Checklist of the Terrestrial and freshwater invertebrate fauna - Unis

The terrestrial and freshwater invertebrate fauna of 

Svalbard: an inventory and bibliography 

26 September 2012

Notes: This is not the definitative inventory! 

This list is based on a collection of over 500 articles concerning the terrestrial invertebrate fauna of 

Svalbard. Nonetheless it should be not be considered as an accurate overview of the complete 

invertebrate fauna of the archipelago. The terrestrial invertebrate fauna is largely only known from 

two localities on Svalbard, Ny-Ålesund (Kongsfjord) and the Longyearbyen region. Little is known 

about the fauna of the east coast or the interior. In addition there are large numbers of doubtful 

identifications and confusion in the taxonomy. 

Wherever possible synonyms occuring in the Svalbard literature are presented in smaller font. 

For further details and comments on the species refer to:- 

Coulson, S.J. 2007 The terrestrial and freshwater invertebrate fauna of the High Arctic 

archipelago of Svalbard. Zootaxa 1448; 41-58. 

Coulson, S.J. Refseth, D. (2004) The terrestrial and freshwater invertebrate fauna of Svalbard 

(and Jan Mayen), pp57-122 in A catalogue of the terrestrial and marine animals of Svalbard 

(eds. Prestrud, P. Strøm H. Goldman H.). Skrifter 201. Norwegian Polar Institute, Tromsø. 

Steve Coulson 

26 September 2012 

Cover map M. Daase; photograph S. Coulson, Tunebreen. 

26 September 2012

The Terrestrial and Freshwater Invertebrate Fauna of Svalbard: 

Sarcomastigophora 

Heliozoea 

_ 

_ 

_ 

Actinophryridae 

Actinophrys sol Ehrenberg 

Clathrulinidae 

Clathrulina elegans Cienk 

Zoomastigophorea 

Cercomonadida 

_ 

_ 


Cercomonadidae 

Cercomonas gigantica Myl'nikov 460 

Cercomonas kiaerdammane Myl'nikov 460 

3

Rhizopoda 

Filosea 

Gromiida 

_ 

_ 



Cyphoderiidae 

Cyphoderia ampulla Leidy 22 

Cyphoderia perlucidus Beyens and Charedez 21 22 

Euglyphidae 

Assulina muscorum Greeff 14 22 24 25 244 260 

Assulina muscorum var. denticulata Chardez 22 

Assulina muscorum var. stenostoma Schönborn 22 260 

Assulina seminulum (Ehrenberg) Leidy 14 260 319 

Euglypha acanthophora Perty 22 25 

4



Euglypha alveolata Dujardin 319 

Euglypha ciliata Ehrenberg 14 22 319 

Euglypha ciliata var. glabra Wailes 260 

Euglypha cristata Leidy 319 

Euglypha compressa Carter 22 260 

Euglypha filifera Penard 22 

Euglypha laevis (Ehrenberg) Perty 14 22 244 260 

Euglypha laevis var. lanceolata Playfair 22 

Euglypha minor Chardez 22 

Euglypha polylepsis Bonnet and Thomas 22 

Euglypha rotunda Wailes and Penard 14 21 22 24 25 260 

5



Euglypha rotunda var. dorsalis Decloitre 14 

Euglypha rotunda var. glabra Wailes 14 

Euglypha rotunda var. minor Wailes 14 

Euglypha strigosa Leidy 22 24 25 

Euglypha strigosa f. glabra Wailes 14 22 24 25 244 260 

Euglypha strigosa var. muscorum Wailes 22 24 

Euglypha tuberculata Dujardin 14 22 25 420 421 

Euglypha tuberculata var. subcylindrica (Playfair) Decloitre 14 

Tracheleuglypha dentata (Vejdovsky) Deflandre 244 

Valkanovia delicatula Valkanov 14 

Gromiidae 

6



Gromia sp. 319 

Nadinella tenella Penard 244 

Pseudodifflugia globulosa Stepanek 244 

Pseudodifflugia gracilis Schlumberger 22 244 

Pseudodifflugia gracilis var. terricola Bonnet and Thomas 244 

Pseudodifflugia senartensis Couteaux 244 

Paulinellidae 

Paulinella chromatophora Lauterborn 22 

Trinematidae 

Corythion aerophila Decoitre 24 

Corythion delamarei Bonnet and Thomas 14 

Corythion dubium Taranek 14 22 24 260 

7



Corythion dubium var. aerophila Decloitre 14 24 

Corythion dubium f. minima Chardez 22 24 

Corythion dubium var. orbicularis Penard 22 24 

Corythion dubium var. terricola Schönborn 244 

Corythion pulchellum Penard 14 

Trinema complanatum Penard 22 25 

Trinema complanatum var. globulosa Chardez 22 

Trinema enchelys (Ehrenberg) Leidy 14 22 24 25 260 319 

Trinema lineare Penard 14 21 22 25 244 260 

Trinema lineare f. minuscula Chardez 14 22 

Trinema lineare f. truncatum Chardez 22 

8



Trinema lineare var. longa Korganova 22 

Trinema navicularis Decloitre 22 

Trinema penardi Thomas and Chardez 22 

9

Lobosea 

Amoebida 

_ 

_ 


Amoebidae 

Amoeba radiosa Ehrenberg 

Amoeba verrucosa Ehrenberg 

Arcellinida 

_ 

_ 


Arcellidae 

Arcella atrocera Leidy 319 

Arcella arenaria Greeff 22 25 260 

Arcella arenaria f. compressa Chardez 22 24 

Arcella arenaria var. sphagnicola Deflandre 22 

Arcella bathystoma Deflandre 22 

10



Arcella discoides Ehrenberg 22 244 

Arcella ovaliformis Chardez and Beyens 22 61 

Arcella rotundata Playfair 22 24 25 

Arcella rotundata var. aplanata Deflandre 22 

Arcella rotundata var. undulata Stepanek 22 25 

Arcella vulgaris Ehrenberg 14 420 421 

Chlamydophrys stercorea Cienk 319 

Cochliopodium erinaceum Penard 14 

Cochliopodium obscurum Penard 22 25 

Cochliopodium opalinum Penard 22 25 

Difflugiella horrida Schönborn 14 

11



Difflugiella oviformis (Penard) Bonnet and Thomas 14 

Difflugiella pusilla (Playfair) Grospietsch 

Microcorycia flava Greef 22 25 

Microcorycia husvikensis Beyens and Chardez 23 

Parmulina louisi Chardez and Beyens 22 

Pseudochlamys patella C. and L. 319 

Pyxidicua cymbalum Penard 25 

Pyxidicula ornata Bartos 14 

Centropyxidae 

Centropyxis aculeata Stein 22 25 244 

Centropyxis aculeata f. compressa 244 

12



Centropyxis acuminata Couteaux and Chardez 22 

Centropyxis aerophila Deflandre 14 21 22 24 25 244 260 

Centropyxis aerophila var. constricta Decloitre 22 

Centropyxis aerophila var. deflandrei Decloitre 22 

Centropyxis aerophila var. sphagnicola Deflandre 14 22 24 25 244 260 

Centropyxis aerophila var. minuta Chardez 24 25 

Centropyxis cassis (Wallach) Deflandre 14 22 

Centropyxis cassis var minima van Oye 22 420 421 

Centropyxis constricta (Ehrenburg) Penard 14 

Centropyxis cordiformis Couteaux 22 

Centropyxis delicatula Penard 22 

13



Centropyxis delicatul var. maior Jung 22 

Centropyxis ecornis Leidy 22 

Centropyxis ecornis var. deflandrei Thomas 14 

Centropyxis gasparella 516 

Centropyxis gasparella var. corniculata Beyens and Chardez 516 

Centropyxis gibba var. gibbosa Godeanu 21 22 23 

Centropyxis globulosa Bonnet and Thomas 22 

Centropyxis laevigata var. maior Jung 22 

Centropyxis lata Jung 22 

Centropyxis minima Deflandre 22 

Centropyxis minuta Deflandre 22 

14



Centropyxis orbicularis Deflandre 14 22 

Centropyxis plagiostoma Bonnet and Thomas 21 22 260 

Centropyxis platystoma (Penard) Deflandre 14 22 244 260 420 421 

Centropyxis pontigulasiformis Beyens and Chardez 21 22 244 

Centropyxis sylvatica Thomas 21 22 24 244 

Centropyxis sylvatica var. minor Bonnet and Thomas 22 

Centropyxis ventricosa Bartos 22 

Cryptodifflugiidae 

Cryptodifflugia compressa f. angustioris Tarnogradsky 22 24 

Difflugiidae 

Difflugia bacillifera Penard 22 

Difflugia bryophila Jung 21 22 24 244 

15



Difflugia compressa var. minima Schönborn 244 

Difflugia constricta Ehrenberg 319 

Difflugia dujardini Chardez 21 22 

Difflugia elegans Penard 244 

Difflugia elegans var. parva Chardez 244 

Difflugia glans Penard 22 24 

Difflugia globularis Leidy 21 22 

Difflugia globulosa Dujardin 22 24 260 319 

Difflugia globulus Hopkinson 21 22 

Difflugia lucida Penard 1902 14 21 22 25 247 

Difflugia lacustris Jung 22 25 

16



Difflugia lata Jung 21 22 

Difflugia linearis (Penard) Gauthier-Lievre and Thomas 22 25 

Difflugia minuta Rampi 21 22 

Difflugia paulii Ogden 21 22 

Difflugia penardi Hopkinson 21 22 244 

Difflugia pulex Penard 14 22 25 247 

Difflugia pulex var. cuneata Playfair 22 

Difflugia pyriformis Perty 244 319 

Difflugia tuberculata Playfair 24 

Difflugia tuberculata var. sphaerica Playfair 22 

Hyalospheniidae 

17



Heleopera petricola Leidy 22 24 319 

Heleopera petricola var amethystea Penard 22 25 

Heleopera sylvatica Penard 22 

Nebela bohemica Taranek 14 22 

Nebela collaris Leidy 22 25 260 244 258 319 

Nebela dentistoma Penard 14 22 25 

Nebela dentistoma var. oblonga Gaut.-L and Thomas 21 22 

Nebela flabellum Leidy 22 

Nebela galeata Penard 22 

Nebela lageniformis Penard 22 25 244 

Nebela militaris Penard 22 24 25 

18



Nebela minor Penard 22 24 

Nebela nobilis Deflandre 22 25 

Nebela parvula Penard 14 

Nebela penardiana Deflandre 21 22 25 

Nebela tincta (Leidy) Awerintzew 14 21 22 24 25 258 420 421 

Nebela tincta f. stenostoma Jung 22 

Nebela tincta var. rotunda Penard 22 

Nebela tubulosa Penard 21 22 

Nebela wailesi Deflandre 21 22 

Lesquereusiidae 

Netzelia labeosa Beyens and Chardez 23 

Microcoryciidae 

19



Microchlamys patella (Claparede and Lachmann) Cockerell 

Microchlamys patella var. arctica Penard 22 25 

Nebelidae 

Schönbornia smithi Beyens and Chardez 23 

Paraquadrulidae 

Paraquadrula discoides (Penard) 260 

Paraquadrula globulosa Ferlan 21 22 

Paraquadrula irregularis Deflandre 21 22 25 

Paraquadrula madarica Valkanov 21 22 

Paraquadrula ogdeni Byens and Chardez 23 

Paraquadrula rotunda Schönborn 22 260 

Phryganelidae 

Phryganella acropodia Hopkinson 14 22 24 25 244 258 260 

20



Phryganella acropodia var. penardi Decloitre 14 22 260 

Phryganella microps Valkanov 14 

Phryganella nidilus Penard 22 

Phryganella paradoxa var. alta Bonnet and Thomas 260 

Phryganella paradoxa Penard 22 

Plagiopyxidae 

Plagiopyxis bathystoma Bonnet 22 

Plagiopyxis callida Penard 14 21 22 25 

Plagiopyxis callida var. grandis Thomas 22 244 

Plagiopyxis declivis Thomas 14 22 

Plagiopyxis labiata Penard 14 21 22 244 

21



Plagiopyxis minuta Bonnet 14 260 

Plagiopyxis oblonga (Bonnet and Thomas) Bonnet 14 

Plagiopyxis penardi Thomas 22 25 

Trigonopyxidae 

Cyclopyxis ambigua Bonnet and Thomas 22 

Cyclopyxis ampulla Leidy 

Cyclopyxis arcelloides var. gibbosa van Oye 22 

Cyclopyxis arcelloides var. minima van Oye 22 

Cyclopyxis eurystoma Deflandre 22 

Cyclopyxis eurystoma var. stenostoma Decloitre 22 

Cyclopyxis kahli Deflandre 

Cyclopyxis leclerqi Beyens and Chardez 21 25 

22



Cyclopyxis perlucidus Beyens and Chardez 

Cyclopyxis plana var. microstoma Schönborn 14 260 

Cyclopyxis puteus Thomas 22 

Trigonopyxis arcula Penard 14 22 420 421 

Vahlkampfiidae 

Naegleria polaris De Jonckheere 2006 386 

Naegleria neopolaris De Jonckheere 2006 386 

Naegleria arctica De Jonckheere 2006 386 

Naegleria spitzbergensis De Jonckheere 2009 386 

23

Ciliophora 


Kinetofragminophorea 

Colpodida 

_ 

_ 

Cyrtolophosis mucicola Stokes 1885 511 

Platyophrya vorax Kahl 1926 511 

Rostrophryides sp. nov. 511 

Colpodidae 

Colpoda cucullus (Müller 1773) 319 

Cyrtophorida 

_ 

_ 


Odontochlamys alpestris Foissner 1981 511 

Chilodonella uncinata (Ehrenberg 1835) 511 

24

Nassulida 

_ 

_ 


Furgasonia cf. trichocystis 511 

Kreyella minuta Foissner 1979 511 

Leptopharynx costatus Mermod 1914 511 

Pleurostomatida 

_ 

_ 

Litonotus lamella (Müller 1773) 511 

Askenasia sp. 440 

Prostomatida 

_ 

_ 


Dileptus margaritifer (Ehrenberg 1833) 511 

25


Fuscheria cf. nodosa 511 

Phialina vertens (Stokes, 1885) 511 

Protospathidium sp. 511 

Trachelophyllum apiculatum (Perty 1852) 511 

Lacrymaria filiformis 511 

Monodinium sp. 469 

Urotrichidae 

Urotricha agilis (Stokes 1886) 511 

Urotricha cf. armata 511 

Suctorida 

_ 

_ 


Sphaerophrya terricola Foissner 1986 511 

26

Synhymeniida 

_ 

_ 



Nassulopsis cf. elegans 511 

27


Oligohymenophorea 

Hymenostomatida 

_ 

_ 

Frontonia acuminata (Ehrenberg 1833) 511 

Frontonia angusta 511 

Tetrahymena rostrata 511 

Cinetochilidae 

Cinetochilum margaritaceum (Perty 1852) 511 

Oxytrichidae 

Paramecium arcticum Doroszewski 511 

Paramecium aurelia complex 511 

Peritrichida 

_ 

_ 


28


Opercularia coarctata Claparede & Lachmann 1854 511 

Pseudovorticella monilata (Tatem 1870) 511 

Mobilidae 

Trichodina tentaculata 511 

Sessilidea 

Ophrydium versatile 511 

Vorticella aquadulcis complex 511 

Vorticella infusionum complex 511 

Vorticella microstoma 511 

Scuticociliatida 

_ 

_ 


Cyclidium glaucoma Müller 1773 511 

29



Sathrophilus muscorum (Kahl 1931) 511 

Protocyclidium muscicola (Kahl 1931) 511 

30


Polyhymenophorea 

Heterotrichida 

_ 

_ 

Blepharisma hyalinum Perty 1849 511 

Bryometopus pseudochilodon Kahl 1932 511 

Climacostomum virens (Ehrenberg 1838) 511 

Hypotrichida 

_ 

_ 


Aspidisca cicada (Müller 1786) 511 

Oxytricha balladyna Song & Wilbert 1989 511 

Oxytricha setigera Stokes 1891 511 

Oxytricha similis Engelmann 1862 511 

31


Sterkiella histriomuscorum Foissner, Blatterer, Berger & 

Kohmann 1991 

Stichotricha aculeata Wrzesniowski 1866 511 

Stylonychia mytilus complex 511 

Uroleptus gallina (Müller 1786) 511 

Oxytrichidae 

Stylonychia pustulata (Müller 1786) 

Oligotrichida 

_ 

_ 


Halteria grandinella (Müller 1773) 440 

Limnostrombidium viride (Stein 1859) 440 

Rimostrombidium sp. 

Stombidium sp 440 469 

511 

32

Rhizopoda 

Lobosea 

Arcellinida 

_ 

_ 



Arcellidae 

Arcella cinctus 420 421 

Centropyxidae 

Cyclopyxis arcelloides van Oye 420 421 

33

Apicomplexa 

Sporozoea 

Coccidia 

_ 

_ 



Sporozoea 

Sarcocystis sp. 

Toxoplasma gondii Nicolle & Manceaux 1909 530 531 544 546 547 549 550 

Isospora plectrophenaxia Dolnik and Loonen 2007 517 

34

Rotifera 

Digononta 

Bdelloidea 

_ 

_ 



Adinetidae 

Adineta barabata Janson 1893 46 305 319 548 

Adineta gracilis Janson 1893 46 305 319 548 

Adineta vaga (Davis 1873) 46 305 319 548 

Adineta vaga minor Bryce 1893 548 

Habrotrochidae 

Habrotrocha angusticollis (Murray 1905) 319 548 

Habrotrocha aspera (Bryce 1892) 

Callidina aspera Bryce 

Habrotrocha bidens (Gosse 1851) 

Habrotrocha milnei Bryce 

319 548 

45 46 319 548 

35



Habrotrocha constricta (Dujardin 1841) 

Callidina constricta Dujardin 

Habrotrocha elegans (Milne 1886) 

Callidina venustra Bryce 

45 46 305 319 548 

319 548 

Habrotrocha insignis Bryce 1915 45 46 305 319 548 

Habrotrocha lata lata (Bryce 1892) 

Callidina lata Bryce 

Habrotrocha ligula ligula Bryce 1913 548 

Habrotrocha novemdens De Koning 1947 548 

Habrotrocha pulchra (Murray 1905) 548 

Habrotrocha pusilla (Bryce 1893) 

Callidina pusilla Bryce 

319 548 

Habrotrocha pussila textrix (Bryce 1897) 319 548 

Habrotrocha roeperi (Milne 1889) 548 

319 

36



Habrotrocha rosa Donner 1949 548 

Philodinidae 

Ceratrocha cornigera (Bryce 1893) 46 319 548 

Didymodactylos carnosus Milne 1916 548 

Macrotrachela aculeata Milne 1886 46 319 548 

Macrotrachela concinna (Bryce 1912) 46 319 548 

Macrotrachela ehrenbergii (Janson 1893) 46 305 319 548 

Macrotrachela insolita de Koning 1947 87 548 

Macrotrachela habita (Bryce 1894) 

Callidina habita Bryce 

Macrotrachela kakkosoma (Schulte 1954) 548 

46 305 319 548 

Macrotrachela multispinosa multispinosa Thomson 1892 305 319 548 

37



Macrotrachela musculosa (Milne 1886) 

Callidina musculosa Milne 

Macrotrachela nana nana (Bryce 1912) 548 

Macrotrachela papillosa (Thomson 1892) 46 319 548 

Macrotrachela plicata hirundinella (Murray 1905) 548 

Macrotrachela plicata plicata (Bryce 1892) 

Callidina plicata Bryce Macrotrachela plicata var hirudinella 

(Murray) 

191 193 194 242 243 319 548 

46 319 

Macrotrachela quadricornifera Milne 1886 46 319 548 

Mniobia bredensis De Koning 1947 548 

Mniobia incrassata (Murray 1905) 319 548 

Mniobia magna (Plate 1889) 548 

Mniobia russeola (Zelinka 1891) 

Callidina russeola Zelenka 

45 46 319 548 

38



Mniobia symbiotica (Zelinka 18886) 548 

Mniobia tentans Donner 1949 548 

Mniobia tetraodon (Ehrenberg 1848) 

Callidina tetraodon Ehrenberg 

Mniobia vargai Donner 1962 548 

Mniobia variabilis Donner 1949 548 

Philodinia acuticornis Murray 1902 548 

Philodinia acuticornis odiosa Milne 1916 

Phiolodinia acuticornis Murray 

319 548 

46 87 319 548 

Philodinia brevipes Murray 1902 46 319 548 

Philodina erhthrophthama Ehrenberg 1832 319 

Philodina flaviceps Bryce 1906 548 

39



Philodinia nemoralis Bryce 1903 46 319 548 

Philodinia proterva Milne 1916 548 

Philodinia rugosa rugosa Bryce 1903 319 548 

Pleuretra alpium (Ehrenberg 1853) 

Callidina alpium Ehrenberg 

46 319 548 

Pleuretra brycei (Weber 1898) 45 46 305 319 548 

Pleuretra hystrix Bartos 1950 548 

Rotaria citrina Ehrenberg 1838 319 

Rotaria rotatoria (Pallas 1766) 

Rotaria vulgaris Schrank Rotaria rotatoria (Pallas) 

Rotaria sordida sordida (Western 1893) 

Rotifer sordida (Western) 

Rotaria tardigrada (Ehrenberg 1832) 

Rotifer tardigradus Ehrenberg Rotifer tardus Ehrenberg 

548 

305 319 548 

319 548 

40



Scepantotrocha sp. 548 

41

Monogononta 

Collothecacea 

_ 

_ 


Collothecidae 

Collotheca campanulata (Dobie 1849) 85 90 

Collotheca libera Zacharias 1894 316 

Collotheca ornata cornuta (Dobie 1842) 84 85 172 

Collotheca ornata ornuta (Ehrenberg 1832) 85 88 171 

Flosculariacea 

_ 

_ 

Filiniidae 

Filinia terminalis (Plate 1886) 246 

Flosculariidae 

Ptygura melicerta (Ehrenberg 1832) 319 

Ploimida 

_ 


42

_ 



Brachionidae 

Kellicottia longispina (Kellicott 1879) 

? Notholca longispina 

Keratella cochlearis (Gosse 1851) 9 87 317 

9 173 276 316 317 

Keratella hiemalis (Carlin 1943) 9 10 90 173 246 316 317 559 

Keratella quadrata (Müller 1786) 

Anuraea aculeata 

Keratella testudo (Ehrenberg 1832) 85 

7 8 9 10 319 

Notholca acuminata (Ehrenberg 1832) 9 10 85 319 

Notholca acuminata var. extensa Olofsson 1918 85 

Notholca foliacea (Ehrenberg 1838) 

Argonotholca foliacea (Ehrenberg) Notholca foliacea var. latistyla 

Olofsson 

Notholca labis Gosse 1887 83 

7 9 83 85 90 171 172 246 316 

43



Notholca latistyla (Olofsson 1918) 9 83 85 90 173 

Notholca latistyla f. amreni De Smet and Bafort 1990 84 85 

Notholca marina (Focke 1991) 85 

Notholca striata (Müller) 

Notholca striata var. bipalium 

(Müller) 

Notholca striata f. extensa 

9 10 319 

Notholca squamala (Müller 1786) 9 10 84 85 88 90 171 172 317 559 

Colurellidae 

Colurella adriatica Ehrenberg 1831 

Colurella amblytelus (Gosse 1851) 319 

Colurella colurus (Ehrenberg 1830) 83 85 89 171 172 319 

Colurella geophila Donner 1981 83 

Colurella hindenburgi Steinecke 1917 83 85 90 317 

44



Colurella obtusa (Gosse 1886) 83 85 319 

Colurella uncinata (Müller 1773) 83 85 90 

Lepadella acuminata (Ehrenberg 1834) 

Metopidia acuminata Ehrenberg 

Lepadella minuta (Montet 1918) 83 85 

Lepadella ovalis (Müller 1786) 

Metopidia lapadella 

Lepadella patella (Müller 1786) 

Metopidia oblonga 

Lepadella quadricarinata (Stenroos 1898) 

Metopidia quadricarinata 

Lepadella triba Myers 1934 85 

83 85 90 319 

83 85 173 172 319 

46 83 84 85 88 90 171 172 305 

83 85 319 

Lepadella triptera Ehrenberg 1830 83 316 

Lophocaris oxysternon (Goose 1851) 319 

Dicranophoridae 

45



Albertia naidis Bousfield 1886 83 

Dicranophorus forcipatus (Müller 1786) 83 85 

Encentrum bidentatum (Lie-Pettersen 1906) 86 

Encentrum dieteri De Smet 1995 86 

Encentrum felis (Müller 1773) 

Encentrum ferox (Western) nom. dub 

Encentrum ferox (Western) nom. dub. 

Encentrum grandi Haring and Myers 1928 

Encentrum grandi (Western) nom. 

dub 

Encentrum marinum (Dujardin 1841) 84 85 

Encentrum mucronatum Wulfert 1936 84 87 88 

Encentrum murrayi Bryce 1922 46 319 

319 

62 

46



Encentrum mustela (Milne 1885) 85 

Encentrum permolle (Gosse 1886) 

Dicranophorus permollis permollis 

(Gosse) 

Encentrum uncinatum (Milne 1886) 

Diglena permollis Gosse 

Encentrum uncinatus (Milne) Dicranophorus uncinatus (Milne) 

Itura aurita aurita (Ehrenberg 1930) 

Itura aurita var. intermedia (Wulfert) 

87 308 

Epiphanidae 

Rhinoglena frontalis Ehrenberg 1858 83 85 

84 88 90 171 172 

Euchlanis deflexa Gosse 1851 83 85 319 

Euchlanis dilatata Ehrenberg 1832 9 83 85 171 172 319 

Euchlanis lyra Hudson 1886 89 

Euchlanis meneta Myers 1930 

Euchlanis oropha Gosse 

85 

83 85 316 319 

47



Catyphna brevis Murray 1913 319 

Catyphna rotundata Olofsson 1918 319 

Lecanidae 

Lecane bulla (Goose 1851) 

Monostyla bulla Gosse 

46 319 

Lecane closterocerca (Schmarda 1859) 83 87 

Lecane cornuta (Müller 1786) 

Monostyla cornuta (Müller) 

Lecane flexilis (Gosse 1886) 

Lecane brevis (Murray) 

Lecane ligona (Dunlop 1901) 317 

Lecane lunaris (Ehrenberg 1832) 

Monostyla lunaris (Ehrenberg) 

Lecane pideis (Harring and Myers 1926) 

Lecane piepelsi de Smet and Bafort 

305 319 

83 319 

46 83 171 172 305 319 

85 171 172 

48



Lecane robertsonae Segers 1993 

Lecane rotundata (Olofsson) 

Lecane rotundata (Oloffson 1918) 

Lindiidae 

Lindia cf. torulosa Dujardin 1841 83 

Mytilinidae 

Lophocharis oxysternon (Gosse 1851) 83 85 

Mytilina bicarinata (Perty 1850) 

62 83 171 319 

Mytilina mucronata (Müller 1773) 83 85 90 171 172 319 

Mytilina ventralis var. brevispina (Ehrenberg 1832) 

Mytilina brevispina Ehrenberg 

Notommatidae 

Cephalodella biungulata Wulfert 1937 85 

46 171 172 305 319 

Cephalodella catellina (O.F. Müller 1786) 83 85 90 171 172 

49



Cephalodella evabroedi De Smet 1988 83 85 

Cephalodella glandulosa Koch-Althaus 1962 85 

Cephalodella gibba (Ehrenberg 1832) 

Diaschiza gibba (Ehrenberg) 

Cephalodella gracilis (Ehrenberg 1832) 

Diaschiza gracilis (Ehrenberg) 

Cephalodella hoodi (Goose 1896) 83 85 

Cephalodella intuta Myers 1924 85 90 

Cephalodella megalocephala (Glascott 1893) 85 

Cephalodella misgurnus Wulfert 1937 84 88 

Cephalodella rotunda Wulfert 1937 85 

Cephalodella sterea (Gosse 1887) 84 88 

83 84 85 88 90 171 172 319 

319 

50



Cephalodella ventripes (Dixon-Nuttall 1901) 171 172 

Cephalodella ventripes var. angustior Donner 1949 83 85 

Diaschiza spp. 

