Invert database Query - UNIS

Notes: This is not a definitative checklist!
This list is based on a collection of over 500 articles concerning the terrestrial invertebrate fauna of
Svalbard. Nonetheless it should be not be considered as an accurate overview of the complete
invertebrate fauna of the archipelago. The terrestrial invertebrate fauna is largely only known from
two localities on Svalbard, Ny-Ålesund (Kongsfjord) and the Longyearbyen region. Little is known
about the fauna of the east coast or the interior. In addition there are large numbers of doubtful
identifications and confusion in the taxonomy.
Coulson, S.J. 2007 The terrestrial and freshwater invertebrate fauna of the High Arctic
archipelago of Svalbard. Zootaxa 1448; 41-58.
Coulson, S.J. Refseth, D. (2004) The terrestrial and freshwater invertebrate fauna of Svalbard
(and Jan Mayen), pp57-122 in A catalogue of the terrestrial and marine animals of Svalbard
(eds. Prestrud, P. Strøm H. Goldman H.). Skrifter 201. Norwegian Polar Institute, Tromsø.
13. april 2011
The
terrestrial
and
freshwater
invertebrate
13. april 2011
fauna of
Svalbard
Page 1 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Sarcomastigophora
Heliozoea
_
_
_
Actinophryridae
Actinophrys sol Ehrenberg
Clathrulinidae
Clathrulina elegans Cienk
Zoomastigophorea
Cercomonadida
_
_
13. april 2011
Cercomonadidae
Cercomonas gigantica Myl'nikov
Cercomonas kiaerdammane Myl'nikov
Page 2 of 52

Rhizopoda
Filosea
Gromiida
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Cyphoderiidae
Cyphoderia ampulla Leidy
Cyphoderia perlucidus Beyens and Charedez
Euglyphidae
Assulina muscorum Greeff
Assulina muscorum var. denticulata Chardez
Assulina muscorum var. stenostoma Schönborn
Assulina seminulum (Ehrenberg) Leidy
Euglypha acanthophora Perty
Euglypha alveolata Dujardin
Euglypha ciliata Ehrenberg
Euglypha ciliata var. glabra Wailes
Euglypha cristata Leidy
Euglypha compressa Carter
Euglypha filifera Penard
Euglypha laevis (Ehrenberg) Perty
Euglypha laevis var. lanceolata Playfair
Euglypha minor Chardez
Euglypha polylepsis Bonnet and Thomas
Euglypha rotunda Wailes and Penard
Euglypha rotunda var. dorsalis Decloitre
Euglypha rotunda var. glabra Wailes
Euglypha rotunda var. minor Wailes
Euglypha strigosa Leidy
Euglypha strigosa f. glabra Wailes
Euglypha strigosa var. muscorum Wailes
Euglypha tuberculata Dujardin
Euglypha tuberculata var. subcylindrica (Playfair) Decloitre
Tracheleuglypha dentata (Vejdovsky) Deflandre
Valkanovia delicatula Valkanov
Gromiidae
Gromia sp.
Nadinella tenella Penard
Pseudodifflugia globulosa Stepanek
Pseudodifflugia gracilis Schlumberger
Pseudodifflugia gracilis var. terricola Bonnet and Thomas
Pseudodifflugia senartensis Couteaux
Paulinellidae
Paulinella chromatophora Lauterborn
Trinematidae
Corythion aerophila Decoitre
Corythion delamarei Bonnet and Thomas
Corythion dubium Taranek
Page 3 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Corythion dubium var. aerophila Decloitre
Corythion dubium f. minima Chardez
Corythion dubium var. orbicularis Penard
Corythion dubium var. terricola Schönborn
Corythion pulchellum Penard
Trinema complanatum Penard
Trinema complanatum var. globulosa Chardez
Trinema enchelys (Ehrenberg) Leidy
Trinema lineare Penard
Trinema lineare f. minuscula Chardez
Trinema lineare f. truncatum Chardez
Trinema lineare var. longa Korganova
Trinema navicularis Decloitre
Trinema penardi Thomas and Chardez
Page 4 of 52

Lobosea
Amoebida
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Amoebidae
Arcellinida
_
13. april 2011
Amoeba radiosa Ehrenberg
Amoeba verrucosa Ehrenberg
_
Arcellidae
Arcella atrocera Leidy
Arcella arenaria Greeff
Arcella arenaria f. compressa Chardez
Arcella arenaria var. sphagnicola Deflandre
Arcella bathystoma Deflandre
Arcella discoides Ehrenberg
Arcella ovaliformis Chardez and Beyens
Arcella rotundata Playfair
Arcella rotundata var. aplanata Deflandre
Arcella rotundata var. undulata Stepanek
Arcella vulgaris Ehrenberg
Chlamydophrys stercorea Cienk
Cochliopodium erinaceum Penard
Cochliopodium obscurum Penard
Cochliopodium opalinum Penard
Difflugiella horrida Schönborn
Difflugiella oviformis (Penard) Bonnet and Thomas
Difflugiella pusilla (Playfair) Grospietsch
Microcorycia flava Greef
Microcorycia husvikensis Beyens and Chardez
Parmulina louisi Chardez and Beyens
Pseudochlamys patella C. and L.
Pyxidicua cymbalum Penard
Pyxidicula ornata Bartos
Centropyxidae
Centropyxis aculeata Stein
Centropyxis aculeata f. compressa
Centropyxis acuminata Couteaux and Chardez
Centropyxis aerophila Deflandre
Centropyxis aerophila var. constricta Decloitre
Centropyxis aerophila var. deflandrei Decloitre
Centropyxis aerophila var. sphagnicola Deflandre
Centropyxis aerophila var. minuta Chardez
Centropyxis cassis (Wallach) Deflandre
Centropyxis cassis var minima van Oye
Centropyxis constricta (Ehrenburg) Penard
Centropyxis cordiformis Couteaux
Page 5 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Centropyxis delicatula Penard
Centropyxis delicatul var. maior Jung
Centropyxis ecornis Leidy
Centropyxis ecornis var. deflandrei Thomas
Centropyxis gasparella
Centropyxis gasparella var. corniculata Beyens and Chardez
Centropyxis gibba var. gibbosa Godeanu
Centropyxis globulosa Bonnet and Thomas
Centropyxis laevigata var. maior Jung
Centropyxis lata Jung
Centropyxis minima Deflandre
Centropyxis minuta Deflandre
Centropyxis orbicularis Deflandre
Centropyxis plagiostoma Bonnet and Thomas
Centropyxis platystoma (Penard) Deflandre
Centropyxis pontigulasiformis Beyens and Chardez
Centropyxis sylvatica Thomas
Centropyxis sylvatica var. minor Bonnet and Thomas
Centropyxis ventricosa Bartos
Cryptodifflugiidae
Cryptodifflugia compressa f. angustioris Tarnogradsky
Difflugiidae
Difflugia bacillifera Penard
Difflugia bryophila Jung
Difflugia compressa var. minima Schönborn
Difflugia constricta Ehrenberg
Difflugia dujardini Chardez
Difflugia elegans Penard
Difflugia elegans var. parva Chardez
Difflugia glans Penard
Difflugia globularis Leidy
Difflugia globulosa Dujardin
Difflugia globulus Hopkinson
Difflugia lucida Penard 1902
Difflugia lacustris Jung
Difflugia lata Jung
Difflugia linearis (Penard) Gauthier-Lievre and Thomas
Difflugia minuta Rampi
Difflugia paulii Ogden
Difflugia penardi Hopkinson
Difflugia pulex Penard
Difflugia pulex var. cuneata Playfair
Difflugia pyriformis Perty
Difflugia tuberculata Playfair
Difflugia tuberculata var. sphaerica Playfair
Hyalospheniidae
Heleopera petricola Leidy
Heleopera petricola var amethystea Penard
Heleopera sylvatica Penard
Nebela bohemica Taranek
Nebela collaris Leidy
Nebela dentistoma Penard
Page 6 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Nebela dentistoma var. oblonga Gaut.-L and Thomas
Nebela flabellum Leidy
Nebela galeata Penard
Nebela lageniformis Penard
Nebela militaris Penard
Nebela minor Penard
Nebela nobilis Deflandre
Nebela parvula Penard
Nebela penardiana Deflandre
Nebela tincta (Leidy) Awerintzew
Nebela tincta f. stenostoma Jung
Nebela tincta var. rotunda Penard
Nebela tubulosa Penard
Nebela wailesi Deflandre
Lesquereusiidae
Netzelia labeosa Beyens and Chardez
Microcoryciidae
Microchlamys patella (Claparede and Lachmann) Cockerell
Microchlamys patella var. arctica Penard
Nebelidae
Schönbornia smithi Beyens and Chardez
Paraquadrulidae
Paraquadrula discoides (Penard)
Paraquadrula globulosa Ferlan
Paraquadrula irregularis Deflandre
Paraquadrula madarica Valkanov
Paraquadrula ogdeni Byens and Chardez
Paraquadrula rotunda Schönborn
Phryganelidae
Phryganella acropodia Hopkinson
Phryganella acropodia var. penardi Decloitre
Phryganella microps Valkanov
Phryganella nidilus Penard
Phryganella paradoxa var. alta Bonnet and Thomas
Phryganella paradoxa Penard
Plagiopyxidae
Plagiopyxis bathystoma Bonnet
Plagiopyxis callida Penard
Plagiopyxis callida var. grandis Thomas
Plagiopyxis declivis Thomas
Plagiopyxis labiata Penard
Plagiopyxis minuta Bonnet
Plagiopyxis oblonga (Bonnet and Thomas) Bonnet
Plagiopyxis penardi Thomas
Trigonopyxidae
Cyclopyxis ambigua Bonnet and Thomas
Cyclopyxis ampulla Leidy
Cyclopyxis arcelloides var. gibbosa van Oye
Cyclopyxis arcelloides var. minima van Oye
Cyclopyxis eurystoma Deflandre
Cyclopyxis eurystoma var. stenostoma Decloitre
Page 7 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Cyclopyxis kahli Deflandre
Cyclopyxis leclerqi Beyens and Chardez
Cyclopyxis perlucidus Beyens and Chardez
Cyclopyxis plana var. microstoma Schönborn
Cyclopyxis puteus Thomas
Trigonopyxis arcula Penard
Vahlkampfiidae
Naegleria polaris De Jonckheere 2006
Naegleria neopolaris De Jonckheere 2006
Naegleria arctica De Jonckheere 2006
Naegleria spitzbergensis De Jonckheere 2009
Page 8 of 52

