octandrum; Galium<br />

hochstetteri; Geranium arabicum; Polygonum spp.; Satureja kilimandschari; Veronica spp.).<br />

The group of tufted hemichryptophytes includes diverse tussock grasses (e.g. Agrostis<br />

quinqueneseta; Festuca richardii, Helichrysum cymosum herbaceous form; Koeleria capensis;<br />

Pentaschistis pictigluma; Poa leptoclada; Pentaschistis schimperiania; Senecio schultzii).<br />

The different vegetation communities support different rodent densities and as a result different<br />

number of <strong>Ethiopian</strong> wolves (Sillero-Zubri et al., 1995). In Afro-alpine ecosystem the density of<br />

<strong>Ethiopian</strong> wolf related with rodent density and Afro-alpine vegetation height. <strong>Wolf</strong> density is<br />

positively correlated with density of rodent prey and short height Afro-alpine vegetation. Highest<br />

wolf density is found in short Afro-alpine herbaceous communities (1.0-1.2 adults/km 2 ) but<br />

lower densities are found in Helichrysum dwarf scrub (0.2/km 2 ), in Ericaceous heathlands and<br />

barren peaks (0.1/km 2 ). Wolves also present at low density (0.1-0.2/km 2 ) in montane grass lands<br />

at lower altitudes. Generally wolves utilize all Afro-alpine but prefer open areas with short<br />

herbaceous and grass land communities where rodents are most abundant, along flat or gently<br />

sloping areas with deep soils and poor drainage in parts.<br />

13


Figure 2: <strong>Ethiopian</strong> wolf sighting on flat slope, short and open grassland<br />

The tops of African mountains have environmental conditions that differ dramatically from the<br />

low lands below. Plants and to a lesser degree animals have evolved some peculiar adaptations in<br />

response to the Afro-alpine climate. Plant gigantism is the most striking example. Giant lobelias,<br />

tree heather (Erica arborea) and giant Saint John's wort (Hyphericum revolutum) are important<br />

elements of the high montane flora of Ethiopia. The everlasting flowers (Helichrysum species)<br />

are conspicuous over large areas of the montanae habitat. Their highly reflective silvery leaves<br />

and dry papery flowers allow them to survive in the desiccating winds up to 4,300 m. The most<br />

conspicuous Afro-alpine vegetation is the giant lobelia whose sentinel forms dot the Afro-alpine<br />

landscape.<br />

14


Figure 3: The endemic giant lobelia from Mt. Abune Yoseph Afro-alpine ecosystem<br />

2.3. North Wollo highlands Afro-alpine ecosystem<br />

The Afro-alpine highlands of North Wollo are the second largest in Northern Ethiopia including<br />

Mt. Abune Yoseph (in Lasta woreda), Mt.Abohoy Gara (in Gidan woreda) Mekulet near Kulf<br />

Amba between Gubalafto and Delanta woreda. It covers a total area of 189.18 km 2 (ESP, 2001).<br />

Growing barley and grazing are the major land use practices in the lower altitude but in the upper<br />

zone agriculture is limited by frost and it provides habitat for wildlife such as <strong>Ethiopian</strong> wolf.<br />

Nowadays due to climate change and population growth these land uses are disturbed in all Afroalpine<br />

ecosystems of the zone.<br />

15


The Afro-alpine habitats in North Wollo are discontinuous. The Abune Yoseph massif is the<br />

largest and highest Afro-alpine area, loosely connected by narrow ridges with the small Abohoy<br />

Gara to the east and the Delanta in the south (Marino, 2003; Ash, 2000). Steep escarpments to<br />

the North of Mt. Abune Yoseph and east of Delanta limit wolf ranges. Elsewhere, overgrazed<br />

meadows and cultivated fields restricted suitable habitat to areas above 3,700-3,800m, or 3,450-<br />

3,500m in Delanta. Sightings and signs of wolves were recorded in all areas, but they were most<br />

abundant in Delanta, where the largest group with 6 individuals, was observed (Marino, 2003).<br />

Research conducted on rodents at Mt. Abune Yoseph (Tekilu, 2008) identified seven species of<br />

which six were endemic to Ethiopia and one insectivore shrew (Crocidura bayileyi) during<br />

survey in wet and dry seasons. The species were A. abyssinicus, S. griseicauda, L.<br />

flavopunctatus, O. typus, Pelomys harringtoni, D. lovati and Mericulus imberbis. These varied in<br />

abundance and density among the different parts of the habitats. Habitats modified by human<br />

activity supported less species diversity and population abundance than those undisturbed or with<br />

better natural vegetation cover. Without adequate management of the habitats supporting small<br />

mammals‟ effective conservation of the endangered <strong>Ethiopian</strong> wolf will be difficult. At the same<br />

time, conservation of the <strong>Ethiopian</strong> wolf in the area can not be successful without the keen<br />

involvement of local communities, whose lives are strictly intertwined with the wildlife and their<br />

habitats.<br />

The Afro-alpine vegetation in Mt. Abune Yoseph is characterized mainly by large extensions<br />

Cherenfe (Euryops pinifolius) Senecio shrubs with scattered patches of grasslands and rocky<br />

areas. On the northern slopes, grasslands of the genus Poa, Agrostis, Carex and Festuca<br />

dominate most lands affected by overgrazing. Only areas with very steep slopes and a high<br />

degree of humidity sustain tall grasslands. The moderately steep and large plateau of the Reem<br />

16


Gedel has abundant giant lobelias (800 to 2500) plants per hectare (Saavedra, 2009). The human<br />

impact is lower at the highest altitudes, where as large areas of the middle and lower altitudinal<br />

belts are highly modified.<br />

Studies of vegetation parameters and environmental factors in association with <strong>Ethiopian</strong> wolf<br />

prey in Mt. Abune Yoseph has multiple purposes, complements previous studies on rodents, help<br />

to sustain food chain among vegetation, rodents and wolf along with attempting to lay<br />

foundation to facilitate the conservation of endemic flora and fauna of this isolated mountain<br />

range considering the various stakeholders involved.<br />

2.4. Community perception and attitude towards Afro-alpine ecosystem<br />

In Africa, the attitudes and perceptions of local communities towards wildlife and areas of<br />

important wildlife habitat has been positive historically (Lewis et al., 1990; Newmank et al.,<br />

1993; Siachoono, 1995). A positive attitude among local communities is essential for successful<br />

wildlife conservation programmes, since wildlife provides physical, emotional, intellectual,<br />

economic and spiritual benefit to human development and well being (Kellert, 1985). An<br />

approach that focuses more on protecting resources by disregarding the needs and interests of the<br />

local resource users is bound to fail (Lewis et al., 1990). The most appealing alternative<br />

approach is to establish agreements, co-operation, partnerships and the equitable distribution of<br />

benefits between communities living in areas with valuable natural resources, or areas adjacent<br />

to the protected areas, and the conservation authority (IIED, 1994). In addition negative attitudes<br />

in recent years have been compounded with potential threats posed by wildlife to economically<br />

important livestock, huntable wildlife and to other forms of land use (Kellert et al., 1996; Bauer,<br />

2003).<br />

17


In Afro-alpine ecosystems the life of the local communities depends on ecosystem services such<br />

as grazing and land for barley cultivation which they perceive positively. In North central<br />

highlands of Ethiopia Guassa-Menz natural resources have been managed for the last hundred<br />

years by traditional practices and these communities have positive perception, an example in<br />

Ethiopia in this regard (Ashenafi, 2001). Negative attitudes towards wolves in Afro-alpine areas<br />

in recent years have been connected with depredation of economically important livestock (sheep<br />

and goat) by wolf (Marino, 2003). For the conservation of natural resources to be effective,<br />

attitudes of local communities towards management of the resource to be protected need to be<br />

studied including what communities perceive as needs and aspirations account (Fiallo and<br />

Jacobson, 1995).<br />

2.5. Impact of the community on the Afro-alpine ecosystem and its wildlife<br />

Africa‟s wildlife is under pressure from habitat loss due to forest clearing, agriculture and<br />

livestock herding; hunting for food and profit; pollution from agricultural and industrial sources;<br />

and disturbance by tourists. As spaces for wildlife shrink and corridors linking areas of habitat<br />

are cut, the survival of healthy species population, especially of larger and highly specialized<br />

animals becomes more threatened. Addressing these problems is a very complex issue, especially<br />

because indigenous peoples have diverse needs for, and claims to, these resources. Thus, the<br />

need for protected spaces for wildlife often seems at odds with human needs (Newmark, 1993).<br />

In North Ethiopia, heavy human pressure impacts on natural resources and biodiversity of the<br />

SMNP including agriculture, livestock grazing, collection of wood for fuel and timber, poaching<br />

and construction of road inside the National Park. Soil erosion, strongly affect the vegetation‟s<br />

potential for regeneration and wildlife habitats for Walia ibex and <strong>Ethiopian</strong> wolf (Keiner, 1999).<br />

18


Contacts and conflicts between wolves and humans are bound to increase with increasing human<br />

density and habitat fragmentation. Moreover, the highlands of Ethiopia are among Africa‟s most<br />

highly dense agricultural areas and the resulting human, agricultural, and wolf conflict has been<br />

devastating to the wolf population.<br />

In the heavily populated Northern highlands for example, livestock graze on high altitude<br />

pastures all year round. Circumstantial evidence also suggests that overgrazing in the densely<br />

populated highlands of the North may have negatively impacted upon the highland rodent fauna<br />

(Nievergelt et al., 1998) and that wolves may predate more frequently on livestock (lambs) or<br />

become nocturnal when human interference is severe (Yalden and Largen 1992). To prove this<br />

11 scat samples (fecal samples) collected from Mt. Abune Yoseph and analyzed by EWCP<br />

between February 1998 and February 1999 clearly revealed composition and diversity of the<br />

wolves diet in the area. Analysis of prey remains in 11 wolf scats (fecal samples), representing<br />

Mt. Abune Yoseph wolf populations (Table 1), confirmed that wolves are specialized hunters of<br />

diurnal rodents and rarely smaller livestock (lambs) across their distribution (Marino, 2003).<br />

Table 1: Composition and diversity of diet from wolves scat analysis at Mt. Abune Yoseph<br />

Prey items<br />

Mt. Abune Yoseph<br />

Otomys typus 36%<br />

Arvicanthis spp 23%<br />

Lophuromys spp 14%<br />

Stenocephalemys albocaudata 14%<br />

Praomys albipes 9%<br />

Livestock 5%<br />

(Source: Marino, 2003)<br />

19


In Northern highlands the livestock predation by wolves has resulted in negative attitudes<br />

amongst the local people, and sporadic killings, at least in South Wollo (Sillero-Zubiri et al.,<br />

1996; Ashenafi, 2001).<br />

In recent time, degradation of Afro-alpine ecosystem by grazing and high-altitude subsistence<br />

agriculture make wolf populations susceptible to extinction hazard due to human persecution,<br />

inbreeding, disease and natural catastrophes (Gottelli and Sillero-Zubiri, 1992; Sillero-Zubiri and<br />