? Cephalodella sp. 

Eosphora najas Ehrenberg 1830 83 85 

Notommata cyrtopus Gosse 1886 83 

Notommata glyphura Wulfert 1935 85 

Resticula nyssa Harring and Myers 1924 83 85 

Proalidae 

Bryceella stylata (Milne 1886) 

Squatinella stylata (Milne) Stephanops stylatus Milne 

Bryceella tenella (Bryce 1897) 

Squatinella tenella (Bryce) Stepanops tenullus Bryce 

Proales decipiens (Ehrenberg 1832) 

Furcularia decipiens Ehrenberg 

319 

85 319 

319 

319 

51



Proales fallaciosa Wulfert 1937 85 

Proales sordida Gosse 1886 319 

Wulfertia ornata Donner 1943 85 

Scaridiidae 

Scaridium longicaudum (Müller 1786) 319 

Synchaetidae 

Polyarthra dolichoptera (Idelson 1925) 7 9 10 83 85 90 173 246 316 317 

Polyarthra dolichoptera f. aptera (Hood 1895) 85 

Polyarthra dolichoptera f. proloba (Albertova 1960) 85 

Polyarthra vulgaris Carlin 1943 319 

Synchaeta lakowitziana Ehrenberg 1832 9 10 

Synchaeta lakowitziana arctica De Smet 1988 883 317 

559 

52



Synchaeta littoralis Rousselet 1902 9 10 

Synchaeta truncata von Hofsten 9 10 

Trichoceridae 

Trichocerca bidens (Lucks 1912) 

Diurella bidens Lucks 

90 319 

Trichocerca. cavia (Gosse 1886) 85 171 172 

Trichocerca cristata Harring 319 

Trichocerca intermedia (Stenroos 1898) 85 

Trichocerca longistyla (Olofsson 1918) 85 

Trichocerca rattus (Müller 1776) 

Rattulus carinatus 

83 85 319 

Trichocerca relicta (Donner 1950) 83 85 

Trichocerca uncinata (Voigt 1902) 

Diurella uncinata (Voigt) 

83 85 90 319 

53



Trichocerca weberi (Jennings 1903) 83 85 90 171 172 

Trichotria truncata (Whitelegge) 191 193 242 

Trichotria pocillum (Müller 1776) 83 

54

Gastrotricha 

_ 


Chaetonotoida 

_ 

_ 


Chaetonotidae 

Chaetonotus sp. 

55

Nematoda 

Adenophorea 

Enoplia 

Stichosomida 

_ 

_ 



Capillariidae 

Capillaria sp 506 

Philometridae 

Philonema oncorhynchi (Kuitenen Ekbaum 1933) 184 

Trichinellidae 

Trichinella spiralis (Owen 1935) 

Trichinella nelsnoni 

Trichinella nativa (Britov and Boev 1972) 180 

36 189 190 207 

56

Penetrantia 

Dorylaimida 

_ 

_ 



Dorylaimidae S.L. 

Enchodelus analatus Thorne 1939 215 

Enchodelus conicaudatus Thorne 1939 215 

Enchodelus cf. macrodoroides (Steiner 1914) 215 

Enchodelus macrodorus Thorne 1939 215 

Enchodelus parvus Loof 1971 215 

Epidorylaimus agilis (de Man 1880) Andrassy 1986 

Eudorylaimus agilis Loof Dorylaimus lugdunensis apud 

Steiner nec. de Man 

Epidorylaimus lugdunensis (de Man 1880) 

Eudorylaimus lugdunensis 

Andrassy, 1959 

215 

215 

57



Eudorylaimus alleni Brzeski1962 215 

Eudorylaimus carteri (Bastian 1865) 

Dorylaimus carteri Bastian 

Eudorylaimus centrocercus (de Man 1880) 

Dorylaimus centrocercus de Man 

Eudorylaimus circulifer (Loof 1961) 215 

191 193 194 243 

Eudorylaimus maksymovi (Altherr 1963) 191 193 215 

Eudorylaimus megodon Loof 1971 215 

Eudorylaimus parvus (de Man 1880) 215 

Eudorylaimus subjunctus Loof 1971 215 

Eudorylaimus vanrosseni Loof 1971 215 

Longidorella magna Loof 1971 215 

6 

58


Mesodorylaimus sp. 171 

Mononchidae 

Prionchulis muscorum (Dujardin 1845) 191 193 

Qudsianematidae 

Labronema sp. 191 193 

Tylencholaimidae 

Tylencholaimus proximus Thorne 1939 215 

Enoplida 

_ 

_ 


Tylencholaimus teres Thorne 1939 215 

Alaimidae 

Alaimus arcuatus Thorne 1939 215 

Alaimus depressus Loof 1971 215 

Alaimus parvus Thorne 1939 215 

Prismatolaimidae 

59



Prismatolaimus dolichurus de Man 1880 6 215 

Prismatolaimus intermedius (Buetschli1873) 6 215 

Prismatolaimus primitivus Loof 1971 215 

Prismatolaimus stenolaimoides Loof 1971 215 

Tobrilidae 

Tobrilus sp. 171 191 193 

Tripylidae 

Tripyla cf. infia Brzeski et Winiszewska-Slipinska 1903 171 

Tripyla glomerans Bastian 1865 

Tripyla papillata Butschli 

215 

60

Secernentia 

Ascaridida 

_ 

_ 


Anisakidae 

Anisakis simplex (Rudolphi 1802) 468 

Contracaecum osculatum (Rudolphi 1801) Baylis 1920 468 

Hysterothylacium aduncum (Rudolphi 1802) 506 

Rhabditida 

_ 

_ 


Bunonematidae 

Bunonema reticulatum Richters 1905 215 

Cephalobidae 

Acrobeles ciliatus von Linstow 1877 215 

Acrobeloides bütschlii Gadea 1954 

Cephalobus bütschlii de Man 

228 

61



Acrobeloides enoploides Loof 1971 215 

Acrobeloides tricornis (Thorne 1925) 38 215 

Cervidellus serratus (Thorne 1925) 215 

Cervidellus spitzbergensis Bostrøm 1987 37 

Chiloplacus saccatus Loof 1971 38 215 

Eucephalobus arcticus Loof 1971 38 215 

Heterocephalobus elongatus (de Man 1880) 

Cephalobus elongatus de Man 

Stegelleta similis (Thorne 1925) 

Stegelleta mucronata 

Panagrolaimidae 

Panagrolaimus papillosus Loof 1971 215 

Teratocephalidae 

215 

6 215 

62


Euteratocephalus crassidens Andrassy 1958 

Metateratocephalus crassidens (de 

Man) 

Teratocephalus costatus Andrassy 1958 39 

Teratocephalus decarinus Anderson 1969 215 

6 39 215 

Teratocephalus lirellus Anderson 1969 39 215 

Teratocephalus terrestris (Butschli1873) 6 

Spirurida 

_ 

_ 

Acuriidae 

Paracuaria adunca (Creplin 1846) Anderson and Wong 1981 468 506 

Stegaphorus stellae-polaris (Parona 1901) 360 468 506 

Strongylida 

_ 


_ 

Trichostrongylidea 

63



Marshallagia marshalli m occidentalis (Ransom, 1907) 

Grosspiculaqia occidentalis 

(Ransom 1907) 

54 131 132 134 385 522 586 

Marshallagia marshalli (Ransom 1907) 5 48 50 53 54 131 132 133 134 369 

384 385 430 431 432 439 489 490 522 586 

Nematodirus helvetianus May 1920 5 133 

Ostertagia arctica Mitzkewitzsch 1929 5 133 134 

Ostertagia grühneri Skrjabin 1929 2 3 5 48 50 53 54 131 132 133 

346 369 383 384 430 431 432 439 488 489 

490 383 522 586 

Ostertagia occidentalis Ransom 1907 5 48 50 53 

Paracuaria adunca (Creplin 1846) 257 

Skrjabinagia arctica (Mitzkevich 1929) 

?Skrjabinagia lyrata (Sjøberg 1926). 

48 49 53 54 131 132 134 522 

Teladorsagia circumcincta (Stadelmann 1894) 48 49 53 54 131 132 133 134 522 

64


Teladorsaga davitiani Andreeva and Satubaldini1984 54 131 134 522 

Teladorsagia trifurcata 522 

Tylenchida 

_ 

_ 


Aphelenchinae 

Aphelenchus nivalis Aurivillius 1883 306 228 

Criconematidae 

Criconemoides annulatus Taylor 1936 

Criconemoides hemispaericaudatus 

Wu 

Criconema sp. 215 

Hoplolaimidae 

Helicotylenchus spitsbergensis Loof 1971 215 

Pratylenchidae 

Pratylenchoides crenicauda Winslow 1958 215 

215 

65



Pratylenchoides magnicauda (Thorne 1935) 

Tylenchorhynchus magnicauda 

Filipijev 

Tylenchidae 

Basiria dolichura Loof 1971 215 

Coslenchus costatus (de Man 1921) 

Tylenchus costatus de Man 

Filenchus thornei (Andrassy 1954) 

Tylenchus thornei Andrassy 

Geocenamus arcticus (Mulvey 1969) 

Tylenchorhynchus arcticus Mulvey 

Geocenamus loofi (Siddiqi1979) 

Tetylenchus joctus Thorne apud Loof Merlinius loofi Siddiqi 

Geocenamus microdorus (Geraert 1966) 

Tylenchorhynchus microdorus 

Geraert 

Lelenchus leptosoma (de Man 1880) 

Tylenchus leptosoma de Man 

215 

215 

215 

215 

215 274 

215 

215 

66



Malenchus bryophilus (Steiner 1914) 

Tylenchus bryophilus Steiner 

Nagelus leptus (Allen 1955) 

Tylenchorhynchus leptus Allen 

Tylenchus davainei Bastien 1865 

Anguillulina davainei (Bastian) 

215 

215 

6 215 

67

Torquentia 

Araeolaimida 

_ 

_ 



Plectidae 

Anaplectus cf. grandepapillatus (Ditlevsen 1928) 171 

Anaplectus granulosus de Coninck and Shürmans Stekhoven 

1933 

Plectus granulosus Bastian 

Anaplectus porosus Allen and Noffsinger 1968 191 193 194 215 

Ceratoplectus communis (Bütschli1873) 

Plectus communis Bütschli 

Ereptonema arcticum Loof 1971 38 215 

Plectus acuminatus Bastian 1865 215 

Plectus cf armatus Butschli1873 215 

Plectus assimilis Butschli1873 215 

228 

68



Plectus aquatilis Andrassy1984 171 

Plectus cirratus Bastian 1865 6 191 193 194 228 

Plectus cornus Maggenti1961 215 

Plectus geophilus de Man 1880 215 

Plectus inquirendus Andrassy 1958 215 

Plectus longicaudatus Butschli1873 215 

Plectus cf. opisthocirculus Andrassy 1952 171 

Plectus parietinus Bastian 1865 171 191 193 194 215 243 

Plectus parvus Bastian 1865 215 

Plectus rhizophilus de Man 1880 215 

Rhabdolaimidae 

69


Rhabdolaimus terrestris de Man 1880 215 

Chromadorida 

_ 

_ 

Achromadoridae 

Achromadora cf. semiarmata Altherr 1952 215 

Achromadora tenax (de Man 1876) 171 215 

Ethmolaimidae 

Ethmolaimus sp. 

Odontolaimus 

Odontolaimus chlorurus de Man 1880 215 

Monohysterida 

_ 

_ 


Monhysteridae 

Eumonhystera dispar (Bastian 1865) 

Monohystera cf. dispar Bastian 

Eumonhystera filiformis (Bastian 1865) 

Monohystera filiformis Butschli 

215 

171 215 

70



Eumonhystera vulgaris (de Man 1880) 

Monohystera vulgaris de Man 

Geomonohystera villosa (Butschli1873) 

Monohystera villosa Butschli 

Monohystera stagnalis Bastian 1865 171 215 

215 

71


Acanthoceophala 

Palaecanthocephala 

Polymorphida 

_ 

_ 


Polymorphidae 

Corynosoma strumosum (Rudolphi 1802) Luhe 1904 468 506 

Notocotylus sp. 506 

Acantocephalan sp. 545 

72

Platyhelminthes 

Cestoda 


Cyclophyllidae 

_ 

_ 


Anoplocephalidae 

Moniezia benedini Moniez 1872 48 49 50 51 131 

Dilepididae 

Anomotaenia campylacantha (Krabbe 1869) 360 

Anomotaenia larina (Krabbe 1869) 

Alcataenia larina 

468 506 

Anomotaenia micacantha (Krabbe 1869) 468 506 

Alcataenia (Rissotaenia) dominicana (Raillet and Henry 1912) 468 506 

Arctotaenia tetrabothriodes 506 

Paricteropaenia porosa (Rudolphi 1810) 468 

Hymenolepididae 

73



Alopparakis larina Furhmann 1921 

Microsomacanthus ductilus 506 

Microsomacanthus microsoma (Creplin, 1829) Lopez-Neyra, 

1942 

Microsomacanthus diorchis (Fuhrmann, 1913). 506 

Taeniidae 

Echinococcus multilocularis Leuckart 1863 399 438 515 545 581 582 588 

Taenia crassiceps 545 

Taenia ovis krabbei (Moniez 1879) Verster 1969 

Cysticercus tarandi 

Taenia polyacantha 545 

Tetrabothrius minor (Loennberg, 1893) 506 

Tetrabothriidae 

Tetrabothrius erostris (Loennberg 1889) 468 506 

506 

48 50 50 131 545 560 

74


Proteocephalidea 

_ 

_ 

Proteocephalidae 

Proteocephalus exiguus Weinland 1858 184 396 

Pseudophyllidea 

_ 


Diphyllobothriidae 

Diphyllobothriidae 

Diphyllobothrium ditremum (Creplin 1825) 184 195 396 491 

Triaenophoridae 

Eubothrium crassum (Bloch 1779) 183 

Eubothrium salvelini Schrank 1790 183 184 277 396 

75

Trematoda 


Opisthorchiida 

Opisthorchiata 

_ 

Heterophyidae 

Cryptocotyle lingua (Creplin 1825) 257 468 506 

Plagiorchiida 

Brachylaimata 

_ 

Microphallidae 

Microphallus pygmaeus (Levinsen,1881) 506 

Microphallus pseudopygmaeus Galaktionov,1980 506 

Microphallus triangulatus 506 

Strigeata 

Plagiorchiata 

_ 

Gymnophallidae 

Gymnophallus somateria (Levinsen,1881) 506 


76



Gymnophallus choledochus Odhner,1900 506 

77

Annelida 

Oligochaeta 

Haplotaxida 

_ 

_ 



Enchytraeidae 

Bryodrilus diverticulatus Cernosvitov 1929 

Bryodrilus ehlersi glandulosa 

30 95 354 351 

Bryodrilus ehlersi Ude 1892 28 30 95 236 237 238 283 

Bryodrilus librus (Neilsen and Christensen 1959) 

Marionina libra Neilsen and 

Christensen 1959 

95 236 238 

Bryodrilus parvus Nurminen 1970 28 30 95 283 351 

Cernosvitoviella immota (Knoller 1935) 30 95 236 237 238 

Cernosvitoviella cf. Pusilla 

Cognettia glandulosa (Michaelsen 1888) 580 

78



Cognettia sphagnetorum (Vejdovsky 1877) 95 237 238 

Enchytraeus albidus Henle 1837 95 236 238 289 290 291 305 306 

Enchytaeus dichaetus Schmelz & Collado 2012 580 

Enchytraeus kincaidi Eisen 1904 

Enchytraeus liefdeensis Stephenson 

Fridericia bulboides Neilsen and Christensen 1959 95 236 238 

30 95 236 237 238 290 291 306 

Fridericia bulbosa (Rosa 1887) 95 236 237 238 266 

Fridericia leydigi (Vejdovsky 1877) 95 236 

Henlea heleotropha Stephenson 1922 augm. Nielsen and 

Christensen 1959 

Henlea heleotrophus Stephenson 

Henlea glandulifera Nurminen 1970 

Henlea similis 

Henlea nasuta (Eisen 1878) 95 

95 237 238 354 351 

79



Henlea perpusilla Friend 1911 

Henlea brucei Stephenson 

28 30 95 236 237 238 266 283 350 351 

373 363 

Henlea similis Nielsen and Christensen 1959 28 30 236 237 238 302 

Henlea ventriculosa (d’Udekem 1954) 28 30 236 237 238 302 351 

Lumbricillus arenarius (Michaelsen 1988) 

Lumbricillus magdalenae Nurminen 

Lumbricillus bülowi Nielsen and Christiensen 1959 95 

Lumbricillus eltoni (Stephenson 1924) 

Enchytraeus eltoni Stephenson 

Lumbricillus muscicolus (Stephenson 1924) 

Enchytraeus muscicolus Stephenson 

Lumbricillus neilseni Nurminen 1965 

as Lumbricillus bülowi Nielsen and 

Christiensen 1959 ref. 30 

Lumbricillus pagenstecheri (Ratzel 1869) 

as Lumbricellus aegialites 

Stepheson 1922 

as Lumbricellus nervosus Ude 1902 

as Lumbricellus necrophagus 

Stephenson 1922 

95 236 238 

95 236 290 

95 236 291 

95 236 237 238 

95 236 238 289 290 291 305 306 

80



Lumbricillus reynoldsoni Backlund 1948 95 236 238 

Lumbricillus rivalis Levinsen 1890 

Lumbricillus semifuscus (Claparede 1861) augm. Stephenson 

1911 

Lumbricillus tuba Stephenson 1911 95 

Marionina aporus (Stephenson 1925) 

Enchytraeus aporus Stephenson 

95 236 291 

Marionina argentea (Michaelsen 1889) 28 30 95 236 237 238 266 

Marionina crymodes (Stephenson 1922) 

Enchytreaus crymodes Stephenson 

95 236 289 305 306 

Mesenchytraeus argentatus Nurminen 1973 28 30 95 283 351 

Mesenchytraeus eltoni Stephenson 1925 95 236 291 306 

Mesenchytraeus flavus (Levinsen 1884) 28 30 95 237 238 283 

81



Mesenchytraeus melanocephalus 

Mesenchytraeus sp. n. 

Lumbricidae 

Dendrodrilus rubidis (Savigny 1826) 580 

Dendrobaena hortensis (Michaelsen 1890) 580 

82

Tardigrada 

Eutardigrada 

Apochela 

_ 

_ 


Milnesiidae 

Milnesium asiaticum Tumanov 2006 576 

Milnesium eurystomum Mauicci 1991 576 

Milnesium tardigradum Doyère 1840 78 221 222 254 319 336 

Parachela 

_ 

_ 


Hypsibiidae 

Acutuncus antarcticus (Richters 1904) 

Hypsibius antarcticus (Richters 

1905) 

Borealibius zetlandicus (Murray 1907) 

Hypsibius (Hypsibius) zetlandicus 

(J. Murray) 

Hypsibius zetlandicus (J.Murray 

1907) 

78 171 221 222 

83



Diphascon (Diphascon) stappersi Richters 1911 

Hypsibius stappersi (Richters) 

Diphascon (Diphascon) alpinum Murray 1906 

Hypsibius alpinus (Murray) 

Diphascon (Diphascon) chilenense Plate 1888 

Diphascon chilenense (Plate) Hypsibius chilenensis Plate 

Diphascon (Diphascon) oculatum oculatum Murray 1906 

Diphascon oculatum Murray 

Diphascon (Diphascon) pingue pingue (Marcus 1936) 

Diphascon pingue (Marcus) 

Diphascon (Diphascon) prorsirostre Thulin 1928 576 

Diphascon (Diphascon) recamieri Richters 1911 

Hypsibius recamieri (Richters) Diphascon recamieri Richters 

Diphascon (Diphascon) tenue Thulin 1928 

Diphascon tenue Thulin Hypsibius tenuis (Thulin) 

Diphascon (Adropion) arduifrons Thulin 1928 

Diphascon arduifrons Thulin 

222 254 319 329 336 

90 171 222 233 251 328 336 

221 222 233 254 336 

78 222 

78 221 222 329 330 576 

78 87 171 221 222 251 330 336 576 

78 221 222 

78 222 

84



Diphascon (Adropion) belgicae Richters 1911 

Fujiscon belgicae (Richters) 

Hypsibius belgicae (Richters) 

F. diphasconiellum Ito 

Diphascon (Adropion) prorsirostre Thulin 1928 

Diphascon prorsirostre Thulin 

Diphascon (Adropion) scoticum J.Murray 1905 

Diphascon belgicae Richters 

Diphascon scoticum J.Murray Hypsibius scoticus (J.Murray) 

78 221 222 254 319 336 

221 222 

Doryphoribius macrodon Binda Pilato and Dastych 1980 27 78 222 

Hebesuncus conjungens (Thulin 1911) 

Diphascon conjugens (Thulin) 

Hypsibius arcticus (Murray 1907) 

Macrobiotus arcticus Murray 

78 90 171 222 233 251 329 330 576 

222 229 233 576 

Hypsibius convergens (Urbanowicz 1925) 78 171 221 222 229 251 329 

Hypsibius dujardini (Doyère 1840) 

Macrobiotus tetradactylus Greef 

Hypsibius microps Thulin 1928 221 222 576 

78 89 90 171 221 222 229 233 576 

85



Hypsibius pallidus Thulin 1911 77 78 222 576 

Isohypsibius bakonyiensis (Iharos 1964) 

Isohypsibius sattleri (Richters, 1902) 

78 222 

Isohypsibius dastychi Pilato Bertolani and Binda 1982 78 222 

Isohypsibius ceciliae Pilato and Binda 1987 248 

Isohypsibius coulsoni 576 

Isohypsibius elegans Binda and Pilato 1971 78 222 330 

Isohypsibius fuscus (Mihelčič 1971/72) 

Hypsibius (Isohypsibius) fuscus 

Mihelcic 

Isohypsibius granulifer granulifer Thulin 1928 

Hypsibius granulifer (Thulin 1928) 

Isohypsibius papillifer papillifer (Murray 1905) 

Hypsibius papillifer (J.Murray) 

222 229 

78 90 87 171 222 251 328 329 

222 328 329 330 

Isohypsibius papillifer bulbosus (Marcus 1928) 222 328 329 336 

86



Isohypsibius prosostomus prosostomus Thulin 1928 

Hypsibius prosostomus (Thulin) 

Isohypsibius pulcher (Mihelčič 1971/72) 

Hypsibius (Hypsibius) pulcher 

Mihelcic 

Isohypsibius schaudinni (Richters 1909) 

Hypsibius schaudinni (Richters) 

Isohypsibius tuberculatus (Plate 1888) 

Macrobiotus tuberculatus Plate Hypsibius tuberculatus Plate 

Itaquascon enckelli (Mihelčič 1971/72) 

Hypsibius (Diphascon) enckelli 

Mihelcic 

Mesocrista spitzbergensis (Richters 1903) 

Hypsibius spitzbergensis Richters Diphascon spitzbergense (Richters) 

Platicrista angustata (Murray 1905) 

Diphascon angustatum Murray Hypsibius angustatus (Murray) 

Ramazzottius cataphractus (Maucci1974) 

Hypsibius cataphractus Maucci 

78 171 221 222 

222 229 

222 336 

222 233 336 319 

222 229 

90 171 191 192 193 194 222 233 

78 221 222 233 336 576 

78 221 222 

87



Ramazzottius montivagus (Dastych 1983) 

Hypsibius montivagus Dastych 1983 

Ramazzottius oberhaeuseri (Doyère 1840) 

Macrobiotus oberhauseri (Doyère) 

Hypsibius oberhauseri Doyère 

Thulinius augusti (Murray 1907) 

Macrobiotus oberhauseri var. 

granulatus Richters 

Pseudobiotus augusti (Murray) Hypsibius (Isohysibius) augusti 

Murray 

78 221 222 329 

78 221 222 233 254 319 336 

222 229 230 

Macrobiotidae 

Macrobiotus ariekammensis Weglarska 1965 78 222 

Macrobiotus brevipes (Mihelčič 1971/72) 210 222 

Macrobiotus crenulatus Richters 1904 26 221 222 233 330 576 

Macrobiotus echinogenitus Richters 1903 

? Macrobiotus crenulatus Richters 

26 78 90 191 192 193 222 233 

Macrobiotus harmsworthi harmsworthi Murray 1907 78 221 222 233 254 319 330 336 576 

Macrobiotus harmsworthi obscurus Dastych 1985 78 192 193 222 576 

88



Macrobiotus hufelandi hufelandi C.A.S. Schultze 1833 

Macrobiotus hufelandi Schultze 

1833 

78 221 222 233 251 319 328 336 576 

Macrobiotus islandicus islandicus Richters 1904 78 191 192 193 221 222 233 576 

Macrobiotus norvegicus Mihelčič 1971/72 222 229 

Macrobiotus spectabilis Thulin 1928 191 192 193 222 

Macrobiotus tenuiformis Tumanov 2007 504 

Macrobiotus voronkovi Tumanov 2007 504 

Macrobiotus willardi Pilato 1977 78 221 222 

Minibiotus intermedius (Plate 1888) 

Macrobiotus intemedius Plate 

Paramacrobiotus areolatus ( Murray 1907) 

Macrobiotus areolatus Murray 1907 

Paramacrobiotus richtersi (Murray 1911) 

Macrobiotus richtersi Murray 1911 

78 221 222 233 319 336 

221 222 336 

78 221 222 229 230 

89



Tenuibiotus danilovi Tumanov 2007 

Macrobiotus danilovi Tumanov 2007 

Richtersius coronifer (Richters 1903) 

Adorybiotus coronifer (Richters) Macrobiotus coronifer Richters 

Calohypsibiidae 

Calohypsibius ornatus (Richters 1900) 

Macrobiotus ornatus var 

spinosissimus Richters 

Eohypsibiidae 

Bertolanius nebulosus (Dastych 1983) 

Amphibolus nebulosus (Dastych 

1983) 

Bertolanius smreczynskii (Weglarska 1970) 

Doryphoribius smreczynskii 

Weglarska 

Bertolanius weglarskae (Dastych 1972) 

Amphibolus weglarskae (Dastych 

1972) 

Murrayidae 

Dactylobiotus ambiguus (Murray 1907) 

