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Resource Inventory of Marine and Estuarine Fishes of the West Coast and Alaska: A Checklist of North Pacific and Arctic Ocean Species from Baja California to the Alaska–Yukon Border OCS Study MMS 2005-030 and USGS/NBII 2005-001 Milton S. Love Marine Science Institute University of California Santa Barbara, CA 93106 love@lifesci.ucsb.edu www.id.ucsb.edu/lovelab Catherine W. Mecklenburg T. Anthony Mecklenburg Pt. Stephens Research P. O. Box 210307 Auke Bay, AK 99821 ptstephens@alaska.com Lyman K. Thorsteinson Western Fisheries Research Center U. S. Geological Survey 6505 NE 65th St. Seattle, WA 98115 lyman_thorsteinson@usgs.gov June 2005 Much of the research was performed under a cooperative agreement between the USGS’s Western Fisheries Research Center and the University of California, Santa Barbara (CA 03WRAG007). Front Cover: (from upper right): Rock Sole, Lepidopsetta bilineata, Janna Nichols; Crescent Gunnel, Pholis laeta, Janna Nichols; Bocaccio, Sebastes paucispinis, Milton Love; Whale Shark, Rhincodon typus, Phillip Cola; Grunt Sculpin, Rhamphocottus richardsonii, Ken Blauvelt; Pacific Spiny Lumpsucker, Eumicrotremus orbis, Ken Blauvelt; Guineafowl Puffer, Arothron meleagris, Phillip Cola; Ocean Sunfish, Mola mola, Phillip Cola Back Cover: Sargo, Anisotremus davidsonii, Phillip Cola Page iv: Ocean Sunfish, Mola mola, Phillip Cola Page x: Bigeye Trevally, Caranx sexfasciatus, Phillip Cola Project Cooperation This research addressed an information need identified by the USGS Western Fisheries Research Center and the University of California, Santa Barbara to the Department of the Interior’s Minerals Management Service, Pacific OCS Region, Camarillo, California. The resource inventory information was further supported by the USGS’s National Biological Information Infrastructure as part of its ongoing aquatic GAP project in Puget Sound, Washington. Report Availability Available for viewing and in PDF at: http://wfrc.usgs.gov http://far.nbii.gov http://www.id.ucsb.edu/lovelab Printed copies available from: Milton Love Marine Science Institute University of California, Santa Barbara Santa Barbara, CA 93106 (805) 893-2935 Lyman Thorsteinson Western Fisheries Research Center U. S. Geological Survey 6505 NE 65th St. Seattle, WA 98115 (206) 526-6569 Minerals Management Service Pacific OCS Region 770 Paseo Camarillo Camarillo, CA 99010 (805) 389-7800 Suggested Citation Love, M. S., C. W. Mecklenburg, T. A. Mecklenburg, and L. K. Thorsteinson. 2005. Resource Inventory of Marine and Estuarine Fishes of the West Coast and Alaska: A Checklist of North Pacific and Arctic Ocean Species from Baja California to the Alaska–Yukon Border. U. S. Department of the Interior, U. S. Geological Survey, Biological Resources Division, Seattle, Washington, 98104, OCS Study MMS 2005-030 and USGS/NBII 2005-001.
TABLE OF CONTENTS Acknowledgments v Introduction vi Species List 1 References Cited 182 Personal Communications 222 Index 224
Page 5 and 6: ACKNOWLEDGMENTS This West Coast inv
Page 7 and 8: 2) Regional guides. In most instanc
Page 9: Some readers may question the capit
Page 12 and 13: Order Chimaeriformes Family Chimaer
Page 14 and 15: Order Squatiniformes Family Squatin
Page 16 and 17: Isurus oxyrinchus Rafinesque, 1810.
Page 18 and 19: Carcharhinus leucas (Müller & Henl
Page 20 and 21: Narcine entemedor Jordan & Starks,
Page 22 and 23: Bathyraja parmifera (Bean, 1881). A
Page 24 and 25: Urotrygon aspidura (Jordan & Gilber
Page 26 and 27: Order Elopiformes Family Elopidae
Page 28 and 29: Gymnothorax castaneus (Jordan & Gil
Page 30 and 31: *Herpetoichthys fossatus (Myers & W
Page 32 and 33: Chiloconger dentatus (Garman, 1899)
Page 34 and 35: Family Monognathidae — Onejaws or
Page 36 and 37: Pliosteostoma lutipinnis (Jordan &
Page 38 and 39: Ariopsis platypogon (Günther, 1864
Page 40 and 41: Family Opisthoproctidae — Barrele
Page 42 and 43: *Maulisia acuticeps Sazonov, 1976.