Macdonald, 1997). Therefore, habitat protection to prevent or reduce further habitat loss would<br />

have to be given great attention to reduce the risk of extinction in particular of those populations<br />

in small areas.<br />

2.6. Socio-economic significance of Afro-alpine ecosystem<br />

The poor and hungry, to a greater extent than others, are directly dependent on functioning<br />

ecosystems and those services that the local ecosystems supply for their survival. Humans<br />

depend on biological resources for their existence. Communities engaged in subsistence<br />

agriculture as a way of life attempt to optimize their production systems by using resources taken<br />

from natural ecosystems (Kothari, 1997). Therefore, biodiversity is critically important to<br />

people‟s livelihoods because it provides diverse subsistence requirements for rural community<br />

including: fodder; fuel; housing; farming and household implements; and spiritual sustenance<br />

(Haverkort and Miller, 1994).<br />

Afro-alpine mountains due to their harsh climatic condition they are not serving as a settlement<br />

area or not occupied as individual property by rural dwellers rather they are managed as<br />

communal resource of adjacent subsistent farmers. In developing countries in general and in<br />

Ethiopia in particular rural communities depend primarily on common property resources for<br />

20


irrigation water, construction material, fuel wood, and grazing land (Berhanu and Scott, 2001;<br />

Hurni, 1986). Similarly, Afro-alpine ecosystems and their resources were exploited under<br />

unrestricted access by their surrounding community for millennia<br />

According to Malcom and Ashenafi (1997), Ethiopia‟s Afro-alpine ecosystem islands are<br />

immensely important with their diverse array of resources present in them for the livelihood of<br />

local communities who used the resources. Resources of Afro-alpine ecosystem that have<br />

significant socio-economic value for the surrounding communities are:<br />

Wild species: the plant and animal species endemic to the ecosystem have immediate economic<br />

value for the society. Grass of the Genus Festuca is used for thatch and basket making and there<br />

are species used for medicine, food and drinking that are harvested by people living around the<br />

ecosystem and sold. So the harvested resources are particularly important for the lower group of<br />

community as a cash crop and a coping mechanism, during severe draught season.<br />

Agricultural resources: even though mostly ended with failed result, the Afro-alpine ecosystem<br />

is used for barley cultivation by subsistence farmers below 3300 m a.s.l. depending on soils,<br />

slope and frost. The domestic animals use for grazing the entire alpine habitat. The area is also<br />

used as a refuge for far adjacent livestock herd during severe draught.<br />

Firewood: the ericaceous heathlands are important sources of firewood. There are Afro-alpine<br />

areas where firewood is gathered on commercial scale. Cherenfe (Euryops pinifolius), Asta<br />

(Erica arborea), Gebera (Lobelia rhynchopetalum), Amija (Hyphericum revolutum), Ashendye<br />

(Kniphopia foliosa) and Tinjut (Helychrisum splendidium) are the dominant shrubs being<br />

collected for household firewood consumption from the ecosystem. Firewood collection usually<br />

21


takes place in the dry season and a large volume is required due to its low calorific value. Cattle<br />

dung is burnt concurrently with these woods.<br />

Tourist values: the Afro-alpine habitats and escarpment provide both spectacular scenery and<br />

natural values with unusual animal life offer great opportunities for tourism. The endemic and<br />

rare species of the ecosystems and the diversified culture and heritage of the highlanders have<br />

potential for tourism. Generally the Afro-alpine ecosystems provide many attractions such as the<br />

scenery, wildlife and a wilderness appeal to potential visitors. In Ethiopia the BMNP and the<br />

SMNP are the two Afro-alpine national parks serve as a tourist destination for wildlife and<br />

highland visitors. Recently, nature based eco-tourism guided by local communities are emerged<br />

around Guassa and Mt. Abune Yoseph Afro-alpine ecosystems as a potential method of<br />

livelihood diversification and income generation for supporting the conservation management of<br />

the areas. Such type of community based tourisms is growing especially around Afro-alpine<br />

areas of Mt. Abune Yoseph by TESFA an <strong>Ethiopian</strong> NGO working with local community for the<br />

improvement of community livelihood through environmental friendly tourism. The Afro-alpine<br />

ecosystems also served as an area of sport hunting for tourists.<br />

Water resources: the high ground in the mountains catches more rain than the surrounding low<br />

lands. This is marked in simen where the low lands less than 35 km east of the peaks receive<br />

only one third of rain fall (Hurni, 1986). Afro-alpine ecosystems of Ethiopia form the watershed<br />

for the low lands of north-east Africa, providing water for millions of people and also used as a<br />

sources of mineral springs.<br />

22


2.7. Sustainable community based Afro-alpine ecosystem management<br />

As the human population grows, demands on natural resources have increased causing<br />

environmental degradation and fragmentation leading to a massive extinction of species (Myers,<br />

1979). To counteract this situation many developing countries design Environmental<br />

conservation policy characterized by a top-down approach that includes the establishment of<br />

state owned biodiversity protected areas. This approach reflects the suspicion of governments<br />

that local communities are incapable of managing their own resources (Ashenafi and N.Leader,<br />

2005). However, conservationists have been searching viable and sustainable alternative solution<br />

in the last 20 years, the approach that involve local communities in the process of resource<br />

management and decision making known as community-based conservation (IIED, 1994).<br />

Decentralization of resource management from the central authority to local communities is<br />

considered a first step forward for successful community-based program. Hence, local people<br />

have become involved more actively in the generation and distribution of benefits. It also<br />

promotes the legal and sustainable use of the wildlife and other natural resources.<br />

Sustainable development is multi-objective process whose goal is to satisfy real human needs ad<br />

infinitum while ensuring the protection of environmental quality, biodiversity and ecosystem<br />

resilience by integrating conservation and management with social and economic objectives at<br />

various social and spatial scales (Okigbo, 1996). For community-based conservation to be<br />

successful, it needs to be flexible enough to cope with a countryside inhabited by a growing<br />

number of extremely poor people who depend on a subsistence existence and whose greatest<br />

goal is to gain economic security (Alcorn, 1997).<br />

23


In Ethiopia the history of resource management reveals, overall, an increasing trend towards<br />

state control, and participatory resource management largely remains something that is talked<br />

about rather than implemented (Alula, 2001). But in contrast to this in Afro-alpine areas, there<br />

are traditional resource management systems practiced by local communities for century that are<br />

essential to the people‟s livelihoods and also to the persistence of biodiversity. For this the<br />

indigenous common property resource management system in the Guassa area of Menz, in the<br />

Central Highlands of Ethiopia, play a great role for the last hundred years in conserving the<br />

threatened Afro-alpine ecosystem biodiversity, including an important population of the<br />

<strong>Ethiopian</strong> wolf (Ashenafi, 2001; Ashenafi et al., 2005). Therefore, common property<br />

management institutions (traditional resource management systems, bylaws, etc.) are crucial for<br />

sustainable resource use and development. Above all, they are a reflection of the existing socioeconomic,<br />

political and cultural situation of the community (Ostrom, 1991). Institutional<br />

arrangements, such as rules and conventions, are clearly important elements in resource<br />

conservation (Murphree, 1993).<br />

In recent time community based protected areas are commonly established to conserve<br />

biodiversity, protected ecosystems, and maintain ecological processes. Many community<br />

protected areas also expected to contribute to sustainable development and poverty reduction<br />

(Nabin et al., 2008).<br />

24


Chapter III: Materials and Methods<br />

3.1. Study area<br />

3.1.1. Location<br />

The study was conducted at Mt. Abune Yoseph above 3700 m a.s.l in Lasta woreda, North Wollo<br />

administrative zone of Amhara National Regional State (ANRS). The administrative center of<br />

North Wollo zone is Woldia. It has mostly rural population of 1.5 million people living in 13<br />

woredas. Lasta Woreda is one district of North Wollo with its administrative center Lalibela.<br />

Mt. Abune Yoseph lies between 12 0 8‟ 7” N and 39 0 15‟ 7” E. It is one of the long isolated<br />

mountains in the Northern massif with a total area of 50 km 2 (EWCP, 2003) and the highest peak<br />

of North Wollo 4286 m a.s.l and the second peak of ANRS next to Ras Dashen 4620 m a.s.l<br />

(ESP, 2001). The massif is limited to the East by the fault escarpment of the Rift valley<br />

depression and separated from the Tigrean Plateau to the North and from the Simen Mountains<br />

(4,600 m of height) to the West by a chain of lower mountain systems (1500-2000 m a.s.l). In<br />

this study, the socio economic survey was conducted on user community of eight villages<br />

selected from four peasant Associations (PAs) found at the foothill and the ecologic survey was<br />

conducted on the mountain plateau above 3700 m a.s.l where the short afro-alpine vegetations<br />

and the wolf range is restricted due to agriculture of subsistent farmers.<br />

25


Lasta Woreda<br />

Figure 4: Location of the study Area<br />

26


3.1.2. Topography<br />

Mt. Abune Yoseph has rugged topography. It is a large peak surrounded by gorges and very<br />

steep slopes, where rocky blocks and stones are scattered all over. Three other high peaks<br />

surround the area: the Big Zigit (4,080 m), the Small Zigit (4,035 m) and the Reem Gedel (4,284<br />

m) a.s.l, providing impressive views over the Afro-alpine plateau and they are fabulously scenic<br />

although their value for tourism is undeveloped.<br />

3.1.3. Climate<br />

The climate of Mt. Abune Yoseph is characterized by a wet season from June to early<br />

September, and a dry season from mid-September to May. Annual rainfall averages 2,000 mm,<br />

mostly falling between July and September. Short rains might fall in any month of the year, but<br />

particularly in March. Generally the area has bi-modal rainfall known as „Meher‟ (long rainfall<br />

period: from June to September) and „Belg‟ (short rainfall period: from February to May). The<br />

mean annual temperature ranges from 7.5 o C to 11 o C which is characterized by moist and very<br />

cold conditions (ESP, 2001).<br />

3.1.4. Soil<br />

The effect of slope on Mt. Abune Yoseph peak is marked in its influence on the run-off, drainage<br />

and consequently upon the depth of the soil. So mountain areas that have steep slopes are liable<br />

to soil erosion and increased surface run-off, which will remove nutrients from the surface<br />

during high rainfall. Most parts of Mt. Abune Yoseph have rocky and shallow soils. The<br />

depression and flat plains are covered by deep black soils. Degraded landscapes are covered by<br />

shallow and stony soils (litho sols). Generally, in Mt. Abune Yoseph the distribution and type of<br />

27


soil differ from place to place based on the geology, physiographic variations and climate which<br />

are clearly observed on vegetation growth of the area.<br />

3.1.5. Vegetations and wild animals<br />

The Abune Yoseph massif shows a complex mosaic of ecosystems, where bush lands,<br />

woodlands, and Afro-alpine grasslands were represented. Remnant natural forests of Erica<br />

arborea patches are seen around the church compounds and at boundaries between mountain and<br />

farmland. Its Afro-alpine vegetations are characterized by giant lobelia, guassa grass and<br />

