Hells Canyon <strong>Chukar</strong> <strong>and</strong> <strong>Gray</strong> <strong>Partridge</strong> Report, Ratti <strong>and</strong> Giudice 18reported a range <strong>of</strong> 49-86%. However, survival <strong>and</strong> reproductive success <strong>of</strong> gray partridgeequipped with radio transmitters should be interpreted with caution because effects <strong>of</strong> h<strong>and</strong>ling<strong>and</strong> radio tagging may be complex, e.g., a function <strong>of</strong> physical condition at time <strong>of</strong> capture, radiotag design, <strong>and</strong> interactions mediated by environmental factors such as weather <strong>and</strong> predatorabundance (Bro et al. 1999). Likewise, mortality estimates derived from seasonal changes indensity may be biased if movements violate the closed-population assumption.4.5. Habitat4.5.1. General4.5.1.1. <strong>Chukar</strong>. Most descriptions <strong>of</strong> chukar habitat include only generalphysiographic characteristics, e.g., cheatgrass (Bromus tectorum)-sagebrush (Artemisia spp.)vegetative type, steep topography, rock outcroppings, talus slopes, <strong>and</strong> arid climate (seeChristensen 1996). The most detailed description was probably given by Galbreath <strong>and</strong>Morel<strong>and</strong> (1953), who specified optimum chukar habitat contained about 50% sage-cheatgrassbunchgrass;45% talus slopes, rock outcroppings, <strong>and</strong> cliffs <strong>and</strong> bluffs; <strong>and</strong> 5% brushy creekbottoms <strong>and</strong> swales. Lindbloom (1998) reported that chukars used rock <strong>and</strong> shrub cover typesmore than expected <strong>and</strong> grass/forb areas less than expected during both spring <strong>and</strong> summer (P
Hells Canyon <strong>Chukar</strong> <strong>and</strong> <strong>Gray</strong> <strong>Partridge</strong> Report, Ratti <strong>and</strong> Giudice 19cover (e.g., fencerows, farmsteads, roadside <strong>and</strong> railroad right-<strong>of</strong>-ways, waterways, idle grass,brush <strong>and</strong> timber, pasture, <strong>and</strong> hay) was preferred during late spring, summer, <strong>and</strong> fall. Plowedstubble was preferred <strong>and</strong> winter wheat was generally avoided during winter (Mendel 1979). Insummer, birds generally used grassl<strong>and</strong>s <strong>and</strong> grain fields, but also used roadsides <strong>and</strong>shelterbelts. In winter, crop stubble (especially cereal grains) <strong>and</strong> woody cover were preferred(Carroll 1993). During severe winter weather with deep snow (>10 cm), woody cover nearfarmsteads may be important (Weig<strong>and</strong> 1980, Schulz 1980, Carroll 1993), although Smith et al.(1982) reported that birds in South Dakota shifted from use <strong>of</strong> row corps to pastures in a winter<strong>of</strong> deep snow (56 cm). Habitat use by gray partridge inhabiting canyons <strong>and</strong> mountainous areas,including Hells Canyon, has not been studied.4.5.2. Nesting4.5.2.1. <strong>Chukar</strong>. Limited research has been conducted on nest-sitecharacteristics (Lindbloom 1998), probably because chukar nests are difficult to find (Galbreath<strong>and</strong> Morel<strong>and</strong> 1953). Nests usually are widely scattered <strong>and</strong> well concealed, but <strong>of</strong>ten arelocated within 183-366 m <strong>of</strong> a water source (Galbreath <strong>and</strong> Morel<strong>and</strong> 1953, Harper et al. 1958).In Nevada, Christensen (1970) found 3 inactive nests, 1 under sagebrush at base <strong>of</strong> hill <strong>and</strong> 2hidden among rocks <strong>and</strong> brush. Harper et al. (1958) found 16 nests in California <strong>and</strong> concludedthat nesting was not limited to any specific cover type. In contrast, Galbreath <strong>and</strong> Morel<strong>and</strong>(1953) reported nest preference for south-facing slopes in Washington. Mackie <strong>and</strong> Buechner(1963) found 24 nests in Washington, but did not describe nest distribution among cover types.In <strong>Idaho</strong>, Lindbloom (1998) found 23 nests (10 renests) <strong>of</strong> radiotagged females. Covertypes used by nesting chukars included grass/forbs (48%), rocks (43%) <strong>and</strong> shrubs (9%).Average slope <strong>and</strong> elevation was 58% (st<strong>and</strong>ard error [SE] = 2) <strong>and</strong> 905 m (SE = 22),respectively. Eighty-seven percent <strong>of</strong> nests were on south-facing slopes (30% southeast, 57%southwest). Rock outcrops were the most prevalent (57%) structure at nest sites, followed bygrass-forbs (26%) <strong>and</strong> low shrubs (17%). <strong>Chukar</strong> nests were well concealed, with visualobstruction <strong>of</strong> 3.9 decimeters (SE = 0.5) <strong>and</strong> overhead canopy averaging 63% (SE = 9).Lindbloom (1998) concluded that nest placement <strong>of</strong> chukars was not affected by presence <strong>of</strong>yellow-star thistle (but see Brood-rearing, below) <strong>and</strong> nest-site quality probably was not limitingthe chukar population inhabiting canyon grassl<strong>and</strong>s along the Salmon River.In Oregon, Walter (2000) found 17 nests <strong>of</strong> radiotagged females <strong>and</strong> 6 incidental nests.Cover types used for nesting included bunchgrass (42%), rock (38%), shrub (8%), <strong>and</strong> annualgrass (8%). Fifty-seven percent <strong>of</strong> nests were concealed by vegetation (bluebunch wheatgrass[35%], basin wildrye [13%], or broom snakeweed [9%]) <strong>and</strong> 43% were located in or under rocks.Most nests were found on relatively steep (19.4° ± 3.3° [ x ± SE]) south-facing slopes.4.5.2.2. <strong>Gray</strong> <strong>Partridge</strong>. No information on gray partridge nests in Hells Canyonor other canyon grassl<strong>and</strong>s is available. Data from other habitats report gray partridge use idlecover (pastures, fence rows, roadsides, <strong>and</strong> shelterbelts) <strong>and</strong> hay fields for nesting (see review byCarroll 1989). Nest-site selection by gray partridge in Europe occurs mainly in linear <strong>and</strong>permanent cover (Potts 1986). In Great Britain, gray partridge will also use cereal grains fornesting, although mainly for renesting (Carroll 1993). In North America, older studies showedgreater use <strong>of</strong> hay fields than in Great Britain. However, in more recent studies, the trend is