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Spotted Knapweed appeared near Missoula before<br />

1925 and is now in every county in Montana.<br />

effect of the herbicide. Figure 2 shows the same spot one year later. <strong>The</strong> herbicide application<br />

eliminated over 99 percent of the seedlings and facilitated establishment of foxtail<br />

barley. Foxtail barley generally is not considered a desired species, but on a lakebed that<br />

is flooded every few years and has alkaline soil when exposed, foxtail barley probably is<br />

the best vegetation one can expect. At this location, cattle consume the foxtail barley quite<br />

readily in the fall and winter.<br />

<strong>The</strong> area shown in Figure 3 is about 200 yards from the location of Figures 1 and 2,<br />

and was inhabited by a very mature stand of perennial pepperweed. This site location was<br />

treated on the same day, and with the same chemical and application rate, as the perennial<br />

pepperweed seedlings shown in Figure 1. One year later there was no effective control of<br />

the mature perennial pepperweed plants (Figure 4). <strong>The</strong> mature stand of perennial pepperweed<br />

was eventually controlled (not eradicated), but it took several more annual treatments.<br />

One critical reason for successful control of the seedlings and unsuccessful control<br />

of the mature perennial pepperweed is the respective root structure of the different age<br />

classes. For about the first 6-8 eight weeks of growth, perennial pepperweed seedlings<br />

behave as an annual, not a perennial plant. <strong>The</strong> roots are short and they have not developed<br />

any perennial buds. <strong>The</strong> buds that develop on the root crown and the roots after eight weeks<br />

of age are what allow this weed to regrow the next year. When the leaves are killed before<br />

the seedling can develop perennial buds on the roots and root crown the plant dies. Mature<br />

perennial pepperweed plants have roots that extend several feet or more into the soil and<br />

can extend laterally for 10-20 feet or more. Every inch of these roots has developed a bud<br />

that can produce a new plant. It is literally impossible to place enough herbicide on the<br />

relatively small amount of leaf area of mature plants to kill all of the buds found on their<br />

very large root system. Some buds will survive and eventually produce stems that emerge<br />

the following growing season. Furthermore, emergence may not occur until the middle<br />

or latter part of the next growing season. Another challenge with applying herbicides to<br />

mature perennial pepperweed is appropriate location of the chemical. <strong>The</strong> lower leaves are<br />

the ones that export most of their carbohydrates from photosynthesis to the roots, and thus<br />

move the herbicide to the roots. <strong>The</strong> upper leaves deliver most of their energy production to<br />

the flowers and seed. It is very difficult to get herbicide, especially from an aerial application,<br />

through the flowers and upper leaves to the lower leaves, where it is most effective. As<br />

Figure 1 shows, all of the seedlings had leaves that were completely exposed and therefore<br />

had the maximum potential for herbicide contact, uptake and movement to the site of action<br />

in the plant. <strong>The</strong> work at Chimney Dam showed that perennial pepperweed up to two years<br />

old could be successfully treated with Cimaron ® Max provided their leaves were readily<br />

accessible to the herbicide. Complete documentation of the effort to control perennial peppewrweed<br />

at Chimeny Dam Reservoir can be found in two publications available on the<br />

internet at the URL addresses found at the end of this article.<br />

Papers described in the text. Please see these for additional discussion of this topic and<br />

photos of the success of the treatments applied.<br />

Schultz. B. W. 2012. <strong>The</strong> Noxious Weed Seedbank: Out of Sight – Out of mind and eventually<br />

out of control. UNCE Fact Sheet 12-01. Available at: http://www.unce.unr.edu/publications/<br />

search/details.asp?searchby=keywordsearch&searchtext=seedbank&submit=Search<br />

Schultz, B. W. 2012. Response of seedling and two-year-old perennial pepperweed (Lepidium<br />

latifolium) plants to herbicide control. Journal of the NACAA, Volume 5, Issue 1. May. Available<br />

at: http://www.nacaa.com/journal/index.php?jid=136.<br />

Schultz, B. W. 2011. Differential Herbicide Effectiveness on Adjacent Populations of Young<br />

(Seedling) And Mature Perennial Pepperweed (Lepidium latifolium). Journal of the NACAA.<br />

Volume 4, Issue 1. November. Available at: http://www.nacaa.com/journal/index.php?jid=103<br />

<strong>The</strong> use of herbicide names in this paper does not imply any recommendation from the University<br />

of Nevada Cooperative Extension. Perennial pepperweed can be treated successfully with a number of<br />

herbicides and any applicator should know the benefits and constraints of each herbicide and make their<br />

selection based upon their specific circumstances.<br />

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April 2013 37

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