Euryops spp. which is locally known as „chifra‟ and „kirshiba‟. Grasslands mostly occur in<br />

association with the Euryops shrub. Its relatively steep slopes are covered by a mosaic of barley<br />

cultures and scattered patches of medium sized Afro-alpine grass meadows, shrubs, bushes and<br />

the endemic giant lobelias.<br />

Mt. Abune Yoseph has 43 species of mammals, from 19 families and nine orders (Saavedra,<br />

2009) and 221 bird species, belonging to 48 families and 16 orders (Lepage, 2006). From the 31<br />

endemic species of mammals known for Ethiopia and Eritrea 7 (22%) and from 16 endemic<br />

species of birds found in Ethiopia 6 (35%) have been found in the massif. Following the data of<br />

the Important Bird Areas of Ethiopia work (EWNHS, 1996) the Abune Yoseph massif could be<br />

considered to be the second most important bird area in the country (Saavedra, 2009).<br />

3.1.6. Water resources<br />

The North Wollo highlands including Mt. Abune Yoseph are the watersheds for the tributaries of<br />

three main river basins of Ethiopia, the Tekeze basin, the Awash basin and the Abay basin.<br />

Several streams intersect the area and feed the Tekeze River.<br />

28


3.2. Methods<br />

3.2.1. Interview<br />

Before formal household interview a preliminary survey aided with check list was conducted to<br />

gather background information about the ecosystem, the natural resources within the ecosystem<br />

and to have an understanding of the relationship between the ecosystem, wildlife, humans and<br />

domestic livestock and it also provide opportunity to contact with key informants present in the<br />

study area. Key informants were nominated from the four PAs surrounding Mt. Abune Yoseph.<br />

They were WEPLAUO experts, development agents, community leaders, elders and Para-<br />

Ecolologists from FZS and ordinary farmers who had lived in the area for long time and had<br />

historic background of the area. Then informal survey together with the key informants was<br />

conducted at the identified PAs. During the survey simple participatory survey techniques (focus<br />

group discussion, transect walk, etc.) that encourage dialogue and initiate informal discussions<br />

between farmers and researcher was selected as a tool. After we made long discussion with the<br />

key informants, based on informal survey results, we select eight villages that were used for<br />

household interview from the four PAs. These eight villages are found at the immediate<br />

proximity of Mt. Abune Yoseph at different distance and they bounded the mountain in all<br />

direction. These villages were Eyebelay and Korit from Wodebye PA, Shegla and Ybaro from<br />

Telfetit PA, Latgie and AbuneYoseph from AbuneYoseph PA, Kassegne and Enjafat from<br />

Enjafat PA.<br />

Finally using the checklist recorded at the preliminary study and the result of informal survey<br />

workable questionnaire for formal interview was developed and pre-tested by selecting two<br />

farmers from each village to check its accuracy and efficiency in eliciting the required data.<br />

29


Based on results of the pre-test, after some questions were revised and modified, it was<br />

administered to a sample of 120 households (6% of the total user households) selected randomly<br />

from the total 2014 households found in the eight villages of the four PAs who frequently visited<br />

the Afro-alpine area, either to herd their livestock, collect grass and firewood, or en route to local<br />

markets.<br />

The aim of the household interview was to generate and verify qualitative and quantitative data<br />

about the socio-economic potential of Mt. Abune Yoseph, and to assess the perception of the<br />

household heads towards the Afro-alpine ecosystem and its wildlife particularly <strong>Ethiopian</strong> wolf<br />

moreover, to evaluate human- wolf conflict in the study area.<br />

3.2.2. Population demography<br />

To get indicator data about human-livestock pressure or interferences on Afro-alpine ecosystem<br />

different years Government statistics (secondary data) on human and livestock population were<br />

collected from Afro-alpine ecosystems surrounding woredas of North Wollo.<br />

3.2.3. Ecological survey<br />

To study the presence/absence and abundance of wolf in the ecosystem, standard EWCP<br />

questionnaires, interviews, and sighting report were employed (Fanshawe et al., 1997; Sillero-<br />

Zubri et al., 2004). Presence /absence and distribution of rodents in relation to vegetation<br />

parameters and environmental factors were recorded along Habitat Assessment Transect (HAT)<br />

laid 1 km apart with the help of Universal Transverse Mercator (UTM) gridlines on a map of<br />

1:50,000 scale that include the study area (EMA, Lalibela, 1999) and Global Positioning System<br />

(GPS) (Burnham et al., 1979; Buckland et al., 1993 and Ashenafi et al., 2005). Measurements<br />

30


were taken within a circular sample plot measuring 5 m radius marked at 200 m interval along<br />

eight transects with a total distance of 22km and 110 sample plots 75 plots at above 3700-4000<br />

m and 35 plots above 4000 m a.s.l. At each sample plot data was recorded on slope, altitude,<br />

habitat type, dominant vegetation species, height and cover of plants, rodent holes, livestock<br />

dung and wolf droppings.<br />

3.3. Data Analysis<br />

All socio-economic and ecologic data were analyzed using the statistical packages SPSS (version<br />

13) and SAS-JMP 5 software. Data collected by interview: to determine wolf presence/absence<br />

and relative abundance, data on socio economic potential obtained from Mt. Abune Yoseph and<br />

data on human-wolf conflict were analyzed using descriptive statistics and the significance was<br />

tested by cross tabulation and chi-square test at (P


Chapter IV: Results and Discussion<br />

4.1. Socio-economic characteristic and economic potential of Mt. Abune<br />

Yoseph<br />

4.1.1. Household characteristics<br />

The characteristics of households 1<br />

(HHs) in the study area were analyzed with descriptive<br />

statistics (Table 2). Of sample household heads 2 (HHHs) 85.2% were male and 14.8% were<br />

females. From the HHHs 85.2% were married and 14.8% were unmarried. The age of<br />

respondents ranged from 18 to 80 years with a mean age of 47.7 years. The family size of the<br />

respondents ranged from a minimum of 1 to a maximum of 10 with a mean of 5.7. Of the HH<br />

respondents 52.5% were illiterate but 47.5% HHs were literate that can only write and read.<br />

From the sample HHs 86.7% had agricultural land holding with mean size of 0.7 ha and the rest<br />

13.3% were landless. Each sample HHs had a minimum of 0 and a maximum of 78 livestock<br />

with the mean number of 13 and standard deviation of 10. The sample HH respondents live<br />

within the range between 1km and 15 km away from Mt. Abune Yoseph.<br />

Table 2: Sample HHs characteristics of Mt. Abune Yoseph<br />

Sex N Marital status N Educational<br />

status<br />

N Land<br />

holding<br />

N Year of residence in<br />

their current location<br />

Male 103 Married 103 Illiterate 63 Landless 16 Min 6<br />

Female 17 Unmarried 71 Literate 57 Land 104 Max 80<br />

holder Mean 42<br />

SD 15.7<br />

1 Households are extended family living under one roof.<br />

2 Household heads are either male or female that are the central decision maker of the household.<br />

32


4.1.2. Socio-economic potential of Mt. Abune Yoseph Afro-alpine ecosystem<br />

The major economic activity of the people living around Mt. Abune Yoseph is subsistence<br />

agriculture, which include crop farming and animal husbandry. They support their livelihood by<br />

exploiting diversified goods and environmental services from the ecosystem. Most HHs (64.2%)<br />

use Mt. Abune Yoseph for 3 months to graze their livestock during the wet season and 61.7%<br />

HHs graze for 9 months during the dry season. From the respondents, 96 (80%) explained they<br />

got economic benefit from Mt. Abune Yoseph. According to the chi-square test there was a<br />

significant difference (p


Communities living closer to Mt. Abune Yoseph exploit effectively the natural resources<br />

throughout the year than those communities living at far distance (Table 4). Therefore, distance<br />

of the households from Mt. Abune Yoseph has a significant (P


(Table 5). The income generating degrees of these environmental goods and services as well as<br />

the HHs involvement vary in Mt. Abune Yoseph depending on some factors.<br />

Table 5: Type of economic benefits obtained from Mt. Abune Yoseph (n =120)<br />

No Benefit from Mt. Abune Yoseph<br />

Response<br />

yes % No %<br />

Firewood 89 74.2 31 25.8<br />

Thatching grass 76 63.3 44 36.7<br />

Hay grass 66 55 54 45<br />

Income from tourism 35 29.2 85 70.8<br />

Other benefit 92 76.7 28 23.3<br />

In cross tabulation analysis a chi-square test showed natural resource utilization from Mt. Abune<br />

Yoseph differ (X 2 =4.62, df =1, P0.05) in economic resource utilization between HHs<br />

based on HHs age, marital status, family size, sex, educational status, livestock number and farm<br />

size.<br />

35


Figure 5:.Land ownership status and economic benefit from Mt. Abune Yoseph<br />

In addition to self consumption and livestock grazing cost each HH got a direct economic benefit<br />

(Table 6) of 20.00 birr per load from firewood selling (Figure 6) and a maximum of 3000.00 birr<br />

from tourism, generated through guiding tourists, renting pack animals and selling locally made<br />

items in 12 months. Generally, 29.2% HHs which distributed across 8 villages got economic<br />

benefit from tourism (Table 5).<br />

Table 6: Price of major economic goods and services from Mt. Abune Yoseph in 1year (n =120)<br />

Item Range Mean Standard Error<br />

Firewood market price in birr per load 0-20 7.06 0.507<br />

Thatching grass market price in birr per load 0-50 19.54 1.639<br />

Hay grass market price in birr per load 0-50 15.25 1.482<br />

Birr obtained from tourism 0-3000 192.25 42.188<br />

36


Figure 6: People selling firewood in the market<br />

HHs also estimated the average economic benefit they got in terms of money including the cost<br />

of grazing and other all ecosystem goods and services (firewood, thatching grass, hay grass,<br />

tourism, construction material etc) from Mt. Abune Yoseph a range of 0 to 20,000 birr with<br />

mean of 1627±207 birr in one year (Table 7).<br />

Table 7: HHs average estimated birr obtained from Mt. Abune Yoseph in one year<br />

N Range Mean ± SE<br />

Average estimated birr per year 120 0-20,000 1627.46± 206.914<br />

37


Even though the pattern of resource utilization differ between HHs, the result of this study has<br />

shown clearly that the community in Mt. Abune Yoseph consider the Afro-alpine ecosystem of<br />

the area to be important for their livelihoods and on many occasions described as it improving<br />

their livelihood. Similar study on ecosystem showed Communities engaged in subsistence<br />

agriculture as a way of life attempt to optimize their production systems by using a diversity of<br />

crop-livestock systems, supplemented by resources taken from natural ecosystems (Kothari,<br />

1997). Therefore, biodiversity from the ecosystem is critically important to people‟s livelihoods.<br />

