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44 TEXTBOOK OF PHARMACOGNOSY AND PHYTOCHEMISTRY<br />

root. It grows further to produce branches in the<br />

acropetal manner known as secondary roots which<br />

further branches to give tertiary roots. These are<br />

all true roots. This system is characteristic <strong>of</strong><br />

dicotyledons.<br />

(b) Adventitious root system: The roots that develop<br />

from any part <strong>of</strong> the plant other than radicle are<br />

termed as adventitious roots. They may develop<br />

from root base nodes or internodes. This type <strong>of</strong><br />

root system is found in monocots <strong>and</strong> in pteridophytes.<br />

(a)<br />

Fig. 4.19 (a) Tap root system <strong>and</strong> (b) Adventitious root system<br />

I. Modification for storage <strong>of</strong> food: This type <strong>of</strong> modification<br />

is shown by both the types <strong>of</strong> roots, i.e. tap roots <strong>and</strong><br />

adventitious roots. They store carbohydrates <strong>and</strong> are used<br />

during early growth <strong>of</strong> successive season.<br />

(i) Tap roots show the following three types <strong>of</strong> modifications:<br />

(a) Conical: These are cone-like, broader at the base<br />

<strong>and</strong> tape-ring at the tip, e.g. carrot.<br />

(b) Fusiform: These roots are more or less spindle<br />

shaped, i.e. tapering at both the ends, e.g. radish.<br />

(c) Napiform: These are spherical shaped <strong>and</strong> very<br />

sharply tapering at lower part, e.g. beat <strong>and</strong> turnip.<br />

(b)<br />

(a) Tuberous roots: These get swollen <strong>and</strong> form single<br />

or isolated tuberous roots which are fusiform in<br />

shape, e.g. sweet potato, jalap, aconite.<br />

(b) Fasciculated tuberous roots: When several tuberous<br />

roots occur in a group or cluster at the base<br />

<strong>of</strong> a stem they are termed as fasciculated tuberous<br />

roots as in dahlia, asparagus.<br />

(c) Palmated tuberous roots: When they are<br />

exhibited like palm with fingers as in<br />

common ground orchid.<br />

(d) Annulated roots: The swollen portion is in the<br />

form <strong>of</strong> a series <strong>of</strong> rings called annules as in<br />

ipecacuanha.<br />

II. Modifications for support: Plant develops special<br />

aerial roots to <strong>of</strong>fer additional support to the plant by way<br />

<strong>of</strong> adventitious roots.<br />

(a) Clinging or Climbing roots: These types <strong>of</strong> roots<br />

are developed by plants like black pepper for<br />

support or for climbing purposes at nodes.<br />

(b) Stilt roots: This type <strong>of</strong> root is observed in maize<br />

<strong>and</strong> screw-pine, which grow vertically or obliquely<br />

downwards <strong>and</strong> penetrate into soil <strong>and</strong> give additional<br />

support to the main plant.<br />

(c) Columnar roots: In certain plants like banyan, the<br />

additional support is given by specially developed<br />

pillars or columnar roots. They even perform the<br />

function <strong>of</strong> regular roots.<br />

(a) (b) (c)<br />

Fig. 4.21 (a) Climbing root (b) Stilt root (c) Columnar root<br />

(a)<br />

(b)<br />

Fig. 4.20 (a) Conical root, (b) Fusiform root <strong>and</strong> (c) Napiform<br />

root<br />

(ii)<br />

Adventitious root show the following types <strong>of</strong> modifications.<br />

They store carbohydrates but do not assume<br />

any special shape.<br />

(c)<br />

III. Modifications for special functions:<br />

(a) Respiratory roots or pneumatophores: The roots<br />

<strong>of</strong> the plant growing in marshy places on seashores<br />

due to continuous water logging are unable<br />

to respire properly. They develop some roots<br />

growing against the gravitational force (in the<br />

air) with minute openings called lenticels. With<br />

the help <strong>of</strong> lenticels they carry on the exchange<br />

<strong>of</strong> gases. They look like conical spikes around<br />

the stems. This type <strong>of</strong> root is observed in case<br />

<strong>of</strong> plants called mangroves found in creeks, i.e.<br />

avicinnia.<br />

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