Macrobiotus ambiguus Murray 

Hypsibius ornatus Richters 

Amphibolus smreczynskii 

(Weglarska 1970) 

504 

78 221 222 319 336 

221 222 233 254 319 336 576 

77 78 222 

90 171 191 192 193 194 222 243 

222 328 329 330 

90 171 222 229 233 336 

90



Dactylobiotus dispar (Murray 1907) 

Macrobiotus dispar Murray 

Dactylobiotus macronyx (Dujardin 1851) 

Macrobiotus macronyx Dujardin 

Murrayon dianeae (Kristensen 1982) 

Macrobiotus dianeae Kristensen 

1982 

Murrayon hibernicus (Murray 1911) 

Macrobiotus hibernicus (Murray 

1911) 

Murrayon pullari (Murray 1907) 

Macrobiotus pullari Murray Macrobiotus pullari (Murray 1907) 

191 192 193 222 233 329 330 

222 234 305 336 

222 329 330 

78 222 

89 90 222 233 336 

91

Heterotardigrada 


Arthrotardigrada 

_ 

_ 


Echiniscidae 

Bryodelphax parvuspolaris 576 

Bryodelphax sinensis (Pilato 1974) 

Echiniscus sinensis Pilato 

78 221 222 

Echiniscus arctomys Ehrenberg 1853 222 233 319 336 

Echiniscus blumi blumi Richters 1903 78 221 222 319 336 

Echiniscus capillatus Ramazzotti1956 78 222 

Echiniscus granulatus (Doyère 1840) 78 221 222 254 

Echiniscus loxophthalamus Richters 1911 222 254 

Echiniscus merokensis Richters 1904 78 221 222 336 576 

92



Echiniscus oihonnae Richters 1903 222 254 336 

Echiniscus quadrispinosus Richters 1902 222 336 576 

Echiniscus spinulosus (Doyère 1840) 222 336 

Echiniscus testudo (Doyère 1840) 78 221 222 336 576 

Echiniscus wendti Richters 1903 78 171 221 222 254 319 336 576 

Pseudechiniscus islandicus (Richters 1904) 222 330 

Pseudechiniscus suillus (Ehrenberg 1853) 78 221 222 329 330 336 576 

Pseudechiniscus victor (Ehrenberg 1853) 78 221 222 254 336 

Testechiniscus spitsbergensis (Scourfield 1897) 

Echiniscus spitsbergensis Scourfield 

78 221 222 254 319 329 330 336 576 

93

Bryozoa 

Phylactolaemata 

_ 

_ 



Statoblasts. Species unknown 519 

94

Chelicerata 

Arachnida 

Acari 

Acariformes 


Oribatida 

Ameronothroidea 

Ameronothridae 

Ameronothrus lineatus (Thorell 1871) 


Eremaeus lineatus Thorell Scutovertex lineatus (Thorell) 

Ameronothrus nidicola Sitnikova 1975 

Probably Ameronothrus lineatus 

(Thorell) 

34 41 42 70 164 181 182 219 254 271 

305 306 318 319 342 343 363 373 442 453 

480 

567 

481 482 483 484 485 499 501 555 563 

56 567 

Brachychthonoidea 

Brachychthoniidae 

Eobrachychthonius borealis (Forsslund 1942) 235 442 567 

Eobrachychthonius latior (Berlese 1910) 

Eobrachychthonius grandis (Sellnick) 

Eobrachychthonius oudemansi van der Hammen 1952 235 442 567 

181 182 219 235 442 567 

95



Liochthonius alpestris (Forsslund 1958) 56 567 

Liochthonius brevis (Michael 1888) 

Brachychthonius perpusillus 

Berlese 

Liochthonius perpusillus (Berlese) 

Liochthonius laetepictus (Berlese 1910) 

Brachychthonius laetepictus Berlese 

Brachychthonius brevis (Michael) 

var. glabra 

181 318 320 442 567 

181 182 235 320 442 567 

Liochthonius lapponicus (Trägårdh 1910) 56 119 181 182 442 563 567 

Liochthonius muscorum Forsslund 1964 56 119 567 

Liochthonius neglectus Moritz 1976 457 567 

Liochthonius sellnicki (Thor 1930) 

Brachychthonius sellnicki Thor 

Liochthonius simplex 563 

Liochthonius strenzki Forsslund 1942 457 567 

Neoliochthonius piluliferus (Forsslund 1942) 457 

181 182 219 235 318 320 445 442 567 

96



Malaconothridae 

Malaconothrus monodactylus (Michael 1888) 

Malaconothrus egregius (Berlese 

1904) 

Phthiacaroidae 

Atropacarus striculus (C.L. Koch 1836) 

Steganacarus striculus (C.L. Koch 

1836) 

441 442 567 

441 442 526 

Carabodoidea 

Carabodidaae 

Carabodes marginatus (Michael, 1884) 526 567 

Tectocepheidae 

Tectocepheus alatus Berlese 1913 235 442 567 

Tectocepheus sarekensis Trägårdh 1910 181 182 442 567 

Tectocepheus velatus (Michael 1880) 56 70 181 182 219 318 320 377 387 420 

421 442 563 567 

Tectocepheus velatus knullei Vanek 1960 442 526 567 

Ceratozetoidea 

Ceratozetidae 

97



Ceratozetes spitsbergensis Thor 1934 70 181 182 272 320 442 567 

Chamobates borealis (Trägårdh 1902) 56 567 

Chamobates birulai (Kulcynski1902) 

Notaspis birulai Kulcynski 

319 567 

Chamobates cuspidatus (Mihael, 1844) 528 567 

Diapterobates notatus (Thorell 1871) 

Oribata notata Thorell 

Trichoribates notatus (Thorell) 

Sphaerozetes notatus Thorell 

Murcia notata (Thorell) 

Edwardzetes edwardsii (Nicolet 1855) 181 182 320 442 567 

Iugoribates gracilis Sellnick 1944 457 567 

Mycobates parlmeliae (Michael 1884) 442 526 567 

Mycobates sarekensis (Trägårdh 1910) 

Calyptozetes sarekensis (Trägårdh) 

13 25 34 56 66 69 70 121 152 155 

157 164 181 191 235 271 182 266 272 305 

306 318 319 320 335 342 363 373 377 387 

411 418 420 421 442 501 526 553 555 563 

69 181 182 219 235 273 318 320 377 411 

420 421 442 501 563 567 

98



Mycobates tridactylus Willmann 1929 442 567 

Cytozetes sp. 563 

Oromurcia lucens (L.Koch 1879) 

Notaspis lucens L. Koch Probably Svalbardia paludicola 

Thor, 1930 

319 567 567 

Punctoribates punctum (C.L. Koch 1839) 442 526 

Sphaerozetes piriformis (Nicolet 1855) 164 305 306 319 567 

Svalbardia paludicola Thor 1930 181 318 442 563 567 

Trichoribates novus (Sellnick 1928) 56 567 

Trichoribates setiger (Trägårdh 1910) 235 442 567 

Trichoribates trimaculatus (C.L.Koch 1836) 56 182 219 555 567 

Cymbaeremaeoidea 

Achipterioidae 

Achipteria sp. 442 

99



Parachipteria punctata (Nicolet 1855) 441 442 526 567 

Micreremidae 

Micreremus brevipes (Michael 1888) 235 442 567 

Damaeoidea 

Damaeidae 

Subbelba montana (Kulcyznski 1992) 

Oribata ursina Thor 1930 

Spatiodamaeus sp. 

?Dameus sp 

219 318 319 567 

235 445 

Damaeus onustus C.L. Koch, 1841 526 567 

Euphthiracaroidea 

Euphthiracaridae 

Microtritia sp. 272 

Galumnoidea 

Galumnidae 

Pergalumna nervosa (Berlese 1914) 272 442 567 

Gustavioidea 

Peloppiidae 

442 

100



Ceratoppia bipilis (Hermann 1804) 

Eremaeus bipilis (Hermann) 

41 42 121 164 305 306 319 555 567 

Ceratoppia hoeli Thor 1930 13 25 66 69 70 152 181 182 235 377 

Ceratoppia sphaerica (L.Koch 1879) 

Notaspis bipilis var. sphaerica (L. 

Koch) 

Gymnodamaeoidea 

Gymnodamaeidae 

Gymnodamaeus sp. 272 

411 420 421 442 501 567 

25 219 306 335 442 526 563 567 

Hermannioidea 

Hermanniidae 

Hermannia reticulata Thorell 1872 13 34 66 69 152 164 181 182 219 235 

305 318 319 320 335 342 377 387 411 420 

421 442 501 526 555 563 567 

Hermannia scabra (L.Koch 1879) 182 319 563 567 

Licneremaeoidea 

Licneremaeidae 

Scutovertex minutus (C.L.Koch 1835) 181 182 235 442 567 

Nanhermannioidea 

101



Nanhermannioidae 

Nanhermannia sellnicki (Forsslund 1958) 442 567 

Nothroidea 

Camisiidae 

Camisia anomia Colloff 1993 13 66 69 335 365 375 377 387 411 418 

420 421 458 563 567 

Camisia biverrucata (C.L.Koch 1839) 

Nothrus biverrucatus C.L.Koch 

181 182 320 442 567 

Camisia borealis (Thorell 1871) 70 150 181 182 235 319 442 526 567 

Camisia dictyna Colloff 1993 567 

Camisia foveolata Hammer 1955 56 121 182 271 280 563 567 

Camisia horrida (Hermann 1804) 

Nothrus horridus (Hermann) Nothrus borealis Thorell 

Camisia invenusta (Michael 1888) 

Nothrus invenusta (Michael) 

Camisia lapponica (Trägårdh 1910) 56 442 567 

34 181 182 219 235 318 320 342 442 567 

567 

102



Camisia spinifer (C.L.Koch 1836) 

Nothrus spinifer C.L.Koch 

181 182 320 442 567 

Nothridae 

Nothrus palustris (C.L.Koch 1839) 441 442 567 

Platynothrus peltifer (C.L.Koch 1839) 

? Platynothrus punctatus (C.L.Koch) 

181 182 235 442 567 

Platynothrus punctatus (C.L.Koch 1839) 441 442 526 563 567 

Platynothrus septentrionalis Sellnick 1944 

Platynothrus capillatus (Berlese) 

var. septentrionalis Sellnick 

181 182 442 

Oppioidea 

Autognetidae 

Autogneta kaisilai Karppinen 1997 181 182 342 442 567 

Oppiidae 

Berniniella sp. 457 

Lauroppia fallax (Paoli 1908) 

Oppiella fallax (Paoli) Oppia fallax (Paoli) 

181 182 320 342 442 567 

103



Dissorhina ornata (Oudemans 1900) 

Cosmoppia ornata (Oudemans) 

Oppia ornata (Oudemans) 

Microppia minus (Paoli 1908) 

Oppiella minus 

Dameosoma ornatum (Oudemans) 

181 182 219 318 320 442 563 567 

272 442 567 

Oppiella neerlandica (Oudemans 1902) 442 457 

Oppiella nova Oudemans 1902 

Dameosoma neerlandicum 

(Oudemans 1900) 

Oppia neerlandica (Oudemans) 

Oppia nova (Oudemans) 

Oppiella (Moritzoppia) unicarinata (Paoli 1908) 442 457 

Moritzoppia neerlandica (Oudemans 1900) 

Oppia translamellata (Willmann) 

Moritzoppia splendens (C.L. Koch 1840) 

Discoppia splendens (C.L. Koch 

1840) 

5 69 157 181 182 219 272 318 320 335 

342 377 387 411 421 441 442 526 567 

70 121 181 182 191 219 235 271 272 320 

342 398 411 441 442 526 563 567 

442 526 567 

Suctobelbella hammeri (Krivolutsky 1966) 441 442 567 

104



Suctobelbella sarekensis (Forsslund 1941) 

Suctobelba sarekensis (Forsslund) 

Suctobelbella subcornigera (Forsslund 1941) 

Suctobelba subcornigera (Forsslund) 

181 182 342 442 567 

181 182 342 441 442 526 567 

Oribatelloidea 

Oribatellidae 

Oribatella arctica Thor 1930 181 318 320 501 561 567 

Oripodoidea 

Oribatulidae 

Oribatula tibialis (Nicolet 1885) 56 219 442 555 563 567 

Oribatula venusta Berlese 1908 

Zygoribatula venusta (Berlese) 

Phauloppia lucorum (C.L.Koch 1841) 

Notaspis lucorum (C.L.Koch) 

Michael 1897 

Zygoribatula exilis (Nicolet 1855) 

Oribatula exilis (Nicolet) 

56 181 182 235 320 342 442 501 567 

181 442 567 

164 182 305 318 319 442 567 

Oripodiidae 

Liebstadia similis (Michael 1888) 219 441 442 567 

105


Trombidiformes (Actinedida) 

Bdelloidea 

Bdellidae 

Bdella longicornis (Linnaeus 1758) 


Bdella decipiens Thorell 

Bdella semiscutata Thor 1930 318 320 

Biscirus lapidarius (Kramer 1881) 

Scirus lapidarius (Kramer) 

159 219 305 306 318 319 320 

20 318 

Cyta latirostris (Hermann 1804) 219 318 320 

Cyta brevirostris (L.Koch 1879) [a variety of C. latirostris] 

Ammonia brevirostris L.Koch sensu 

Trägårdh 

Neomolgus capillatus (Kramer 1881) 

Molgus capillatus (Kramer) 

Neomolgus littoralis (Linnaeus 1758) 

Bdella arctica Thorell 1872 

Bdella groenlandicus Trägårdh 

Bdella sanguinea (Trouessart) 

Bdella littoralis (Linnaeus) 

Molgus littoralis (L.) 

318 

12 20 41 159 219 305 306 318 411 465 

558 563 

Neomolgus pallipes (L.Koch) 159 305 306 

Cunaxidae 

106



Cunaxoides croceus (C.L.Koch 1838) 

Eupalus croceus (C.L.Koch) 

Eupodoidea 

Eupodidae 

Cocceupodes clavifrons (R. Canestrini1886) 

Coccorhagidia clavifrons 

(Canestrini) 

Eupodes clavifrons Canestrini 

320 

219 318 320 

Eupodes variegatus C.L.Koch 1838 219 318 320 

Protereunetes borneri Thor 1934 219 320 

Penthaleidae 

Penthaleus maior (Duges 1834) 

Penthaleus insulanus Thorell 

Penthalodidae 

Penthalodes ovalis (Dugès 1834) 

Chromotydeus arcticus Trägårdh 

1905 

Penthaleus arcticus Trägårdh 

Rhagidiidae 

Coccorhagidia clavifrons (Canestrini1886) 

Rhagidia clavifrons (Canestrini) 

Penthalodes ovalis Dugès 

219 318 319 320 

219 306 318 319 320 

318 

107



Rhagidia gelida Thorell 1872 20 41 159 219 305 306 318 319 320 

Lebertioidea 

Sperchonidae 

Sperchon brevirostris (Koenike 1895) 

Sperchon lineatus Thor 1899 

Pachygnathoidea 

Alicorhagiidae 

Alicorhagia clavipilus (Thor 1931) 

Epistomalycus clavipilus Thor 

Alicorhagia plumipilis (Thor 1931) 

Epistomalycus plumipilis Thor 

Nanorchestidae 

Nanorchestes arboriger (Berlese 1904) 

Alycus arboriger Berlese 

Pygmephoroidea 

Pygmephoridae 

Bakerdania arcticus (Thor 1934) 

Pediculoides arcticus (Thor) 

Raphignathoidea 

Stigmaeidae 

20 305 318 

320 

320 

219 318 320 

320 

108



Eustigmaeus oudemansi (Thor 1930) 

Ledermülleria oudemansi (Thor) 

Eustigmaeus pulchellus (Thor 1930) 

Liostigmaeus pulchellus Thor 

Tetranychoidea 

Tetranychidae 

Bryobia borealis Oudemans 1930 306 318 320 

Trombidioidea 

Podothrombiidae 

Podothrombium bicolor (Hermann 1804) 318 

Podothrombium curtipalpe (Thor 1900) 318 

Podothrombium svalbardense Oudemans 1930 318 

Tydeoidea 

Tydeiidae 

Coccotydeus globifer (Thor 1931) 320 

Microtydeus constans Thor 1931 219 320 

318 

318 

109


Triophtydeus pinicolus (Oudemans 1929) 320 

Tydaeolus tenuiclaviger (Thor 1931) 

Coccotydeus tenuiclaviger (Thor) 

Tydeus croceus (Linnaeus 1758) 

Tydeus langei Thor 1934 320 

Tydeus svalbardensis Thor 1932 219 320 

Parasitiformes 

Astigmata 

Analgoidea 

Avenzoariidae 

Zachvatkinia isolata Mironov, 1989 565 

Gamasida (Mesostigmata) 

Ascoidea 

Ascidae 

Antennoseius (Vitzthumia) oudemansi (Thor 1930) 


Vitzthumia oudemansi Thor 

320 

56 318 320 543 564 563 570 

Arctoseius babenkoi Makarova 1999 450 451 452 543 

110



Arctoseius cetratus (Sellnick 1940) 452 543 

Arctoseius haalovi Lindquist 1963 452 564 563 571 575 

Arctoseius laterincisus Thor 1930 318 320 543 563 

Arctoseius multidentatus Evans 1955 56 452 543 564 570 

Arctoseius ornatus Evans 1955 450 452 543 

Arctoseius pristinus Karg 1992 56 543 

Arctoseius tajnnricus Petrova and Makarova 1991 56 450 452 462 543 570 

Arctoseius tschernovi Makarova 2000 564 

Arctoseius weberi Evans 1955 56 452 543 564 

Melichares sp. 56 543 

111



Proctolaelaps parvanalis (Thor 1930) 

Lasioseuis parvanalis Thor 

Zerconopsis sp. 56 543 

Phytoseiidae 

Amblyseius magnanalis Thor 1930 

Lasioseuis magnanalis Thor 

Dermanyssoidea 

Haemogamasidae 

Haemogamasus ambulans (Thorell 1872) 

Eulaelaps ambulans (Thorell) 

318 320 542 543 564 

318 320 543 564 

201 318 320 564 

Hypoaspididae 

Hypoaspis ovalis L.Koch 159 305 319 543 

Laelapidae 

Laelaps hilaris (Koch) 201 543 

Eviphidoidea 

Eviphididae 

Thinoseius spinosus (Willmann, 1939) 525 563 573 

Parasitoidea 

Parasitidae 

112



Pergamasus sp. 219 

Paragamasus (Aclerogamasus) insertus (Micherdziński, 1969) 580 

Vulgarogamasus immanis (Berlese, 1904) 579 

Vulgarogamasus remberti (Oudemans 1912) 580 

Rhodacaroidea 

Ologamasidae 

Gamasellus borealis (Koch 1879) 306 543 

Veigaioidea 

Veigaiidae 

Zercon andrei Sellnick, 1958 574 

Zerconoidea 

Zerconidae 

Zercon forsslundi Sellnick 1958 56 543 564 570 

Zercon solenites Haarløv 1942 564 563 572 570 

Zercon spinosus 

113


Zercon triangularis C.L.Koch 1836 318 320 543 

Zercon zelawaensis Sellnick 1944 56 543 

Zerconopsis muestairi (Schweizer, 1949) 574 

Ixodida (Metastigmata) 

Ixodoidea 

Ixodidae 

Ixodes (Ceratixodes) uriae White 1852 

Araneae 

_ 

_ 

_ 


Ixodes putus (Pickard-Cambridge) 

Gnaphosidae 

Micaria constricta Emerton 1894 

Micaria eltoni Jackson 

41 554 

Hahniidae 

Hahnia helveola Simon 1875 138 169 312 

Linyphiidae 

138 146 160 166 167 169 306 314 

114



Halorates holmgreni (Thorell 1871) 

Coryphaeolanus holmgreni Thor 

Coryphaeus mendicus L. Koch 

Collinsia holmgreni (Thorell 1871) 

Halorates spetsbergensis (Thorell 1871) 

Collinsia spetsbergensis (Thorell 

1871) 

? Typhochraestus borealis Thor 

Microeriogone spetsbergensis 

Halorates thulensis (Jackson 1934) 

Collinsia thulensis (Jackson 1934) 

Lophomma holmgreni (Thorell) 

Coryphaeus holmgreni Thor 

Typhochraestus spetsbergensis 

(Thor) 

Erigone spetsbergensis Thorell 

Erigone longipalpis Sundevall 1830 319 

Erigone arctica palearctica Braendegaard 1934 

Erigone arctica White 

41 42 43 44 120 137 146 160 163 166 

167 168 169 305 306 312 322 570 

43 56 120 137 138 146 160 161 129 163 

166 167 168 169 199 271 305 306 312 314 

318 319 345 378 420 421 535 570 

146 160 312 

12 43 55 56 120 137 138 146 160 129 

163 166 167 168 169 199 271 305 306 312 

314 318 345 378 411 420 421 535 570 

Erigone psychrophila (Thorell 1871) 137 129 138 146 160 163 166 167 168 199 

306 312 345 378 420 421 535 570 

Erigone tirolensis L. Koch 1872 20 41 42 43 44 160 166 167 169 305 

306 312 322 535 570 

115



Hilaria glacialis (Thorell 1871) 

Erigone glacialis Thorell 

Improphantes complicatus (Emerton 1882) 

Lepthyphantes hyperboreus Strand 

Lepthyphantes complicatus 

(Emerton 1882) 

Mughiphantes sobrius (Thorell 1871) 

Micryphantes hyperboreus C.L.Koch 

Linyphia sobria Thorell Lepthyphantes sobrius (Thorell 1871) 

Mecynargus borealis (Jackson 1930) 

Rhaebothorax borealis (Jackson) Conigerella borealis (Jackson) 

Agyneta nigripes (Simon 1884) 

Agyneta nigripes (Simon) 

Meioneta nigripes (Simon 1884) 

Micryphantes nigripes Simon 

Oreonetides vaginatus (Thorell 1872) 312 

Walckenaeria karpinskii (O.P.-Cambridge 1873) 

Cornicularia karpinskii (Cambridge) 

43 55 56 120 137 138 146 160 129 163 

166 167 168 169 199 305 306 312 314 318 

319 345 378 420 421 535 570 

43 120 160 169 312 345 

43 120 129 137 138 146 160 163 166 167 

168 

570 

169 199 305 306 312 314 319 345 535 

138 146 160 191 193 194 199 

41 42 44 146 160 161 163 167 169 199 

306 312 314 322 345 378 420 421 535 570 

146 160 322 

116



Tapinocyba insecta (L.Koch 1869) 

Colobocyba insecta L.Koch 

Philodromidae 

Thanatus formicinus (Clerck 1757) 345 

169 312 

117

Mandibulata 

Collembola 

Arthropleona 

_ 

_ 



Arrhopalitidae 

Arrhopalites principalis Stach 1945 

Arrhopalites binoculatus (Borner) 

Entomobryidae 

Entomobrya nivalis (L. 1758) 

Entomobrya subarctica Stach 1962 

Lepidocyrtus lignorum Fabricus 1793 

Lepidocyrtus lanuginosus (Gmelin) 

Hypogastruridae 

Hypogastrura assimilis Krausbauer, 1898 580 

Bonetogastrura nivalis (Martynova 1973) 117 

56 112 114 116 318 326 327 387 562 

33 112 116 286 326 327 562 

58 60 69 116 112 117 211 284 286 306 

318 319 326 342 387 562 587 

118



Ceratophysella longispina (Tullberg 1876) 

Hypogastrura longispina (Tullberg) 

Hypogastrura armata (Nicolet) 

Ceratophysella succinea (Gisin 1949) 

Hypogastrura hirsuta Valpas 

Achorutes longispinus 

Hypogastrura succinea Gisen Hypogastrura armata (Nicolet) 

Hypogastrura concolor (Carpenter 1900) 112 116 117 563 570 

Hypogastrura purpurescens (Lubbock 1868) 580 

Hypogastrura sensilis (Folsom 1919) 112 116 570 

Hypogastrura tullbergi (Schäffer 1900) 

Achorutes dubius Tullberg 

Hypogastrura spitsbergensis Stach 

Hypogastrura viatica (Tullberg 1871) 

Achorutes viaticus Tullberg 

Achorutes tullergi Schäffer 

58 60 107 112 116 117 140 141 142 143 

211 263 266 278 286 306 318 319 326 327 

358 400 418 420 421 443 444 470 537 562 

33 

562 

112 114 116 122 123 219 284 326 327 

13 29 31 32 33 56 58 66 68 69 

70 98 112 116 117 123 150 152 155 156 

157 271 211 241 262 263 264 266 268 286 

296 306 284 326 327 352 354 358 363 373 

20 56 58 60 109 112 114 116 117 123 

140 141 142 143 191 193 194 211 212 256 

263 266 278 284 286 305 306 308 318 319 

326 327 332 353 358 411 418 420 421 429 

119



Podura hyperborea Boheman 1865 

Hypogastrura or Ceratophysella. 

Species unknown. 

112 116 327 562 

Willemia anophthalma Börner 1901 112 114 116 122 219 263 266 318 326 327 

562 570 

Willemia scandinavica Stach 1949 112 

570 

114 116 117 141 143 266 284 562 563 

Willemia similis Mills 1934 114 116 117 562 570 

Xenylla humicola (Fabicius 1780) 20 41 42 56 58 60 110 112 114 116 

117 122 141 143 211 263 266 278 284 286 

305 306 318 319 326 327 332 358 470 537 

562 570 

Isotomidae 

Agrenia bidenticulata (Tullberg 1876) 

Isotoma bidenticulata Tullberg 

Archisotoma besselsi (Packard 1877) 

?Isotoma spitzbergensis Lubbock 

Isotoma janmayensis (Wahlgren) 

?Isotoma arctica Stscherbakow 

15 20 56 58 60 112 114 116 117 208 

211 212 263 266 271 278 286 306 318 319 

326 327 537 562 570 

58 60 112 114 116 117 122 211 217 219 

266 278 305 306 327 562 

120



Archisotoma megalops (Bagnall 1939) 112 114 116 127 208 209 562 

Archisotoma polaris Fjellberg and Poinsot 1975 

Isotoma arctica Stscherbakow 

Folsomia coeruleogrisea (Hammer, 1938) 

Isotomina gracilis Stach 

Folsomia alpha Grow and 

Christiansen 1976 

Folsomia bisetosa Gisin 1953 

Folsomia fimetaria (L.) 