Ciliophora
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Kinetofragminophorea
Colpodida
_
_
Cyrtolophosis mucicola Stokes 1885
Platyophrya vorax Kahl 1926
Rostrophryides sp. nov.
Colpodidae
Colpoda cucullus (Müller 1773)
Cyrtophorida
_
_
Nassulida
_
_
Odontochlamys alpestris Foissner 1981
Chilodonella uncinata (Ehrenberg 1835)
Furgasonia cf. trichocystis
Kreyella minuta Foissner 1979
Leptopharynx costatus Mermod 1914
Pleurostomatida
_
_
Prostomatida
_
_
13. april 2011
Litonotus lamella (Müller 1773)
Askenasia sp.
Dileptus margaritifer (Ehrenberg 1833)
Fuscheria cf. nodosa
Phialina vertens (Stokes, 1885)
Protospathidium sp.
Trachelophyllum apiculatum (Perty 1852)
Lacrymaria filiformis
Monodinium sp.
Urotrichidae
Urotricha agilis (Stokes 1886)
Urotricha cf. armata
Page 9 of 52

Suctorida
_
_
Synhymeniida
_
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Sphaerophrya terricola Foissner 1986
Nassulopsis cf. elegans
Oligohymenophorea
Hymenostomatida
_
_
Frontonia acuminata (Ehrenberg 1833)
Frontonia angusta
Tetrahymena rostrata
Cinetochilidae
Cinetochilum margaritaceum (Perty 1852)
Oxytrichidae
Paramecium arcticum Doroszewski
Paramecium aurelia complex
Peritrichida
_
_
Opercularia coarctata Claparede & Lachmann 1854
Pseudovorticella monilata (Tatem 1870)
Mobilidae
Trichodina tentaculata
Sessilidea
Ophrydium versatile
Vorticella aquadulcis complex
Vorticella infusionum complex
Vorticella microstoma
Scuticociliatida
_
_
13. april 2011
Cyclidium glaucoma Müller 1773
Sathrophilus muscorum (Kahl 1931)
Protocyclidium muscicola (Kahl 1931)
Page 10 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Polyhymenophorea
Heterotrichida
_
_
Hypotrichida
_
_
Blepharisma hyalinum Perty 1849
Bryometopus pseudochilodon Kahl 1932
Climacostomum virens (Ehrenberg 1838)
Aspidisca cicada (Müller 1786)
Oxytricha balladyna Song & Wilbert 1989
Oxytricha setigera Stokes 1891
Oxytricha similis Engelmann 1862
Sterkiella histriomuscorum Foissner, Blatterer, Berger & Kohmann 1991
Stichotricha aculeata Wrzesniowski 1866
Stylonychia mytilus complex
Uroleptus gallina (Müller 1786)
Oxytrichidae
Stylonychia pustulata (Müller 1786)
Oligotrichida
_
_
13. april 2011
Halteria grandinella (Müller 1773)
Limnostrombidium viride (Stein 1859)
Rimostrombidium sp.
Stombidium sp
Page 11 of 52

Rhizopoda
Lobosea
Arcellinida
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Arcellidae
Arcella cinctus
Centropyxidae
Cyclopyxis arcelloides van Oye
Page 12 of 52

Apicomplexa
Sporozoea
Coccidia
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Sporozoea
Sarcocystis sp.
Toxoplasma gondii Nicolle & Manceaux 1909
Isospora plectrophenaxia Dolnik and Loonen 2007
Page 13 of 52

Rotifera
Digononta
Bdelloidea
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Adinetidae
Adineta barabata Janson 1893
Adineta gracilis Janson 1893
Adineta vaga (Davis 1873)
Adineta vaga minor Bryce 1893
Habrotrochidae
Habrotrocha angusticollis (Murray 1905)
Habrotrocha aspera (Bryce 1892)
Habrotrocha bidens (Gosse 1851)
Habrotrocha constricta (Dujardin 1841)
Habrotrocha elegans (Milne 1886)
Habrotrocha insignis Bryce 1915
Habrotrocha lata lata (Bryce 1892)
Habrotrocha ligula ligula Bryce 1913
Habrotrocha novemdens De Koning 1947
Habrotrocha pulchra (Murray 1905)
Habrotrocha pusilla (Bryce 1893)
Habrotrocha pussila textrix (Bryce 1897)
Habrotrocha roeperi (Milne 1889)
Habrotrocha rosa Donner 1949
Philodinidae
Ceratrocha cornigera (Bryce 1893)
Didymodactylos carnosus Milne 1916
Macrotrachela aculeata Milne 1886
Macrotrachela concinna (Bryce 1912)
Macrotrachela ehrenbergii (Janson 1893)
Macrotrachela insolita de Koning 1947
Macrotrachela habita (Bryce 1894)
Macrotrachela kakkosoma (Schulte 1954)
Macrotrachela multispinosa multispinosa Thomson 1892
Macrotrachela musculosa (Milne 1886)
Macrotrachela nana nana (Bryce 1912)
Macrotrachela papillosa (Thomson 1892)
Macrotrachela plicata hirundinella (Murray 1905)
Macrotrachela plicata plicata (Bryce 1892)
Macrotrachela quadricornifera Milne 1886
Mniobia bredensis De Koning 1947
Mniobia incrassata (Murray 1905)
Mniobia magna (Plate 1889)
Mniobia russeola (Zelinka 1891)
Mniobia symbiotica (Zelinka 18886)
Mniobia tentans Donner 1949
Mniobia tetraodon (Ehrenberg 1848)
Page 14 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Mniobia vargai Donner 1962
Mniobia variabilis Donner 1949
Philodinia acuticornis Murray 1902
Philodinia acuticornis odiosa Milne 1916
Philodinia brevipes Murray 1902
Philodina erhthrophthama Ehrenberg 1832
Philodina flaviceps Bryce 1906
Philodinia nemoralis Bryce 1903
Philodinia proterva Milne 1916
Philodinia rugosa rugosa Bryce 1903
Pleuretra alpium (Ehrenberg 1853)
Pleuretra brycei (Weber 1898)
Pleuretra hystrix Bartos 1950
Rotaria citrina Ehrenberg 1838
Rotaria rotatoria (Pallas 1766)
Rotaria sordida sordida (Western 1893)
Rotaria tardigrada (Ehrenberg 1832)
Scepantotrocha sp.
Page 15 of 52