4.2. Community perception towards Mt. Abune Yoseph Afro-alpine ecosystem<br />

The perception of HHs towards Mt. Abune Yoseph Afro-alpine ecosystem was assessed and<br />

analyzed with logistic regression by taking the HHs response on need to protect Mt. Abune<br />

Yoseph dependent variable. The dependent variable was taken as dummy: 0 if the response on<br />

need to protect Mt. Abune Yoseph was negative and 1 if the response was positive. The<br />

explanatory variables examined during the logistic regression were age; sex; length of residence<br />

in the area; educational status; marital status; family size; distance from Mt. Abune Yoseph, land<br />

access and livestock number. In addition the use or nonuse of natural resources such as firewood,<br />

thatching grass, hay grass and the presence/absence of wild animal danger at Mt. Abune Yoseph<br />

were taken as explanatory variables.<br />

38


Table 8: Distribution of Nominal variables employed in logistic regression<br />

Nominal Variables<br />

Number Percent<br />

Sex Male 103 85.8<br />

Female 17 14.2<br />

Marital status Married 103 85.8<br />

Single 17 14.2<br />

Educational status Illiterate 63 52.5<br />

Literate 57 47.5<br />

Land access Yes 104 86.7<br />

No 16 13.3<br />

Firewood Yes 89 74.2<br />

No 31 25.8<br />

Thatch grass yes 76 63.3<br />

No 44 36.7<br />

Hay grass yes 66 55<br />

No 54 45<br />

Wild animal danger at Mt. Abune Yoseph Yes 108 90<br />

No 12 10<br />

Need to protect Mt. Abune Yoseph Yes 85 70.8<br />

No 35 29.2<br />

<strong>Ethiopian</strong> wolf-human existence relation Yes 78 65<br />

No 42 35<br />

Local attitude towards <strong>Ethiopian</strong> wolf Positive 79 65.8<br />

Negative 41 34.2<br />

Table 9: Descriptives of continuous variables employed in logistic regression (n=120)<br />

Continuous Variable<br />

Min Max Mean Std Deviation<br />

Age of sample HH 18 80 47.5 12.268<br />

Family size of the HH 1 10 5.7 1.827<br />

Distance of HH from AY 2 15 7.72 4.199<br />

Livestock possession of the HH 0 78 13.69 10.509<br />

39


Table 10: Factors determining the perception of respondents towards Mt. Abune Yoseph Afroalpine<br />

ecosystem and its management<br />

Explanatory variable Estimate Std Error Chi-Square P-value<br />

Intercept 1.683332 1.560696 1.16 0.2808<br />

Age 0.007933 0.0346202 0.05 0.8188<br />

Sex[female] 0.36493 0.5590733 0.43 0.5139<br />

Marital status[married] 0.414794 0.5574251 0.55 0.4568<br />

Family size -0.23529 0.1752716 1.8 0.1795<br />

Educational status[illiterate] 1.38216 0.3684552 14.07 0.0002<br />

Living length at Mt. Abune Yoseph -0.01641 0.0243986 0.45 0.5011<br />

Distance from Mt. Abune Yoseph -0.24994 0.0959698 6.78 0.0092<br />

Land access [NO] 1.308343 0.4655066 7.9 0.0049<br />

Livestock number 0.085859 0.0366595 5.49 0.0192<br />

Firewood[NO] 0.043504 0.4976789 0.01 0.9303<br />

Thatching grass[NO] 0.096351 0.4358704 0.05 0.825<br />

Hay[NO] 0.216391 0.3881591 0.31 0.5772<br />

Wild animal danger at Mt. Abune Yoseph[NO] -0.37451 0.4248673 0.78 0.3781<br />

R Square (U) 0.317<br />

Observations (or Sum Wgts) 120<br />

The community support the protection of Mt. Abune Yoseph Afro-alpine ecosystem for various<br />

reasons but the perception towards the need to protect its natural resources differ between<br />

communities based on different socio-economic characteristic and pattern of environmental<br />

goods and services harvested from the ecosystem and wild animal danger occurred in the area<br />

(Table 10).<br />

There was a significant difference (P


the habitat when compared with illiterate households. The support of literate households might<br />

be due to the concept they got on conservation of natural resources from formal and informal<br />

education during their stay at school. Therefore, non educated HHs had negative perception<br />

towards Mt. Abune Yoseph Afro-alpine ecosystem and they did not also support its sustainable<br />

management when compared with educated HHs of the area. Study conducted on Masindi<br />

District CBFM by (Kugonza et al., 2009) in Uganda showed the same result, i.e. higher<br />

percentage of illiterate respondents were unwilling to participate in CBFM mainly because they<br />

miss out necessary information and training on sustainable tree conservation.<br />

Similarly, there was a significant difference (P


management (Table 10). This means that landless HHs did not show positive perception for the<br />

conservation of the ecosystem. This might be associated with their interest to get private<br />

farmland from the ecosystem and as well as to exploit the resources of the ecosystem freely for<br />

their basic needs. Study in Masindi District confirmed, farmers that have secured private land<br />

ownership showed participation in CBFM than farmers using government land or land not<br />

claimed secured private ownership (Kugonza et al., 2009).<br />

In Mt. Abune Yoseph besides farming, livestock rearing is the other economic activity that<br />

supports crop cultivation and HH‟s economy. The major livestock kept by the community in the<br />

area are cattle, sheep, donkey, horse and mules. In the area cattle are responsible for ploughing<br />

and threshing harvested crops. They are also important for providing meat, milk and milk<br />

products.<br />

Logistic regression results, showed the presence of significant difference (P< 0.05) between HHs<br />

by livestock number in their perception towards Mt. Abune Yoseph Afro-alpine ecosystem<br />

conservation (Table 10). So HHs that has more heard size support or have positive perception<br />

towards Mt. Abune Yoseph Afro-alpine ecosystem and its conservation management than those<br />

had less and none livestock number. Based on informal survey and discussion made with key<br />

informants, since HHs with more heard size did not have enough private grazing land, if<br />

conserved they could compensate their livestock feed shortage by cut and carry system.<br />

4.3. Community perception towards <strong>Ethiopian</strong> wolf<br />

To analyze the perception of community towards <strong>Ethiopian</strong> wolf, logistic regression was used to<br />

model responses, as it provided a convenient way to undertake categorical data analysis. In<br />

practice the analysis and interpretation are quite similar to the well-known procedure of multiple<br />

42


egressions (Freeman, 1987). To find out how the community perceives the rare and critically<br />

endangered wildlife of the area, a dummy of the respondent‟s attitude, 0 if the local community<br />

attitude response towards <strong>Ethiopian</strong> wolf was negative and 1 if the response was positive taken<br />

as dependent variable. The explanatory variables age, sex, marital status, family size, educational<br />

status, distance from Mt. Abune Yoseph, land access, livestock number, response on wildlife<br />

danger and <strong>Ethiopian</strong> wolf human coexistence relation at Mt. Abune Yoseph were related to the<br />

perception response to the wolf (Table 8 and 9).<br />

The respondents‟ distance from the Mt. Abune Yoseph associated with different (P


Table 11: Factors determining the perception of respondents towards <strong>Ethiopian</strong> wolves<br />

Explanatory variable Estimate Std Error Chi-Square P-value<br />

Intercept -2.49328 1.3014543 3.67 0.0554<br />

Age 0.007224 0.0243306 0.09 0.7665<br />

Sex[female] -0.77332 0.4805182 2.59 0.1075<br />

Marital status[married] -0.59374 0.4760479 1.56 0.2123<br />

Family size 0.258119 0.1512715 2.91 0.0879<br />

Educational status[illiterate] 0.381671 0.2883903 1.75 0.1857<br />

Distance from Mt. Abune Yoseph -0.14242 0.0580992 6.01 0.0142<br />

Land access [NO] 0.103772 0.4061492 0.07 0.7983<br />

Livestock number 0.134623 0.0445717 9.12 0.0025<br />

Wild animal danger at Mt. Abune Yoseph[NO] -1.19366 0.3766712 10.04 0.0015<br />

<strong>Ethiopian</strong> wolf -human existence Relation[No] 0.103766 0.2356485 0.19 0.6597<br />

R Square (U) 0.2335<br />

Observations (or Sum Wgts) 120<br />

Among the people who thought the <strong>Ethiopian</strong> wolf was a bad species, most gave sheep predation<br />

by the wolf as their main reason (Table 12). Carnivores commonly generate negative attitude<br />

among the rural residents in many regions of the world since they prey upon domestic animals<br />

(Bauer, 2003; Woodroffe, 2000). On the other hand, respondents that considered <strong>Ethiopian</strong><br />

wolves a good species mentions various reasons including their potential to attract tourism,<br />

entertainment, endemic and heritage value and their utilitarian value in controlling rodents<br />

(Table 12). According to Ashenafi (2001), large number of residents from Guassa area knew that<br />

the <strong>Ethiopian</strong> wolf is an endemic species. In addition, as reported by Deresse (2003), the<br />

<strong>Ethiopian</strong> wolf has the potential to attract tourists and bring income to their respective areas. A<br />

study in Minnesota, U.S.A. showed that there was strong positive attitude towards timber wolf<br />

44


(Canis lupus) and agreed that timber wolf was symbolic of nature‟s wonder and beauty (Kellert,<br />

1985).<br />

Table 12: HHs reason why they perceive wolves good or bad species at Mt. Abune Yoseph<br />

snoRaeR Number Percent<br />

<strong>Ethiopian</strong> wolf is not good species due to sheep predation 35 29.2<br />

Since means of tourism income 22 18.3<br />

Since endemic, heritage, and visited by tourists for entertainment 49 40.8<br />

Since rodent hunter protect crop damage 14 11.7<br />

Total 120 100.0<br />

The perception towards <strong>Ethiopian</strong> wolf also associated with livestock number of HHs. There was<br />

a significant difference (P


indicated in Table 8 than the study time of (Marino, 2003) in which (90%) of people reported<br />

negative attitude towards <strong>Ethiopian</strong> wolf in North Wollo highlands. The decrease of local<br />

community negative attitude towards <strong>Ethiopian</strong> wolf might be associated with the education and<br />

awareness raising done by EWCP and still by FZS in partner with GOs for the last decades<br />

through out the country in general and in Afro-alpine ecosystems in particular.<br />

4.4. Human-livestock interference to Afro-alpine ecosystem and the resulting<br />

conflict<br />

4.4.1. Human-livestock interference pressure and its impact<br />

The trend of human and livestock population has increased continuously in and around the Afroalpine<br />

ecosystem of North Wollo highlands (Table13). At present the human population density<br />

in and around Afro-alpine ecosystem of North Wollo woredas on average is greater than 120<br />

people per km 2 .<br />

Table 13: Human population in woredas that have Afro-alpine ecosystem in North Wollo<br />