Folsomia sensibilis Kseneman 

Folsomia gracilis (Stach 1962) 

Folsomia bisetosella Fjellberg 2005 523 562 563 

Folsomia ciliata Babenko & Bulavintsev, 1993 562 

Folsomia quadrioculata (Tullberg 1871) 

Isotoma quadrioculata Tullberg 

Folsomia binoculata (Wahlgren, 1899) 

Isotoma binoculata Wahlgren Folsomia regularis Hammer 1953 

13 116 117 118 562 570 

33 42 69 70 112 116 117 122 141 143 

219 

570 

249 286 326 327 537 112 114 562 563 

41 58 60 69 112 114 116 117 122 123 

141 143 219 249 263 266 271 284 306 318 

319 326 327 358 377 387 420 421 470 562 

563 570 

16 20 29 32 37 56 58 60 68 69 

109 111 112 114 116 117 123 140 141 142 

143 191 193 194 211 212 217 243 249 262 

266 268 271 278 284 305 306 318 319 326 

33 58 60 112 116 117 249 271 284 286 

318 319 326 327 562 570 

121



Folsomia fimetaria (Linnaeus, 1758) 580 

Folsomia sexoculata (Tullberg 1871) 20 58 60 114 116 122 140 141 142 143 

Folsomia stella Grow and Christiansen 1976 69 114 116 562 

219 249 286 305 306 318 326 327 400 429 

443 444 539 562 570 

Folsomia taimyrica Martnova 1973 56 114 116 326 562 570 

Parisotoma notabilis Schäffer 1896 

Isotoma notabilis Schäffer 1896 

Isotoma anglicana Lubbock 1862 

Isotoma viridis Bourlet Isotoma palustris (Gmelin) 

Desoria grisea (Lubbock, 1869) 580 

Desoria multisetis (Carpenter and Phillips 1922) 

Isotoma multisetis Carpenter and 

Phillips 1922 

112 114 116 122 219 284 562 

15 20 41 42 56 58 60 69 70 112 

114 116 117 122 211 212 219 263 266 278 

284 286 305 306 318 319 326 327 332 358 

377 387 411 418 420 421 470 502 537 558 

20 58 60 112 116 305 327 562 

122



Desoria neglecta (Schäffer 1900) 

Isotoma fennica (Reuter) 

Isotoma neglecta Schäffer 1900 

Desoria olivacea (Tullberg 1871) 

Isotoma olivacea Tullberg 1871 

Isotoma regularis Hammer 1957 

Isotoma binoculata Wahlgren 

Desoria fennica (Reuter, 1895) 

Isotoma ruseki Fjellberg 1979 

Desoria tshernovi Martynova 1974 

Isotoma fennica (Reuter) 

Isotoma olivacea (Tullberg) 

Isotoma violacea Tullberg 

Isotoma nanseni Fjellberg 

Isotoma tshernovi 

Pseudanurophorus inoculatus Bödvarsson 1958 562 

20 33 58 60 112 114 116 117 263 286 

318 319 327 562 

58 112 114 116 263 266 306 318 562 

112 562 

112 116 562 

56 69 112 116 117 326 327 418 537 562 

Pseudanurophorus psammophilus Potapov & Stebaeva, 2002 56 112 114 116 117 122 326 562 

Tetracanthella arctica Cassagnau 1959 

Tetracanthella wahlgreni 

Linnaniemi: Thor 

Tetracanthella coerulea (Haller) 

Tetracanthella pilosa Schött: 

Wahlgren 

Vertagopus arcticus Martynova 1969 112 117 562 

20 58 60 112 114 116 123 141 143 271 

305 318 319 327 332 358 375 387 418 458 

470 562 587 

123



Vertagopus pseudocinereus Fjellberg 1975 112 116 562 

Neanuridae 

Anurida granaria (Nicolet 1847) 58 112 116 306 318 327 562 

Anurida maritima (Guerin 1836) 117 562 

Anurida polaris (Hammer 1954) 

Anurida fridgida Fjellberg 

Friesea quinquespinosa Wahlgren 1900 

Friesea nauroisi Cassagnau 

Micranurida pygmaea Börner 1901 

Anurida pygmaea (Börner) 

Odontellidae 

Xenyllodes armatus Axelson 1903 112 116 318 327 562 

Onychuiridae 

Deuteraphorura variablis (Stach 1954) 580 

56 69 70 112 114 116 117 141 143 263 

266 284 326 537 562 570 

112 

570 

116 117 284 326 327 358 470 562 563 

41 56 112 114 116 117 122 219 263 284 

326 387 537 562 570 

124



Megaphorura arctica (Tullberg 1876) 

Aphorura arctica Tullberg 

Onychuirus armatus (Tullberg) var. 

arcticus Tullberg 

Protaphorura arctica 

Lipura arctica Tullberg 

Onychiurus arcticus (Tullberg) 

Mesaphorura jirii Rusek 1982 112 114 116 562 

Mesaphorura macrochaeta Rusek 1976 

Tullbergia krausbaueri Börner: Gisin 

12 13 20 35 41 42 57 58 60 66 

69 109 112 114 116 117 122 150 151 152 

153 155 156 157 162 194 211 217 219 243 

263 266 281 282 284 286 296 305 306 307 

112 114 116 117 122 219 562 

Mesaphorura tenuisensillata Rusek 1974 112 114 116 562 

Oligaphorura groenlandica (Tullberg 1876) 

Onychiurus groenlandicus (Tullberg) Lipura groenlandica (Tullberg) 

Oligaphorura ursi (Fjellberg 1984) 

Onychiurus ursi (Fjellberg) 

Thalassaphorura duplopunctatus (Strenzke 1954) 

Onychiurus duplopunctatus 

(Strenzke) 

Onychuirus neglectus Schäffer 

Protaphorura duplopunctatus 

(Strenzke 1954) 

Onychiurus debilis Moniez: (Gisin) 

Aphorura neglecta Schäffer: 

Wahlgren 

13 56 58 60 66 69 112 114 115 116 

117 122 123 140 141 143 152 156 157 193 

211 217 219 263 266 281 282 284 306 318 

319 326 327 358 373 375 377 400 411 418 

114 115 116 117 387 562 563 570 

60 112 114 116 117 122 219 284 319 326 

327 332 562 

125


Protaphorura macfadyeni (Gisin 1953) 

Aphorura armata Tullberg: Wahlgren 

Onychiurus duodecimpunctatus 

Onychuirus macfadyyeni (Gisin) 

Protaphorura subuliginata Gisin 1956 562 563 

41 42 112 114 116 122 219 326 332 562 

Tullbergia arctica Wahlgren 1900 112 114 116 122 219 326 327 562 

Chaetophorura simplex (Gisin 1958) 

Neelipleona 

_ 

_ 

Tullbergia simplex Gisin 1958 

Neelidae 

Megalothorax minimus (Willem 1900) 

Neelus (Megalothorax) minimus 

Willem: Gisin 

Symphypleona 

_ 

_ 


112 114 116 117 562 

112 114 116 117 122 219 263 326 562 

Katiannidae 

Sminthurinus aureus (Lubbock 1862) 20 112 318 327 562 

126



Sminthurinus concolor (Meinert 1896) 

Sminthurinus niger (Lubbock) 

Sminthurididae 

Sminthurides inaequalis armatus Bretfeld 2000 359 420 421 562 

Sminthurides malmgreni (Tullberg 1876) 

Sminthurus malmgreni Tullberg 

Sphaeridia pumilis (Krausbauer 1898) 112 116 562 

20 33 41 42 58 60 69 112 114 116 

117 122 211 219 263 286 305 306 284 318 

326 327 377 387 420 421 562 570 

15 20 56 58 60 112 116 117 200 266 

278 286 305 318 319 327 358 418 470 562 

127

Insecta 

Endopterygota 

Siphonaptera 

_ 

_ 


Ceratophyllidae 

Ceratophyllus vagabundus vagabundus (Boheman 1866) 

Pulex vagabunda Boheman Ceratophyllus digitalis (Wahlgren) 

Mioctenopsylla arctica arctica Rothschild 1922 

Coleoptera 

_ 

_ 


Mioctenopsylla arctica Rothschild 

Anthicidae 

Anthicus flavipes (Panzer 1797) 179 

75 105 176 223 319 563 

71 128 176 223 555 563 

Byrrhidae 

Simplocaria metallica (Sturm 1807) 113 128 261 293 303 305 

Carabidae 

Amara quenseli (Schönherr 1806) 17 113 302 303 305 411 

Chrysomelidae 

128



? Gonioctena arctica Mannerheim 1853 104 

Phytodecta ? 104 306 

Coccinellidae 

Coccinella septempunctata L. 1758 

Cryptophagidae 

Atomaria atricapilla Stephens 1830 113 179 

Atomaria lewisi Reitter 1877 113 178 

Curculionidae 

Orchestris saliceti Fabricus 

Tachyerges (Orchestris) saliceti 

Fabricus 

Isochnus flagellum (Erichson 1902) 

? Rhynchaenus foliorum (Müller) Rhynchaenus flagellum (Ericson 

1902) 

Dermestidae 

Reesa vespulae (Milliron, 1939) 

Dytiscidae 

76 104 108 198 306 319 

113 128 146 178 303 305 318 387 

129



Hydroporus striola (Gyllenhal 1827) 196 

Lathridiidae 

Latrididius minutus (L. 1767) 

Enicmus minutus L. 

Silvanidae 

Oryzaephilus mercator (Fauvel 1889) 370 

Staphylinidae 

Atheta graminicola (Gravenhorst 1806) 

? Atheta prope frigida Sahlberg 1806 

Boreophilia subplana (J. Sahlberg 1880) 

Atheta subplana J. Sahlberg 

Eucnecosum brachypterum (Gravenhorst 1802) 

Arpedium brachypterum 

(Gravenhorst) 

20 105 205 

12 56 76 104 128 146 156 157 408 411 

501 558 

76 113 128 146 198 319 

20 113 198 205 

Micralymma marinum (Strøm 1783) 20 76 113 146 198 205 319 

Olophrum boreale (Paykull 1792) 20 41 113 205 

130

Diptera 


Omalium caesum Gravenhorst 1806 113 

Omalium septentrioneis Thomson 1857 20 205 

Brachycera 

_ 

Anthomyiidae 

Fucellia focorum (Fallén 1819) 


Scatomyza hyperborea Boheman 

Zaphne frontata (Zetterstedt 1838) 

Acroptena frontata Zetterstedt 

Scatomyza obscura Boheman 

Calliphoridae 

Calliphora vicina Robineau-Desvoidy 1830 

Calliphora erythrocephala (Meigen 

1826) 

Aricia frontata Zetterstedt 

Hylemyia frontata Zetterstedt 

146 321 

42 65 120 305 306 308 319 321 

Cynomyia mortuorum (L. 1761) 20 41 42 101 219 553 

Protophormia atriceps (Zetterstedt, 1845) 

Boreellus atriceps (Zetterstedt, 1845) 

306 

306 529 

131



Protophormia terranovae (Robineau-Desvoidy 1830) 

Phormia terrae-novae Robineau- 

Desvoidy 1830 

Phormia groenlandica (Zetterstedt) 

Coelopidae 

Coelopa eximia Stenhammer 1854 319 

Fucomyia frigida Fallén 1805 

Coelopa frigida (Fabricus) 

Drosophilidae 

Drosophila funebris (Fabricus 1787) 125 281 388 

105 146 158 239 306 423 505 

59 65 105 125 146 306 319 505 

Empididae 

Rhamphomyia caudata Zetterstedt 1838 65 125 146 185 187 305 306 319 505 

Heleomyzidae 

Heleomyza modesta (Meigen, 1878) 156 232 319 341 

Heleomyza serrata (L. 1758) 20 101 105 185 187 

Heleomyza borealis (Boheman 1865) 

Leria modesta Meigen (det. Collin) ? Leria geniculata Zetterstedt 

Helomyzidae 

125 497 498 505 541 552 

132



Leria geniculata Zetterstedt 1847 120 319 

Neolaria prominens (Becker 1897) 125 267 505 

Neoleria septentrionalis (Collin 1923) 

Leria septentrionalis Collin (det. 

Collin) 

Muscidae 

Spilogona brunneifrons Ringdahl 1931 313 321 

Spilogona denudata (Holmgren 1869) 

Aricia denudata Holmgren Aricia ranunculi Holmgren 

Spilogona dorsata (Zetterstedt 1845) 

Aricia conspurcata Holmgren 

Aricia fuliginosa Holmgren 

Aricia illota Holmgren 

Limnophora hyperborea Boheman 

Spilogona megastoma (Boheman 1866) 

Aricia ludibunda Holmgren 

Limnophora arctica Becher 

Aricia dorsata Zetterstedt 

Aricia hyperborea Boheman 

Aricia labiosa Boheman 

Aricia pauxilla Holmgren 

Limnophora megastoma (Boheman) 

20 59 65 101 319 

319 321 

Spilogona quinquelineata (Zetterstedt 1838) 313 321 

20 65 101 102 219 305 306 319 505 

59 65 305 319 321 505 

133



Spilogona triangulifera (Zetterstedt 1838) 

Aricia triangulifera Zetterstedt 

Limnophora triangulifera Zetterstedt 

Aricia vitticollis Zetterstedt 

Spilogona varsviensis Schnabl in Schnabl and Dziedzicki 1911 

Piophilidae 

Parapiophila vulgaris (Fallén 1820) 

Piophila vulgaris Fallén Piophila sp. cf. vulgaris Fallén 

Scatophagidae 

Allomyella frigida (Holmgren 1883) 

Microprosopa frigida (Holmgren) 

Scatophaga furcata (Say 1823) 

Scatophaga squalida Meigen 

Scatophaga litorea (Fallén 1819) 

Scatophaga nigripes (Holmgren) 

59 120 185 187 319 321 

20 101 213 505 

20 101 

59 125 146 319 411 420 

20 59 101 125 146 219 

Scatophaga varipes (Holmgren 1883) 65 219 305 306 

Sphaeroceidae 

Copromyza (Borborus) uncinatus (Duda 1923) 125 313 

134


Norrbomia fumipennis (Stenhammar 1855) 

Copromyza (Borborus) fumipennis 

Stenhammer 

125 319 505 

Syrphidae 

Syrphus ribesii (L. 1758) 65 105 120 146 156 174 177 185 

Syrphus torvus Osten-Sacken 1875 105 106 281 282 

Parasyrphus tarsatus (Zetterstedt 1838) 

Scaeva dryadis Holmgren 

Syrphus tarsatus Zetterstedt 

Syrphus tarsatus Zetterstedt 

(dryadis Holmgren) 

Tachinidae 

Tachina glacialis Boheman 319 

Nematocera 

_ 

Culicidae 

Aedes (Ochlerotatus) nigripes (Zetterstedt 1838) 


Aedes alpinus L. Culex nigripes Zetterstedt 

Mycetophilidae 

Allodida sp. 146 

265 420 421 

59 96 146 147 282 296 305 319 422 

135



Boletina maculata Holmgren 1870 

Boletina apicalis (Walker) 

Coelosia tenella (Zetterstedt 1852) 

Boletina setipennis (Holmgren) 

Exechia frigida (Boheman 1865) 

Mycetophila frigida Boheman 

Phronia egregia Dziedzicki1889 

Phronia melaena Edwards 

Rhymosia sp. 146 

Scatopsidae 

Scatopse notata (L. 1758) 

Tipula notata L 

Sciaridae 

Bradysia nervosa (Meigen 1818) 

Sciara variabilis Zetterstedt 

Bradysia praecox (Meigen 1818) 

Sciara praecox Meigen 

20 59 96 98 101 146 319 323 

59 96 98 146 319 323 

20 59 96 97 101 121 219 305 

98 146 306 323 

99 

308 

96 98 305 306 

136



Camptochaeta consimilis (Holmgren 1869) 

Corynoptera consimilis (Holmgren) Sciara consimilis Holmgren 

Camptocheata delicata (Frey 1948) 

Neosciara delicata Lengersdorf Sciara pallidiventris Holmgren 

Corynoptera röderi Lengersdorf 1931 

Sciara röderi Lengersdorf 

Lycoriella (Hemineurina) abbrevinervis (Holmgren 1869) 

Sciara abbrevinervis Holmgren 

Lycoriella (Hemineurina) cochleata (Rübsaamen 1898) 323 

Lycoriella (Hemineurina) modesta (Staeger 1840) 

Lycoriella arctica (Holmgren) Frey 

Sciara holmgreni Rübsaamen 

(frigida Holmgren) 

Sciara arctica Holmgren 

Sciara frigida Holmgren 

Sciara groenlandica Rübsaamen 

Sciara ecalcarata Holmgren 

Lycoriella postconspicua Mohrig, 1985 505 

Lycoriella (Hemineurina) vitticollis (Holmgren 1883) 

Sciara glacialis Rübsaamen Sciara permutata (Lundbeck) 

Lycoriella (Lycoriella) parva (Holmgren 1869) 

Sciara parva Holmgren 

20 96 101 323 

20 96 98 99 101 305 306 319 505 

20 101 

96 323 

20 96 97 99 101 306 318 323 

318 323 

137



Schwenckfeldina tridentata (Rübsaamen 1898) 

Rhynchosciara laguncularis 

Lengersdorf 

Simulidae 

Prosimulium ursinum (Edwards 1935) 

Simulium ursinum Edwards 

Sciara tridentata Rübsaamen 

20 96 97 101 102 305 306 318 505 

20 101 100 

Trichoceridae 

Trichocera borealis Lackschewitz 1934 73 74 296 310 

Trichocera hiemalis (De Geer 1760) 

incisurata 

Trichocera lutea (Becher 1886) 

Metatrichocera lutea (Becher) 

319 

20 73 74 96 97 98 101 110 219 

Trichocera maculipennis Meigen 1818 20 97 101 110 219 

Trichocera parva Meigen 1804 319 

Trichocera saltator (Harris 1776) 96 98 102 

Chironomidae 

Chironominae 

138



Chironomus (Chironomus) hyperboreus Staeger 1841 210 

Chironomus (Chironomus) lugubris Zetterstedt 1850 154 210 513 551 

Sergentia coracina (Zetterstedt 1850) 210 

Micropsectra insignilobus Kieffer 1924 557 

Micropsectra logani Johannsen 1928 557 

Micropsectra radialis Goetghebuer 1939 210 557 

Paratanytarsus austriacus (Kieffer 1924) 210 513 557 

Tanytarsus heliomesonyctios Langton 1999 557 

Diamesiinae 

Diamesa aberrata Lundbeck 1889 154 210 513 551 

Diamesa arctica (Boheman 1865) 154 210 513 563 

139



Diamesa arctica sensu Willassen and Velle (unpubl.) 551 

Diamesa bertrami Edwards 1935 154 210 513 551 

Diamesa bohemani Goetghebuer 1932 210 513 551 

Diamesa hyperborea Holmgren 1869 210 

Diamesa incallida (Walker) 513 

Diamesa lindrothi Goetghebuer 1931 154 210 

Diamesa lundstroemi Kieffer 1911 210 

Orthocladiinae 

Camptocladius stercorarius (De Geer 1776) 210 

Chaetocladius dissipatus (Edwards), 513 551 

Chaetocladius dentiforceps (Edwards) 513 551 

140



Chaetocladius holmgreni (Jabcobson 1898) 210 513 551 

Chaetocladius laminatus Brundin 1947 154 551 

Chaetocladius perennis (Meigen 1830) 210 513 551 

Cricotopus (Cricotopus) ephippium (Zetterstedt 1838) 210 

Cricotopus (Cricotopus) lestralis (Edwards 1924) 210 513 

Cricotopus (Cricotopus) pilosellus Brund 1956 210 

Cricotopus (Cricotopus) polaris Keiffer 1926 513 551 

Cricotopus (Cricotopus) tibialis (Meigen 1804) 210 

Cricotopus (Cricotopus) villosus Hirvenoja 1973 210 

Cricotopus (Isocladius) glacialis Edwards 1922 210 

Heterotrissocladius subpilosus (Kieffer 1911) 210 

141



Hydrobaenus conformis (Holmgren 1869) 210 513 551 563 

Limnophyes brachytomus (Kieffer 1922) 154 210 513 551 

Limnophyes edwardsi Sæther 1990 210 513 551 

Limnophyes eltoni (Edwards 1921) 210 513 

Limnopheys lapponicus Goetghebuer 1939 210 

Limnophyes pumilio (Holmgren 1869) 210 551 

Metriocnemus cataractarum Kieffer 1919 210 

Metriocnemus eurynotus (Holmgren 1883) 210 513 

Metriocnemus obscuripes (Holmgren 1869) 154 

Metriocnemus picipes (Meigen 1818) 210 

Metriocnemus similis Kieffer 1922 210 

142



Metriocnemus ursinus (Holmgren 1869) 210 513 551 

Oliveridia tricornis (Oliver 1976) 491 

Orthocladius (Eudactylocladius) gelidus Kieffer 1922 210 513 

Orthocladius (Euorthocladius) telochaetus Langton 1985 154 210 513 551 

Orthocladius (Euorthocladius) thienemanni Keiffer 513 551 

Orthocladius (Orthocladius) decoratus (Holmgren 1869) 210 513 551 

Orthocladius (Orthocladius) knuthi Soponis 1977 210 

Orthocladius (Orthocladius) obumbratus Johannsen 1905 210 

Orthocladius (Orthocladius) rhyacobius Kieffer 1911 513 

Orthocladius (Orthocladius) trigonolabis Edwards 1924 

Orthocladius (Pogonocladius) consobrinus (Holmgren 1869) 513 551 

143



Parachlus kiefferi 563 

Paraphaenocladius brevinervis (Holmgren 1869) 210 513 

Psectrocladius (Monopsectrocladius) calcaratus (Edwards 1929) 

Psectrocladius (Psectrocladius) barbimanus Edwards 1929 

Psectrocladius (Psectrocladius) borealis Kieffer 1919 210 

Psectrocladius (Psectrocladius) limbatellus (Holmgren 1869) 513 551 

Psectrocladius (Psectrocladius) octomaculatus Wülker 1956 

Psectrocladius (Psectrocladius) oxyura Langton 1985 21 

Psectrocladius (Psectrocladius) ventricosus Kieffer 1925 21 

Psectrocladius psilopterus (Keiffer 1906) 513 

Pseudosmittia oxoniana (Edwards 1922) 210 

144


Smittia brevipennis (Boheman 1856) 210 

Smittia extrema (Holmgren 1869) 210 

Smittia flexinervis (Kieffer 1911) 210 

Smittia lasiops (Malloch 1915) 210 

Zalutschia tatrica (Pagast 1935) 210 

Chironomus anthracinus - type 362 

Tanypodinae 

Arctopelopia barbitarsis (Zetterstedt 1850) 210 

Arctopelopia melanosoma (Goetghebuer 1933) 210 

Procladius crassinervis (Zetterstedt 1838) 

Hymenoptera 

Apocrita 

_ 

Braconidae 


145



Aphidius leclanti 578 

Biosteres sp. 20 255 

Diaeretellus svalbardicum Chaubert and Tomanović 2012 578 

Diaertiella rapae (M'Intosh 1855) 420 421 

Dendrocerus bifoveatus (Keiffer 1907) 378 387 417 420 421 

Ichneutes hyperboreus Holmgren 1869 305 306 319 333 378 420 421 

Trioxys sp. 378 

Ceraphronidae 

Conostigmus spetsbergensis (Holmgren 1869) 

Ceraphron spetsbergensis Holmgren 

Lagynodes pallidus Boheman 1832 331 

Cynipidae 

Alloxysta sp. 156 378 387 420 421 

319 

146



Ichneumonidae 

Aclastus borealis (Boheman 1866) 

Hemiteles septentrionalis Holmgren 

Aclastus gracilis (Thomson 1884) 

? Aclastus gracilis minutus 

Bridgman 1886 

Aptesis erratica (Holmgren 1969) 

Microcryptus erraticus Holmgren Phygadeuon erraticus Holmgren 

Atractodes bicolor Gravenhorst 1829 var. arcticus Holmgren 

1872 

128 146 319 331 378 387 420 421 

20 255 

20 255 319 

Atractodes pusillus (Förester 1876) 374 378 387 420 421 

Gelis glacialis (Holmgren 1869) 

Hemiteles glacialis Holmgren 

Hypamblys leucopgus (Holmgren 1869) 

Mesoleius leucopgus Holmgren 

Mesochorus palanderi Holmgren 1869 308 

Orthocentrus nigricornis Boheman 1866 

Stenomacrus nigricornis (Boheman) 

146 319 378 420 421 

319 

59 128 318 319 331 

147


Stenomacrus groenlandicus Jussila 1996 374 378 387 420 421 

Stenomacrus nigricornis (Boheman) 

Phygadeuon sp. 

Plectiscidea hyperborea (Holmgren 1869) 

Plectiscus hyperboreus Holmgren 

Stenomacrus pedestris (Holmgren 1869) 

Orthocentrus pedestris Holmgren 

Stenomacrus validicornis (Boheman 1866) 

Orthocentrus validicornis Boheman 

146 319 331 378 387 420 421 

Synodites lineiger (Thompson 1893) 378 380 420 421 

Tymmophorus obscuripes (Holmgren 1858) 

Diplazon arcticus (Holmgren) 

Symphyta 

_ 

Tenthredinidae 

Amauronematus hyperboreus (Thomson 1875) 259 319 


20 128 146 305 318 319 331 333 

331 

148


Amauronematus alberich Benson 1934 18 20 

Amauronematus villosus (Thomson 1862) 

Nematus villosus Thomson 

Amauronematus rubiginosus Hellén 1951 447 

Euura sp.? 318 

18 20 231 305 319 

Pontopristia amentorum (Förster 1854) 331 378 420 421 

Pristiphora frigida (Boheman 1865) 

Pristiphora adelungi Kenow Nematus frigidus Boheman 

Pristiphora stavdingeri (Ruthe 1859) 

Lepidoptera 

_ 

_ 


Pristiphora asperlatus Benson 

Noctuidae 

Apamea exulis (Lefèbvre, 1836) 

Apamea exulis (ef.) Apamea maillardi (Geyer 1834) 

18 20 59 128 146 259 305 306 

59 

4 175 204 455 

149



Syngrapha interrogationsis (L. 1758) 

Plusia interrogationsis (L.) 

Nymphalidae 

Nymphalis antiopa (L. 1758) 282 

204 269 

Vanessa cardui (L. 1758) 204 214 281 282 

Oecophoridae 

Hofmannospila pseudospretella (Stainton 1849) 

Papilionidae 

Parnassius apollo (L.1758) 72 

Pieridae 

Pieris napi (L.1758) 

Artogeia napi (L.) 