Monogononta
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Collothecacea
_
_
Collothecidae
Collotheca campanulata (Dobie 1849)
Collotheca libera Zacharias 1894
Collotheca ornata cornuta (Dobie 1842)
Collotheca ornata ornuta (Ehrenberg 1832)
Flosculariacea
_
_
Filiniidae
Filinia terminalis (Plate 1886)
Flosculariidae
Ptygura melicerta (Ehrenberg 1832)
Ploimida
_
_
Brachionidae
Kellicottia longispina (Kellicott 1879)
Keratella cochlearis (Gosse 1851)
Keratella hiemalis (Carlin 1943)
Keratella quadrata (Müller 1786)
Keratella testudo (Ehrenberg 1832)
Notholca acuminata (Ehrenberg 1832)
Notholca acuminata var. extensa Olofsson 1918
Notholca foliacea (Ehrenberg 1838)
Notholca labis Gosse 1887
Notholca latistyla (Olofsson 1918)
Notholca latistyla f. amreni De Smet and Bafort 1990
Notholca marina (Focke 1991)
Notholca striata (Müller)
Notholca squamala (Müller 1786)
Colurellidae
Colurella adriatica Ehrenberg 1831
Colurella amblytelus (Gosse 1851)
Colurella colurus (Ehrenberg 1830)
Colurella geophila Donner 1981
Colurella hindenburgi Steinecke 1917
Colurella obtusa (Gosse 1886)
Colurella uncinata (Müller 1773)
Lepadella acuminata (Ehrenberg 1834)
Lepadella minuta (Montet 1918)
Lepadella ovalis (Müller 1786)
Lepadella patella (Müller 1786)
Lepadella quadricarinata (Stenroos 1898)
13. april 2011
Page 16 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Lepadella triba Myers 1934
Lepadella triptera Ehrenberg 1830
Lophocaris oxysternon (Goose 1851)
Dicranophoridae
Albertia naidis Bousfield 1886
Dicranophorus forcipatus (Müller 1786)
Encentrum bidentatum (Lie-Pettersen 1906)
Encentrum dieteri De Smet 1995
Encentrum felis (Müller 1773)
Encentrum ferox (Western) nom. dub.
Encentrum grandi Haring and Myers 1928
Encentrum marinum (Dujardin 1841)
Encentrum mucronatum Wulfert 1936
Encentrum murrayi Bryce 1922
Encentrum mustela (Milne 1885)
Encentrum permolle (Gosse 1886)
Encentrum uncinatum (Milne 1886)
Itura aurita aurita (Ehrenberg 1930)
Epiphanidae
Rhinoglena frontalis Ehrenberg 1858
Euchlanis deflexa Gosse 1851
Euchlanis dilatata Ehrenberg 1832
Euchlanis lyra Hudson 1886
Euchlanis meneta Myers 1930
Catyphna brevis Murray 1913
Catyphna rotundata Olofsson 1918
Lecanidae
Lecane bulla (Goose 1851)
Lecane closterocerca (Schmarda 1859)
Lecane cornuta (Müller 1786)
Lecane flexilis (Gosse 1886)
Lecane ligona (Dunlop 1901)
Lecane lunaris (Ehrenberg 1832)
Lecane pideis (Harring and Myers 1926)
Lecane robertsonae Segers 1993
Lecane rotundata (Oloffson 1918)
Lindiidae
Lindia cf. torulosa Dujardin 1841
Mytilinidae
Lophocharis oxysternon (Gosse 1851)
Mytilina bicarinata (Perty 1850)
Mytilina mucronata (Müller 1773)
Mytilina ventralis var. brevispina (Ehrenberg 1832)
Notommatidae
Cephalodella biungulata Wulfert 1937
Cephalodella catellina (O.F. Müller 1786)
Cephalodella evabroedi De Smet 1988
Cephalodella glandulosa Koch-Althaus 1962
Cephalodella gibba (Ehrenberg 1832)
Cephalodella gracilis (Ehrenberg 1832)
Cephalodella hoodi (Goose 1896)
Cephalodella intuta Myers 1924
13. april 2011
Page 17 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Cephalodella megalocephala (Glascott 1893)
Cephalodella misgurnus Wulfert 1937
Cephalodella rotunda Wulfert 1937
Cephalodella sterea (Gosse 1887)
Cephalodella ventripes (Dixon-Nuttall 1901)
Cephalodella ventripes var. angustior Donner 1949
Diaschiza spp.
Eosphora najas Ehrenberg 1830
Notommata cyrtopus Gosse 1886
Notommata glyphura Wulfert 1935
Resticula nyssa Harring and Myers 1924
Proalidae
Bryceella stylata (Milne 1886)
Bryceella tenella (Bryce 1897)
Proales decipiens (Ehrenberg 1832)
Proales fallaciosa Wulfert 1937
Proales sordida Gosse 1886
Wulfertia ornata Donner 1943
Scaridiidae
Scaridium longicaudum (Müller 1786)
Synchaetidae
Polyarthra dolichoptera (Idelson 1925)
Polyarthra dolichoptera f. aptera (Hood 1895)
Polyarthra dolichoptera f. proloba (Albertova 1960)
Polyarthra vulgaris Carlin 1943
Synchaeta lakowitziana Ehrenberg 1832
Synchaeta lakowitziana arctica De Smet 1988
Synchaeta littoralis Rousselet 1902
Synchaeta truncata von Hofsten
Trichoceridae
Trichocerca bidens (Lucks 1912)
Trichocerca. cavia (Gosse 1886)
Trichocerca cristata Harring
Trichocerca intermedia (Stenroos 1898)
Trichocerca longistyla (Olofsson 1918)
Trichocerca rattus (Müller 1776)
Trichocerca relicta (Donner 1950)
Trichocerca uncinata (Voigt 1902)
Trichocerca weberi (Jennings 1903)
Trichotria truncata (Whitelegge)
Trichotria pocillum (Müller 1776)
Page 18 of 52

Gastrotricha
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Chaetonotoida
_
_
13. april 2011
Chaetonotidae
Chaetonotus sp.
Page 19 of 52

Nematoda
Adenophorea
Stichosomida
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Capillariidae
Capillaria sp
Philometridae
Philonema oncorhynchi (Kuitenen Ekbaum 1933)
Trichinellidae
Trichinella spiralis (Owen 1935)
Trichinella nativa (Britov and Boev 1972)
Page 20 of 52

Penetrantia
Dorylaimida
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Dorylaimidae S.L.
Enchodelus analatus Thorne 1939
Enchodelus conicaudatus Thorne 1939
Enchodelus cf. macrodoroides (Steiner 1914)
Enchodelus macrodorus Thorne 1939
Enchodelus parvus Loof 1971
Epidorylaimus agilis (de Man 1880) Andrassy 1986
Epidorylaimus lugdunensis (de Man 1880)
Eudorylaimus alleni Brzeski1962
Eudorylaimus carteri (Bastian 1865)
Eudorylaimus centrocercus (de Man 1880)
Eudorylaimus circulifer (Loof 1961)
Eudorylaimus maksymovi (Altherr 1963)
Eudorylaimus megodon Loof 1971
Eudorylaimus parvus (de Man 1880)
Eudorylaimus subjunctus Loof 1971
Eudorylaimus vanrosseni Loof 1971
Longidorella magna Loof 1971
Mesodorylaimus sp.
Mononchidae
Prionchulis muscorum (Dujardin 1845)
Qudsianematidae
Labronema sp.
Tylencholaimidae
Tylencholaimus proximus Thorne 1939
Tylencholaimus teres Thorne 1939
Enoplida
_
_
Alaimidae
Alaimus arcuatus Thorne 1939
Alaimus depressus Loof 1971
Alaimus parvus Thorne 1939
Prismatolaimidae
Prismatolaimus dolichurus de Man 1880
Prismatolaimus intermedius (Buetschli1873)
Prismatolaimus primitivus Loof 1971
Prismatolaimus stenolaimoides Loof 1971
Tobrilidae
Tobrilus sp.
Tripylidae
Tripyla cf. infia Brzeski et Winiszewska-Slipinska 1903
Tripyla glomerans Bastian 1865
13. april 2011
Page 21 of 52

Secernentia
Ascaridida
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Anisakidae
Rhabditida
Anisakis simplex (Rudolphi 1802)
Contracaecum osculatum (Rudolphi 1801) Baylis 1920
Hysterothylacium aduncum (Rudolphi 1802)
_
_
Bunonematidae
Bunonema reticulatum Richters 1905
Cephalobidae
Acrobeles ciliatus von Linstow 1877
Acrobeloides bütschlii Gadea 1954
Acrobeloides enoploides Loof 1971
Acrobeloides tricornis (Thorne 1925)
Cervidellus serratus (Thorne 1925)
Cervidellus spitzbergensis Bostrøm 1987
Chiloplacus saccatus Loof 1971
Eucephalobus arcticus Loof 1971
Heterocephalobus elongatus (de Man 1880)
Stegelleta similis (Thorne 1925)
Panagrolaimidae
Panagrolaimus papillosus Loof 1971
Teratocephalidae
Euteratocephalus crassidens Andrassy 1958
Teratocephalus costatus Andrassy 1958
Teratocephalus decarinus Anderson 1969
Teratocephalus lirellus Anderson 1969
Teratocephalus terrestris (Butschli1873)
Spirurida
_
_
Acuriidae
Paracuaria adunca (Creplin 1846) Anderson and Wong 1981
Stegaphorus stellae-polaris (Parona 1901)
Strongylida
_
_
13. april 2011
Trichostrongylidea
Marshallagia marshalli m occidentalis (Ransom, 1907)
Marshallagia marshalli (Ransom 1907)
Nematodirus helvetianus May 1920
Ostertagia arctica Mitzkewitzsch 1929
Page 22 of 52