Woreda<br />

Area Population Per Year And Density Per Km 2<br />

Inkm 2 2000 Density 2002 Density 2004 density 2007 density<br />

Lasta* 2844 185395 65.2 191938 67.5 196468 69 201653 70.9<br />

Delanta* 1529 165717 108.4 171300 112 178448 116.7 203385 133<br />

Gidan 1236 158602 128.3 158602 128.3 160558 129.9 172537 139.6<br />

G/laftu 1037 144506 139.4 160357 154.7 167446 161.5 170289 164<br />

Lasta* includes the new Bugna woreda<br />

(Source: ANRS BoFAED different years Statistical Bulletins)<br />

Delanta* includes the new Dawnt woreda<br />

In Africa, the major problem facing protected areas today is the increase in human settlement of<br />

adjacent lands and the unauthorized harvesting of resources within the protected areas<br />

46


(Newmark, et al., 1993). This high number of human population has its own impact on the<br />

wildlife population and their ecosystem. As the number of human population increases,<br />

encroachment increases; then the extent of resource exploitation also increases.<br />

This can easily be observed by increase livestock grazing in Mt. Abune Yoseph Afro-alpine<br />

ecosystem. The mean period of utilization of Mt. Abune Yoseph Afro-alpine ecosystem as a<br />

grazing land was 8.11 ± 3 months in the year and 2.34 ± 1 months during wet season and the<br />

overall range of grazing period in the ecosystem was 0 -11 months (Table 14).<br />

Table 14: Livestock grazing period at Mt. Abune Yoseph (n =120)<br />

Grazing Period Range Mean ± SD<br />

Grazing months in the year 0-11 8.11± 3.092<br />

Wet season grazing months 0-3 2.34 ± 1.017<br />

47


Table 15: Livestock population growth trend in woredas that have Afro-alpine ecosystem<br />

Woreda Area/ Km 2 / Livestock 2000 2002 2004 2007<br />

Lasta* 2844 Cattle 136924 153875 166197 187381<br />

Sheep and goat 137321 144186 165600 205539<br />

Pack animal 13001 20918 24819 31618<br />

Total livestock 287246 318979 356616 424538<br />

Density /km 2 101 112.2 125.4 149.3<br />

Delanta* 1529 Cattle 93141 108283 114173 123335<br />

Sheep and goat 130280 182648 188719 196636<br />

Pack animal 16709 26392 27639 44480<br />

Total livestock 240130 317322 330531 364451<br />

Density /km 2 157 207.5 216.2 238.4<br />

Gidan 1236 Cattle 72287 82341 85422 87500<br />

Sheep and goat 157149 161900 171430 225639<br />

Pack animal 13861 21471 22316 25321<br />

Total livestock 243297 265712 279168 338460<br />

Density /km 2 196.8 215 225.9 273.8<br />

Gubalaftu 1037 Cattle 81260 99134 105020 115025<br />

Sheep and goat 93888 114404 116579 139619<br />

Pack animal 12198 13693 18293 19862<br />

Total livestock 187346 227231 239892 247506<br />

Density /km 2 180.7 219 231.3 238.7<br />

Lasta* includes the new Bugna woreda Delanta* includes the new Dawnt woreda<br />

(Source: North Wollo zone ARDD)<br />

The year to year abrupt increase of human and livestock population in North Wollo especially in<br />

woredas that have Afro-alpine ecosystem resulted in increase in settlement and encroachment of<br />

livestock and dogs that poses serious threats to the <strong>Ethiopian</strong> wolf within the ecosystem. Similar<br />

study in BMNP by Stephens et al. (2001) proved that the abrupt increasing of settlement and<br />

livestock number around protected area has to be given major concern for three reasons. First for<br />

the erosive effect of livestock and its detrimental effect on natural flora and fauna of the<br />

ecosystem, second the presence of large number of domestic stock in the area impaired and has<br />

negative visual impact on tourist activities. Third, result in interference competition with<br />

wildlife, such as dogs compete with wolves for prey, spread disease and hybridize with wolves<br />

(Sillero-Zubiri, 1994). Therefore, the rapid increase of human (Table 13) and livestock<br />

48


population (Table 15) in Afro-alpine woredas of North Wollo in general and around Mt. Abune<br />

Yoseph in particular indicate a clear anthropogenic threat on wildlife. In addition to this, the<br />

cattle and human population continuously devastated the habitat of the <strong>Ethiopian</strong> wolf. As a<br />

result, according to 75% of sample HHs report the <strong>Ethiopian</strong> wolf number at Mt. Abune Yoseph<br />

decreased (Table 20).<br />

The reason forwarded (Table 16) by the sample HHs for wolf decrease in Mt. Abune Yoseph<br />

were migration of <strong>Ethiopian</strong> wolf to the areas where the human activity is less, human-livestock<br />

encroachment to the ecosystem, drought due to deforestation and the competition between the<br />

<strong>Ethiopian</strong> wolf and common jackal have shown clearly the human livestock interference impact<br />

on the ecosystem and the resulted conflict with wildlife.<br />

Table 16: Reasons for change in wolf number trend at Mt. Abune Yoseph (n=120)<br />

snoRaeR Number %<br />

Increased since protected 16 13.3<br />

Increased since personal or dog disturbance is forbidden 14 11.7<br />

Decreased by draught due to deforestation 21 17.5<br />

Decreased due to migration 26 21.7<br />

Decreased due to encroachment 22 18.5<br />

Decreased due to competition with common jackal 37 7121<br />

Total 120 100<br />

In Mt. Abune Yoseph Afro-alpine ecosystem grazing competition between livestock and gelada<br />

baboon which leads to overgrazing is also a common phenomenon (personal observation).<br />

49


Fire wood collection was another type of community activity which has a detrimental impact in<br />

the study area. The community living around the Mt. Abune Yoseph Afro-alpine ecosystem used<br />

different types of Afro-alpine plant species from the ecosystem for firewood (Figure 6). Fifty<br />

seven percent sample HHs report Cherenfe (Euryphos pinifolius), Asta (Erica arborea) and<br />

Amija (Hyphericum revolutum) were dominant species used for firewood in the area while the<br />

rest 43% respondents reported Cherenfe, Asta and Gebera (Lobelia rhynchopetalum) were the<br />

most common. Similarly, many (64.5%) respondents indicated that they collect shrubby<br />

vegetation in Menz-Guassa for firewood. The most commonly collected plant species in Guassa<br />

was Cherenfe (E. pinifolius). Other species including: Erica or Asta, (E. arborea); Nechilo (H.<br />

splendidum); St.John‟s wort or Ameja (H. revolutum); and, dry giant Lobelia or Gebera (L.<br />

rhynchopetalum) are also collected (Ashenafi, 2001). In our study Most (63.3%) of the<br />

respondents mentioned the reason why these plant species preferred was, since these were the<br />

only growing plant species in the area where as 36.7% explained these species since they are the<br />

highly flammable firewood species in the ecosystem. Such uncontrolled firewood exploitation<br />

throughout the year had a great impact on the quality of habitats for rodent species, which are the<br />

prime diet of the <strong>Ethiopian</strong> wolf.<br />

In addition, from discussion with key informants, this firewood collection activity has resulted in<br />

ecosystem degradation due to extreme erosion and gully formation. In turn due to erosion, the<br />

land fertility around the ecosystem has been decreasing gradually from year to year, the crop<br />

production in the area decreasing from time to time and finally framers surrounding the<br />

ecosystem obliged to push their farm in to the ecosystem to cultivate more land. As a result,<br />

there were more farmlands inside the Mt. Abune Yoseph Afro-alpine ecosystem and its<br />

50


surrounding. This has resulted in continuous land clearing leading to habitat fragmentation and<br />

loss of biodiversity (Figure 7).<br />

Figure 7: Farming inside Afro-alpine ecosystem by clearing lobelia<br />

From the community living at the hill foot of Mt. Abune Yoseph 90% of sample HHs report as<br />

they faced conflict with wildlife (Table 17). Almost similar findings were observed from the<br />

study conducted in Tanzania on the conflict between wildlife and local people living adjacent to<br />

protected areas. Over 71% of local people surveyed, reported conflict with wildlife (Newmark et<br />

al., 1994).<br />

Table 17: HHs response on whether or not faced wild animal problem<br />

snRoaeRn Number Percent<br />

Not faced problem 12 10.0<br />

Faced problem 108 90.0<br />

Total 120 100.0<br />

51


The conflict between wildlife and local people in and around the Mt. Abune Yoseph Afro-alpine<br />

ecosystem involved livestock predation and crop damage. From those sample HHs who reported<br />

causes of conflict to wildlife 30.8% reported livestock predation, 9.2% reported crop damage,<br />

while 50% reported both livestock predation and crop damage (Table 18).<br />

Based on descriptive statistics and chi-square test the problem caused by wildlife differed<br />

(P


4.4.2. Human conflict with <strong>Ethiopian</strong> wolf at Mt. Abune Yoseph<br />

In Mt. Abune Yoseph <strong>Ethiopian</strong> wolf (C. simenesis) and common jackal (Canis aureus) was the<br />

most common sheep predators singled out by the sample HHs. Leopard (Panthera pardus) and<br />

spotted hyena (Crocuta crocuta) were also predating sheep in the area (Table 19).<br />

Table 19: Mean ±SE number of sheep preyed in the last 10 years at Mt. Abune Yoseph (n=120)<br />

Range Mean ± SE<br />

Sheep preyed by <strong>Ethiopian</strong> wolf in 10 years 0-10 4.26±0.357<br />

Sheep preyed by Common jackal in 10 years 0-15 5.14±0.539<br />

Sheep preyed by Leopard in 10 years 0-4 0.83±0.137<br />

Sheep preyed by Spotted hyena in 10 years 0-7 1.15±0.21<br />

Comparing the most sheep predators of the area 25.8% HHs responded sheep loss due to the<br />

<strong>Ethiopian</strong> wolf, 15% lost due to the common jackals, 36.7% lost sheep due to both the <strong>Ethiopian</strong><br />

wolf and/or common jackal and the rest 22.5% HHs reported no sheep loss due to these two<br />

predators in the last 10 years (Figure 8). Therefore, according to the HHs response both<br />

<strong>Ethiopian</strong> wolf and Common jackal are the main sheep predator of the area.<br />

53


Sheep predators<br />

Figure 8: HHs response on sheep predators at Mt. Abune Yoseph for the last 10 years<br />

Most HHs responded <strong>Ethiopian</strong> wolf was the main sheep predator of the area when compared<br />

with Common jackal in the last 10 years (Figure 8). The same result was reported by (Marino,<br />

2003) in that since the communities have lost many animals, they perceived <strong>Ethiopian</strong> wolves as<br />

the main predators of livestock in North Wollo highlands. Even though more HHs blamed<br />

<strong>Ethiopian</strong> wolf as a major sheep predator in their response illustrated in Figure 8, the number of<br />

sheep killed in the indicated years showed more sheep was preyed by Common jackal than<br />

<strong>Ethiopian</strong> wolf (Table 19). This is due to <strong>Ethiopian</strong> wolf attack less sheep number from most<br />