Margaritia sticticalis (L 1761) 

Pyralidae 

Pempelia dilutella Hubner 1796 

Pempelia subornatella Duponchel 

175 204 281 

175 204 

150


Plutella xylostella (L. 1758) 

Plutella cruciferarum var. nivella ? Plutella maculipennis (Curtis) 

20 41 104 110 175 204 205 214 253 281 

282 287 306 319 376 411 420 568 

Plutella polaris Zeller 1880 175 204 214 566 568 

Matilella fusca (Haworth 1811) 

Pyla fusca Haworth 1811 

Ephestia kuehniella Zeller 1879 

Trichoptera 

_ 

_ 

Limnophilidae 

Apatania zonella Zetterstedt 1840 

Exopterygota 

Anoplura 

_ 

_ 


Apatania arctica (Boheman) 

Goniataulius arcticus Boheman 

Apatelia zonella Zetterstedt 

Echinophthiriidae 

Antarctophthirus tricechi (Boheman 1865) 176 

204 382 379 

20 82 173 195 196 205 279 281 305 306 

319 324 325 378 491 538 

151


Echinophthirius horridus (Olfers 1816) 

Dermaptera 

_ 

_ 

Echinophthirius groenlandicus 

Becher 

Forficulidae 

Forficula auricularia L 1758 

Dictyoptera 

_ 

_ 

Blattellidae 

Blattella germanica (L. 1767) 

Ephemeroptera 

_ 

_ 

Leptophlebiidae 

Leptophlebia vespertina (L. 1758) 173 

Hemiptera 

_ 


_ 

Aphidoidea 

110 176 

152


Sitobion calvulum (Ossiannilsson 1958) 

Acyrthosiphon calvulum 

Ossiannilsson 1958 

139 245 411 419 508 507 

Acyrthosiphon svalbardicum Heikinheimo 1968 139 150 156 294 295 296 297 298 299 300 

301 334 344 347 348 387 408 411 415 420 

421 454 509 514 552 569 

Cinara abieticola (Cholodkovsky 1899) 

?Cinara piceae (Panzer 1801) ?Dilachnus piceae (nec Walker) 

105 139 

Pemphigus groenlandicus (Rübsamer 1898) 139 145 

Phthiraptera 

_ 

_ 


Amblycera 

Ancistrona vagelli (J.C. Fabricius 1787) 224 

Austromenopon brevifimbriatum (Piaget 1880) 224 

Austromenopon corporosum (Kellogg and Kuwana 1901) 224 

Austromenopon fuscofasciatum (Piaget 1880) 

Menopon fuscofasciatum Piaget 

126 224 

153



Austromenopon lutescens (Burmeister 1838) 

Menopon lutescens Burmeister 

Austromenopon merguli Timmermann 1954 224 

Austromenopon uriae Timmermann 1954 224 

110 126 224 319 

Trinoton anserinum (J.C. Fabricius 1805) 126 224 308 

Ischnocera 

Anaticola anseris (L. 126 224 

Anaticola crassicornis rubromaculatus (Rudow 1869) 224 

Carduiceps zonarius (Nitzsch [in Giebel],1866) 

Carduiceps complexivus (Kellogg 

and Chapman) 

Goniodes lagopi (L. 1758) 224 

Lagopoecus pallidovittatus (Grube 1851) 

Nirmus quadrulatus Nitzsch Lagopoecus cameratus (Nitzsch and 

Lyonet) 

126 224 

126 224 

154



Lunaceps neiris Timmermann 1954 

? Degeegeriella actophilus (Kellogg 

and Chapman 1899) 

126 224 319 

Lunaceps phaeopi (Denny 1842) 126 224 

Ornithobius hexophtalmus (Nitzsch 1871) 126 224 

Perineus nigrolimbatus (Giebel 1874) 

Esthiopterum nigrolimatum (Giebel) Lipeurus mutabilis Piaget 

Quadraceps aethereus (Giebel 1874) 

Cummingsiella aetherea klatti 

(Timmermann 1954) 

Mjöberginirmus klatti (Timmermann) 

Quadraceps hiaticulae (O. Fabricius 1780) 

Chadraceps hiaticulae (O. Fabricius) 

1780 

Quadraceps normifer (Grube 1851) 

Cummingsiella normifer (Grube 

1851) 

Quadraceps obliquus (Mjöberg 1910) 

Cummingsiella obliqua aquilonis 

Timmermann 1974 

Cummingsiella klatti (Timmerman 

1954) 

Cummingsiella hiaticulae (O. 

Fabricius 1780) 

110 126 124 224 319 

224 

124 224 

224 

224 

155



Quadraceps ornatus striolatus (Nitzsch [in Giebel], 1866) 

Cummingsiella ornatus (Grube 

1851) 

Quadraceps phalaropi (Denny 1842) 

Cummingsiella phalaropi (Denny 

1842) 

Quadraceps strepsilaris (Denny 1842) 

Cummingsiella strepsilaris (Denny 

1842) 

Saemundssonia (Saemundssonia) calva (Kellogg 1896) 124 224 

Saemundssonia (Saemundssonia) cephalus (Denny 1842) 126 224 

Saemundssonia (Saemundssonia) grylle (O. Fabricius 1780) 126 224 

Saemundssonia (Saemundssonia) inexspectata Timmermann 

1951 

Saemundssonia (Saemundssonia) lari (O. Fabricius 1780) 

Docophorus gonothorax (Giebel) 

Docophorus lari Denny 

Philopterus sp. 

Saemundssonia (Saemundssonia) lockleyi Clay 1949 

? Saemundssonia sternae (L.) 

224 

224 

224 

224 

110 126 124 224 

126 224 

156


Saemundssonia (Saemundssonia) merguli (Denny 1842) 126 124 224 

Saemundssonia (Saemundssonia) occidentalis (Kellogg 1896) 124 224 

Saemundssonia (Saemundssonia) scolopacisphaeopodis 

(Schrank 1803) 

Saemundssonia (Saemundssonia) tringae tringae (O. Fabricius 

1780) 

Philopterus fusiformis (Denny) 

Saemundssonia (Saemundssonia) tringae variabilis (Denny 1842) 

Saemundssonia (Saemundssonia) tringae lobata Martens 1974 224 

Thysanoptera 

_ 

_ 


Thripidae 

Aptinothrips rufus Haliday 1836 410 

126 224 

124 224 319 

157

Crustacea 

Branchiopoda 

Cladocera 

_ 

_ 



Bosminidae 

Bosmina longirostris Müller 1785 

Eubosmina longispina (Müller 1785) 

Bosmina longispina Müller Bosmina obstusirostris Sars 

Chydoridae 

Acroperus harpae (Baird 1835) 19 20 165 170 466 

Alona guttata Sars 1862 165 173 466 

Chydorus sphaericus (Müller 1785) 19 20 130 165 170 171 173 227 305 316 

318 319 466 559 

Daphniidae 

Daphnia pulex 

Species complex 

Daphnia pulicaria 

Daphnia longiremis 

Daphnia (Hyalodaphnia) umbra 

Daphnia middendorffiana 

Daphnia tenebrosa 

Daphnia longispina 

7 9 19 20 64 92 130 144 148 149 

165 170 171 173 191 195 196 202 218 225 

227 305 316 318 319 337 338 339 356 357 

395 390 391 401 402 403 404 406 435 466 

158


Macrothricidae 

Macrothrix hirsuticornis Norman and Brady 1867 130 165 170 171 173 226 275 276 305 311 

319 466 556 559 

Macrothrix hirsuticornis arctica (Sars 1890) 

Ctenopoda 

_ 

_ 

Macrothrix arctica Sars 

20 19 227 318 319 

Sididae 

Sida crystallina (Müller 1776) 19 20 173 

Notostraca 

_ 

_ 


Triopsidae 

Lepidurus arcticus (Pallas 1793) 19 20 130 171 173 188 195 196 305 318 

319 391 405 466 479 512 510 536 

159

Copepoda 

Calanoida 

_ 

_ 


Centropagidae 

Limnocalanus macrurus Sars 1863 173 311 

Temoridae 

Eurytemora raboti Richard 1897 9 130 171 305 318 319 527 559 

Cyclopoida 

_ 

_ 


Cyclopidae 

Acanthocyclops gigas (Claus 1857) 

Megacyclops gigas (Claus) Cyclops gigas Claus 

Cyclops abyssorum Sars 1863 

Cyclops strenuus Fischer Cyclops strenuus medianus 

19 20 197 319 305 

9 19 20 103 130 173 195 197 318 319 

390 391 491 559 

Cyclops vicinus Ulianine 1875 19 20 103 197 305 319 317 

Diacyclops crassicaudis (Sars 1863) 

Cyclops crassicaudis Sars 

130 171 173 197 311 318 

160

Harpacticoida 

_ 

_ 


Canthocamptidae 

Maraenobiotus brucei brucei (Richard 1898) 

Maraenobiotus brucei (Richard 

1898) 

311 536 

Harpacticidae 

Harpacticus uniremis Kröyer 1845 305 319 

Tachidiidae 

Tachidius discipes Giesbrecht 1882 

Tachidius brevicornis Lilljeborg ?Tachidius spitsbergensis Olofsson 

Siphonostomatoida 

_ 

_ 


305 315 318 

Lernaeopodidae 

Salmincola edwardsii Olsson 1869 184 277 

Salmincola carpionis (Krøyer 1837) 396 

161

Malacostraca 

Amphipoda 

_ 

_ 


Gammaridae 

Gammaracanthus lacustris (Sars 1867) 396 

Gammaracanthus loricatus (Sars 1896) 135 

Mysidacea 

_ 

_ 


Mysidae 

Mysis relicta Loven 1861 

Mysis oculata (Fabricus) var. relicta 

305 496 

162

Ostracoda 

Podocopida 

_ 

_ 



Candonidae 

Candona candida Müller 1776 20 319 

Eucandona rectangulata (Alm 1914) 

Candona rectangulata Alm 

Cyprididae 

Cyclocypris ovum (Jurine 1820) 319 

Eucypris arctica Olofsson 1918 305 319 

Eucypris glacialis (Sars 1890) 

Prionocypris glacialis (Sars) 

Heterocypris fretensis (Brady and Robertson 1870) 136 

Tonnacypris glacialis (G.O.Sars, 1890) 524 

Cytherideidae 

136 305 311 319 318 

19 20 136 319 305 496 

163



Cytherissa lacustris (Sars 1863) 311 491 

Limnocytheridae 

Limnocythere inopinata (Baird 1843) 136 

164

To be classified 

_ 

_ 

_ 

_ 

_ 



Cestoda 

Nadejdolepsis nitida 506 

Aploparakis leonovi 506 

Aploparakis crassipenis 506 

165

Svalbard Invertebrate references 

1 

2 

3 

4 

5 

6 

7 

8 

9 

10 

11 

12 

13 

14 

15 

16 

17 

18 

19 

20 

21 

22 

23 

24 

25 

26 

27 

The Terrestrial Invertebrate Fauna of Svalbard: references 

Aagaard, K., Bergvik, T. & Sendstad, E. 1975: Pyla (Salebria) fusca Haw. Lepidoptera found at 

Spitsbergen. Nor. J. Entomol. 22, 163–164. 

Albon, S. D. & Langvatn, R. 1997: Parasites and the regulation of Svalbard reindeer. Final report 

1996. Trondheim: Norwegian Institute for Nature Research. 

Albon, S. D., Langvatn, R., Halvorsen, O., Irvine, R. J. & Ropstad, E. 1996: Parasites, body condition 

and reproduction in Svalbard reindeer. In: Ecosystem studies in the Ny-Ålesund area. Proceedings 

from the second Ny-Ålesund seminar, Cambridge, 28–29 February 1996. P. 78. Oslo: Norwegian 

Polar Institute. 

Alendal, E. & Helle, O. 1983: Helminth parasites of muskoxen Ovibos moschatus in Norway including 

Spitsbergen and Sweden with a synopsis of parasites reported from this host. Fauna Nor. Ser. A 4, 

41–52. 

Alendal, E., Andersen, T. & Fjeldså, A. 1980: New records of Apamea maillardi (Geyer, 1932) 

(Lepidoptera: Noctuidae) in Adventdalen, Spitsbergen. Fauna Nor. Ser. B 27, 78. 

Allgén, C. 1953: Terrestrial nematodes from Jan Mayen. Ann. Mag. Nat. Hist. 6, 665–689. 

Amren, H. 1964: Ecological and taxonomical studies on zooplankton from Spitsbergen. Zool. Bidr. 

Upps. 36, 209–277. 

Amren, H. 1964: Ecological studies of zooplankton populations in some ponds on Spitsbergen. Zool. 

Bidr. Upps. 36, 161–191. 

Amren, H. 1964: Studies on the ecology and taxonomy of zooplankton, especially the Rotifers of 

Spitsbergen. Acta Univ. Upps. 42, 1–10. 

Amren, H. 1964: Temporal variation of the rotifers Keratella quadrata (Müller) in some ponds on 

Spitsbergen. Zool. Bidr. Upps. 36, 193–208. 

Arnemo, R. 1974: The food of char on Bjørnøya. Nor. Polarinst. Årb. 1972, 71. Oslo: Norwegian 


Aunaas, T., Baust, J. G. & Zachariassen, K. E. 1983: Ecophysiological studies on arthropods from 

Spitsbergen. Polar Res. 1, 235–240. 

Bale, J. S., Hodkinson, I. D., Block, W., Webb, N. R., Coulson, S. J. & Strathdee, A. T. 1997: Life 

history strategies of Arctic terrestrial arthropods. Pp. 137–165. In S. J. Woodin & M. Marquiss (eds.): 

Ecology of Arctic environments. Br. Ecol. Soc. Spec. Publ. 13. Oxford: Blackwell. 

Balik, V. 1994: On the soil testate amoebae fauna (Protozoa: Rhizopoda) of the Spitsbergen is lands 

(Svalbard). Arch. Protistenkd. 144, 365– 372. 

Bengtson, S. A. & Fjellberg, A. 1975: Summer food of the purple sandpiper Calidris maritima in 

Spitsbergen, Norway. Astarte 8, 1–6. 

Bengtson, S. A., Fjellberg, A. & Solhøy, T. 1974: Abundance of tundra arthropods in Spitsbergen, 

Norway. Entomol. Scand. 5, 137–142. 

Bengtson, S. A., Fjellberg, A. & Solhøy, T. 1975: Amara quenselii new record (Coleoptera: 

Carabidae) new to Svalbard, Norway. Nor. J. Entomol. 22, 81–82. 

Benson, R. B. 1934: Sawfies from Bear Island. Ann. Mag. Nat. Hist. 14, 207–213. 

Bertram, G. C. L. & Lack, D. 1938: Notes on the animal ecology of Bear Island. J. Anim. Ecol. 7, 

27–52. 

Bertram, G. C. L. 1933: The freshwater Crustacea of Bear Island. Ann. Mag. Nat. Hist. 12, 538–543 

Beyens, L. & Chardez, D. 1995: An annotated list of testate amoebae observed in the Arctic 

between the longitudes 27° E and 168° W. Arch. Protistenkd. 146, 219–233. 

Beyens, L. & Chardez, D. 1997: New testate amoebae taxa from the polar regions. Acta Protozool. 

36, 137–142. 

Beyens, L. & Chardez, D. with collaboration of De-Bock, P. 1986: Some new and rare testate 

amoebae from the Arctic. Acta Protozool. 25, 81– 91 

Beyens, L., Chardez, D., De Landtsheer, R. with collaboration of Baere, D. 1986: Testate amoebae 

communities from aquatic habitats in the Arctic. Polar Biol. 6, 197–205. 

Beyens, L., Chardez, D., De Landtsheer, R. with collaboration of De-Bock, P. & Jacques, E. 1986: 

Testate amoebae populations from moss and lichen habitats in the Arctic. Polar Biol. 5, 165–173. 

Binda, M. A., Pilato, G. & Dastych, H. 1980: Descrizione di una nuova specie di Eutardigrado: 

Doryphoribus macrodon. Animalia 7, 23–27. 

Binda, M. G. 1988: Ridescrizione di Macrobiotus echinogenitus Richters, 1904 e sulvalore di buona 

specie di Macrobiotus crenulatus Richters, 1904 (Eutardigrada). Animalia 15, 201–210. 

166

28 

29 

30 

31 

32 

33 

34 

35 

36 

37 

38 

39 

40 

41 

42 

43 

44 

45 

46 

47 

48 

49 

50 

51 

52 

53 

54 

55 

56 

57 

58 


Birkemoe, T. & Dózsa-Farkas, K. 1994: New records of Enchytraeidae (Oligochaeta) from 

Spitsbergen, Svalbard. Fauna Nor. Ser. A. 15, 35–44. 

Birkemoe, T. & Leinaas, H. P. 1999: Reproductive biology of the Arctic collembolan Hypogastrura 

tullbergi. Ecography 22, 31–39. 

Birkemoe, T. & Sømme, L. S. 1998: Population dynamics of two collembolan species in an Arctic 

tundra. Pedobiologia 42, 131–145. 

Birkemoe, T. 1998: Population dynamics and life history strategies of Arctic Collembola. Dr. Scient. 

thesis, University of Oslo. 

Birkemoe, T.1995: Population dynamics of Enchytraeidae at the Arctic tundra at Spitsbergen, 

Svalbard. Newslett. Enchytraeidae 4, 42–52. 

Birkenmajer, K. (ed.) 1968: Polish Spitsbergen Expeditions, 1957–1960. Warsaw: Polish Academy 

of Sciences. 

Block W. C. 1966: Some Arctic Oribatei. Acarolgia 8, 161–162. 

Block, W., Webb, N. R., Coulson, S., Hodkinson, I. D. & Worland, M. R. 1994: Thermal adaptation in 

the Arctic collembolan Onychiurus arcticus (Tullberg). J. Insect Physiol. 40, 715–722. 

Bolas-Fernadez, F., Armas-Serra, C. Fernandez de Gabriel, D. & Gomez-Barrio, A. 1987: 

Resistencia a bajas temperaturas de larvas musculares de aislamientos de Trichinella spp. Rev. Ibe 

Parasitol. Vol. Extraordin. 1987, 159–163. 

Boström, S. 1987: A new terrestrial nematode species (Rhabditida: Cephalobidae) from 

Spitzbergen. Polar Biol. 7, 375–378. 

Boström, S. 1987: A scanning electron microscope study of some species of terrestrial nematodes 

from Spitzbergen. Nematologica 33, 366– 377 

Boström, S. 1989: The taxonomic position of some teratocephalid nematodes—a scanning electron 

microscope study. Rev. Nematol. 12, 181–190. 

Brehm, V. 1917: Die v. Dr. A. Koch gesam. Entomostraken aus Spitzbergen. Arch. Hydrobiol. 11, 

609–623. 

Bristowe, W. S. 1922: The insects and arachnids of Jan Mayen. Proc. Camb. Philol. Soc. 21, 38–43. 

Bristowe, W. S. 1925: The fauna of the Arctic island of Jan Mayen and its probable origin. Ann. Mag. 

Nat. Hist. 40, 480–485. 

Bristowe, W. S. 1933: The spiders of Bear Is land. Medd. 21. Oslo: Norwegian Polar Institute. 

Bristowe, W. S. 1948: Spiders from the Arctic island of Jan Mayen. Proc. Zool. Soc. Lond. 118, 

223–225. 

Bryce, D. 1897: Contibutions to the non-marine fauna of Spitsbergen. Part II. Report on the Rotifera. 

Proc. Zool. Soc. Lond. 1897, 793–799. 

Bryce, D. 1922: On some Rotifera from Spitsbergen. The Oxford University Expedition to Spits 

bergen, 1921. J. Quekett Microsc. Club 14, 24–27. 

Brzoska, W. 1976: Productivitat un Energiegehalte von Getossp?anzen in Adventdalen 

(Spitsbergen). Oecologia 22, 387–398. 

Bye, K. & Halvorsen, O. 1983: Abomasal nematodes of the Svalbard reindeer (Rangifer tarandus 

platyrhynchus Vrolik). J. Wildl. Dis. 19, 101–105. 

Bye, K. 1980: Parasitter hos Svalbard-rein (Rangifer tarandus platyrhynchus Vrolik), med vekt på 

gastrointestinale nematoder og deres epi demi ologi. Cand. Real. thesis, University of Tromsø. 

Bye, K. 1983: Parasites of the Svalbard reindeer Rangifer tarandus platyrhynchus. Fauna 36, 82– 

86. Oslo 

Bye, K. 1985: Cestodes of reindeer (Rangifer tarandus platyrhynchus Vrolik) on the Arctic islands of 

Svalbard. Can. J. Zool. 63, 2885–2887. 

Bye, K. 1986: Parasitter i svalbardrein 2. Bendelmark. In N. A. Øritsland (ed.): Svalbard rein og dens 

livsgrunnlag. Pp. 134–138. Oslo: Universitets forlaget. 

Bye, K. 1986: Parasitter i svalbardrein 3. Encellede parasitter. In N. A. Øritsland (ed.): Svalbard-rein 

og dens livsgrunnlag. Pp. 139–141. Oslo: Universitetsforlaget. 

Bye, K., Halvorsen, O.& Nilssen, K. 1987: Immigration and regional abomasal nematodes of the 

Svalbard reindeer. J. Biogeogr. 14, 451–458. 

Byrkjedal, I. 1975: Svalbadrypas føde i august. Sterna 14, 87–90. 

Byzova, J. B., Uvarov, A. V. & Petrova, A. D. 1995: Seasonal changes in communities of soil 

invertebrates in tundra ecosystems of Hornsund, Spitsbergen. Polar Res. 16, 245–266. 

Callaghan, T. V., Sonesson, M. & Sømme, L. 1992: Response of terrestrial plants and invertebrates 

to environmental change at high latitudes. Philos. Trans. R. Soc. Lond. Ser. B 338, 279–288. 

Carpenter, G. D. H. & Philips, J. K. C. 1922: The Collembola of Spitsbergen and Bear Island. Proc. 

R. Ir. Acad. 37B, 11–21. 

167

59 

60 

61 

62 

63 

64 

65 

66 

67 

68 

69 

70 

71 

72 

73 

74 

75 

76 

77 

78 

79 

80 

81 

82 


Carpenter, G. D. H. 1927: Further records of Collembola from Spitsbergen. Proc. R. Ir. Acad. 37B, 

193–200. 

Carpenter, G. D. H. 1937: Notes on a collection of insects made by the Oxford University Arctic 

Expedition in West Spitsbergen. Ann. Mag. Nat. Hist. 20, 411–415. 

Chardez, D. & Beyens, L. with the collaboration of De Bock, P. 1987: Arcella ovaliformis sp. nov., a 

new testate amoeba from Edgeøya, a High Arctic island. Arch. Protistenkd. 134, 297–301. 

Chengalath, R. & Koste, W. 1989: Composition and distributional patterns in Arctic rotifers. 

Hydrobiologia 186/87, 191–200. 

Chesunov, A. V. & Pletnikova, M. V. 1986: New data on commensal nematodes from the genus 

Gammarinema Kinne et Gerlach, 1953 (Chrom a doria, Monhysterida). Byulleten Moskovskogo 

Obshchestva Ispytatelei Prirody Otdel Biologicheskii 91, 73–83. 

Colbourne, J. K., Crease, T. J., Weider, L. J., Hebert, P. D. N., Dufresne, F. & Hobæk, A. 1998: 

Phylo genetics and evolution of a circumarctic species complex (Cladocera: Daphnia pulex). Biol. J. 

Linn. Soc. 65, 347–365. 

Collin, J. E. 1923: Diptera (Orthorrhapha, Brachy cera and Cyclorrhapha) from Spitsbergen and 

Bear Island. Results of the Oxford University Expedition to Spitsbergen, 1921. Ann. Mag. Nat. Hist. 

11, 116–123. 

Coulson, S. J., Hodkinson, I. D., Block, W., Webb, N. R. & Worland, M. R. 1995: Low summer 

temperatures: a potential mortality factor for High Arctic soil microarthropods? J. Insect Physiol. 41, 

783–792. 

Coulson, S. J., Hodkinson, I. D., Strathdee, A. T., Block, W., Webb, N. R., Bale, J. S. & Worland, M. 

R. 1995: Thermal environments of Arctic soil organisms during winter. Arct. Alp. Res. 27, 365– 371 

Coulson, S. J., Hodkinson, I. D., Webb, N. R., Block, W., Bale, J. S., Strathdee, A. T., Worland, M. 

R. & Wooley, C. 1996: Effects of experimental temperature elevation on high-Arctic soil microarthropod 

populations. Polar Biol. 16, 147–153. 

Coulson, S. J., Leinaas, H. P., Ims, R. A. & Søvik, G. 2000: Experimental manipulation of the winter 

surface ice layer: the effects on a High Arctic soil microarthropod community. Ecography 23, 

299–306. 

Coulson, S., Hodkinson, I. D., Strathdee, A., Bale, J. S. & Block, W. 1993. Simulated climate 

change: the interaction between vegetation type and microhabitat temperatures at Ny-Alesund, 

Svalbard. Polar Biol. 13, 67–70. 

Cyprich, D. & Krumpal, M. 1991: Incidence and taxonomy of Mioctenopsylla arctica arctica 

Rothschild 1922 (Siphonaptera) in the Spitsbergen. Biologia 46, 545–549. 

Czekelius, D. 1929: Uber das Vorkommen und die Verbreitung von Parnassius apollo in Spitzbergen 

X. Internat. Congr. Entomol. Vol. 2. Pp. 1461–1467. Budapest. (Note: unable to locate this 

reference; cited in M. S. Mani, 1962: Introduction to high altitude entomology, London: Methuen.) 

Dahl, C. 1957: Die Gattung Trichocera in Spitzbergen, Bäreninsel und Jan Mayen (Dipt.). Opusc. 

Entomol. 22, 227–237. 

Dahl, C. 1973: Notes on the arthropod fauna of Spitsbergen, III: 1. Trichoceridae Diptera of 

Spitsbergen. Ann. Entomol. Fenn. 39, 49–59. 

Damph, A. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen gesammelt in den Jahren 

1907 und 1908. Aphaniptera. In A. Koenig (ed.): Avifauna Spitzbergensis. Forschungsreisen nach 

der Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und ? oristischen Ergebnisse 

Pp. 276–279. Bonn: printed privately. 

Daniel, K. & Bernhauer (no initial given) 1911: Land-Arthropoden von der Bären-Insel und 

Spitzbergen gesammelt in den Jahren 1907 und 1908. Coleoptera. In A. Koenig (ed.): Avifauna 

Spitzbergensis. Forschungsreisen nach der Bären-Insel und dem Spitzbergen Archipel, mit ihren 

faunistischen und ? oristischen Ergeb nissen. Pp. 271–272. Bonn: printed privately. 

Dastych, H. 1983: Two new Eutardigrada species from West Spitsbergen and the Tatra Mountains. 

Bull. Soc. Am. Sci. Lett. Pozn. Ser. D Sci. Biol. 23, 195–200. 

Dastych, H. 1985: West Spitsbergen Tardigrada. Acta Zool. Crac. 28, 169–214. 

Dastych, H. 1987: Two new species of Tardigrada from the Canadian subartic with some notes on 

sexual dimorphism in the family Echiniscidae. Entomol. Mitt. Zool. Staatsinst. Zool. Mus. Hambg. 8, 

319–334. 

Dastych, H. 1991: Isohypsibius sattleri (Richters, 1902), a valid species (Tardigrada). Senckenb. Bio 

71, 181–189. 

Dastych, H. 1991: Redescription of Hypsibius antarticus (Richters, 1904), with some notes on 

Hypsibius articus (Murray, 1907) (Tardigrada). Mitt. Hambg. Zool. Mus. Inst. 88, 141–159. 

De Smet, W. H. & Van Rompu, E. A. 1994. Rotifera and Tardigrada from some Cryoconite holes on 

a Spitsbergen (Svalbard) glacier. Belg. J. Zool. 124, 27–37. 

168

83 

84 

85 

86 

87 

88 

89 

90 

91 

92 

93 

94 

95 

96 

97 

98 

99 

100 

101 

102 

103 

104 

105 

106 

107 

108 

109 

110 

111 

112 


De Smet, W. H. & Van Rompu, E.A. 1996: Freshwater tardigrades from Hopen, Svalbard (76° 31' 

N). Fauna Nor. Ser. A 17, 1–9. 

De Smet, W. H. 1988: Rotifers from Bjørnøya (Svalbard), with the description of Cephalodella 

evaboedi n. sp. and Synchaeta lakowitziani arctica n. subsp. Fauna Nor. Ser. A 9, 1–18. 

De Smet, W. H. 1990: Notes on the monogonont rotifers from submerged mosses collected on 

Hopen (Svalbard). Fauna Nor. Ser. A, 11, 1–8. 