Tylenchida
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Ostertagia grühneri Skrjabin 1929
Ostertagia occidentalis Ransom 1907
Paracuaria adunca (Creplin 1846)
Skrjabinagia arctica (Mitzkevich 1929)
Teladorsagia circumcincta (Stadelmann 1894)
Teladorsaga davitiani Andreeva and Satubaldini1984
Teladorsagia trifurcata
Aphelenchinae
Aphelenchus nivalis Aurivillius 1883
Criconematidae
Criconemoides annulatus Taylor 1936
Criconema sp.
Hoplolaimidae
Helicotylenchus spitsbergensis Loof 1971
Pratylenchidae
Pratylenchoides crenicauda Winslow 1958
Pratylenchoides magnicauda (Thorne 1935)
Tylenchidae
Basiria dolichura Loof 1971
Coslenchus costatus (de Man 1921)
Filenchus thornei (Andrassy 1954)
Geocenamus arcticus (Mulvey 1969)
Geocenamus loofi (Siddiqi1979)
Geocenamus microdorus (Geraert 1966)
Lelenchus leptosoma (de Man 1880)
Malenchus bryophilus (Steiner 1914)
Nagelus leptus (Allen 1955)
Tylenchus davainei Bastien 1865
Page 23 of 52

Torquentia
Araeolaimida
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Plectidae
Anaplectus cf. grandepapillatus (Ditlevsen 1928)
Anaplectus granulosus de Coninck and Shürmans Stekhoven 1933
Anaplectus porosus Allen and Noffsinger 1968
Ceratoplectus communis (Bütschli1873)
Ereptonema arcticum Loof 1971
Plectus acuminatus Bastian 1865
Plectus cf armatus Butschli1873
Plectus assimilis Butschli1873
Plectus aquatilis Andrassy1984
Plectus cirratus Bastian 1865
Plectus cornus Maggenti1961
Plectus geophilus de Man 1880
Plectus inquirendus Andrassy 1958
Plectus longicaudatus Butschli1873
Plectus cf. opisthocirculus Andrassy 1952
Plectus parietinus Bastian 1865
Plectus parvus Bastian 1865
Plectus rhizophilus de Man 1880
Rhabdolaimidae
Rhabdolaimus terrestris de Man 1880
Chromadorida
_
_
Achromadoridae
Achromadora cf. semiarmata Altherr 1952
Achromadora tenax (de Man 1876)
Ethmolaimidae
Ethmolaimus sp.
Odontolaimus
Odontolaimus chlorurus de Man 1880
Monohysterida
_
_
Monhysteridae
Eumonhystera dispar (Bastian 1865)
Eumonhystera filiformis (Bastian 1865)
Eumonhystera vulgaris (de Man 1880)
Geomonohystera villosa (Butschli1873)
Monohystera stagnalis Bastian 1865
13. april 2011
Page 24 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Acanthoceophala
Palaecanthocephala
Polymorphida
_
_
13. april 2011
Polymorphidae
Corynosoma strumosum (Rudolphi 1802) Luhe 1904
Notocotylus sp.
Acantocephalan sp.
Page 25 of 52

Platyhelminthes
Cestoda
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Cyclophyllidae
_
_
Anoplocephalidae
Moniezia benedini Moniez 1872
Dilepididae
Anomotaenia campylacantha (Krabbe 1869)
Anomotaenia larina (Krabbe 1869)
Anomotaenia micacantha (Krabbe 1869)
Alcataenia (Rissotaenia) dominicana (Raillet and Henry 1912)
Arctotaenia tetrabothriodes
Paricteropaenia porosa (Rudolphi 1810)
Hymenolepididae
Alopparakis larina Furhmann 1921
Microsomacanthus ductilus
Microsomacanthus microsoma (Creplin, 1829) Lopez-Neyra, 1942
Microsomacanthus diorchis (Fuhrmann, 1913).
Taeniidae
Echinococcus multilocularis Leuckart 1863
Taenia crassiceps
Taenia ovis krabbei (Moniez 1879) Verster 1969
Taenia polyacantha
Tetrabothrius minor (Loennberg, 1893)
Tetrabothriidae
Tetrabothrius erostris (Loennberg 1889)
Proteocephalidea
_
_
Proteocephalidae
Proteocephalus exiguus Weinland 1858
Pseudophyllidea
_
Diphyllobothriidae
Diphyllobothriidae
Diphyllobothrium ditremum (Creplin 1825)
Triaenophoridae
Eubothrium crassum (Bloch 1779)
Eubothrium salvelini Schrank 1790
13. april 2011
Page 26 of 52

Trematoda
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Opisthorchiida
Opisthorchiata
_
Heterophyidae
Cryptocotyle lingua (Creplin 1825)
Plagiorchiida
Brachylaimata
_
Microphallidae
Microphallus pygmaeus (Levinsen,1881)
Microphallus pseudopygmaeus Galaktionov,1980
Microphallus triangulatus
Strigeata
Plagiorchiata
_
Gymnophallidae
Gymnophallus somateria (Levinsen,1881)
Gymnophallus choledochus Odhner,1900
13. april 2011
Page 27 of 52

Annelida
Oligochaeta
Haplotaxida
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Enchytraeidae
Bryodrilus diverticulatus Cernosvitov 1929
Bryodrilus ehlersi Ude 1892
Bryodrilus parvus Nurminen 1970
Cernosvitoviella immota (Knoller 1935)
Cernosvitoviella cf. Pusilla
Cognettia sphagnetorum (Vejdovsky 1877)
Enchytraeus albidus Henle 1837
Enchytraeus kincaidi Eisen 1904
Fridericia bulboides Neilsen and Christensen 1959
Fridericia bulbosa (Rosa 1887)
Fridericia leydigi (Vejdovsky 1877)
Henlea heleotropha Stephenson 1922 augm. Nielsen and Christensen 1959
Henlea glandulifera Nurminen 1970
Henlea nasuta (Eisen 1878)
Henlea perpusilla Friend 1911
Henlea similis Nielsen and Christensen 1959
Henlea ventriculosa (d’Udekem 1954)
Lumbricillus arenarius (Michaelsen 1988)
Lumbricillus bülowi Nielsen and Christiensen 1959
Lumbricillus eltoni (Stephenson 1924)
Lumbricillus muscicolus (Stephenson 1924)
Lumbricillus neilseni Nurminen 1965
Lumbricillus pagenstecheri (Ratzel 1869)
Lumbricillus reynoldsoni Backlund 1948
Lumbricillus rivalis Levinsen 1890
Lumbricillus semifuscus (Claparede 1861) augm. Stephenson 1911
Lumbricillus tuba Stephenson 1911
Marionina aporus (Stephenson 1925)
Marionina argentea (Michaelsen 1889)
Marionina crymodes (Stephenson 1922)
Marionina libra Neilsen and Christensen 1959
Mesenchytraeus argentatus Nurminen 1973
Mesenchytraeus eltoni Stephenson 1925
Mesenchytraeus flavus (Levinsen 1884)
Mesenchytraeus melanocephalus
Mesenchytraeus sp. n.
Page 28 of 52