HHs where as Common jackal killed more sheep number from less HHs. Similar result was<br />

observed by (Marino, 2003) in SMNP, cases of livestock losses to <strong>Ethiopian</strong> wolves were<br />

reported, but common jackal (C. aureus) were indicated to be the main predator.<br />

54


Based on HHs response, on predation rate of <strong>Ethiopian</strong> wolf, Common jackal and both, the<br />

present finding differ from previous works conducted in Menz by Ashenafi (2001) and around<br />

SMNP by Messele (2006). According to Ashenafi‟s finding only 14.5% of Menz-Guassa sample<br />

community reported sheep loss to the <strong>Ethiopian</strong> wolf and 40.3% to Common jackal. Where as<br />

according to Messele‟s study only 18.3% of Simen sample community reported sheep loss to the<br />

<strong>Ethiopian</strong> wolf and 8% to both the <strong>Ethiopian</strong> wolf and Common jackal. But in present study<br />

most sample HHs from Mt. Abune Yoseph explained as they lost more sheep by both (<strong>Ethiopian</strong><br />

wolf and Common jackal) and <strong>Ethiopian</strong> wolf alone respectively (Figure 8). This difference<br />

might be associated with lack of comparing the degree of sheep loss within 10 years at Mt.<br />

Abune Yoseph by sample HHs between common sheep predator or due to minimized livestock<br />

encroachment to the wolf habitats in Simen (protected as National Park) and Guassa protected<br />

for more than hundred years by Qero system of Menz community and still managed by<br />

committee of community (Ashenafi, 2001, and Ashenafi et al., 2005). In the same study Messele<br />

(2006) mentioned the reason for the less number of sheep loss to the <strong>Ethiopian</strong> wolf in SMNP is<br />

that the distribution of the <strong>Ethiopian</strong> wolf is very limited in contrast to the Common jackal. At<br />

the same time, its habitat is far from the human settlement area. However, Messele also reported<br />

in his finding humans attack the <strong>Ethiopian</strong> wolf in some parts of the study area.<br />

4.5. Ecological assessment of Mt. Abune Yoseph Afro-alpine ecosystem<br />

4.5.1. Households response on <strong>Ethiopian</strong> wolf presence/ absence<br />

All HHs (100%) without any difference responded as they observed wolf at Mt. Abune Yoseph<br />

before and had enough knowledge about it (Table 20).<br />

55


Table 20: HHs response on wolf observation, observation time and trend at Mt. Abune Yoseph<br />

Response Number %<br />

HHs seen wolf at Mt. Abune Yoseph 120 100<br />

<strong>Wolf</strong> observation time at Mt. Abune Yoseph Dawn 49 40.8<br />

Dawn & dusk 71 59.2<br />

<strong>Wolf</strong> trend at Mt. Abune Yoseph Increasing 30 25<br />

Decreasing 90 75<br />

Each sample HH had seen <strong>Ethiopian</strong> wolf in a mean number of 6 ± 0.7 times in the last 12<br />

months and 13 ±1.7 times in the last 5 years (Table 21). Most 59.2% of respondents observed<br />

<strong>Ethiopian</strong> wolf at dawn and dusk while 40.8% of sample HHs observed only during dawn time at<br />

Mt. Abune Yoseph (Table 20). The large mean number of <strong>Ethiopian</strong> wolves seen at once in Mt.<br />

Abune Yoseph by respondents was 4±0.2 (Table 21).<br />

Table 21: HHs wolf observation frequency, wolf seen at once and wolf estimation at Mt. Abune<br />

Yoseph (n =120)<br />

Range Mean ±SE<br />

<strong>Ethiopian</strong> wolf observation frequency in the last 12 months 0-60 6.01±0.670<br />

<strong>Ethiopian</strong> wolf observation frequency in the last 5 years 1-150 12.96±1.703<br />

Large number of wolf seen by HHs at once in Abune Yoseph 1-12 4.20±0.209<br />

HHs estimate number of <strong>Ethiopian</strong> wolf at Abune Yoseph 1-60 18.93±1.225<br />

Most (75%) HHs reported the wolf number was decreased in comparison with the past times; in<br />

the contrary 25 % HHs reported as the wolf number increased in Mt. Abune Yoseph (Table 20).<br />

Opposite result was reported by (Marino, 2003) many respondents (80%) at that time believed<br />

that wolves were recovering from past persecution in North Wollo highlands. This might be<br />

considered from persecution to wolves by dog and local community rather than relating the trend<br />

with its habitat loss, because in our study the few respondents that supported the increasing trend<br />

56


of <strong>Ethiopian</strong> wolf in Mt. Abune Yoseph gave absence of wolf persecution from local people and<br />

dogs as reason, i.e. protection of the ecosystem by the guard and the restriction law by the local<br />

authority on farmers and herders not to let in with dogs into the ecosystem were mentioned for<br />

wolf increasing trend as reason (Table 16).<br />

Based on sample HH‟s estimate the current time <strong>Ethiopian</strong> wolf number at Mt. Abune Yoseph<br />

was analyzed with descriptive statistics and gave mean number of 18.9 ±1.3 wolves (Table 21).<br />

These numbers are consistent with recent estimations (Sillero-Zubri et al., 2000; Marino, 2003)<br />

that suggest an amount of wolves ranging between 19 and 23 for the three massifs of North<br />

Wollo with wolf‟s presence (Mt. Abune Yoseph, Mt.Abohoy Gara and Delanta) totaling 140 km 2<br />

of available habitat.<br />

In descriptive statistics with cross tabulation chi-square test indicated the presence of significant<br />

difference(X 2 =33.5, df =21, P


time on wolf number enabled the HHs to estimate the nearest wolf number during the interview.<br />

There was also a difference (X 2 =773.9, df =714, P0.05) between HHs based on sample villages, age, marital<br />

status, length of living years, distance, land ownership status and land size in wolf number<br />

estimation between HHs at Mt. Abune Yoseph.<br />

4.5.2. Determination of wolf distribution by rodent burrow<br />

In this study one way analysis of variance showed slope has significant effect (P


40<br />

35<br />

30<br />

Rodent burrow<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

-5<br />

flat moderate steep<br />

S<br />

All Pairs<br />

Tukey-Kramer<br />

0.05<br />

Figure 9: One way Analysis of Rodent burrows by Slope<br />

Table 22: Analysis of rodent burrows by slope and mean comparison using Tukey-Kramer HSD<br />

Source DF Sum of<br />

Squares<br />

Mean<br />

Square<br />

F Ratio P-value Comparisons for all pairs<br />

using Tukey-Kramer HSD<br />

Levels*<br />

Mean<br />

Slope 2 162.62 81.31 3.2244 0.0437 Flat A 4241<br />

Error 107 2698.15 25.22 eafneoon A B 3223<br />

C. Total 109 2860.76 steep B 7274<br />

*Levels not connected by the same letter are significantly different<br />

There was a significant linear association (P


Table 23: Bivariate fit analysis of rodent burrow by altitude<br />

Factor Estimate Std Error t Ratio P-value<br />

Intercept -97.82648 19.36096 -5.05 0.0001<br />

Alt 0.0253655 0.00489 5.19 0.0001<br />

R Square 0.199464<br />

Observations ( Sum Wgts) 771<br />

Afro-alpine plant height showed significant effect (P


Table 24: Analysis of rodent burrow by vegetation height and mean comparison using Tukey-<br />

Kramer HSD<br />

Source DF Sum of<br />

Squares<br />

Mean<br />

Square<br />

F Ratio P-value Comparisons for all pairs<br />

using Tukey-Kramer HSD<br />

lnvnl *<br />

enoe<br />

Vegetation 3 390.8559 130.285 5.5914 0.0013 0-10 cm A 5.00<br />

height<br />

Error 106 2469.9078 23.301 11-30 cm A B 1.47<br />

C. Total 109 2860.7636 31-50 cm B 1.31<br />

>50 cm B 0.58<br />

*Levels not connected by the same letter are significantly different<br />

Figure 11: Short Afro-alpine vegetation (E. pinifolius) with rodent burrow from Mt. Abune<br />

Yoseph<br />

Afro-alpine vegetation cover did not show significant effect (P> 0.05) in determining rodent<br />

burrow distribution at Mt. Abune Yoseph (Table 25). This is different from the literature section<br />

61


in 2.2.2; this difference could be due to different rodent communities from other Afro-alpine<br />

ecosystem such as from Bale or due to high impact by livestock at Mt. Abune Yoseph.<br />

Table 25: Analysis of rodent burrow by vegetation cover and means of rodent burrow at each<br />

level of vegetation cover<br />

Source DF Sum of Mean F P-value Mean for one way ANOVA<br />

Squares Square Ratio<br />

Level* N Mean SDerror<br />

Vegetation cover 3 38.86 12.95 0.4866 0.6923 26-50% 17 2.53 1.2514<br />

Error 106 2821.90 26.62 5-25% 11 3.91 1.5557<br />

C. Total 109 2860.76 51-75% 45 2.00 0.7692<br />

76-100% 37 2.92 0.8482<br />

4.5.3. Determination of <strong>Ethiopian</strong> wolf abundance by sighting at Mt. Abune<br />

Yoseph<br />

The one month sighting of <strong>Ethiopian</strong> wolf at Mt. Abune Yoseph was used to estimate the current<br />

status of <strong>Ethiopian</strong> wolf and also proved the result of habitat assessment transect with similar<br />

analysis using one way ANOVA and bivariate analysis with the same explanatory variable<br />

Altitude, slope, vegetation height.<br />

Bivariate analysis showed the presence of significant linear relation (P


observed in the Afro-alpine belt on 13 different days during the visits to this habitat from 1998 to<br />

2002. On November 30 th 2002 they counted four different individuals, three in the Reem Gedel<br />

plateau and another one in a valley east of Reem Gedel. They were seen walking and resting in<br />

open moorlands and grasslands of the massif, mainly in the Reem Gedel plateau (between 3,950<br />

and 4,150 m a.s.l.).<br />

Table 26: Bivariate analysis of wolf sighting by altitude<br />

Term Estimate SDError t Ratio P-value<br />

Intercept -18.14916 6.529746 -2.78 0.0096<br />

Altitude 0.0048631 0.001638 2.97 0.0061<br />

R square 0.239464<br />

Observations ( Sum Wgts) 21<br />

There was also a significant difference (P


Table 27: Analysis of wolf sighting by slope and mean comparisons using Tukey-Kramer HSD<br />

Source<br />

DF<br />

Sum of<br />

Squares<br />

Mean<br />

Square<br />

F Ratio P-value Comparisons for all<br />

pairs using Tukey-Kramer HSD<br />

Levels*<br />

Mean<br />

Slope 2 5.966667 2.98333 3.7640 0.0361 flat A 1.53<br />

Error 27 21.400000 0.79259 moderate A B 1.17<br />

C. Total 29 27.366667 steep B 0.00<br />

*Levels not connected by the same letter are significantly different<br />

Afro-alpine vegetation height also showed significant effect (P50 cm<br />

vegetation Height<br />

All Pairs<br />

Tukey-Kramer<br />

0.05<br />

Figure 13: One way analysis of wolf sighting by vegetation height<br />

64


Table 28: Analysis of wolf sighting by vegetation height and mean comparisons using Tukey-<br />