De Smet, W. H. 1993: Report on rotifers from Barentsøya, Svalbard (78° 30' N). Fauna Nor. Ser. A, 

14, 1–26. 

De Smet, W. H. 1995: Description of Encentrum dieteri sp. nov. (Rotifera: Dicranophoridae) from the 

High Arctic with a redescription of E. bidentatum (Lie-Pettersen, 1906) and E. murrayi Bryce, 1922. 

Belg. J. Zool. 125, 349–361. 

De Smet, W. H., Van Rompu, E. A. & Beyens, L. 1987: Rotifera, Gastrotricha en Tardigrada uit 

Shetland, de Faroër en Spitsbergen. (Rotifera, Gastrotricha et Tardigrada provenant du Shetland, les 

îles Féroé et du Spitsberg). Natuurwet. Tijdschr. 69, 81–102. 

De Smet, W. H., Van Rompu, E. A. & Beyens, L. 1988: Contribution to the Rotifers and aquatic 

Tardigrada of Edgeoya (Svalbard). Fauna Nor. Ser. A, 9, 19–30. 

Décamps, H. & Voisin, J. F. 1971: On the occurence of Apatania zonella Zett. (Trichoptera: 

Limnophilidae) in Svalbard. Nor. Entomol. Tidsskr. 18, 135. 

Doncaster, J. P & Stroyan, H. L. G. 1952: A new subgenus and new species of aphid from Jan 

Mayen Island. Ann. Mag. Nat. Hist. 5, 984–987. 

Doroszewski, M. 1958. About studies on Hornsund Protozoa. Przegl. Geo?z. 3, 147–150. 

Doroszewski, M. 1958: Microfauna of high north. Problemy 2, 145–146. 

Doroszewski, M. 1958: Paramecium arcticum sp. nov. Bull. Acad. Pol. Sci. Ser. Biol. 7, 73–78. 

Dózsa-Farkas, K. 1999: Taxonomical problems in enchytraeids (Oligocheata) from Spitsbergen. 

Newslett. Enchytraeidae 6, 21–30. 

Edwards, F. W. 1924: Results of the Merton College Expedition to Spitsbergen 1923. Diptera 

Nematocera. Ann. Mag. Nat. Hist. 14, 162–174. 

Edwards, F. W. 1925: Diptera (Nematocera) from Spitsbergen. Results of the Oxford University 

Expedition to Spitsbergen, 1924. Ann. Mag. Nat. Hist. 16, 354–357. 

Edwards, F. W. 1935: Diptera from Bear Island. Ann. Mag. Nat. Hist. 15, 531–543. 

Edwards, F. W. 1935: Diptera Nematocera from East Greenland. Ann. Mag. Nat. Hist. 15, 467–473. 

Edwards, F. W. 1937: Oxford University Expedition to North-East Land, 1936: Diptera. With a note 

on the pupa of Orthocladius conformis Holmgren by Aug. Thienemann. Ann. Mag. Nat. Hist. 20, 

360–364. 

Edwards, F.W. 1922: Results of the Oxford University Expedition to Spitsbergen, 1921. Diptera 

Nematocera. Ann. Mag. Nat. Hist. 10, 193– 215 

Edwards, F.W. 1923: On the Nematocerous Diptera of Jan Mayen Island. Ann. Mag. Nat. Hist. 10, 

235–240. 

Einsle, U. 1974: Planctonic Crustacea from Bjørnøya. Nor. Polarinst. Årb. 1972, 70. Oslo: Norwegian 


Elton, C. S. 1925: Coleoptera and Lepidoptera from Spitsbergen. Results of the Oxford University 

Expedition to Spitsbergen, 1924. Ann. Mag. Nat. Hist. 16, 357–359. 

Elton, C. S. 1925: The dispersal of insects to Spitsbergen. Trans. Entomol. Soc. Lond. 1925, 

289–299. 

Elton, C. S. 1929: Aphids and hover? ies in Northeast Land (Spitzbergen) in 1924: an additional note 

Proc. Entomol. Soc. Lond. 4, 76–77. 

Endal, L. J. 1979: Cryptostigmata (Acari) i tre plantesamfunn på Svalbard og virkninger av råolespill 

og simulert beitetrykk på denne fauna-en. Thesis, University of Trondheim. 

Ericson, J. B. 1902: Orchestes ? agellum Erics., ein Kafer von Spitzbergen. Verh. Zool.– Bot. Ges. 

Wein. 52, 15–17. 

Eriksen, B. E. 1998: Effects of desiccation and rehydration on Collembola from Spitsbergen, 

Svalbard. Thesis, University of Oslo. 

Fischer, F. & Becher, E. 1886: Insecten von Jan Mayen. Österr. Polarstat. Jan Mayen. Int. 

Polarforsch. Beob. Erg. K. Akad. Wiss. III, F, 59–66. 

Fischer, Z. 1990: The in?uence of humidity and tempertaure upon the rate of soil metabolism in the 

area of Hornsund (Spitsbergen). Pol. Polar Res. 11, 17–24. 

Fjellberg, A. & Poinsot, N. 1975: Archisotoma polaris new species of Collembola Isotomidae from 

Spitsbergen, Norway. Nor. J. Entomol. 22, 109–112. 

169

113 

114 

115 

116 

117 

118 

119 

120 

121 

122 

123 

124 

125 

126 

127 

128 

129 

130 

131 

132 

133 

134 

135 

136 

137 

138 

139 

140 

141 

142 


Fjellberg, A. 1980: Identi?cation keys to Norwegian Collembola. Ås: Norsk Entomologisk Forening 

(Norwegian Entomology Society). 

Fjellberg, A. 1983: Three species of staphylinid Coleoptera new to Spitsbergen, Norway. Fauna Nor. 

Ser. B 30, 110–111. 

Fjellberg, A. 1984: Collembola from Jan Mayen, Bjørnøya and Hopen Norway with additions to the 

species list from Spitsbergen. Fauna Nor. Ser. B 31, 69–76. 

Fjellberg, A. 1987: Northern species of Onychiurus Gervais, 1841, subgenus Archaphorura Bagnall, 

1941 (Collembola: Onychiuridae). Entomol. Scand. 18, 279–288. 

Fjellberg, A. 1994: The Collembola of the Norwegian Arctic islands. Medd. 133. Oslo: Norwegian 


Fjellberg, A. 1997: Collembola from Nordaustlandet, Svalbard. Fauna Nor. Ser. B 44, 71–75. 

Forsslund, K.-H. 1964: Liochthonius muscorum n.sp. und L. lapponicus (Träg.) (Acari, Oribatei). 

Medd. Grønl. 91, 1–18. 

Freiherr von Geyr, H. & le Roi, O. 1911: Land-Arthropoden von der Bären-Insel und Spitz bergen 

gesammelt in den Jahren 1907 und 1908. Chironomidae. In A. Koenig (ed.): Avifauna 

Spitzbergensis. Forschungsreisen nach der Bären-Insel und dem Spitzbergen Archipel, mit ihren fau 

istischen und ? oristischen Ergeb nissen. Pp 272–274. Bonn: printed privately. . 

Frénot, Y. & Van Vliet-Lanoë, B. 1992: Distribution, abundance and activity of invertebrates in a 

hummocky soil in Svalbard. In: Proceedings of the Arctic Research Seminar, Paris, Oct. 15.–16. 

1992. Pp. 1–10. Paris: Franco-Norwegian Foundation. 

Gisin, H. 1953: Collembola from Jan Mayen. Ann. Mag. Nat. Hist. 6, 228–234. 

Gisin, H. 1961: Collembolen aus der Sammlung C. Börner des deutschen Entomologishen Instituts 

(Apterygota). II. Spitzbergen und Faroer. Beitr. Entomol. 11, 540–545. 

Hackman, W. & Nyholm, E. S. 1968: Notes on the arthropod fauna of Spitsbergen, II: 9. Mallophaga 

from Spitsbergen and Bear Island. Ann. Entomol. Fenn. 34, 75–82. 

Hackman, W. 1968: Notes on the arthropod fauna of Spitsbergen, II: 11. Diptera: Empididae, 

Acalyptratae families and Scatophagidae. Ann. Entomol. Fenn. 34, 93–98. 

Halvorsen, G. & Gullestad, N. 1976: Freshwater Crustacea in some areas of Svalbard. Arch. 

Hydrobiol. 78, 383–395. 

Halvorsen, O. & Bye, K. 1986: Parasitter i svalbardrein 1. Rundmark i løpen. Pp. 120–133. In N. A. 

Øritsland (ed.): Svalbardrein og dens livsgrunnlag. Oslo: Universitetsforlaget. 

Halvorsen, O. & Bye, K. 1999: Parasites, biodiversity, and population dynamics in an ecosystem in 

the High Arctic. Vet. Parasitol. 84, 205– 227 

Halvorsen, O. 1986: Epidemiology of reindeer parasites. Parasitol. Today 2, 334–339. 

Halvorsen, O., Stien, A., Irvine, J., Langvatn, R. & Albon, S. 1999: Evidence for continued 

transmission of parasitic nematodes in reindeer during the Arctic winter. Int. J. Parasitol. 29, 

567–579. 

Hammar, J. 1982: The Arctic char in Arctic regions. Fauna och Flora 72, 85–92. 

Hartmann, G. 1992: Zur Kenntnis der rezenten und subfossilen Ostracoden des Liefdefjords 

(Nordspitzbergen, Svalbard). I. Teil. Mitt. Hambg. Zool. Mus. Inst. 89, 181–225. 

Hauge, E. & Sømme, L. 1997: Records of spiders (Araneae) from Nordaustlandet and Sjuøyane, 

Svalbard. Fauna Nor. Ser. A 18, 17–20. 

Hegstad, A. 1978: Araneae på Svalbard. Thesis, University of Trondheim. 

Heikinheimo, O. 1968: Notes on the arthropod fauna of Spitsbergen. II: 10. The aphid fauna of 


Hertzberg, K. & Leinaas, H. P. 1998: Drought stress as a mortality factor in two pairs of sympatric 

species of Collembola at Spitsbergen. Polar Biol. 19, 302–306. 

Hertzberg, K. 1997: Migration of Collembola in a patchy environment. Pedobiologia 41, 494–505. 

Hertzberg, K. 1998: Population response to habitat patchiness: Arctic Collembola as a study system 

Dr. Scient. thesis, University of Oslo. 

Hertzberg, K., Leinaas, H. P. & Ims, R. A. 1994: Patterns of abundance and demography: 

Collembola in a habitat patch gradient. Ecography 17, 349–359. 

Hessen, D. O. 1996: Competitive trade-off strategies in Arctic Daphnia linked to melanism and UV-B 

stress. Polar Biol. 16, 573–579. 

Hille Ris Lambers, D. 1955: Hemipterdidae. The zoology of Iceland. Vol. III. Part 52a, 1–29. 

Copenhagen: Munksgaard. 

Hinz, W. 1976: Zur Okologie der Tundra Zentralspitsbergen. Nor. Polarinst. Skr. 163. Oslo: 

Norwegian Polar Institute. 

170

143 

144 

145 

146 

147 

148 

149 

150 

151 

152 

153 

154 

155 

156 

157 

158 

159 

160 

161 

162 

163 

164 

165 

166 

167 

168 

169 

170 


Hirvenoja, M. 1967: Notes on the arthropod fauna of Spitsbergen. I, 7. Chironomidae and Culicidae 

(Diptera) from Spitsbergen. Ann. Entomol. Fenn. 33, 52–61. 

Hobæk, A. & Weider, L. J. 1999: A circumpolar study of Arctic biodiversity: phylogeographic patterns 

in the Daphnia pulex complex. Ambio 28, 245–250. 

Hobæk, A., Weider, L. J. & Wolf, H. G. 1993: Ecological genetics of Norwegian Daphnia. 3. Clonal 

richness in an Arctic apomictic complex. Heredity 71, 323–330. 

Hodkinson, I. D. & Webb, N. R. 1996: Temperature, climate change and the environmental 

constraints on the life history strategies of Arctic terrestrial invertebrates. In: Ecosystem studies in the 

Ny-Ålesund area. Proceedings from the second Ny-Ålesund seminar, Cambridge 28–29 February 

1996. Pp. 27–30. Oslo: Norwegian Polar Institute. 

Hodkinson, I. D. & Wookey, P. A. 1999: Functional ecology of soil organisms in tundra ecosystems: 

towards the future. Appl. Soil Ecol. 11, 111–126. 

Hodkinson, I. D. 1994: The British Research Project on Svalbard. Arct. Insect News 5, 8–12. 

Hodkinson, I. D., Coulson, S. J., Webb, N. R. & Block, W. 1996: Can Arctic soil microarthropods 

survive elevated summer temperatures? Funct. Ecol. 10, 314–321. 

Hodkinson, I. D., Coulson, S. J., Webb, N. R., Block, W., Strathdee, A. T., Bale, J. S. & Worland, M. 

R. 1996: Temperature and the biomass of ?ying midges (Diptera: Chironomidae) in a High Arctic 

ecosystem. Oikos 75, 241–248 

Hodkinson, I. D., Coulson, S., Webb, N. R., Block, W., Strathdee, A. T. & Bale, S. J. 1994: Feeding 

studies on Onychiurus arcticus (Tullberg) (Collembola: Onychiuridae) on West Spitsbergen. Polar 

Biol. 14, 17–19. 

Hodkinson, I. D., Healey, V. & Coulson, S. 1994: Moisture relationships of the High Arctic 

collembolan Onychiurus arcticus. Physiol. Entomol. 19, 109–114. 

Hodkinson, I. D., Webb, N. R., Bale, J. S., Block, W., Coulson, S. J. & Strathdee, A. T. 1998: Global 

change and Arctic ecosystems: conclusions and predictions from experiments with terrestrial 

inverstebrates on Spitsbergen. Arct. Alp. Res. 30, 306–313. 

Hoffmann, K. H. 1976: Organic body constituants of Protophormia terrae-novae (Dipt.) from 

Spitsbergen compared with ?ies from a laboratory stock. Oecologia 23, 13–16. 

Holm, Å. 1937: Zur Kenntnis der Spinnenfauna Spitzbergens und der Baren Insel. Ark. Zool. 29A 

(18), 1–13. 

Holm, Å. 1958: The spiders of the Isfjord region of Spitsbergen. Zool. Bidr. Upps. 33, 29– 67 

Holm, Å. 1959: Notes on Arctic spiders. Ark. Zool. 12, 511–515. 

Holmstrup, M. & Sømme, L. 1998: Dehydration and cold hardiness in the Arctic collembolan 

Onychuirus arcticus Tullberg, 1876. J. Comp. Physiol. B. 168, 197–203. 

Huhta, V. 1967: Notes on the arthropod fauna of Spitsbergen. I: 3. Some notes of the spider fauna of 

Spitsbergen. Ann. Entomol. Fenn. 33, 27. 

Hull, J. E. 1921: On some land-mites (Acari) from Spitsbergen and Bear Island. Results of the 

Oxford University Expedition to Spitsbergen, 1921. No. 23. Ann. Mag. Nat. Hist. Lond. 10, 621–623. 

Husman, S., Jacobi, H. U., Meijering, M. P. D. & Reise, B. 1978: Distribution and ecology of 

Svalbards Cladocera. Verh. Int. Ver. Limnol. 20, 2452–2456. 

Hågvar, S. & Hegstad, A. 1969: A sample of spiders (Araneida) from Svalbard. Nor. Polarinst. Årb. 

1969, 218–220. Oslo: Norwegian Polar Institute. 

Hågvar, S. 1970: Collemboler som næring for vadere på Svalbard. Nor. Polarinst. Årb. 1968, 

130–131. Oslo: Norwegian Polar Institute. 

Hågvar, S. 1971: Some observations on Coleoptera, Hymenoptera and Siphonaptera in Svalbard 

1968. Nor. Polarinst. Årb. 1969, 101–106. Oslo: Norwegian Polar Institute. 

Haarlov, N. 1977: Ectoparasites (Mallophaga, Siphonaptera, Acarina) from birds of Jan Mayen 

Island, Norway. Nor. J. Entomol. 24, 37–41. 

Jackson, A. R. 1922: On the spiders captured by Mr. C. S. Elton at Spitsbergen and Bear Island in 

1921. Results of the Oxford University Expedition to Spitsbergen. Ann. Mag. Nat. Hist. Lond. 9, 

163–169 

Jackson, A. R. 1924: On the spiders of Spitsbergen. Results of the Merton College Expedition to 

Spitsbergen in 1923. No. 1. Ann. Mag. Nat. Hist. Lond. 9, 77–79. 

Jackson, A. R. 1925: On the third collection of spiders captured by Mr. C. S. Elton in Spitsbergen. 

Results of the Oxford University Arctic Expedition, 1924. Ann. Mag. Nat. Hist. Lond. 9, 536–538. 

Jackson, A. R. 1934: On the collection of spiders made in 1928 by Dr. Sig Thor in Spitsbergen. Nor. 

Entomol. Tidsskr. 3, 332–356. 

Jacobi, H. U. & Meijering, M. 1978: On the limnology of Bear Island with special reference to 

Cladocera. Astarte 11, 79–88. 

171

171 

172 

173 

174 

175 

176 

177 

178 

179 

180 

181 

182 

183 

184 

185 

186 

187 

188 

189 

190 

191 

192 

193 

194 

195 

196 


Janiec, K. & Salwicka, K. 1996: Monogonont composition in different freshwater habitats on 

Spitsbergen (Arctic) and King George Island (Antarctica). Mem. Natl. Inst. Polar Res. Spec. Issue 51 

91–98. 

Janiec, K. 1997: The comparison of freshwater invertebrates of Spitsbergen (Arctic) and King 

George Island (Antarctic). Pol. Polar Res. 17, 173–202. 

Jørgensen, I. & Eie, J. A. 1993: The distribution of zooplankton, zoobenthos and ? sh in lakes and 

ponds of the Mosself peninsula, Svalbard. NINA Forsk.rapp. 45, 1–25. Trondheim: Norwegian 

Institute for Nature Research. 

Kaisila, J. 1967: Notes on the arthropod fauna of Spitsbergen. I: 1. Travel report. Ann. Entomol. 

Fenn. 33, 13–18. 

Kaisila, J. 1973: Notes on the arthropod fauna of Spitsbergen. III: 15. The Lepidoptera of 


Kaisila, J. 1973: Notes on the arthropod fauna of Spitsbergen. III: 16. The Anoplura and 

Siphonaptera of Spitsbergen. Ann. Entomol. Fenn. 39, 63–66. 

Kanervo, E. 1968: Notes on the arthropod fauna of Spitsbergen. II: 12 Hover?ies (Dipt. Syrphidae) 

from Spitsbergen. Ann. Entomol. Fenn. 34, 98–99. 

Kangas, E. 1967: Notes on the arthropod fauna of Spitsbergen. I: 5. Identi?cation of the Coleoptera 

collected by the Finnish Spitsbergen expeditions. Ann. Entomol. Fenn. 33, 41–43. 

Kangas, E. 1973: Notes on the arthropod fauna of Spitsbergen. I: 2. On the Coleoptera fauna of 


Kapel, C. M. O., Pozio, E., Sacchi, L. & Prestrud, P. 1999: Freeze tolerance, morphology, and 

RAPD–PCR identi? cation of Trichinella nativa in naturally infected Arctic foxes. J. Parasitol. 85, 

144–147. 

Karppinen, E. & Krivolutsky, D. A. 1982: List of oribatid mites (Acarina, Oribatei) of the northern 

paleoarctic region. I. Europe. Acta Entomol. Fenn. 41, 1–18. 

Karppinen, E. 1967: Notes on the arthropod fauna of Spitsbergen. Data on the Oribatids (Acari) from 


Kennedy, C. R. 1978: Studies on the biology of Eubothrium salvelini and E. crassum in resident and 

migratory Salvelinus alpinus and Salmo trutta and in S. salar in north Norway and the islands of 

Spitsbergen and Jan Mayen. J. Fish Biol. 12, 147–162. 

Kennedy, C. R. 1978: The parasite fauna of resident char Salvelinus alpinus from Arctic islands, with 

special reference to Bear Island. J. Fish Biol. 13, 457–466. 

Kieffer, J. J. & Lundbeck, W. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen 

gesammelt in den Jahren 1907 und 1908. Diptera. In A. Koenig (ed.): Avifauna Spitzbergensis. 

Forschungsreisen nach der Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und 

?oristischen Ergeb nissen. P. 274. Bonn: printed privately. 

Kieffer, J. J. & Theinemann, A. 1919: Chironomiden gesammelt von Dr. A Koch (Münster i. W.) auf 

den Lofoten, der Bäreninsel und Spitzbergen (Dipt.). Entomol. Mitt. 8, 38–48. 

Kieffer, J. J. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen gesammelt in den Jahren 

1907 und 1908. Diptera. In A. Koenig (ed.): Avifauna Spitzbergensis. Forschungsreisen nach der 

Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und ? oristischen Ergeb nissen. P 

275. Bonn: printed privately. 

Kieffer, J. J. 1919: Chironomiden der nördlichen Polarregion. Entomol. Mitt. 8, 110–124. 

Kjos-Hanssen, B. 1983: Freeze-resistance of Trichinella cysts in polar bears from the High Arctic 

region of Norway (Svalbard). Acta Vet. Scand. 24, 244–246. 

Kjos-Hanssen, B. 1984: Trichinella isolates from polar bears in Svalbard, freeze resistance and 

infectivity in rats and swine. Nord. Vet.med. 36, 57–61. 

Klekowski, R. Z. & Opalinski, K. W. 1986: Matter and energy ?ow in Spitsbergen ornigenic tundra. 

Polar Res. 4, 187–197. 

Klekowski, R. Z. & Opalinski, K. W. 1989: Oxygen consumption in Tardigrada from Spitsbergen. 

Polar Biol. 9, 299–303. 

Klekowski, R. Z. & Opalinski, K. W. 1990: Die Erkundung der Mechanisme von 

Umwandlungsprozessen von Energie und Materie im Okologischen System der Tundra 

Spitsbergens. Milu 7, 49–67. Berlin. 

Klekowski, R. Z. & Opalinski, K. W. 1992: Polish biological investigations in Spitsbergen. In R. Z. 

Klekowski & K. W. Opalinski. (eds.): Landscapes, life world and man in High Arctic. Pp. 139–158. 

Warsaw: Institute of Ecology, Polish Academy of Sciences. 

Klemetsen, A. 1994: The Arctic char of Bear Island. Fauna 47, 81–88. Oslo. 

Klemetsen, A., Grotnes, P. E., Holthe, H. & Kristoffersen, K. 1985: Bear Island (Barents Sea) charr. 

Inst. Freshw. Res. Drottningholm Rep. 62, 98–119. 

172

197 

198 

199 

200 

201 

202 

203 

204 

205 

206 

207 

208 

209 

210 

211 

212 

213 

214 

215 

216 

217 

218 

219 

220 

221 

222 

223 

224 

225 

226 


Koch, K. D. & Meijering, M. P. D. 1985: On the distribution and ecology of the Cyclopidae on Bear 

Island (74°30' N, 19° E). Verh. Int. Ver. Limnol. 22, 3144–3148. 

Koenig, A. 1911: Avifauna Spitzbergensis. Forschung sreisen nach der Bären-Insel und dem 

Spitzbergen Archipel, mit ihren faunistischen und ? oristischen Ergeb nissen. Bonn: printed privately 

Koponen, S. 1980: Spider fauna in the Adventfjorden area, Spitsbergen. Rep. Kevo Subarct. Res. 

Stn. 16, 13–16. Turku and Ivalo, Finland: Subarctic Research Institute. 

Kristensen, R. M. & Vestergaard, K. 1975: Døgnaktivitet under arktiske sommerbetingelser hos 

springhalen Sminthurides malgreni Tullberg (Collembola). Entomol. Medd. 43, 21–32 

Krumpàl, M., Cyprich, D., Zejda, J. & Ambros, M. 1991: The occurance of ? eld vole (Micro tus 

arvalis Pallas 1778) and its acarofauna on Spitsbergen (Svalbard). Biologia 46, 881– 885. Bratislava 

Kubicek, H. & Terek, J. 1991: Zooplankton of Svalbard (Spitsbergen). Biologia 46, 873–879. 

Bratislava. 

Kureck, A. 1966: Schlupfrytmus von Diamesa arctica (Diptera: Chironomidae) auf Spitzbergen. 

Oikos 17, 276–277. 

Lack, D. 1933: Trichoptera, Lepidoptera and Coleoptera from Bear Island. Ann. Mag. Nat. Hist. 

Lond. 12, 205–210. 

Langton, P. H. 1984: Review of type specimens of the limbatellus group with a provisional key to 

known females of Psectrocladius Keiffer (Diptera: Chironimidae). Entomol. Scand. 15, 477–485. 

Larsen, T. & Kjos-Hanssen, B. 1983: Trichinella sp. in polar bears from Svalbard in relation to hide 

length and age. Polar Res. 1, 89–96. 

Leinaas, H. P. & Ambrose Jr., W. G. 1992: Utilisation of different foraging habitats by the purple 

sandpiper, Caladris maritima, on a Spitsbergen beach. Fauna Nor. Ser. C 15, 85–91. 

Leinaas, H. P. & Ambrose Jr., W. G. 1999: Decision between small and large prey: reduced energy 

acquisition by pre-migratory purple sandpipers, Calidris maritima, on Svalbard. Polar Biol. 22, 

264–270. 

Lindegaard, C. 1997: Diptera Chironomidae, non-biting midges. In A. N. Nilsson (ed.): Aquatic 

insects of north Europe. A taxonomic handbook. Vol. 2. Pp. 266–294. Stensrup, Denmark: Apollo 

Books. 

Linnaniemi, W. M. 1935: Beitrag zur Kenntnis der Collembolenfauna Spitsbergens. Ann. Entomol. 

Fenn. 1, 137–141. 

Linnaniemi, W. M. 1935: Collembolen aus Spitsbergen, Insel Hopen, Kong Karls Land und Jan 

Mayen, eingesammelt von norwegischen arktischen Expeditionen. Nor. Entomol. Tidskr. 3, 379–381 

Lohm, U. 1978: Piophila vulgaris new-record (Diptera: Piophilidae) new to Spitsbergen, Svalbard. 

Nor. J. Entomol. 25, 224. 

Lokki, J., Malmstrom, K. K. & Suomalainen, E. 1978: Migration of Vanessa cardui new rec ord and 

Plutella xylostella (Lepidoptera) to Spits bergen in the summer 1978. Not. Entomol. 58, 121–123. 

Loof, P. A. A. 1971: Freeliving and plant parasitic nematodes from Spitzbergen collected by Mr. H. 

von Rossen. Meded. Landbouwhogesch. Wageningen 71, 1–86. 

Losos, B. & Kubicek, F. 1988: Contribution to the Chironomidae Diptera from Spitsbergen, Norway. 

Scr. Fac. Sci. Nat. Univ. Purkynianae Brun. 18, 75–88. 

Lubbock, J. 1898: On some Spitsbergen Collembola. J. Linn. Soc. 26, 616–619. 

Luecke, C. & O’Brien, W. J. 1983: Photoprotective pigments in a pond morph of Daphnia 

middendorf?ana. Arctic 36, 365–368. 