Tardigrada
Eutardigrada
Apochela
_
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Milnesiidae
Parachela
_
_
13. april 2011
Hypsibiidae
Milnesium tardigradum Doyère 1840
Acutuncus antarcticus (Richters 1904)
Borealibius zetlandicus (Murray 1907)
Diphascon (Diphascon) stappersi Richters 1911
Diphascon (Diphascon) alpinum Murray 1906
Diphascon (Diphascon) chilenense Plate 1888
Diphascon (Diphascon) oculatum oculatum Murray 1906
Diphascon (Diphascon) pingue pingue(Marcus 1936)
Diphascon (Diphascon) recamieri Richters 1911
Diphascon (Diphascon) tenue Thulin 1928
Diphascon (Adropion) arduifrons Thulin 1928
Diphascon (Adropion) belgicae Richters 1911
Diphascon (Adropion) prorsirostre Thulin 1928
Diphascon (Adropion) scoticum J.Murray 1905
Doryphoribius macrodon Binda Pilato and Dastych 1980
Hebesuncus conjungens (Thulin 1911)
Hypsibius arcticus (Murray 1907)
Hypsibius convergens (Urbanowicz 1925)
Hypsibius dujardini (Doyère 1840)
Hypsibius microps Thulin 1928
Hypsibius pallidus Thulin 1911
Isohypsibius bakonyiensis (Iharos 1964)
Isohypsibius dastychi Pilato Bertolani and Binda 1982
Isohypsibius ceciliae Pilato and Binda 1987
Isohypsibius elegans Binda and Pilato 1971
Isohypsibius fuscus (Mihelčič 1971/72)
Isohypsibius granulifer granulifer Thulin 1928
Isohypsibius papillifer papillifer (Murray 1905)
Isohypsibius papillifer bulbosus (Marcus 1928)
Isohypsibius prosostomus prosostomus Thulin 1928
Isohypsibius pulcher (Mihelčič 1971/72)
Isohypsibius schaudinni (Richters 1909)
Isohypsibius tuberculatus (Plate 1888)
Itaquascon enckelli (Mihelčič 1971/72)
Mesocrista spitzbergensis (Richters 1903)
Platicrista angustata (Murray 1905)
Ramazzottius cataphractus (Maucci1974)
Ramazzottius montivagus (Dastych 1983)
Page 29 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Ramazzottius oberhaeuseri (Doyère 1840)
Thulinius augusti (Murray 1907)
Macrobiotidae
Macrobiotus ariekammensis Weglarska 1965
Macrobiotus brevipes (Mihelčič 1971/72)
Macrobiotus crenulatus Richters 1904
Macrobiotus echinogenitus Richters 1903
Macrobiotus harmsworthi harmsworthi Murray 1907
Macrobiotus harmsworthi obscurus Dastych 1985
Macrobiotus hufelandi hufelandi C.A.S. Schultze 1833
Macrobiotus islandicus islandicus Richters 1904
Macrobiotus norvegicus Mihelčič 1971/72
Macrobiotus spectabilis Thulin 1928
Macrobiotus tenuiformis Tumanov 2007
Macrobiotus voronkovi Tumanov 2007
Macrobiotus willardi Pilato 1977
Minibiotus intermedius (Plate 1888)
Paramacrobiotus areolatus ( Murray 1907)
Paramacrobiotus richtersi (Murray 1911)
Tenuibiotus danilovi Tumanov 2007
Richtersius coronifer (Richters 1903)
Calohypsibiidae
Calohypsibius ornatus (Richters 1900)
Eohypsibiidae
Bertolanius nebulosus (Dastych 1983)
Bertolanius smreczynskii (Weglarska 1970)
Bertolanius weglarskae (Dastych 1972)
Murrayidae
Dactylobiotus ambiguus (Murray 1907)
Dactylobiotus dispar (Murray 1907)
Dactylobiotus macronyx (Dujardin 1851)
Murrayon dianeae (Kristensen 1982)
Murrayon hibernicus (Murray 1911)
Murrayon pullari (Murray 1907)
Page 30 of 52

Heterotardigrada
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Arthrotardigrada
_
_
13. april 2011
Echiniscidae
Bryodelphax sinensis (Pilato 1974)
Echiniscus arctomys Ehrenberg 1853
Echiniscus blumi blumi Richters 1903
Echiniscus capillatus Ramazzotti1956
Echiniscus granulatus (Doyère 1840)
Echiniscus loxophthalamus Richters 1911
Echiniscus merokensis Richters 1904
Echiniscus oihonnae Richters 1903
Echiniscus quadrispinosus Richters 1902
Echiniscus spinulosus (Doyère 1840)
Echiniscus testudo (Doyère 1840)
Echiniscus wendti Richters 1903
Pseudechiniscus islandicus (Richters 1904)
Pseudechiniscus suillus (Ehrenberg 1853)
Pseudechiniscus victor (Ehrenberg 1853)
Testechiniscus spitsbergensis (Scourfield 1897)
Page 31 of 52

Bryozoa
Phylactolaemata
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Statoblasts. Species unknown
Page 32 of 52