Kramer HSD<br />

Source DF Sum of<br />

Squares<br />

Mean<br />

Square<br />

F<br />

Ratio<br />

P-value Comparisons for all pairs<br />

using Tukey-Kramer HSD<br />

lnvnl *<br />

Mean<br />

Vegetation height 3 8.816667 2.93889 4.1192 0.0162 0-10 cm A 1.90<br />

Error 26 18.550000 0.71346 11-30 cm A B 1.10<br />

C. Total 29 27.366667 31-50 cm B 1.00<br />

>50 cm B 0.25<br />

*Levels not connected by the same letter are significantly different<br />

In this study similar parameters altitude, slope and vegetation height determined rodent burrow<br />

distribution and actual wolf sighting across Mt. Abune Yoseph Afro-alpine ecosystem.<br />

65


Chapter V: Conclusion and Recommendations<br />

5.1. Conclusion<br />

This study was designed to develop an understanding of the existing community management and<br />

status of Mt. Abune Yoseph Afro-alpine ecosystem by taking <strong>Ethiopian</strong> wolf as a flagship species<br />

of the ecosystem. Accordingly, efforts have been made to explore socio-economic and natural<br />

resources assessment of the mountain for the improvement of user community livelihood, their<br />

perception towards its management and its endemic wild (<strong>Ethiopian</strong> wolf) as well as tried to show<br />

the human- livestock interference impact and the resulting conflict with wolf in the ecosystem.<br />

The methods employed, which consist of socio-economic interview, ecologic survey and discussions<br />

with key informants were useful in addressing the aims and leading to the following conclusions.<br />

The results of this study clearly showed Mt. Abune Yoseph Afro-alpine ecosystem provide essential<br />

ecosystem economic goods and services vitally important for the livelihoods of local communities<br />

within the limit of sustainability. Due to this fact, majority of the local community had positive<br />

perception towards the ecosystem and its flagship species the <strong>Ethiopian</strong> wolf. A good or positive<br />

perception towards communal Afro-alpine ecosystem helps to protect or conserve its resources<br />

from the increased human and livestock interference impact by designing sustainable ecosystem<br />

management together with community which can avoid the existing human-wildlife conflict<br />

particularly human-wolf conflict in the area.<br />

Currently, tremendous pressure of the swelling human and livestock population expressed<br />

through unsustainable resource exploitation from the ecosystem in the form of long time grazing,<br />

frequent firewood collection and agricultural encroachment leads to erosion, gulley formation<br />

66


and low productivity that ends in resource depletion and habitat fragmentation causing<br />

decreasing trend of <strong>Ethiopian</strong> wolf at Mt. Abune Yoseph. However, the presence of 19 wolves<br />

according to local community estimation and well distribution of rodent burrow observed in<br />

habitat assessment transect along with good opportunity of seeing up to 3 wolves at once during<br />

wolf sighting as well as an exhausted natural resources exploitation from the ecosystem indicate<br />

the existing status and socio economic potential of Mt. Abune Yoseph Afro-alpine ecosystem.<br />

5.2. Recommendations<br />

Global warming and the increased human and livestock population pose their pressure on Afroalpine<br />

ecosystems during the last decades and result in the confinement of this ecosystem to the<br />

highest mountain above 3200 m a.s.l. But nowadays in Ethiopia land above 3000 m a.s.l. has<br />

been converted rapidly to farmland. Areas previously considered too cold for agriculture and too<br />

inhospitable for habituation even though ends in failure today all are farmlands and settlement<br />

areas.<br />

In Mt. Abune Yoseph, cereals are cultivated even higher, with land ploughed above 3,600 m.<br />

Much of the Afro-alpine habitat has already disappeared. Nowadays, the line of 3,700 m a.s.l.<br />

contains the overall Afro-alpine ecosystem of the Abune Yoseph massif, and only a few small<br />

degraded patches of this ecosystem remain below this altitude.<br />

As the results of this study indicated the most exceptional resource values of this ecosystem<br />

which together give the area its conservation significance, the rare, endemic and endangered<br />

species of flora and fauna as well as the cultural and economic benefits fetched by the local and<br />

downstream users from the ecosystem are in a great danger of extinction. To cut this scenario the<br />

following recommendations are given based on the research findings.<br />

67


Since the attitude and action of human towards something can be shaped by<br />

understanding and knowledge great attention should have to be given for education and<br />

awareness creation starting from the local farmers, school children up to local, regional,<br />

and national leaders in order to discharge their conservation responsibility for Mt. Abune<br />

Yoseph Afro-alpine ecosystem protection.<br />

According to this study poor HHs that are landless, proximate to the ecosystem and none<br />

or less number livestock owners showed negative attitude towards Mt.AbuneYoseph<br />

Afro-alpine ecosystem conservation due to economic reason to exploit the resource of the<br />

ecosystem freely in unsustainable way. So in order to reduce the pressure on natural<br />

resources by this part of the community, it has paramount importance to create an activity<br />

that diversifies the livelihood of the local poor community. In this regard using the<br />

location advantage of Mt. Abune Yoseph existing near to Lalibela, the city that holds one<br />

of Ethiopia‟s top attractions: the rock-hewn churches which are considered the greatest<br />

historical sites of the Christian world, development of eco-tourism for Mt.AbuneYoseph<br />

will be a good opportunity. According to the city tourism office, in Lalibela tourism is<br />

already developed and visited by 10,000-20,000 tourists yearly by using the comfortable<br />

plane of <strong>Ethiopian</strong> air lines and car trip that give chance to see large part of the country.<br />

Therefore in Mt. Abune Yoseph using its rare and endemic flag ship species <strong>Ethiopian</strong><br />

wolf, Gelada baboon and birds as well as the scenic attraction of the Afro-alpine<br />

ecosystem as a comparative advantage nature based tourism is recommended to support<br />

the ecosystem protection as well as to improve the livelihood of the local poor<br />

community.<br />

68


The Lasta woreda administration should have to take the front line with other GO and<br />

NGO,s to implement conservation based development that consider Mt. Abune Yoseph<br />

and Lalibela the natural and cultural heritage as a good opportunity by proposing<br />

development link between the two hotspots. According to this research finding to reduce<br />

the livestock and firewood collection burden on the ecosystem and to avoid humanlivestock<br />

and wildlife conflict happen due to resource competition alternative livestock<br />

fodder system that avoid free grazing and encourage zero-grazing technology as well as<br />

ecologically sound, socially acceptable and economically affordable fuel sources such as<br />

bio-fuel and solar technology side by side planting well identified tree species in terms of<br />

the area context must be promoted by mobilizing the local community.<br />

It is also advisable if the future management of Mt. Abune Yoseph Afro-alpine<br />

ecosystem will be legally recognized community based protected area, in which its<br />

boundary is clearly known and that has resource management plan developed by the<br />

involvement of the local community, in order to include their long time custom and<br />

tradition of sustainable resource protection and utilization and also to encourage the<br />

development of their own by-law which go hand in hand with the regional land and<br />

biodiversity management strategy.<br />

69


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74


Appendix<br />

1: Interview questionnaire<br />

Part-1 Questionnaire on socio-Economic survey<br />

Date of interview ______________________ Enumerator's Name________________________<br />

1. Name of household head_______________________________________________________<br />

Region________Zone_________Wereda________kebele_________Village ____________<br />

2. Age _______<br />

3. Sex: Male _______Female_______<br />

4. Educational status:<br />

1. Can‟t read and write 2. Elementary school<br />

3. High school 4.Vocational<br />

5. Marital status: 1. Married 2. Single 3. Divorced 4.Widow<br />

6. Family size____________<br />

7. How long have you lived at your current Got (Months/years) ______<br />

8. How far is your residence from Mt. Abune Yoseph (in km) _________<br />

9. Do you have access to land 1.Yes 2. No<br />

10. If yes to question 9,what is the size of your land holding by land use type(in timad)<br />

N0 Land use type Size in Timad *Equivalent size in ha.<br />

1 Farm land<br />

2 Woodlot<br />

3 Grazing land<br />

4 homested<br />

5 Other<br />

Total<br />

* 1ha.=4 Timad<br />

11. How many livestock do you own at this time oxen______ cows_____ sheep_______<br />

goats_____mules _____horses______donkeys_______ hens‟ _____ Beehives _____<br />

12. For how long do you graze at Mt. Abune Yoseph (AY) during dry and wet season<br />

During dry season for ___________ months.<br />

75


During wet season for ___________ months.<br />

13. Does AY bring economic benefits to your household 1. Yes 2. No<br />

14. Based on question 13, which type of benefit<br />

1. Household firewood 1.Yes 2. No, if yes, how many loads of fire wood do you collect<br />

in the last 12months _____________________<br />

Market price of firewood per load__________<br />

2. Thatching grass 1.Yes 2. No if yes, how many loads of thatching grass do you cut<br />

in the last 12months _______________________<br />

Market price of thatching grass per load________<br />

3. Hay 1.Yes 2. No if yes how many loads of hay do you harvest<br />

in the last 12 months ______________________<br />

Market price of hay per load_________________<br />

4. Income from tourists 1. Yes 2. No if yes in what type of tourist activity did you<br />

involved and how much birr did you get from tourism in past years _____ ETB.<br />

5. Others __________________________________________________________<br />

15. What estimated benefits have you made in birr from mt. AY on average in a year _______<br />

Part-2 Perception towards Afro-alpine Ecosystem, Wildlife and Wildlife conflict<br />

16. Is there a need to protect the natural resources of AY Afro-alpine ecosystem 1. Yes 2. No<br />

17. Who do you think is responsible for the protection of AY Afro-alpine ecosystem<br />

1. Government 2. Local community 3. NGO<br />

18. What changes have you observed in the vegetation of AY in the last 20 years Intact /<br />

improving / degrading / no idea, Why do you think so __________________________<br />