Laarsonen, E. 1985, Butter?ies from Spitsbergen. Baptria 10, 69–72. Helsinki. 

MacFadyen, A. 1954: The invertebrate fauna of Jan Mayen Island (East Greenland). J. Anim. Ecol. 

23, 261–297. 

Marshall, A. J. 1948: The Oxford University expedition to Jan Mayen, 1947: Bull. Oxf. Univ. Explor. 

Club 1, 7–10. 

Maucci, W. 1996: Tardigrada of the Arctic tundra with descriptions of two new species. Zool. J. Linn. 

Soc. 116, 185–204. 

McInnes, S. J. 1994: Zoogeographic distribution of terrestrial and freshwater tardigrades from curren 

literature. J. Nat. Hist. 28, 257–352. 

Mehl, R. 1992: Fleas (Siphonaptera) from seabirds and their nests in mainland Norway and 

Spitzbergen. NIPH Ann. 15, 3–15. Oslo: Norwegian Institute of Public Health. 

Mehl, R., Bang, C., Kjos-Hanssen, B. & Lie, H. 1982: Mallophaga from Svalbard. Fauna Nor.Ser. B 

29, 19–23. 

Meijering, M. P. D. & Jacobi, H. U. 1981: Timing of Cladocera in waters of Bear Island (74° 30' N, 

19° E) and Spiekeroog (53° 46' N, 7° 42' E). Verh. Int. Ver. Limnol. 21, 1545–1549. 

Meijering, M. P. D. 1972: Herzfrequenz und Lebensablauf von Daphnia pulex de Geer in 

Spitsbergen. Z. Wiss. Zool. 184, 395–412. 

173

227 

228 

229 

230 

231 

232 

233 

234 

235 

236 

237 

238 

239 

240 

241 

242 

243 

244 

245 

246 

247 

248 

249 

250 

251 

252 

253 

254 

255 

256 


Meijering, M. P. D. 1979: Lifecycle, ecology and timing of Macrothrix hirsuticornis in Svalbard. 

Polarforschung 49, 151–171. 

Menzel, R. 1920: Ueber freilebende Nematoden aus der Arktis. Festschr. Zschokke. 17. 

Milhelcic, C. J. 1969: Ein Beidrag zur Kenntnis der Susswassertardigraden Nordeuropas. Verh. 

Zool.–Bot. Ges. Wien 110–111, 37–45. 

Milhelcic, C. J. 1971: Susswassertardigraden aus Nordeuropa. Entomol. Scand. 2, 205–214. 

Morice, F. D. 1922: Results of the Oxford University Expedition to Spitsbergen, 1921. No. 15. Saw? 

ies. Ann. Mag. Nat. Hist. 9, 219–220. 

Morton-Firth, C. J., Block, W. & Worland, M. R. 1996: Organism survival and cell integrity in a freeze 

tolerant insect. Cryo-Letters 17, 25–30. 

Murray, J. 1907: Arctic Tardigrada. Trans. R. Soc. Edinb. 45, 669–681. 

Murray, J. 1907: Encystment of Tardigrada. Trans. R. Soc. Edinb. 45, 837–854. 

Niedbala, W. 1971: Oribatei (Acari) of Spitsbergen. Bull. Acad. Pol. Sci. 11, 737–742. 

Numinen, M 1965: Enchytraeid and lumbricid records (Oligochaeta) of Spitsbergen Ann Zool 

Fenn 2, 1–10 

Numinen, M 1967: Faunistic notes on north-European enchytraeids (Oligochaeta) Ann Zool Fenn 

4, 567–587 

Numinen, M. 1966: Further notes on Enchytraeidae (Oligochaeta) of Spitsbergen. Ann. Zool. Fenn. 

3, 68–69. 

Nuorteva, P. 1967: Notes on the arthropod fauna of Spitsbergen. I: 8. Observations on the blow?ies 

(Diptera; Calliphoridae) of Spitsbergen. Ann. Entomol. Fenn. 33, 62–64. 

Oliver, D. R. 1962: A review of the subfamily Orthocladiinae (Chironomidae, Diptera) of Bear Island. 

Astarte 20, 1–19. 

Opalinski, K. W. & Klekowski, R. Z. 1989: Oxygen consumption in Macrotrachela musculaosa and 

Trichotria truncata (Rotatoria) from the High Arctic. Polar Res. 7, 133–137. 

Opalinski, K. W. & Klekowski, R. Z. 1992: Metabolic adaptations to temperature in Spitsbergen 

invertebrates. In K. E. Opalinski & R. Z. Klekowski (eds.): Landscape, life world and man in High 

Arctic. Pp. 209–306. Warsaw: Institute of Ecology, Polish Academy of Science. 

Opalinski, K. W. 1991: Primary production and organic matter destruction in Spitsbergen tundra. Pol 

Polar Res. 12, 419–434. 

Opravilova, V. 1989: Some information on testate amoebae from Spitsbergen. Fauna Nor. Ser. A 

10, 33–37. 

Ossiannilsson, F. 1958: Acyrthosiphon calvulus, n. sp. A new aphid (Hemiptera, Homoptera) from 

Spitzbergen. Entomol. Tidskr. 79, 66–68. 

Pejler, B. 1974: Planktonic Rotatoria from Bjørn øya. Nor. Polarinst. Årb. 1972, 69. Olso: Norwegian 


Penard, E. 1903: Notice sur les Rhizopodes du Spitzberg. Arch. Protistenkd. 2, 238–282. 

Pilato, G. & Binda, M. G. 1987: Isohyp sibius ceciliae, nouva specie di eutardigrada (Hypsibiidae) 

dello Spitsbergen. Animalia 14, 125–129. Catania. 

Potapov, M. B. & Babenko, A. B. 2000: Species of the genus Folsomia (Collembola, Isotomidae) of 

northern Asia. Eur. J. Entomol. 97, 51–74. 

Prestrud, P., Stuve, G. & Holt, G. 1993: The prevalence of Trichonella sp. in Arctic foxes (Alopex 

lagopus) in Svalbard. J. Wildl. Dis. 29, 337–340. 

Pugh, P. J. A. & McInnes, S. J. 1998: The origin of Arctic terrestrial and freshwater tardigrades. Pola 

Biol. 19, 177–182. 

Rakusa-Suszczewski, S. & Styczynski, B. 1968: Tendipedidae of selected freshwater habitats in the 

Hornsund region, Vestspitsbergen. In K. Birkenmajer (ed.): Polish Spitsbergen Expeditions 

1957–1960. Pp. 151–158. Warsaw: Polish Academy of Sciences. 

Rebel, H. 1925: Ueber das Auf? nden einer Noctuide auf Spitzbergen. Nor. Entomol. Tidsskr. 2, 53. 

Richters, F. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen gesammelt in den Jahren 

1907 und 1908. Mioosfauna. In A. Koenig (ed.): Avifauna Spitzbergensis. Forschungsreisen nach de 

Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und ? oristischen Ergebnissen. 

Pp. 283–286. Bonn: printed privately. Also printed as Richters, F. 1911. Faune des Mousses 

Tardigrades. Campagne Arctique de 1907. P. 20. Brussels: Imprimerie Scienti?c. 

Roman, A. 1934: Parasitic Hymenoptera from Bear Island. Ann. Mag. Nat. Hist. 14, 203–207. 

Ruppel, G. 1968: Uber Okologie und Tagesrhythmus von Bodenarthropoden eutrophierter 

Tundragebeite Westspitzbergens. Pedobiologia 8, 150–157. 

174

257 

258 

259 

260 

261 

262 

263 

264 

265 

266 

267 

268 

269 

270 

271 

272 

273 

274 

275 

276 

277 

278 

279 

280 

281 

282 

283 

284 


Sagerup, K., Gabrielsen, G. W., Skorping, A. & Skaare J. U. 1998: Association between 

polychlorinated biphenyl (PCB) concentrations nematodes in glaucous gulls, Larus hyperboreus, from 

Bear Island. Organohal. Comp. 39, 449–451. 

Sandon, H. 1924: Some protozoa from the soils and mosses of Spitsbergen. Zool. J. Linn. Soc. 

Lond. 35, 449–475. 

Schmiedeknecht, O. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen gesammelt in 

den Jahren 1907 und 1908. Hymenoptera. In A. Koenig (ed.): Avifauna Spitzbergensis. 

Forschungsreisen nach der Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und 

?oristischen Ergeb nissen. P. 275. Bonn: printed privately. 

Schörnborn, W. 1966: Beitrag zur Okologie und Systematik det Testaceen Spitzbergens. 

Limnologica 4, 463–470. Berlin. 

Sendstad, E. & Sveum, P. 1985: Jordbunnsøkologi. Svalbardreinen og dens næringsgrunnlag. 

Avslutningrapport for MAB Svalbardprosjeket 1975–1985. Pp. 35–61. Oslo: Norwegian Polar 

Institute. 

Sendstad, E. 1974: New ? nd of Simplocaria metallica (Coleoptera, Byrrhidae) in Svalbard, Norway. 

Nor. Entomol. Tidsskr. 21, 111–111. 

Sendstad, E. 1975: En økologisk studie av Svalbards jordbunnsfauna med spesiell vekt på 

jordlevende Collembola. Thesis, University of Trondheim. 

Sendstad, E. 1976: Notes on the Collembola communities in some major plant–sociological 

associations at Spitsbergen Svalbard. Nor. J. Entomol. 23, 143–148. 

Sendstad, E. 1976: Notes on the soil fauna of Svalbard, in relation to its abiotic environment. Fauna 

29, 41–44. Oslo. 

Sendstad, E. 1976: Syrphus torvus (Diptera: Syrphidae). Records from Spitsbergen, Svalbard, 

Norway. Nor. J. Entomol. 23, 90–91. 

Sendstad, E. 1977: Jordbunnsøkologi på Svalbard. MAB i Norge. Svalbardprosjektet Rapport 3. 

Oslo: Norwegian Polar Institute. 

Sendstad, E. 1977: Notes on the biology of an Arctic bird rock. Nor. Polarinst. Årb. 1977, 265– 270. 

Oslo: Norwegian Polar Institute. 

Sendstad, E. 1981: Soil ecology of a lichen heath at Spitsbergen, Svalbard, Norway. Effects of 

arti?cial removal of the lichen plant cover. J. Range Manage. 34, 442–445. 

Sendstad, E., Bergvik, T. & Hegstad, A. 1976: Plusia interrogationis new-record (Lepidoptera, 

Noctuidae) found at Svalbard, Norway. Nor. J. Entomol. 23, 91–92. 

Sendstad, E., Solem, J. O. & Aagaard, K. 1976: Studies of terrestrial chironomids (Diptera) from 

Spitsbergen. Nor. J. Entomol. 24, 91–98. 

Seniczak, S. & Plichta, W. 1978: Structural dependence of moss-mites populations (Acari: Oribatei) 

on patchiness of vegetation in moss– lichen tundra at the north coast of Hornsund, West 

Spitsbergen. Pedobiologia 18, 145–152. 

Seyd, E. L. 1979: The evolution and distribution of Calyptozetes sarekensis (Acari: Oribatei). Biol. J. 

Linn. Soc. 12, 1–18. 

Siddiqi, M. R. 1979: Taxonomy of the plant nematode subfamily Merliniinae Siddiqi, 1970, with 

descriptions of Merlinius processus n. sp., M. loo? n. sp. and Amplimerlinius globigerus n. sp. from 

Europe. Syst. Parasitol. 1, 43–60. 

Skreslet, S. & Foged, N. 1970: The ecosystem of the Arctic lake Nordlaguna, Jan Mayen Island. II 

Plankton and benthos. Astarte, 3, 53–61. 

Skreslet, S. 1973: The ecosystem of an Arctic lake Nordlaguna, Jan Mayen Island. III. Ecology of 

Arctic char, Salvelinus alpinus (L.). Astarte 6, 43–54. 

Sobecka, E. & Piasecki, W. 1993: Parasitic fauna of Arctic charr, Salvelinus alpinus (L., 1758) from 

the Hornsund region (Spitsbergen). Acta Ichthyol. Piscatoria 23 Suppl.. 99–106. 

Solem, J. O. & Sendstad, E. 1978: Diversity in diel periodicity of Collembola communities at 

Spitsbergen Svalbard Norway. Nor. J. Entomol. 25, 9–14. 

Solem, J. O., Sendstad, E., Bergvik, T. & Hegstad, A. 1990: New record of Apatania zonella Zett. 

(Trichoptera: Limniphilidae) from Svalbard. Nor. J. Entomol. 24, 85. 

Solhøy, T. 1976: Camisia foveolata Hammer, 1955 (Acari: Oribatei) found in Norway and on 

Svalbard. Nor. J. Entomol. 23, 89. 

Soot-Ryen, T. 1942: Some Tendipedids from Spitsbergen. Norsk Entomol. Tidsskr. 6, 82–83. 

Stach, J. 1962: On the fauna of Collembola from Spitsbergen. Acta Zool. Crac. 7, 1–20. 

Stainton, H. T. 1880: On the singular new species of Plutella collected in Spitsbergen. Entomol. 

Monogr. Mag. 17, 108–109. 

Steiner,G. 1916: Beitrage zur geophischen Verbreitung freilebender Nematoden. Zool. Anz. 47, 

50–74. 

175

285 

286 

287 

288 

289 

290 

291 

292 

293 

294 

295 

296 

297 

298 

299 

300 

301 

302 

303 

304 

305 

306 

307 

308 

309 

310 

311 

312 

313 


Stephenson, J. 1922: The Oligochaeta of the Oxford University Spitsbergen expedition. Proc. Zool. 

Soc. Lond. 1922, 1109–1138. 

Stephenson, J. 1924: On some Oligochaete worms from Spitsbergen. Results of the Merton College 

Expedition to Spitsbergen, 1923. No. 3. Ann. Mag. Nat. Hist. 13, 210–216. 

Stephenson, J. 1925: The Oligochaeta of Spitsbergen and Bear Island: some additions and 

summary. Proc. Zool. Soc. Lond. 1925, 1293–1322. 

Strand, A. 1942: Die Kaferfauna von Svalbard. Medd. 54. Oslo: Norwegian Polar Institute. 

Strand, A. 1969: Simplocaria metallica Sturm new to Svalbard (Col., Byrrhidae). Nor. Entomol. 

Tidsskr. 16, 23–24. 

Strathdee, A T , Howling, G G & Bale, J S 1995: Cold hardiness of overwintering aphid eggs J 

Insect Physiol 41, 653–657 

Strathdee, A. T. & Bale, J. S. 1995: Factors limiting the distribution of Acyrthosiphon svalbardicum 

(Hemiptera: Aphididae) on Spitsbergen. Polar Biol. 15, 375–380. 

Strathdee, A. T. & Bale, J. S. 1996: Life cycle and morph production in the Arctic aphid 

Acyrthosiphon brevicorne. Polar Biol. 16, 293–300. 

Strathdee, A. T. & Bale, J. S. 1998: Life on the edge: insect ecology in Arctic environments. Ann. 

Rev. Entomol. 43, 85–106. 

Strathdee, A. T., Bale, J. S., Block, W. C., Coulson, S. J., Hodkinson, I. D. & Webb, N. R. 1993: 

Effects of temperature elevation on a ?eld population of Acyrthosiphon svalbardicum (Hemiptera: 

Aphididae) on Spitsbergen. Oecologia 96, 457–465. 

Strathdee, A. T., Bale, J. S., Hodkinson, I. D., Block, W., Webb, N. R. & Coulson, S. J. 1993: 

Identi?cation of three previously unknown morphs of Acyrthosiphon svalbardicum Heikinheimo 

(Hemiptera: Aphididae) on Spitsbergen. Entomol. Scand. 24, 43–47. 

Strathdee, A. T., Bale, J. S., Strathdee, F. C., Block, W. C., Coulson, S. J. & Webb, N. R. 1995: 

Climatic severity and the response to temperature elevation of Arctic aphids. Glob. Change Biol. 1, 

23–28. 

Strathdee, A.T., Bale, J. S., Block, W. C., Webb, N. R., Hodkinson, I. D. & Coulson, S. J. 1993: 

Extreme adaptive life-cycle in a High Arctic aphid, Acyrthosiphon svalbardicum. Ecol. Entomol. 18, 

254–258. 

Strømme, J. A. 1989: Metabolic rates of alpine and High Arctic Amara quenseli (Col., Carabidae) 

beetles. Fauna Nor. Ser. B 36, 57–58. 

Strømme, J. A., Ngari, T. W. & Zachariassen, K. E. 1986: Physiological adaptations in Coleoptera on 

Spitsbergen. Polar Res. 4, 199–204. 

Styczynski, B. & Rakusa-Suszczewski, S. 1963: Tendipedidae of selected water habitats of 

Hornsund region (Spitzbergen). Pol. Arch. Hydrobiol. 11, 237–247. 

Summerhayes, V. S. & Elton, C. S. 1923: Contributions to the ecology of Spitsbergen and Bear 

Island. J. Ecol. 11, 214–286. 

Summerhayes, V. S. & Elton, C. S. 1928: Further contributions to the ecology of Spitsbergen. J. 

Ecol. 16, 193–268. 

Sustr, T. & Block, W. 1998: Temperature dependence and acclimatory response of amylase in the 

High Arctic springtail Onychiurus arcticus (Tullberg) compared with the temperate species 

Protaphora armata (Tullberg). J. Insect Physiol. 10, 991–999. 

Sveum, P. 1987: The in?uence of grazing by Hypogastrura viatica (Insecta: Collembola) on microbia 

activity in decomposing kelp on Spitsbergen. Polar Res. 5, 71–77. 

Syrjämäki, S. 1968: A peculiar swarming mechanism of an Arctic chironomid (Diptera) at 

Spitzbergen. Ann. Zool. Fenn. 5, 151–152. 

Syrjämäki, S. 1968: Diel patterns of swarming and other activities of two Arctic dipterans 

(Chironomidae and Trichoceridae) on Spitsbergen. Oikos 19, 250–258. 

Sywula, T., Namiotko, T., Sell, J., Witkowski, A. & Zajaczkowski, M. 1994: Crustacean species new 

to Spitsbergen with notes on the polymorphism and the subfossil preservation of Cyther issa lacustri 

(G. O. Sars). Polar Res. 13, 233–235. 

Sømme, L. & Birkemoe, T. 1997: Cold tolerance and dehydration in Enchytraeidae from Svalbard. J. 

Comp. Physiol. B 167, 264–269. 

Sømme, L. & Birkemoe, T. 1999: Demography and population densities of Folsomia quadrioculata 

(Collembola, Isotomidae) on Spitsbergen. Nor. J. Entomol. 46, 35–45. 

Sømme, L. 1979: Insect life on Svalbard. Fauna 32, 137–144. Oslo. 

Sømme, L. 1993: The terrestrial arthropod fauna of Svalbard. Arct. Insect News 4, 2–4. 

Tambs-Lyche, H. 1967: Notes on the distribution of some Arctic spiders. Astarte 28, 1–13. 

Teschner, D. 1965: Beitrag zur Dipterenfauna von Spitzbergen. Zool. Anz. 175, 364–366. 

176

314 

315 

316 

317 

318 

319 

320 

321 

322 

323 

324 

325 

326 

327 

328 

329 

330 

331 

332 

333 

334 

335 

336 

337 

338 

339 


Thaler, K. 1975: Über einige Spinnen aus Spitzbergen (Arachnida, Aranei). Mitt. Schweiz. Entomol. 

Ges. 48, 437–442. 

Thomas, W. 1986: Accumulation of airborne trace pollutants by Arctic plants and soil. Water Sci. 

Technol. 18, 47–58. 

Thomasson, K. 1958: Zur Planktonkunde Spitz bergens. Hydrobiologia 12, 226–236 

Thomasson, K. 1961: Zur Planktonkunde Spitzbergens. 2. Hydrobiologia 18, 192–198. 

Thor, S. 1930a: Beitrage zur Kenntnis der invertebraten Fauna von Svalbard. Skrifter om Svalbard 

og Ishavet 27. Oslo: Norwegian Polar Institute. (This is Thor’s list of species, based on his own 

collection. Cf. Thor 1930b, below, which appeared in the same publication.) 

Thor, S. 1930b: Beitrage zur Kenntnis der invertebraten Fauna von Svalbard. Skrifter om Svalbard 

og Ishavet 27. Oslo: Norwegian Polar Institute. (These are species listed from previous expeditions 

listed by Thor but not checked by him. Cf. Thor 1930a, above, which appeared in the same 

publication.) 

Thor, S. 1934: Neue Beitrage zur Kenntnis der invertebraten Fauna von Svalbard. Zool. Anz. 107, 

114–139. 

Tiensuu, L. 1968: Notes on the arthropod fauna of Spitsbergen. II–13: Muscidae (Diptera) of 


Toft, S. 1979: Spiders from Jan Mayen. Fauna Nor. Ser. B 26, 24–25. 

Tuomikoski, R. 1967: Notes on the arthropod fauna of Spitsbergen. I: 6. Mycetophilidae and 

Sciaridae from Spitsbergen, collected by Dr. J. Kaisila in 1965: Ann. Entomol. Fenn. 33, 43–51. 

Ulmer, G. 1911: Land-Arthropoden von der Bären-Insel und Spitzbergen gesammelt in den Jahren 

1907 und 1908. Trichoptera. In A. Koenig (ed.): Avifauna Spitzbergensis. Forschungsreisen nach de 

Bären-Insel und dem Spitzbergen Archipel, mit ihren faunistischen und ? oristischen Ergebnissen. P 

276. Bonn: printed privately. 

Ulmer, G. 1932: Die Trichopteren, Ephemeropteren und Plecopteren des arktischen Gebeite. Fauna 

Arct. 6, 209–226. 

Uvarov, A. V. & Byzova, J. B. 1995: Species diversity and distribution of Collembola in the vicinity of 

the Polish Polar Station, Hornsund area, Spitsbergen. Pol. Polar Res. 16, 233–243. 

Valpas, A. 1967: Notes on the arthropod fauna of Spitsbergen. I: 4. Collemboles of Spitsbergen. 

Ann. Entomol. Fenn. 33, 28–40. 

Van Rompu, E. A. & De Smet, W. H. 1988: Some aquatic Tardigrada from Bjørnøya (Svalbard). 

Fauna Nor. Ser. A 9, 31–36. 

Van Rompu, E. A. & De Smet, W. H. 1991: Contribution to the freshwater Tardigrada from 

Barentsøya, Svalbard (78° 30' N). Fauna Nor. Ser. A 12, 29–39. 

Van Rompu, E. A. & De Smet, W. H. 1996: Freshwater tardigrades from Hopen, Svalbard (76° 31' 

N). Fauna Nor. Ser. A 17, 1–9. 

Vikberg, V. 1973: Notes on the arthropod fauna of Spitsbergen. III: List of the Hymenoptera collected 

by the Finnish Spitsbergen Expeditions. Ann. Entomol. Fenn. 39, 67–68. 

Wahlgren, E. 1900: Beitrage zur Fauna der Baren-Insel: 4. Collembola. Svenska 

Veteskapsakademien Stockholm Handlingar Bih. 26, 1–8. 

Waterston, J. 1923: Results of the Oxford University Expedition to Spitsbergen, 1921. No. 25. 

Hymenoptera, Parasitica; Ichneumonidae. Ann. Mag. Nat. Hist. 11, 31–33. 

Webb, N. R., Coulson, S. J., Hodkinson, I. D., Block, W., Bale, J. S. & Strathdee, A. T. 1998: The 

effects of experimental temperature elevation on populations of cryptostigmatic mites in High Arctic 

soils. Pedobiologia 42, 298–308. 

Webb, N. R., Hodkinson, I. D., Coulson, S. J., Block, W., Strathdee, A. T. & Bale, J. S. 1995: Life 

history and ecophysiological responses to temperature in Arctic terrestrial invertebrates. In T. V. 

Callaghan et al. (eds.): Global change and Arctic terrestrial ecosystems. Ecosystem Research 

Report 10. Pp. 49–55. Brussels: Directorate-General, Science, Research and Development, 

European Commission. 

Weglarska, B. 1966: Die Tardigraden (Tardigrada) Spitzbergens. Acta Zool. Crac. 11, 43–51. 

Weider, L. J. & Hobæk, A. 1994: Molecular bio geography of clonal lineages in a High-Arctic 

apomictic Daphnia complex. Mol. Ecol. 3, 497– 506 

Weider, L. J., Hobæk, A., Paul, D.-N. & Crease, T. J. 1999: Holarctic phylogeography of an asexual 

species complex—II. Allozymic variation and clonal structure in Arctic Daphnia. Mol. Ecology 8, 1–13 

Weider, L.J., Hobæk, A., Colbourne, J. K., Crease, T. J., Dufresne, F. & Herbert, P. D. N. 1999: 

Holarctic phylogeography of an asexual species complex—I. Mitochondrial DNA variation in Arctic 

Daphnia. Evolution 53, 777–792. 

177

340 

341 

342 

343 

344 

345 

346 

347 

348 

349 

350 

351 

352 

353 

354 

355 

356 

357 

358 

359 

360 

361 

362 

363 


Worland M. R., Grubor-Lajsic, G. & Montiel, P. O. 1998: Partial desiccation induced by sub-zero 

temperatures as a component of the survival strategy of the Arctic collembolan Onychiurus arcticus 

(Tullberg). J. Insect Physiol. 44, 211–219. 

Worland, M. R. 1996: The relationship between water content and cold tolerance in the Arctic 

collembolan Onychiurus arcticus (Collembola: Onychiuridae). Eur. J. Entomol. 93, 341–348. 

Wuthrich, C. 1989: Die Bodenfauna in der Arctischen Umwelt des Kongsfjords 

(Spitzbergen)—Versuch einer intergrativen Betrachtung eines Okosystems. Basl. Beit. Physiogeogr. 

12. 

Wuthrich, C. 1991: Landschaftsokologische Umweltforschung: beitrage zur den Wechselwirkungen 

zwischen biotischen und abiotischen Faktoren im hocharktischen Okosystem (Spitzbergen). Die Erd 

122, 335–352. 

Young, S. 1994: Insects that carry a global warning. New Sci. 142, 30 April 1994, 32–35. 

Aakra, K., & Hauge, E. (2003) Checklist of Norwegian Spiders (Arachnida: Araneae) including 

Svalbard and Jan Mayen. Norwegian Journal of Entomology, 50, 109-129. 

Albon, S.D., Stien, A.M., Irvine, R.J., Langvatn, R., Ropstad, E., & Halvorsen, O. (2002) The role of 

parasites in the dynamics of a reindeer population. Proceedings of the Royal Society of London. 

Series B, 269,1625-1632. 

Bale, J.S. (1999) Impacts of climate warming on Arctic Aphids: a comparative analysis. Ecological 

Bulletins, 47, 38-47. 

Bale, J.S., Masters, G.J., Hodkinson, I.D., Awmack, C., Bezemer, T.M., Brown, V.K., Butterfield, J., 

Buse, A., Coulson, J.C., Farrar, J., Good, J.E.G., Harrington, R., Hartley, S., Jones, T.H., Lindroth, 

R.L., Press, M.C., Symrnioudis, I., Wat,t A.D., & Whittaker, J.B. (2002) Herbivory in global climate 

change research: direct effects of rising temperature on insect herbivores. Global Change Biology, 

81, 1-16. 