Chelicerata
Arachnida
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Gnaphosidae
Micaria constricta Emerton 1894
Hahniidae
Hahnia helveola Simon 1875
Linyphiidae
Halorates holmgreni (Thorell 1871)
Halorates spetsbergensis (Thorell 1871)
Halorates thulensis (Jackson 1934)
Erigone longipalpis Sundevall 1830
Erigone arctica palearctica Braendegaard 1934
Erigone psychrophila (Thorell 1871)
Erigone tirolensis L. Koch 1872
Hilaria glacialis (Thorell 1871)
Improphantes complicatus (Emerton 1882)
Mughiphantes sobrius (Thorell 1871)
Mecynargus borealis (Jackson 1930)
Agyneta nigripes (Simon 1884)
Oreonetides vaginatus (Thorell 1872)
Walckenaeria karpinskii (O.P.-Cambridge 1873)
Tapinocyba insecta (L.Koch 1869)
Philodromidae
Thanatus formicinus (Clerck 1757)
Acariformes
Oribatida
Ameronothroidea
Ameronothridae
Ameronothrus lineatus (Thorell 1871)
Ameronothrus nidicola Sitnikova 1975
Brachychthonoidea
Brachychthoniidae
Eobrachychthonius borealis (Forsslund 1942)
Eobrachychthonius latior (Berlese 1910)
Eobrachychthonius oudemansi van der Hammen 1952
Liochthonius alpestris (Forsslund 1958)
Liochthonius brevis (Michael 1888)
Liochthonius laetepictus (Berlese 1910)
Liochthonius lapponicus (Trägårdh 1910)
Liochthonius muscorum Forsslund 1964
Liochthonius neglectus Moritz 1976
Liochthonius sellnicki (Thor 1930)
Liochthonius strenzki Forsslund 1942
Neoliochthonius piluliferus (Forsslund 1942)
13. april 2011
Page 33 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Malaconothridae
Malaconothrus egregius (Berlese 1904)
Phthiacaroidae
Steganacarus striculus (C.L. Koch 1836)
Carabodoidea
Carabodidaae
Carabodes marginatus (Michael, 1884)
Tectocepheidae
Tectocepheus alatus Berlese 1913
Tectocepheus sarekensis Trägårdh 1910
Tectocepheus velatus (Michael 1880)
Tectocepheus velatus knullei Vanek 1960
Ceratozetoidea
Ceratozetidae
Ceratozetes spitsbergensis Thor 1934
Chamobates borealis (Trägårdh 1902)
Chamobates birulai (Kulcynski1902)
Chamobates cuspidatus (Mihael, 1844)
Diapterobates notatus (Thorell 1871)
Edwardzetes edwardsii (Nicolet 1855)
Luoribates gracilis Sellnick 1944
Mycobates parlmeliae (Michael 1884)
Mycobates sarekensis (Trägårdh 1910)
Mycobates tridactylus Willmann 1929
Oromurcia lucens (L.Koch 1879)
Punctoribates punctum (C.L. Koch 1839)
Sphaerozetes piriformis (Nicolet 1855)
Svalbardia paludicola Thor 1930
Trichoribates novus (Sellnick 1928)
Trichoribates setiger (Trägårdh 1910)
Trichoribates trimaculatus (C.L.Koch 1836)
Cymbaeremaeoidea
Achipterioidae
Achipteria sp.
Parachipteria punctata (Nicolet 1855)
Micreremidae
Micreremus brevipes (Michael 1888)
Damaeoidea
Damaeidae
Oribata ursina Thor 1930
Spatiodamaeus sp.
Damaeus onustus C.L. Koch, 1841
Euphthiracaroidea
Euphthiracaridae
Microtritia sp.
Galumnoidea
Galumnidae
Pergalumna nervosa (Berlese 1914)
Gustavioidea
Page 34 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Peloppiidae
Ceratoppia bipilis (Hermann 1804)
Ceratoppia hoeli Thor 1930
Ceratoppia sphaerica (L.Koch 1879)
Gymnodamaeoidea
Gymnodamaeidae
Gymnodamaeus sp.
Hermannioidea
Hermanniidae
Hermannia reticulata Thorell 1872
Hermannia scabra (L.Koch 1879)
Licneremaeoidea
Licneremaeidae
Scutovertex minutus (C.L.Koch 1835)
Nanhermannioidea
Nanhermannioidae
Nanhermannia sellnicki (Forsslund 1958)
Nothroidea
Camisiidae
Camisia anomia Colloff 1993
Camisia biverrucata (C.L.Koch 1839)
Camisia borealis (Thorell 1871)
Camisia foveolata Hammer 1955
Camisia horrida (Hermann 1804)
Camisia invenusta (Michael 1888)
Camisia lapponica (Trägårdh 1910)
Camisia spinifer (C.L.Koch 1836)
Nothridae
Nothrus palustris (C.L.Koch 1839)
Platynothrus peltifer (C.L.Koch 1839)
Platynothrus punctatus (C.L.Koch 1839)
Platynothrus septentrionalis Sellnick 1944
Oppioidea
Autognetidae
Autogneta kaisilai Karppinen 1997
Oppiidae
Berniniella sp.
Discoppia splendens (C.L. Koch 1840)
Lauroppia fallax (Paoli 1908)
Lauroppia translamellata (Willmann 1923)
Dissorhina ornata (Oudemans 1900)
Microppia minus (Paoli 1908)
Oppiella neerlandica (Oudemans 1902)
Oppiella nova Oudemans 1902
Oppiella (Moritzoppia) unicarinata (Paoli 1908)
Suctobelbella hammeri (Krivolutsky 1966)
Suctobelbella sarekensis (Forsslund 1941)
Suctobelbella subcornigera (Forsslund 1941)
Oribatelloidea
Oribatellidae
13. april 2011
Page 35 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Oribatella arctica Thor 1930
Oripodoidea
Oribatulidae
Oribatula tibialis (Nicolet 1885)
Oribatula venusta Berlese 1908
Phauloppia lucorum (C.L.Koch 1841)
Zygoribatula exilis (Nicolet 1855)
Oripodiidae
Liebstadia similis (Michael 1888)
Trombidiformes (Actinedida)
Bdelloidea
Bdellidae
Bdella longicornis (Linnaeus 1758)
Bdella semiscutata Thor 1930
Biscirus lapidarius (Kramer 1881)
Cyta latirostris (Hermann 1804)
Cyta brevirostris (L.Koch 1879) [a variety of C. latirostris]
Neomolgus capillatus (Kramer 1881)
Neomolgus littoralis (Linnaeus 1758)
Neomolgus pallipes (L.Koch)
Cunaxidae
Cunaxoides croceus (C.L.Koch 1838)
Eupodoidea
Eupodidae
Cocceupodes clavifrons (R. Canestrini1886)
Eupodes variegatus C.L.Koch 1838
Protereunetes borneri Thor 1934
Penthaleidae
Penthaleus maior (Duges 1834)
Penthalodidae
Penthalodes ovalis (Dugès 1834)
Rhagidiidae
Coccorhagidia clavifrons (Canestrini1886)
Rhagidia gelida Thorell 1872
Lebertioidea
Sperchonidae
Sperchon brevirostris (Koenike 1895)
Pachygnathoidea
Alicorhagiidae
Alicorhagia clavipilus (Thor 1931)
Alicorhagia plumipilis (Thor 1931)
Nanorchestidae
Nanorchestes arboriger (Berlese 1904)
Pygmephoroidea
Pygmephoridae
Bakerdania arcticus (Thor 1934)
Raphignathoidea
Stigmaeidae
Eustigmaeus oudemansi (Thor 1930)
Page 36 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Eustigmaeus pulchellus (Thor 1930)
Tetranychoidea
Tetranychidae
Bryobia borealis Oudemans 1930
Trombidioidea
Podothrombiidae
Podothrombium bicolor (Hermann 1804)
Podothrombium curtipalpe (Thor 1900)
Podothrombium svalbardense Oudemans 1930
Tydeoidea
Tydeiidae
Coccotydeus globifer (Thor 1931)
Microtydeus constans Thor 1931
Triophtydeus pinicolus (Oudemans 1929)
Tydaeolus tenuiclaviger (Thor 1931)
Tydeus croceus (Linnaeus 1758)
Tydeus langei Thor 1934
Tydeus svalbardensis Thor 1932
Parasitiformes
Gamasida (Mesostigmata)
Ascoidea
Ascidae
Antennoseius (Vitzthumia) oudemansi (Thor 1930)
Arctoseius babenkoi Makarova 1999
Arctoseius cetratus (Sellnick 1940)
Arctoseius haalovi Lindquist 1963 ms
Arctoseius laterincisus Thor 1930
Arctoseius multidentatus Evans 1955
Arctoseius ornatus Evans 1955
Arctoseius pristinus Karg 1992
Arctoseius tajnnricus Petrova and Makarova 1991
Arctoseius weberi Evans 1955
Melichares sp.
Proctolaelaps parvanalis (Thor 1930)
Zerconopsis sp.
Phytoseiidae
Amblyseius magnanalis Thor 1930
Dermanyssoidea
Haemogamasidae
Haemogamasus ambulans (Thorell 1872)
Haemogamasus nidi Michael 1892
Hypoaspididae
Hypoaspis ovalis L.Koch
Laelapidae
Laelaps hilaris (Koch)
Eviphidoidea
Eviphididae
Thinoseius spinosus (Willmann, 1939)
Rhodacaroidea
13. april 2011
Page 37 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Ologamasidae
Gamasellus borealis (Koch 1879)
Veigaioidea
Veigaiidae
Zercon andrei Sellnick, 1958
Zerconoidea
Zerconidae
Zercon forsslundi Sellnick 1958
Zercon spinosus
Zercon triangularis C.L.Koch 1836
Zercon zelawaensis Sellnick 1944
Zerconopsis muestairi (Schweizer, 1949)
Ixodida (Metastigmata)
Ixodoidea
Ixodidae
Ixodes (Ceratixodes) uriae White 1852
Page 38 of 52

Mandibulata
Collembola
Arthropleona
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Arrhopalitidae
Arrhopalites principalis Stach 1945
Entomobryidae
Entomobrya nivalis (L. 1758)
Lepidocyrtus lignorum Fabricus 1793
Hypogastruridae
Bonetogastrura nivalis (Martynova 1973)
Ceratophysella longispina (Tullberg 1876)
Ceratophysella succinea (Gisin 1949)
Hypogastrura concolor (Carpenter 1900)
Hypogastrura sensilis (Folsom 1919)
Hypogastrura tullbergi (Schäffer 1900)
Hypogastrura viatica (Tullberg 1871)
Podura hyperborea Boheman 1865
Willemia anophthalma Börner 1901
Willemia scandinavica Stach 1949
Willemia similis Mills 1934
Xenylla humicola (Fabicius 1780)
Isotomidae
Agrenia bidenticulata (Tullberg 1876)
Archisotoma besselsi (Packard 1877)
Archisotoma megalops (Bagnall 1939)
Archisotoma polaris Fjellberg and Poinsot 1975
Folsomia coeruleogrisea (Hammer, 1938)
Folsomia bisetosa Gisin 1953
Folsomia bisetosella Fjellberg 2005
Folsomia ciliata Babenko & Bulavintsev, 1993
Folsomia quadrioculata (Tullberg 1871)
Folsomia binoculata (Wahlgren, 1899)
Folsomia sexoculata (Tullberg 1871)
Folsomia stella Grow and Christiansen 1976
Folsomia taimyrica Martnova 1973
Parisotoma notabilis Schäffer 1896
Isotoma anglicana Lubbock 1862
Desoria multisetis (Carpenter and Phillips 1922)
Desoria neglecta (Schäffer 1900)
Desoria olivacea (Tullberg 1871)
Isotoma regularis Hammer 1957
Desoria fennica (Reuter, 1895)
Desoria tshernovi Martynova 1974
Pseudanurophorus inoculatus Bödvarsson 1958
Pseudanurophorus psammophilus Potapov & Stebaeva, 2002
Tetracanthella arctica Cassagnau 1959
Page 39 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Vertagopus arcticus Martynova 1969
Vertagopus pseudocinereus Fjellberg 1975
Neanuridae
Anurida granaria (Nicolet 1847)
Anurida maritima (Guerin 1836)
Anurida polaris (Hammer 1954)
Friesea quinquespinosa Wahlgren 1900
Micranurida pygmaea Börner 1901
Odontellidae
Xenyllodes armatus Axelson 1903
Onychuiridae
Megaphorura arctica (Tullberg 1876)
Mesaphorura jirii Rusek 1982
Mesaphorura macrochaeta Rusek 1976
Mesaphorura tenuisensillata Rusek 1974
Oligaphorura groenlandicus (Tullberg 1876)
Oligaphorura ursi (Fjellberg 1984)
Thalassaphorura duplopunctatus (Strenzke 1954)
Protaphorura macfadyeni (Gisin 1953)
Protaphorura subuliginata Gisin 1956
Tullbergia arctica Wahlgren 1900
Chaetophorura simplex (Gisin 1958)
Neelipleona
_
_
Neelidae
Megalothorax minimus (Willem 1900)
Symphypleona
_
13. april 2011
_
Katiannidae
Sminthurinus aureus (Lubbock 1862)
Sminthurinus concolor (Meinert 1896)
Sminthurididae
Sminthurides inaequalis armatus Bretfeld 2000
Sminthurides malmgreni (Tullberg 1876)
Sphaeridia pumilis (Krausbauer 1898)
Page 40 of 52