19. Are populations of wild animals in AY stable/ increasing/ decreasing / no idea in the last 20<br />

years Why do you think so ______________________________________________<br />

20. Is the conservation of wild animals and plants a good thing 1. Yes 2. No<br />

Why do you think so ____________________________________________<br />

76


21. How do you know the concept of Natural resource conservation<br />

1. Form training given on NR of AY<br />

2. through extension education given by DA<br />

3. By own experience<br />

4. from formal school<br />

5. Other sources (specify) __________________________________<br />

22. Do you think that the presence of bylaw is important and helped the conservation of<br />

Mt. AY<br />

1. Yes 2. No if yes how ____________________________________<br />

23. Name wild animals that you know at AY __________________________________<br />

____________________________________________________________________<br />

24. Do you face problems because of wildlife in your area 1. Yes 2. No<br />

25. If yes, to question number 24, what are the problems you face<br />

_________________________________________________________________________<br />

_________________________________________________________________________<br />

26. Which wild animals are the most problematic in terms of livestock predation at AY<br />

No Predator Prey Extent (No of animals killed from your herd)<br />

Last year In last 5 years In last 10 years<br />

1<br />

2<br />

3<br />

27. What is the trend of livestock predation by wildlife 1. Stable 2. Increasing 3.decreasing<br />

28. Which wild animals are most problematic in terms of crop damage<br />

77


No Wild animal Type of crop damaged Extent of damage in the last 12 months<br />

(H*, M*, L*)<br />

1<br />

2<br />

3<br />

* H=heavy, M=medium, L=light<br />

29. What is the tendency of crop damage by wildlife 1. Stable 2. Increasing 3.decreasing<br />

30. Describe techniques you use to minimize the impacts of livestock predation and crop damage<br />

______________________________________________________________________________<br />

______________________________________________________________________________<br />

31. Have you personally seen <strong>Ethiopian</strong> wolves (EW) in AY 1. Yes 2. No<br />

32. How many times have you seen EW in the last 12 month‟s ______and past 5 years ______<br />

33. What times of the day you are more likely to see EW ______________________<br />

34. What is the largest group of EW you have seen in AY ____________<br />

35. What do you think about the number of EW in AY 1. Stable 2. Increasing 3. Decreasing<br />

Explain the reasons for thinking so: ___________________________________________<br />

36. How many wolves do you think are present at AY currently ________________________<br />

37. Do you think the presence of EW in AY bad for people and their animals 1. Yes 2. No if<br />

no why ____________________________________________________________<br />

38. In your opinion, can wolves live harmoniously in the same area with people 1. Yes 2. No<br />

39. What is the attitude of local people towards wolves 1. Positive 2. Indifferent 3.negative<br />

40. Have you ever seen wolves taking any livestock other than sheep and goat 1. Yes 2. No<br />

Part-3 Sustainable Natural resource utilization and Management<br />

41. List the dominant tree/ shrub species you use as a fire wood from mt. AY in order of<br />

importance 1._________________ 2._________________ 3._______________<br />

42. Why do you prefer to use the tree/ shrub species mentioned in Q.41<br />

_____________________________________________________________________<br />

43. Do you experience the shortage of fuel wood 1. Yes 2. No<br />

78


44. If yes to question 43, what mechanism do you use to solve the shortage<br />

1. Using energy saving stove 2. Using livestock dung 3. Other specify_______<br />

45. Do you have your own tree planted at your homestead 1. Yes 2. No<br />

46. Is there any traditional system established by the community to manage AY 1. Yes 2. No<br />

if yes, name it _________________________________________________________<br />

47. Is there any local bylaw that binds your activity at AY 1. Yes 2. No<br />

48. Do you know soil and water conservation activities constructed by local community in AY<br />

1. Yes 2. No if yes, which type 1. Different terraces 2. Planting seedlings<br />

3. Gully treatment 4. Other specify _____<br />

49. Which would be your future choice of management option for natural resources in AY<br />

Why<br />

1. If managed as community protected area 2. If divided amongst users<br />

3. If managed as closure area 4. If managed under investors<br />

5. if managed as public reserve/ state national park/<br />

______________________________________________________________________________<br />

______________________________________________________________________________<br />

50. Are there any problems in sharing the resources at AY 1. Yes<br />

2. No if yes, what are those problems<br />

___________________________________________________________________________<br />

___________________________________________________________________________<br />

79


2: Contingency tables<br />

Table 1: Chi-square test of HHs land ownership status and economic resource utilization from<br />

Mt. Abune Yoseph<br />

HHs land ownership<br />

status<br />

sample HH got economic benefit from Mt.<br />

Abune Yoseph<br />

not benefited<br />

Total<br />

benefited<br />

0 16 16<br />

not own farmland<br />

own farm land 24 80 104<br />

Total 24 96 120<br />

Chi-Square Tests<br />

Value df<br />

Asymp. Sig. (2-<br />

sided)<br />

Exact Sig.<br />

(2-sided)<br />

Exact Sig.<br />

(1-sided)<br />

Pearson Chi-Square 4.615(b) 1 .032<br />

Continuity Correction(a) 3.286 1 .070<br />

Likelihood Ratio 7.734 1 .005<br />

Fisher's Exact Test .039 .021<br />

Linear-by-Linear<br />

Association<br />

4.577 1 .032<br />

N of Valid Cases 120<br />

Table 2: Chi-square test of HHs sex for wolf number estimation at Mt. Abune Yoseph<br />

<strong>Ethiopian</strong> <strong>Wolf</strong> estimate number at Mt. Abune Yoseph by sample HHs Total<br />

1 2 3 4 5 6 7 8 9 10 11 12 15 16 17 20 25 30 35 40 50 60<br />

sex male 1 0 1 3 4 3 1 5 0 18 1 2 11 4 2 7 8 18 1 4 8 1 103<br />

female 0 1 2 2 1 2 0 0 1 5 0 0 1 0 0 1 0 1 0 0 0 0 17<br />

Total 1 1 3 5 5 5 1 5 1 23 1 2 12 4 2 8 8 19 1 4 8 1 120<br />

Chi-Square Tests<br />

Value df Asymp. Sig. (2-sided)<br />

Pearson Chi-Square 33.495(a) 21 .041<br />

Likelihood Ratio 30.799 21 .077<br />

Linear-by-Linear Association 10.329 1 .001<br />

N of Valid Cases<br />

120<br />

80


Table 3: Chi-square test of HHs family size to estimate wolf number at Mt. Abune Yoseph<br />

family <strong>Ethiopian</strong> <strong>Wolf</strong> estimate number at Mt. Abune Yoseph by sample HHs Total<br />

size 1 2 3 4 5 6 7 8 9 10 11 12 15 16 17 20 25 30 35 40 50 60<br />

1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 0 0 0 0 3<br />

3 0 0 0 0 0 0 0 0 0 1 0 0 3 0 1 1 1 3 0 0 1 0 11<br />

4 0 0 0 2 0 1 0 0 1 5 0 1 3 1 0 0 0 2 0 1 1 0 18<br />

5 0 0 1 0 1 1 0 1 0 5 0 0 1 1 0 2 0 3 0 1 1 1 19<br />

6 1 1 1 2 0 2 1 0 0 6 0 0 3 0 0 1 2 4 0 1 3 0 28<br />

7 0 0 0 1 0 1 0 1 0 4 0 0 1 1 0 1 2 5 0 0 1 0 18<br />

8 0 0 0 0 3 0 0 3 0 2 1 1 1 0 0 1 2 1 0 1 1 0 17<br />

9 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1 0 0 0 3<br />

10 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 2<br />

Total 1 1 3 5 5 5 1 5 1 23 1 2 12 4 2 8 8 19 1 4 8 1 120<br />

Chi-Square Tests<br />

Value df Asymp. Sig. (2-sided)<br />

Pearson Chi-Square 222.280(a) 189 .049<br />

Likelihood Ratio 132.744 189 .999<br />

Linear-by-Linear Association .037 1 .847<br />

N of Valid Cases<br />

120<br />

Table 4: Chi-square test of HHs educational status to estimate wolf number at Mt. Abune<br />

Yoseph<br />

Educational <strong>Ethiopian</strong> <strong>Wolf</strong> estimate number at Mt. Abune Yoseph by sample HHs Total<br />

status of<br />

HHs 1 2 3 4 5 6 7 8 9 10 11 12 15 16 17 20 25 30 35 40 50 60<br />

illiterate 1 1 3 5 5 4 1 4 1 15 0 0 5 3 0 2 2 7 0 1 3 0 63<br />

literate 0 0 0 0 0 1 0 1 0 8 1 2 7 1 2 6 6 12 1 3 5 1 57<br />

Total 1 1 3 5 5 5 1 5 1 23 1 2 12 4 2 8 8 19 1 4 8 1 120<br />

Chi-Square Tests<br />

Value df Asymp. Sig. (2-sided)<br />

Pearson Chi-Square 37.674(a) 21 .014<br />

Likelihood Ratio 47.441 21 .001<br />

Linear-by-Linear Association 16.736 1 .000<br />

N of Valid Cases<br />

120<br />

81


Table 5: Chi-square test of HHs based on livestock number to estimate wolf number at Mt.<br />

Abune Yoseph<br />

HHs<br />

livestock<br />

<strong>Ethiopian</strong> <strong>Wolf</strong> estimate number at Mt. Abune Yoseph by sample HHs Total<br />

number 1 2 3 4 5 6 7 8 9 10 11 12 15 16 17 20 25 30 35 40 50 60<br />

0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1<br />

3 0 0 0 1 0 1 0 0 0 2 0 0 0 1 0 1 0 0 0 0 0 0 6<br />

4 0 1 1 1 1 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 6<br />

5 0 0 0 0 1 1 0 0 1 3 0 0 0 0 1 0 0 1 0 0 0 0 8<br />

6 0 0 0 0 0 0 0 0 0 2 0 0 3 0 0 1 0 0 0 0 1 0 7<br />

7 0 0 0 0 0 0 0 1 0 3 0 0 1 1 1 0 0 1 0 1 0 0 9<br />

8 0 0 0 1 1 0 0 0 0 0 0 0 1 0 0 0 0 2 0 0 0 0 5<br />

9 1 0 0 0 0 1 0 1 0 3 0 0 0 0 0 0 1 1 0 0 2 0 10<br />

10 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 2 0 0 0 1 5<br />

11 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 2 0 1 0 0 5<br />

12 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 4<br />

13 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 2 0 1 0 0 5<br />

14 0 0 0 0 0 0 0 0 0 2 0 1 0 0 0 1 0 0 0 0 1 0 5<br />

15 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 2<br />

16 0 0 0 0 0 1 0 0 0 0 0 0 2 0 0 1 1 1 0 0 1 0 7<br />

17 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 2<br />

19 0 0 1 0 0 0 0 1 0 1 0 0 0 0 0 0 1 1 0 0 1 0 6<br />

20 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 2<br />

21 0 0 0 1 0 0 0 0 0 0 1 0 1 0 0 0 0 2 0 1 0 0 6<br />

22 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 2<br />

24 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1<br />

25 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1<br />

26 0 0 0 0 0 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 3<br />

27 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1<br />

28 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1<br />

29 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1<br />

31 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

32 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1<br />

33 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 1<br />

34 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

36 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

40 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

42 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1<br />

78 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1<br />

Total 1 1 3 5 5 5 1 5 1 23 1 2 12 4 2 8 8 19 1 4 8 1 120<br />

82


Chi-Square Tests<br />

Value df Asymp. Sig. (2-sided)<br />

Pearson Chi-Square 773.891(a) 714 .059<br />

Likelihood Ratio 322.989 714 1.000<br />

Linear-by-Linear Association 2.374 1 .123<br />

N of Valid Cases<br />

120<br />

83

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