Page 44 and 45: Coregonus pidschian (Gmelin, 1789).
Page 46 and 47: Diplophos taenia Günther, 1873. To
Page 48 and 49: Bathophilus brevis Regan & Trewavas
Page 50 and 51: Benthalbella linguidens (Mead & Bö
Page 52 and 53:
Stemonosudis macrura (Ege, 1933). P
Page 54 and 55:
Diaphus anderseni Tåning, 1932. Lo
Page 56 and 57:
Nannobrachium hawaiiensis Zahuranec
Page 58 and 59:
Tarletonbeania taylori Mead, 1953.
Page 60 and 61:
Family Ophidiidae — Cusk-eels Bas
Page 62 and 63:
Family Bythitidae — Livebearing B
Page 64 and 65:
comm. to C. W. M.) to southern Cali
Page 66 and 67:
British Columbia (Peden 1975), and
Page 68 and 69:
Porichthys myriaster Hubbs & Schult
Page 70 and 71:
Family Oneirodidae — Dreamers Ber
Page 72 and 73:
Family Gigantactinidae — Whipnose
Page 74 and 75:
Baja California (Collette in Fische
Page 76 and 77:
Oxyporhamphus micropterus (Valencie
Page 78 and 79:
Melamphaes macrocephalus Parr, 1931
Page 80 and 81:
Herald 1983; max.: Hayashi in Nakab
Page 82 and 83:
Syngnathus californiensis Storer, 1
Page 84 and 85:
Scorpaena sonorae Jenkins & Everman
Page 86 and 87:
Sebastes crameri (Jordan, 1897). Da
Page 88 and 89:
Sebastes melanostomus (Eigenmann &
Page 90 and 91:
Sebastes rufus (Eigenmann & Eigenma
Page 92 and 93:
Prionotus ruscarius Gilbert & Stark
Page 94 and 95:
Family Cottidae — Sculpins Archis
Page 96 and 97:
Clinocottus globiceps (Girard, 1858
Page 98 and 99:
Gulf of Alaska at Prince William So
Page 100 and 101:
Myoxocephalus niger (Bean, 1881). W
Page 102 and 103:
Rastrinus scutiger (Bean, 1890). Ro
Page 104 and 105:
Family Hemitripteridae — Sailfin
Page 106 and 107:
Agonomalus mozinoi Willimovsky & Wi
Page 108 and 109:
authors do not recognize the aix fo
Page 110 and 111:
Eumicrotremus asperrimus (Tanaka, 1
Page 112 and 113:
Careproctus colletti Gilbert, 1896.
Page 114 and 115:
*Crystallichthys mirabilis Jordan &
Page 116 and 117:
Liparis rutteri (Gilbert & Snyder,
Page 118 and 119:
108 recently been collected from th
Page 120 and 121:
*Cephalopholis panamensis (Steindac
Page 122 and 123:
Mycteroperca xenarcha Jordan, 1888.
Page 124 and 125:
Family Apogonidae — Cardinalfishe
Page 126 and 127:
Family Carangidae — Jacks Alectis
Page 128 and 129:
Hemicaranx zelotes Gilbert, 1898. B
Page 130 and 131:
Family Bramidae — Pomfrets Brama
Page 132 and 133:
Family Lobotidae — Tripletails Lo
Page 134 and 135:
124 Haemulon scudderi Gill, 1862. G
Page 136 and 137:
Family Polynemidae — Threadfins P
Page 138 and 139:
Gulf of California (Chao in Fischer
Page 140 and 141:
Family Chaetodontidae — Butterfly
Page 142 and 143:
Cirrhitus rivulatus Valenciennes, 1
Page 144 and 145:
Zalembius rosaceus (Jordan & Gilber
Page 146 and 147:
Cabo San Lucas, southern Baja Calif
Page 148 and 149:
*Scarus compressus (Osburn & Nichol
Page 150 and 151:
Eucryphycus californicus (Starks &
Page 152 and 153:
Lycodapus fierasfer Gilbert, 1890.
Page 154 and 155:
Lycodes raridens Taranetz & Andrias
Page 156 and 157:
Anisarchus medius (Reinhardt, 1837)
Page 158 and 159:
Lumpenus fabricii Reinhardt, 1836.