Insecta
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Siphonaptera
_
_
Ceratophyllidae
Ceratophyllus vagabundus vagabundus (Boheman 1866)
Mioctenopsylla arctica arctica Rothschild 1922
Anoplura
_
_
Echinophthiriidae
Antarctophthirus tricechi (Boheman 1865)
Echinophthirius horridus (Olfers 1816)
Coleoptera
_
_
Anthicidae
Anthicus flavipes (Panzer 1797)
Byrrhidae
Simplocaria metallica (Sturm 1807)
Carabidae
Amara quenseli (Schönherr 1806)
Chrysomelidae
Phytodecta ?
Coccinellidae
Coccinella septempunctata L 1758
Cryptophagidae
Atomaria atricapilla Stephens 1830
Atomaria lewisi Reitter 1877
Curculionidae
Orchestris saliceti Fabricus
Isochnus flagellum (Ericson 1902)
Dermestidae
Reesa vespulae (Milliron, 1939)
Dytiscidae
Hydroporus striola Gyllenhal 1827
Lathridiidae
Latrididius minutus (L. 1767)
Silvanidae
Oryzaephilus mercator Fauvel
Staphylinidae
Atheta graminicola (Gravenhorst 1806)
Boreophilia subplana (J. Sahlberg 1880)
Eucnecosum brachypterum (Gravenhorst 1802)
Micralymma marinum (Strøm 1783)
13. april 2011
Page 41 of 52

Dermaptera
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Olophrum boreale (Paykull 1792)
Omalium caesum Gravenhorst 1806
Omalium septentrioneis Thompson 1857
_
_
Forficulidae
Forficula auricularia L 1758
Dictyoptera
_
_
Blattellidae
Diptera
13. april 2011
Blattella germanica (L. 1767)
Brachycera
_
Anthomyiidae
Fucellia focorum (Fallén 1819)
Zaphne frontata (Zetterstedt 1838)
Calliphoridae
Calliphora vicina Robineau-Desvoidy 1830
Cynomyia mortuorum (L. 1761)
Protophormia atriceps (Zetterstedt, 1845)
Protophormia terranovae (Robineau-Desvoidy 1830)
Coelopidae
Coelopa eximia Stenhammer 1854
Fucomyia frigida Fallén 1805
Drosophilidae
Drosophila funebris (Fabricus 1787)
Empididae
Rhamphomyia caudata Zetterstedt 1838
Heleomyzidae
Heleomyza modesta (Meigen, 1878)
Heleomyza serrata (L. 1758)
Heleomyza borealis (Boheman 1865)
Helomyzidae
Leria geniculata Zetterstedt 1847
Neolaria prominens (Becker 1897)
Neoleria septentrionalis (Collin 1923)
Muscidae
Spilogona brunneifrons Ringdahl 1931
Spilogona denudata (Holmgren 1869)
Spilogona dorsata (Zetterstedt 1845)
Spilogona megastoma (Boheman 1866)
Spilogona quinquelineata (Zetterstedt 1838)
Spilogona triangulifera (Zetterstedt 1838)
Spilogona varsviensis Schnabl in Schnabl and Dziedzicki 1911
Piophilidae
Page 42 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Parapiophila vulgaris (Fallén 1820)
Scatophagidae
Allomyella frigida (Holmgren 1883)
Scatophaga furcata (Say 1823)
Scatophaga litorea (Fallén 1819)
Scatophaga varipes (Holmgren 1883)
Sphaeroceidae
Copromyza (Borborus) uncinatus (Duda 1923)
Norrbomia fumipennis (Stenhammar 1855)
Syrphidae
Syrphus ribesii (L. 1758)
Syrphus torvus Osten-Sacken 1875
Parasyrphus tarsatus (Zetterstedt 1838)
Tachinidae
Tachina glacialis Boheman
Nematocera
_
Culicidae
Aedes (Ochlerotatus) nigripes (Zetterstedt 1838)
Mycetophilidae
Allodida sp.
Boletina maculata Holmgren 1870
Coelosia tenella (Zetterstedt 1852)
Exechia frigida (Boheman 1865)
Phronia egregia Dziedzicki1889
Rhymosia sp.
Scatopsidae
Scatopse notata (L. 1758)
Sciaridae
Bradysia nervosa (Meigen 1818)
Bradysia praecox (Meigen 1818)
Camptochaeta consimilis (Holmgren 1869)
Camptocheata delicata (Frey 1948)
Corynoptera röderi Lengersdorf 1931
Lycoriella (Hemineurina) abbrevinervis (Holmgren 1869)
Lycoriella (Hemineurina) cochleata (Rübsaamen 1898)
Lycoriella (Hemineurina) modesta (Staeger 1840)
Lycoriella postconspicua Mohrig, 1985
Lycoriella (Hemineurina) vitticollis (Holmgren 1883)
Lycoriella (Lycoriella) parva (Holmgren 1869)
Schwenckfeldina tridentata (Rübsaamen 1898)
Simulidae
Prosimulium ursinum (Edwards 1935)
Trichoceridae
Trichocera borealis Lackschewitz 1934
Trichocera hiemalis (De Geer 1760)
Trichocera lutea (Becher 1886)
Trichocera maculipennis Meigen 1818
Trichocera parva Meigen 1804
Trichocera saltator (Harris 1776)
Page 43 of 52

13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Chironomidae
Chironominae
Chironomus (Chironomus) hyperboreus Staeger 1841
Chironomus (Chironomus) lugubris Zetterstedt 1850
Sergentia coracina (Zetterstedt 1850)
Micropsectra insignilobus Kieffer 1924
Micropsectra logani Johannsen 1928
Micropsectra radialis Goetghebuer 1939
Paratanytarsus austriacus (Kieffer 1924)
Tanytarsus heliomesonyctios Langton 1999
Diamesiinae
Diamesa aberrata Lundbeck 1889
Diamesa arctica (Boheman 1865)
Diamesa arctica sensu Willassen and Velle (unpubl.)
Diamesa bertrami Edwards 1935
Diamesa bohemani Goetghebuer 1932
Diamesa hyperborea Holmgren 1869
Diamesa incallida (Walker)
Diamesa lindrothi Goetghebuer 1931
Diamesa lundstroemi Kieffer 1911
Orthocladiinae
Camptocladius stercorarius (De Geer 1776)
Chaetocladius dissipatus (Edwards),
Chaetocladius dentiforceps (Edwards)
Chaetocladius holmgreni (Jabcobson 1898)
Chaetocladius laminatus Brundin 1947
Chaetocladius perennis (Meigen 1830)
Cricotopus (Cricotopus) ephippium (Zetterstedt 1838)
Cricotopus (Cricotopus) lestralis (Edwards 1924)
Cricotopus (Cricotopus) pilosellus Brund 1956
Cricotopus (Cricotopus) polaris Keiffer 1926
Cricotopus (Cricotopus) tibialis (Meigen 1804)
Cricotopus (Cricotopus) villosus Hirvenoja 1973
Cricotopus (Isocladius) glacialis Edwards 1922
Heterotrissocladius subpilosus (Kieffer 1911)
Hydrobaenus conformis (Holmgren 1869)
Limnophyes brachytomus (Kieffer 1922)
Limnophyes edwardsi Sæther 1990
Limnophyes eltoni (Edwards 1921)
Limnopheys lapponicus Goetghebuer 1939
Limnophyes pumilio (Holmgren 1869)
Metriocnemus cataractarum Kieffer 1919
Metriocnemus eurynotus (Holmgren 1883)
Metriocnemus obscuripes (Holmgren 1869)
Metriocnemus picipes (Meigen 1818)
Metriocnemus similis Kieffer 1922
Metriocnemus ursinus (Holmgren 1869)
Oliveridia tricornis (Oliver 1976)
Orthocladius (Eudactylocladius) gelidus Kieffer 1922
Orthocladius (Euorthocladius) telochaetus Langton 1985
Orthocladius (Euorthocladius) thienemanni Keiffer
Orthocladius (Orthocladius) decoratus (Holmgren 1869)
Page 44 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Orthocladius (Orthocladius) knuthi Soponis 1977
Orthocladius (Orthocladius) obumbratus Johannsen 1905
Orthocladius (Orthocladius) rhyacobius Kieffer 1911
Orthocladius (Orthocladius) trigonolabis Edwards 1924
Orthocladius (Pogonocladius) consobrinus (Holmgren 1869)
Paraphaenocladius brevinervis (Holmgren 1869)
Psectrocladius (Monopsectrocladius) calcaratus (Edwards 1929)
Psectrocladius (Psectrocladius) barbimanus Edwards 1929
Psectrocladius (Psectrocladius) borealis Kieffer 1919
Psectrocladius (Psectrocladius) limbatellus (Holmgren 1869)
Psectrocladius (Psectrocladius) octomaculatus Wülker 1956
Psectrocladius (Psectrocladius) oxyura Langton 1985
Psectrocladius (Psectrocladius) ventricosus Kieffer 1925
Psectrocladius psilopterus (Keiffer 1906)
Pseudosmittia oxoniana (Edwards 1922)
Smittia brevipennis (Boheman 1856)
Smittia extrema (Holmgren 1869)
Smittia flexinervis (Kieffer 1911)
Smittia lasiops (Malloch 1915)
Zalutschia tatrica (Pagast 1935)
Chironomus anthracinus - type
Tanypodinae
Arctopelopia barbitarsis (Zetterstedt 1850)
Arctopelopia melanosoma (Goetghebuer 1933)
Procladius crassinervis (Zetterstedt 1838)
Ephemeroptera
_
_
Leptophlebiidae
Leptophlebia vespertina (L. 1758)
Hemiptera
_
_
Aphidoidea
Sitobion calvulum (Ossiannilsson 1958)
Acyrthosiphon svalbardicum Heikinheimo 1968
Cinara abieticola (Cholodkovsky 1899)
Pemphigus groenlandicus (Rübsamer 1898)
Hymenoptera
13. april 2011
Apocrita
_
Braconidae
Biosteres sp.
Diaertiella rapae (M'Intosh 1855)
Dendrocerus bifoveatus (Keiffer 1907)
Ichneutes hyperboreus Holmgren 1869
Trioxys sp.
Ceraphronidae
Conostigmus spetsbergensis (Holmgren 1869)
Page 45 of 52

