134 C.F. Jaarsma, G.P.A. Willems / Landscape and Urban Planning 58 (2002) 125–135Half of this percentage can be attributed to MRRs,according Derckx (1995).Lower <strong>traffic</strong> flows do not necessarily lead to fewerroad kills. Huijser et al. (1998) discussed this ideathat higher <strong>traffic</strong> flows do not always lead to moreroad kills, because a bigger barrier effect can, in fact,prevent species from attempting to cross the road.Conversely, lower <strong>traffic</strong> flows, leading to a smallerbarrier effect, might in fact encourage species to tryto cross the road more frequently, resulting in moreroad kills.Nevertheless, according Forman and Alexander(1998), road kills have only a minimal impact onthe overall population size. Once you minimise theeffects of road kills, the barrier effects can beincreased and vice verse. Greater disturbance has agreater impact on the fauna than either the road kills orthe destruction of suitable <strong>habitat</strong>, because this impactaffects far more animals within a larger area.Fragmentation does not automatically lead to adecrease of biodiversity. The new spatial heterogenitythat results can provide space for new species. Formany species the landscape is not an archipel of<strong>habitat</strong> islands, but rather a mosaic of different ecotopesin a characteristic pattern (RMNO, 1990).Finally, the impacts of (de-)<strong>fragmentation</strong> on theecology and the concept of the TCRA can also affectthe local inhabitants and outdoor recreationists eitheron bike or foot. They may feel themselves safer and/orare less hampered <strong>by</strong> noise and emissions. For farmers,road crossings with cattle or tractor can becomeeasier. Thus, an improvement of the liveability of localresidents and recreationists can go hand in hand withde-<strong>fragmentation</strong> of <strong>habitat</strong>.7. ConclusionThe concept of the TCRA was developed for TheNetherlands (Jaarsma, 1997), a small industrialisedcountry with a dense population and a dense <strong>rural</strong> roadnetwork. The idea behind <strong>rural</strong> <strong>traffic</strong> calming thatincludes the concentration of <strong>through</strong> <strong>traffic</strong> on alimited number of <strong>rural</strong> highways and leaving the<strong>minor</strong> <strong>roads</strong> to smaller flows of <strong>rural</strong> bound <strong>traffic</strong>,is believed to be equally relevant in other countries.The relevancy is greatest where present diffuse <strong>traffic</strong>flows are hampering local people, recreationists, and/or the fauna. To implement the concept alternativeroutes within the region must be available, usuallyafter adaptation of existing <strong>roads</strong>. In countries with adense public road network, such alternatives are generallyavailable.The concept of TCRA was developed to improve<strong>rural</strong> <strong>traffic</strong> safety (Jaarsma, 1994). The concept hasalso proven useful for reducing <strong>habitat</strong> <strong>fragmentation</strong>along MRRs (Jaarsma and van Langevelde, 1997). Toreduce the <strong>habitat</strong> <strong>fragmentation</strong> of the complementing<strong>roads</strong>, such as the <strong>rural</strong> highways and motorways,various engineering and design solutions are available.These include fences, green bridges, and underpasses.Finally, <strong>by</strong> decreasing the current concentration ofdiffuse flows on the <strong>minor</strong> <strong>roads</strong> in favour of a limitednumber of major <strong>roads</strong>, the effects of the large-scalesolutions for the <strong>fragmentation</strong> of <strong>habitat</strong> will beincreased.AcknowledgementsThe authors are grateful to their colleague ChrisBaltjes for preparing and executing the GIS presentationsand the calculation of the acreage in both pilotstudies. Furthermore, we like to thank Donna Isaac,research fellow at the Land Use Planning Group, forcorrecting and improving the language of the paper.ReferencesCuperus, R., Kanters, K.J., Schaik, A.W.J., 1993. 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De dassenpopulatie in Nederland in 1960–1990(the badgerpopulation in The Netherlands in 1960–1990).Rapport 91/6, Rijksinstituut voor natuurbeheer, Leersum (inDutch).Catharinus F. Jaarsma is a senior lecturer at the WageningenUniversity, Land Use Planning Group. His research and his coursesfocus on <strong>rural</strong> road and <strong>traffic</strong> planning for several types of landuse. Growing attention for environmental aspects and <strong>habitat</strong><strong>fragmentation</strong> have given new impulses to <strong>rural</strong> <strong>traffic</strong> research,within the wider perspective of environmental circumstances.Geert P.A. Willems graduated in Land Use and Spatial Planning atWageningen University in 1999. In 2000, he joined the Land UsePlanning Group of Wageningen University as a research fellow. Hisresearch focuses on the ecological, environmental and safetyimpacts of infrastructure in <strong>rural</strong> areas.