Page 160 and 161:
Rhodymenichthys dolichogaster (Pall
Page 162 and 163:
Axoclinus storeyae (Brock, 1940). C
Page 164 and 165:
Kali macrura (Parr, 1933). Longnose
Page 166 and 167:
Starksia guadalupae Rosenblatt & Ta
Page 168 and 169:
Family Blenniidae — Combtooth Ble
Page 170 and 171:
*Tomicodon boehlkei Briggs, 1955. C
Page 172 and 173:
Eucyclogobius newberryi (Girard, 18
Page 174 and 175:
Family Ephippidae — Spadefishes C
Page 176 and 177:
Family Trichiuridae — Cutlassfish
Page 178 and 179:
Scomber japonicus Houttuyn, 1782. C
Page 180 and 181:
Family Nomeidae — Driftfishes or
Page 182 and 183:
Citharichthys gordae Beebe & Tee-Va
Page 184 and 185:
Family Pleuronectidae — Righteye
Page 186 and 187:
Lyopsetta exilis (Jordan & Gilbert,
Page 188 and 189:
178 Lavenberg 1997). Benthic, at de
Page 190 and 191:
Family Ostraciidae — Boxfishes La
Page 192 and 193:
REFERENCES CITED Aasen, G. A., D. A
Page 194 and 195:
Andriashev, A. P. 1958. An addition
Page 196 and 197:
Bertelsen, E., T. W. Pietsch, and R
Page 198 and 199:
Castro-Aguirre, J. L., H. Espinosa
Page 200 and 201:
Craig, P. and L. Haldorson. 1986. P
Page 202 and 203:
Evseenko, S. A. 1999. Early develop
Page 204 and 205:
Gartman, R. and A. Groce. 1998. Dem
Page 206 and 207:
Harold, A. S. 1993. Phylogentic rel
Page 208 and 209:
Jordan, D. S. and B. W. Evermann. 1
Page 210 and 211:
Kushner, D. J., D. Lerma, J. Shaffe
Page 212 and 213:
Lissner, A. L. and J. H. Dorsey. 19
Page 214 and 215:
Mecklenburg, C. W. 2003. Family Zap
Page 216 and 217:
Myers, R. F. 1999. Micronesian reef
Page 218 and 219:
Orr, J. W., M. A. Brown, and D. C.
Page 220 and 221:
Peden, A. E. and D. E. Wilson. 1976
Page 222 and 223:
Richards, D. V. and J. M. Engle. 20
Page 224 and 225:
Schmidt, P. J. 1950. Fishes of the
Page 226 and 227:
Stein, D. L. 1978. A review of the
Page 228 and 229:
Tokranov, A. M. and A. M. Orlov. 20
Page 230 and 231:
Wilimovsky, N. J. 1954. List of the
Page 232 and 233:
PERSONAL COMMUNICATIONS Alisa A. Ab
Page 234 and 235:
Index A Ablennes hians, 63 Abudefdu
Page 236 and 237:
Index Apogonidae, 114 Apogon atrica
Page 238 and 239:
Index Basketweave Cusk-eel, 51 Bask
Page 240 and 241:
Index Black (continued) Bobtail Eel
Page 242 and 243:
Index C C-O Sole, 176 C-O Turbot, 1
Page 244 and 245:
Index Cheilotrema saturnum, 126 Che
Page 246 and 247:
Index Cowcod, 77 Cow Sharks, 2 Craf
Page 248 and 249:
Index Dipline Eelpout, 144 Diplobat
Page 250 and 251:
Index Eulachon, 33 Euleptorhamphus
Page 252 and 253:
Index Giant (continued) Sea Bass, 1
Page 254 and 255:
Index Halfspotted Tonguefish, 178 H
Page 256 and 257:
Index Ichthyapus selachops, 20 Icht
Page 258 and 259:
Index Lestidium johnfitchi, 41 nudu
Page 260 and 261:
Index Lycenchelys (continued) jorda
Page 262 and 263:
Index Microlophichthys microlophus,
Page 264 and 265:
Index Netdevils, 59 Netdevil Angler
Page 266 and 267:
Index P Pachycara bulbiceps, 145 gy
Page 268 and 269:
Index Paralonchurus goodei, 128 Par
Page 270 and 271:
Index Popeye Blacksmelt, 29 Cataluf
Page 272 and 273:
Index Red (continued) Scorpionfish,
Page 274 and 275:
Index Samoa Sawtooth Eel, 23 Sandfi
Page 276 and 277:
Index Seriola dorsalis, 119 lalandi
Page 278 and 279:
Index Smalleye Snipe Eel, 21 Square
Page 280 and 281:
Index Spotted (continued) Grouper,
Page 282 and 283:
Index Tan Bristlemouth, 35 Tapertai
Page 284 and 285:
Index U Ulca, 94 Ulcina olrikii, 98
Page 286:
Index Y Y-Prickleback, 147 Yellow-m
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