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Lagynodes pallidus Boheman 1832
Cynipidae
Alloxysta sp.
Ichneumonidae
Aclastus borealis (Boheman 1866)
Aclastus gracilis (Thomson 1884)
Aptesis erratica (Holmgren 1969)
Atractodes bicolor Gravenhorst 1829 var. arcticus Holmgren 1872
Atractodes pusillus (Förester 1876)
Gelis glacialis (Holmgren 1869)
Hypamblys leucopgus (Holmgren 1869)
Mesochorus palanderi Holmgren 1869
Orthocentrus nigricornis Boheman 1866
Stenomacrus groenlandicus Jussila 1996
Stenomacrus nigricornis (Boheman)
Phygadeuon sp.
Plectiscidea hyperborea (Holmgren 1869)
Stenomacrus pedestris (Holmgren 1869)
Stenomacrus validicornis (Boheman 1866)
Synodites lineiger (Thompson 1893)
Tymmophorus obscuripes (Holmgren 1858)
Symphyta
_
Tenthredinidae
Amauronematus hyperboreus (Thomson 1875)
Amauronematus alberich Benson 1934
Amauronematus villosus (Thomson 1862)
Amauronematus rubiginosus Hellén 1951
Euura sp.?
Pontopristia amentorum (Förster 1854)
Pristiphora frigida (Boheman 1865)
Pristiphora stavdingeri (Ruthe 1859)
Lepidoptera
_
_
Noctuidae
Apamea zeta Haliday 1836
Syngrapha interrogationsis (L. 1758)
Nymphalidae
Nymphalis antiopa (L. 1758)
Vanessa cardui (L. 1758)
Oecophoridae
Hofmannospila pseudospretella (Stainton 1849)
Papilionidae
Parnassius apollo (L.1758)
Pieridae
Pieris napi (L.1758)
Margaritia sticticalis (L 1761)
Pyralidae
Pempelia dilutella Hubner 1796
13. april 2011
Page 46 of 52

Mallophaga
_
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Amblycera
Ischnocera
Thysanoptera
_
13. april 2011
_
Thripidae
Plutella xylostella (L. 1758)
Plutella polaris Zeller 1880
Pyla fusca Haworth 1811
Ephestia kuehniella Zeller 1879
Ancistrona vagelli (Fabricus 1787)
Austromenopon brevifimbriatum (Piaget 1880)
Austromenopon corporosum (Kellogg and Kuwana 1901)
Austromenopon fuscofasciatum (Piaget 1880)
Austromenopon lutescens (Burmeister 1838)
Austromenopon merguli Timmermann 1954
Austromenopon uriae Timmermann 1954
Trinoton anserinum (Fabricus 1805)
Anaticola anseris (L.
Anaticola rubrumaculatus (Rudow 1869)
Carduiceps zonarius (Nitzsch 1866)
Cummingsiella aetherea klatti (Timmermann 1954)
Cummingsiella hiaticulae (Fabricus 1780)
Cummingsiella normifer (Grube 1851)
Cummingsiella obliqua aquilonis Timmermann 1974
Cummingsiella ornatus (Grube 1851)
Cummingsiella phalaropi (Denny 1842)
Cummingsiella strepsilaris (Denny 1842)
Gonoides lagopi (L. 1758)
Lagopoecus affinis (Children 1836)
Lunaceps neiris Timmermann 1954
Lunaceps phaeopi (Denny 1842)
Ornithobius hexophtalmus (Nitzsch 1871)
Perineus nigrolimbatus (Giebel 1874)
Saemundssonia calva (Kellogg 1896)
Saemundssonia cephalus (Denny 1842)
Saemundssonia grylle (Fabricus 1780)
Saemundssonia inexpectata Timmermann 1951
Saemundssonia lari (Fabricus 1780)
Saemundssonia lockleyi Clay 1949
Saemundssonia merguli (Denny 1842)
Saemundssonia occidentalis (Kellogg 1896)
Saemundssonia scolopacisphaeopodis (Schrank 1803)
Saemundssonia tringae tringae (Fabricus 1780)
Saemundssonia tringae variabilis (Denny 1842)
Saemundssonia tringae lobata Martens 1974
Aptinothrips rufus Haliday 1836
Page 47 of 52

Trichoptera
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
Limnophilidae
Apatania zonella Zetterstedt 1840
Page 48 of 52

Crustacea
Branchiopoda
Cladocera
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Bosminidae
Bosmina longirostris Müller 1785
Eubosmina longispina (Müller 1785)
Chydoridae
Acroperus harpae (Baird 1835)
Alona guttata Sars 1862
Chydorus sphaericus (Müller 1785)
Daphniidae
Daphnia pulex
Macrothricidae
Macrothrix hirsuticornis Norman and Brady 1867
Macrothrix hirsuticornis arctica (Sars 1890)
Ctenopoda
_
_
Sididae
Sida crystallina (Müller 1776)
Notostraca
_
13. april 2011
_
Triopsidae
Lepidurus arcticus (Pallas 1793)
Page 49 of 52

Copepoda
Calanoida
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Centropagidae
Limnocalanus macrurus Sars 1863
Temoridae
Eurytemora raboti Richard 1897
Cyclopoida
_
_
Cyclopidae
Acanthocyclops gigas (Claus 1857)
Cyclops abyssorum Sars 1863
Cyclops vicinus Ulianine 1875
Diacyclops crassicaudis (Sars 1863)
Harpacticoida
_
_
Canthocamptidae
Maraenobiotus brucei brucei (Richard 1898)
Harpacticidae
Harpacticus uniremis Kröyer 1845
Tachidiidae
Tachidius discipes Giesbrecht 1882
Siphonostomatoida
_
_
Lernaeopodidae
Salmincola edwardsii Olsson 1869
Salmincola carpionis (Krøyer 1837)
13. april 2011
Page 50 of 52

Malacostraca
Amphipoda
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
_
Gammaridae
Gammaracanthus lacustris (Sars 1867)
Gammaracanthus loricatus (Sars 1896)
Mysidacea
_
_
Mysidae
Mysis relicta Loven 1861
Ostracoda
Podocopida
13. april 2011
_
_
Candonidae
Candona candida Müller 1776
Eucandona rectangulata (Alm 1914)
Cyprididae
Cyclocypris ovum (Jurine 1820)
Eucypris arctica Olofsson 1918
Eucypris glacialis (Sars 1890)
Heterocypris fretensis (Brady and Robertson 1870)
Tonnacypris glacialis (G.O.Sars, 1890)
Cytherideidae
Cytherissa lacustris (Sars 1863)
Limnocytheridae
Limnocythere inopinata (Baird 1843)
Page 51 of 52

To be classified
_
_
_
13. april 2011
The Terrestrial and Freshwater Invertebrate Fauna of Svalbard:
_
Cestoda
Nadejdolepsis nitida
Aploparakis leonovi
Aploparakis crassipenis
Page 52 of 52