Bang, C., Gullested, B., & Nordenhaug, M. (1962) Notes on Svalbard fauna 2. The Norwegian 

Ornithological Spitsbergen Expedition in the summer of 1962. Norsk Polarinstitutett Årbok, 1962:93- 

119. 

Bauer, R. (2002) Survival of frost and drought conditions in the soil by enchytraeids (Annelida; 

Oligochaeta) in Arctic alpine and temperate areas. European Journal of Soil Biology, 38:251-254. 

Birkemoe, T, Coulson, S.J., & Sømme, L.(2000) Life cycles and population dynamics of enchytraeids 

Oligochaeta from the High Arctic. Canadian Journal of Zoology, 78, 2079-2086. 

Birkemoe, T., & Leinaas, H.P. (2000) Effects of temperature on the development of an arctic 

Collembola Hypogastrura tullbergi. Functional Ecology, 14, 693-700. 

Birkemoe, T., & Liengen, T. (2000) Does collembolan grazing influence nitrogen fixation by 

cyanobacteria in the high Arctic? Polar Biology, 23, 589-592. 

Birkemoe, T., & Leinaas, H.P. (2001) Growth and development in a high Arctic Collembola: adaptive 

variation in local populations living in contrasting thermal environments Ecological Entomology, 26, 

100-105. 

Block, W. (2002) Interactions of water ice nucleators and desiccation in invertebrate cold survival. 

European Journal of Entomology, 99, 259-266. 

Borgeraas, J., & Hessen, D.O. (2002) Variations of antioxidant enzymes in Daphnia species and 

populations as related to ambient UV exposure. Hydrobiologia, 477, 15-30. 

Borgeraas, J., & Hessen, D.O. (2002) Diurnal patterns of antioxidant activity in alpine and arctic 

Daphnia under in situ UV-radiation. Archiv für Hydrobiologie, 156, 83-95. 

Brattbakk, I., Rønning, O.I., & Sensted, E. (1978) Økologiske studier på Svalbard 1 Økologiske 

effekter av nedbryting av et simulert råoleutslipp i strandsonene på Svalbard 1976 – 1977. Rapport 

STF21 A78064. SINTEF, Trondheim. p 31. 

Bretfeld, G. (2000) Third report on Symphypleona from Russia and also Georgia Kazakhstan 

Kirghizia and the Ukraine Insecta Collembola. Abhandlungen und Berichte des Naturkundemuseums 

Görlitz, 72, 1-57. 

Broek van der, E. (1968) Bird Parasites Collected by GJ van Oordt on Spitsbergen. Ardea, 56, 286- 

287. 

Brooks, S.J., & Birks, H.J.B. (2001) Chironomid-inferred air temperatures from Lateglacial and 

Holocene sites in north-west Europe: progress and problems. Quaternary Science Reviews, 20, 

1723-1741. 

Brooks, S.J., & Birks, H.J.B. (2004) The dynamics of Chironomidae (Insecta: Diptera) assemblages 

in response to environmental change during the past 700 years on Svalbard. Journal of 

Paleolimnology, 31, 483-498. 

Carstens, T., & Sendstad, E. (1979) Oil spill on the shore of an ice-covered fjord in Spitsbergen. 

International Conference on Port and Ocean Engineering Under Arctic Conditions, Norwegian 

Institute of Technology, University of Trondheim, Trondheim. p 1227-1242. 

178

364 

365 

366 

367 

368 

369 

370 

371 

372 

373 

374 

375 

376 

377 

378 

379 

380 

381 

382 

383 

384 

385 

386 


Castella, E., Adalsteinsson, H., Brittain, J.E., Gislason, G.M., Lehmann, A., Lencioni, V., Lods- 

Crozet, B., Maiolini, B., Milner, A.M., Olafsson, J.S., Saltveit, S.J., & Snook, D.L. (2001) 

Macrobenthic invertebrate richness and composition along a latitudinal gradient of European glacierfed 

streams. Freshwater Biology, 46, 1811-1831. 

Colloff, M. (1993) A taxonomic revision of the oribatid genus Camisia (Acari: Oribatida). Journal of 

Natural History, 1993, 1325-1408. 

Convey, P., Greenslade, P., Arnold, R.J. & Block, W. (1999) Collembola of sub-Antarctic South 

Georgia. Polar Biology, 22, 1-6 

Convey, P., Greenslade, P., & Pugh, P.J.A. (2000a) The terrestrial micro-arthropod fauna of the 

South Sandwich Islands. Journal of Natural History, 34, 597-609. 

Convey, P., Smith, R.I.L., Peat, H.J. & Pugh, P.J.A. (2000b). The terrestrial biota of Charcot Island, 

eastern Bellingshausen Sea, Antarctica: an example of extreme isolation. Antarctic Science 12, 406- 

413. 

Cote, S.D., Stien, A., Irvine, R.J., Dallas, J.F., Marshall, F., Halvorsen, O., Langvatn, R., & Albon, 

S.D. (2005) Resistance to abomasal nematodes and individual genetic variability in reindeer. 

Molecular Ecology, 14, 4159-4168. 

Coulson, S.J. (2007) On the occurrence of Oryzaephilus mercator (Fauvel, 1889) (Coleoptera: 

Silvanidae) on Svalbard, Norway. Norwegian Journal of Entomology. 

Coulson, S.J. (2000) A review of the terrestrial and freshwater invertebrate fauna of the High Arctic 

archipelago of Svalbard. Norwegian Journal of Entomology, 47, 41-63. 

Coulson S.J., Hodkinson I.D., Webb N.R., Block W., Bale J.S., Strathdee A.T., Worland M.R. & 

Wooley C. (1996) Effects of experimental temperature elevation on high-arctic soil microarthropod 

populations. Polar Biology 16; 147-153. 

Coulson, S.J., & Birkemoe, T. (2000) Long-term cold tolerance in Arctic invertebrates: recovery after 

4 years at below -20°C. Canadian Journal for Zoology, 78, 2055-2058. 

Coulson, S.J., Hodkinson, I.D., Webb, N.R., & Harrison, J.A. (2001) Atractodes pusillus Forster 

1876 and Stenomacrus groenlandicus jussila 1996 (Hymenoptera: Ichneumonidae) new records to 

Svalbard Norway. Norwegian Journal of Zoology, 48, 328. 

Coulson, S.J., Hodkinson, I.D., Webb, N.R., & Harrison, J.A. (2002). Survival of terrestrial soildwelling 

arthropods on and in seawater: implications for trans-oceanic dispersal Functional Ecology, 

16, 353-356. 

Coulson, S.J., Hodkinson, I.D., Webb, N.R., Mikkola, K., Harrison, J.A., & Pedgley, D.E. (2002) 

Aerial colonization of high Arctic islands by invertebrates: the diamondback moth Plutella xylostella 

(Lepidoptera : Yponomeutidae) as a potential indicator species. Diversity and Distribution, 86, 327- 

334. 

Coulson, S.J., Hodkinson, I.D., & Webb, N.R. (2003) Microscale distribution patterns in high Arctic 

soil microarthropod communities: the influence of plant species within the vegetation mosaic. 

Ecography, 26, 801-809. 

Coulson, S.J., Hodkinson, I.D., & Webb, N.R. (2003) Aerial dispersal of invertebrates over a high- 

Arctic glacier foreland: Midtre Lovenbreen Svalbard. Polar Biology, 26, 530-537. 

Coulson, S.J., Hodkinson, I.D., Webb, N.R., & Convey, P. (2003) A high-Arctic population of Pyla 

fusca (Lepidoptera: Pyralidae) on Svalbard? Polar Biology, 26, 283-285. 

Coulson, S.J., Jussila, R., Hodkinson, I.D., & Webb, N.R. (2003) Synodites lineiger Thomson 1893 

(Hymenoptera: Ichneumonidae) new to Svalbard with an updated list of ichneumonid species from 

the archipelago. Norwegian Journal of Entomology, 50, 50-55. 

Coulson, S.J., & Refseth, D. (2004) The terrestrial and freshwater invertebrate fauna of Svalbard 

(and Jan Mayen). In Prestrud P, Strøm H, Goldman HA, editors. A catalogue of the terrestrial and 

marine animals of Svalbard. Skrifter 201. Norwegian Polar Institute, Tromsø. p 57-122. 

Coulson, S.J. (2007) On the occurrence of Oryzaephilus mercator (Fauvel, 1889) (Coleoptera: 

Silvanidae) on Svalbard, Norway. Norwegian Journal of Entomology. 

Dallas, J.F.R., Irvine, R.J., & Halvorsen, O. (2000) DNA evidence that Ostertagia gruehneri and 

Ostertagia arctica (Nematoda: Ostertagiinae) in reindeer from Norway and Svalbard are conspecific. 

International Journal of Parasitology 30, 655-658. 

Dallas, J.F.R., Irvine, R.J., Halvorsen, O., & Albon, S.D. (2000) Identification by polymerase chain 

reaction PCR of Marshallagia marshalli and Ostertagia gruehneri from Svalbard reindeer. 

International Journal of Parasitology, 30, 863-866. 

Dallas, J.F.R., Irvine, R.J., & Halvorsen, O. (2001) DNA evidence that Marshallagia marshalli 

Ransom 1907 and M. occidentalis Ransom 1907 (Nematoda: Ostertagiinae) from Svalbard reindeer 

are conspecific. Systematic Parasitology, 50, 101-103. 

De Jonckheere, J.F. (2006). Isolation and molecular identification of free-living amoebae of the 

genus Naegleria from Arctic and sub-Antarctic regions. European Journal of Protistology, 42, 115- 

123. 

179

387 

388 

389 

390 

391 

392 

393 

394 

395 

396 

397 

398 

399 

400 

401 

402 

403 

404 

405 

406 

407 

408 

409 

410 

411 

412 


Dollery, R., Hodkinson, I.D., & Jónsdóttir, I.S. (2006) Impact of warming and timing of snow melt on 

soil microarthropod assemblages associated with Dryas -dominated plant communities on Svalbard. 

Ecography, 29, 111-119. 

Ehrenroth, B., & Lohm, U. (1967) Faunistic and floristic observations in central Spitsbergen in the 

summer of 1966. Fauna and Flora, 62, 263-278. 

Elven, R. & Elevebakk A. (1996) Vascular plants. In Elvebakk, A. & Prestrud, P. editors. A catalogue 

of Svalbard plants, fungi and cyanobacteria. Skrifter 198, Norwegian Polar Institute, Tromsø. p. 9-55 

Evenset, A., Christensen, G.N., Skotvold, T., Fjeld, E., Schalabach, M., Wartena, E. & Gregor, D. 

(2004) A comparison of the contaminants in two high Arctic lake ecosystems, Bjønøya (Bear Island), 

Norway. The Science of the Total Environment, 318, 125-141. 

Evenset, A., Christensen, G.N. & Kallenborn, R. (2005). Selected chlorobornes, polychlorinated 

napthalenes and brominated flame retardents in Bjørnøya (Bear Island) freshwater biota. 

Environmental Pollution, 136, 419-430. 

Frénot, Y. Chown, S..L, Whinam, J., Selkirk, P.M., Convey, P., Skotnicki, M., Bergstrom, D.M. 

(2005) Biological invasions in the Antarctic: extent, impacts and implications. Biological Reviews, 80 

45-72. 

Glick, P.A. (1939) The distribution of insects, spiders and mites in the air. Technical Bulletin 673, 

USDA, Washington. 

Greenslade, P. (2002) Assessing the risk of exotic Collembola invading subantarctic islands: 

prioritising quarantine management. Pedobiologia, 46, 338-344 

Geest van, G.J., Spierenburg, P., Van Donk, E. & Hessen, D.O. (2007). Daphnia growth rates in 

arctic ponds: limitation by nutrients or carbon? Polar Biology, 30, 235-242. 

Hammar, J. (2000) Cannibals and parasites: conflicting regulators of bimodality in high latitude Arctic 

char Salvelinus alpinus. Oikos, 88, 33-47. 

Hawes, T.C., Couldridge, C.E., Bale, J.S., Worland, M.R., & Convey, P. (2006) Habitat temperature 

and the temporal scaling of cold hardening in the high Arctic collembolan, Hypogastrura tullbergi 

(Schäffer). Ecological Entomology, 31, 450-459. 

Hayward, S.A.L., Worland, M.R., Bale, J.S., & Convey, P. (2000) Temperature and the 

hygropreference of the Arctic Collembolan Onychiurus arcticus and mite Lauroppia translamellata. 

Physiological Entomology, 25, 266-272. 

Henttonen, H., Fuglei, E., Gower, C.N., Haukisalmi, V., Ims, R.A., Niemimaa, J., & Yoccoz, N.G. 

(2001) Echinococcus multilocularis on Svalbard: introduction of an intermediate host has enabled the 

local life-cycle. Parasitology, 12, 547-552. 

Hertzberg, K., Yoccoz, N.G., Ims, R.A., & Leinaas, H.F. (2000) The effects of spatial habitat 

configuration on recruitment growth and population structure in arctic Collembola. Oecologia, 12, 381 

390. 

Hessen, D.O., Borgeraas, J., Kessler, K., & Refseth, U.H. (1999) UV-B susceptibility and 

photoprotection of Arctic Daphnia morphotypes. Polar Research, 18, 345-352. 

Hessen, D.O., & Rukke, N.A. (2000) UV radiation and low calcium as mutual stressors for Daphnia. 

Limnology and Oceanography, 45, 1834-1838. 

Hessen, D.O., Borgeraas J., & Orbaek, J.B. (2002) Responses in pigmentation and anti-oxidant 

expression in Arctic Daphnia along gradients of DOC and UV exposure. Journal of Plankton 

Research, 24, 1009-1017. 

Hessen, D.O., Blomqvist, P., Dahl-Hansen, G., Drakare, S., & Lindstrom, E.S. (2004) Production 

and food web interactions of Arctic freshwater plankton and responses to increased DOC. Archiv für 

Hydrobiologie, 1593, 289-307. 

Hessen, D.O., Rueness, E.K., & Stabell, M. (2004) Circumpolar Analysis of morphological and 

genetic diversity in the Notostracan Lepidurus arcticus. Hydrobiologia, 51, 73-84. 

Hessen, D.O. & Leu, E. (2006) Trophic transfer and trophic modification of fatty acids in high Arctic 

lakes. Freshwater Biology, 51, 1987-1998. 

Hisdal V 1998 Svalbard nature and history. Norwegian Polar Institute. Oslo. 

Hodkinson, I.D. (1999) Species response to global environmental change or why ecophysiological 

models are important: a reply to Davis et al. Journal of Animal Ecology, 68, 1259-1262. 

Hodkinson, I.D. (2003) Metabolic cold adaptation in arthropods: a smaller-scale perspective. 

Functional Ecology, 17, 562-567. 

Hodkinson, I.D. (2004) Aptinothrips rufus Haliday,Thysanoptera and Apamea zeta Treitschke, 

Lepidoptera on Svalbard. Norwegian Journal of Entomology, 51, 127-128. 

Hodkinson, I.D. (2005) Adaptations of invertebrates to terrestrial Arctic environments. Transactions o 

the Royal Norwegian Society of Sciences and Letters, 2005, 1-44. 

Hodkinson, I.D., & Bird, J.M. (1998). Host-specific insect herbivores as sensors of climate change in 

arctic and Alpine environments. Arctic Alpine Research, 30, 78-83. 

180

413 

414 

415 

416 

417 

418 

419 

420 

421 

422 

423 

424 

425 

426 

427 

428 

429 

430 

431 

432 

433 

434 

435 

436 


Hodkinson, I.D., Webb, N.R., Bale, J.S., Coulson, S.J. & Strathdee A.T. (1998) Global change and 

high arctic ecosystems: conclusions and predictions from experiments with terrestrial invertebrates o 

Spitsbergen. Arctic and Alpine Research, 30, 306-313. 

Hodkinson, I.D., Coulson, S.J., Harrison, J., & Webb, N.R. (2001) What a wonderful web they 

weave: spiders nutrient capture and early ecosystem development in the high Arctic - some counterintuitive 

ideas on community assembly. Oikos, 95, 349-352. 

Hodkinson, I.D., Coulson, S.J., & Bird, J.M. (2002) Discovery of a rare alate morph of Acyrthosiphon 

svalbardicum Heikinheimo (Homoptera; Aphididae) a hitherto 'apterous' aphid species: description 

and implications for species ecology. Norwegian Journal of Entomology, 49, 77-80. 

Hodkinson, I.D., Webb N.R., & Coulson, S.J. (2002) Primary community assembly on land - the 

missing stages: why are the heterotrophic organisms always there first? Journal of Ecology, 90, 569- 

577. 

Hodkinson, I.D., Buhl, P.N., Coulson, S.J., & Webb, N.R. (2003) First record of Dendrocerus 

bifoveatus Kieffer 1907 (Hymenoptera: Ceraphronoidea) from Svalbard. Norwegian Journal of 

Entomology, 50, 105-106. 

Hodkinson, I.D., & Bird, J.M. (2004) Anoxia tolerance in high Arctic terrestrial microarthropods. 

Ecological Entomology, 29, 506-509. 

Hodkinson, I.D., Bird, J.M., Cooper, E.J., & Coulson, S.J. (2004a) The sexual morphs of the 

endemic Svalbard aphid Acyrthosiphon calvulus Ossiannilsson with notes on species biology. 

Norwegian Journal of Entomology, 51, 131-135. 

Hodkinson, I.D., & Coulson, S.J. (2004) Are high Arctic terrestrial food chains really that simple? The 

Bear Island food web revisited. Oikos, 106, 427-431. 

Hodkinson, I.D., Coulson, S.J., & Webb, N.R. (2004b) Invertebrate community assembly along 

proglacial chronosequences in the high Arctic. Journal of Animal Ecology, 73, 556-568. 

Hoel, A. (1967) Svalbard. Svalbards Historie 1596 – 1965. Sverre Kildahl’s Boktrykkeri, Oslo 

Hoffmann, K.H. (1976) Organic body constituents of Protophormia terrae novae Dipt. from 

Spitsbergen compared with flies from a laboratory stock. Oecologia, 23, 13-16. 

Holmgren, A.E. (1869). Bidrag till kannedomen om Beeren Eilands och Spetsbergens insekt fauna. 

Kungliga Svenska Vetenskapsakademiens Handlingar, 8, 1-56. 

Holmstrup, M. (2002) Strategies for cold and drought tolerance in permeable soil invertebrates. 

Doctor’s dissertations (Dsc). National Environmental Research Institute, Silkeborg, Denmark. 196 pp 

Holmstrup, M., Bayley, M., & Ramlov, H. (2002) Supercool or dehydrate? An experimental analysis 

of overwintering strategies in small permeable arctic invertebrates. Proceedings of the National 

Academy of Sciences USA, 99, 5716-5720. 

Hoset, K.S., Espmark, Y., Moksnes, A.., Haugan, T., Ingebrigtsen, M. & Lier, M. (2004) Effect of 

ambient temperature on food provisioning and reproductive success in snow buntings Plectrophenax 

nivalis in the high arctic. Ardea, 92, 239-246. 

Hoxmark, C. (2000) Temperature responses of egg development and post-embryonic growth in a 

High Arctic collembolan Folsomia quadrioculata (Collembola: Isotomidae). Hovedoppgave i biologi 

(Cand.scient.) University of Oslo, Oslo. p 26 

Ims, R.A., Leinaas H.P., & Coulson, S.J. (2004) Spatial and temporal variation in patch occupancy 

and population density in a model system of an arctic Collembola species assemblage. Oikos, 105, 

89-100. 

Irvine, R.J. (2000) Use of moxidectin treatment in the investigation of abomasal nematodiasis in wild 

reindeer Rangifer tarandus platyrhynchus. Veternary Record, 147, 570-573. 

Irvine, R.J., Stien, A., Halvorsen, O., Langvatn, R., & Albon, S.D. (2000) Life-history strategies and 

population dynamics of abomasal nematodes in Svalbard reindeer Rangifer tarandus platyrhynchus. 

Parasitology, 120, 297-311. 

Irvine, R.J., Stien, A., Dallas, J.F., Halvorsen, O., Langvatn, R., & Albon, S.D. (2001) Contrasting 

regulation of fecundity in two abomasal nematodes of Svalbard reindeer Rangifer tarandus 

platyrhynchus. Parasitology, 122, 673-681. 

Jensen, D.B., & Christensen, K.D. (2003) Species Diversity in Greenland. In The biodiversity of 

Greenland – a country study. Technical Report 55. Grønlands Naturinstitut, Nuuk. p 41-96. 

Jensen, T.C., Leinaas, H.P. & Hessen, D.O. (2006). Age dependent shift in response to food 

element composition in Collembola: contrasting effects of dietary nitrogen. Oecologia, 149, 583-592 

Jeppesen, E., Jensen, J.P., Jensen, C., Faafeng, B., Hessen, D.O., Sondergaard, M., Lauridsen, T., 

Brettum, P., & Christoffersen, K. (2003) The impact of nutrient state and lake depth on top-down 

control in the pelagic zone of lakes: A study of 466 lakes from the temperate zone to the arctic. 

Ecosystems, 64, 313-325. 

Johansen. S. (1999) Origin of driftwood in North Norway and its relevance for transport routes of drift 

ice and pollution to the Barents Sea. Science of the Total Environment, 231, 201-225. 

181

437 

438 

439 

440 

441 

442 

443 

444 

445 

446 

447 

448 

449 

450 

451 

452 

453 

454 

455 

456 

457 

458 

459 

460 

461 

462 


Karppinen, E., & Krivolutsky, D.A. (1982) List of oribatid mites (Acarina, Oribatei) of northern 

palaearctic region. I. Europe. Acta Entomologica Fennici, 41, 1-18. 

Kruse, H., Kirkemo, A.M., & Handeland, K. (2004) Wildlife as source of zoonotic infections. 

Emerging Infectious Diseases, 10, 2067-2072. 

Langvatn, R., Albon, S.D., Irvine, R.J., Halvorsen, O., & Ropstad, E. (1999) Parasitter kondisjon og 

reproduksjon hos svalbardrein. In Bengtson SA, Mehlum F, Severinsen T, editors. Terrestrisk 

økologisk forskningsprogram på Svalbard, Svalbardtundraens Økologi. Meddelelser 150 in Tromsø 

Norsk Polarinstitutt. p 139-148. 

Laybourn-Parry, J. & Marshall, W.A. (2003) Photosynthesis mixotrophy and microbial plankton 

dynamics in two high Arctic lakes during summer. Polar Biology, 26, 17-524 

Lebedeva, N.V. & Krivolutsky, D.A. (2003) Birds spread soil microarthropods to Arctic Islands. 

Doklady Biological Sciences, 391, 392-332. 

Lebedeva, N.V., Lebedev, V.D. and Lelekhina, E.N. (2006) New data on the oribatid mite (Oribatei) 

fauna of Svalbard. Doklady Biological Sciences, 407, 182-186. 

Leinaas, H.P. (1999) Spredning og kolonisering Betydning av habitatheterogenitet og 

populasjonsdynamikk. In Bengtson SA, Mehlum F, Severinsen T, editors. Terrestrisk økologisk 

forskningsprogram på Svalbard, Svalbardtundraens Økologi. Meddelelser 150 in Tromsø Norsk 

Polarinstitutt. p 175-180. 

Leinaas, H.P. (1999) Collemboler i arktisk. In Bengtson SA, Mehlum F, Severinsen T, editors. 

Terrestrisk økologisk forskningsprogram på Svalbard, Svalbardtundraens Økologi. Meddelelser 150 

in Tromsø Norsk Polarinstitutt. p 55-66. 

Leinaas, H.P. (1999). The springtail Hypogastrura tullbergi. Arctic Insect News, 10, 4-6 

Lengersdorf, F. (1935) Neosciara delicata nov spec. Norsk Entomology Tidskrift, 4,75. 

Lindqvist, E. (1961) To the knowledge of some sawflies (Hymenoptera: Tenthredinidae). Notulae 

Entomologica, 41, 8-15. 

Lods-Crozet, B.V., Lencioni, V., Olafsson, J.S., Snook, D.L., Velle, G., Brittain, J.E., Castella, E., & 

Rossaro, B. (2001) Chironomid (Diptera: Chironomidae) communities in six European glacier-fed 

streams. Freshwater Biology 46, 1791-1809. 

McGeoch, M.A,, Le Roux, P.C., Hugo, E.A. & Chown, S.L. (2006) Species and community 

responses to short-term climate manipulation: Microarthropods in the sub-Antarctic. Austral Ecology 

31, 719-731 

Makarova, O.L.. (2000a) To studing mites of the genus Arctoseius Thor (Parasitiformes: Ascidae) on 

the Far North 1. Morphometric analysis of the taxonomical features of the arctic species and the 

description of Arctoseius tschernovi sp n and A. miranalis sp n. Zoologichesky Zhurnal, 79, 800-817 

Makarova, O.L. (2000b) To studying mites of the genus Arctoseius (Parasitiformes: Ascidae) from 

the far north 2. Description of Arctoseius productus sp n and A. babenkoi sp n and the keys to 

identifying High Arctic species. Zoologichesky Zhurnal, 79, 907-917. 

Makarova, O.L.. (2000c) To studying mites of the genus Arctoseius (Parasitiformes: Ascidae) from 

the Far North 3. Species areas and ecological preference. Zoologichesky Zhurnal, 79, 1045-1052. 

Marshall, D.J., & Convey, P. (2004) Latitudinal variation in habitat specificity of ameronothrid mites 

Oribatida. Experimental and Applied Acarology, 34, 21-35. 

Mehlum, F. (1999) Tilpasninger hos arktiske organismer til kort sommersesong Tiden som 

begrensende faktor. In Bengtson SA, Mehlum F, Severinsen T, editors. Terrestrisk økologisk 

forskningsprogram på Svalbard, Svalbardtundraens Økologi. Meddelelser 150 in Tromsø Norsk 

Polarinstitutt. p 161-170. 

Mikkola, K. & Goater, B. 1988. The taxonomic status of Apamea exulis Duponchel and A. assimilis 

Doubleday in relation to A. maillardi Geyer and A. zeta Treitschke (Lepidoptera: Noctuidae). 

Entomologists Gazette, 39, 117-257. 

Milner, A.M., Brittain, J.E., Castella, E., & Petts, G.E. (2001) Trends of macroinvertebrate community 

structure in glacier-fed rivers in relation to environmental conditions: a synthesis. Freshwater Biology 

46, 1833-1847. 

Monson, F. Unpublished data 

Moore, P.D. (2002) Biogeography - Springboards for springtails. Nature, 418, 381-381. 

Munch, H. (1962) Neue brutvogel der inselgruppe Sptzbergen. Sterna, 5, 80-88. 

Myl'nikov, A.P. (2002) New amoeboid flagellates of the genus Cercomonas (Cercomonadida: 

Protozoa) from lakes of Spitzbergen Island. Zoologichesky Zhurnal, 8110, 1187-1192. 

Nathan, R. (2006) Long-distance dispersal of plants. Science, 313 786-788. 

Petrova, A.D. & Makorava, O.L. (1991) Arctoseius tajmyricus - a new species of gamasid mites 

(Mesostigmata: Aceosejidae) phoretic on trichocerid flies, Diptera. Entomological Review, 70, 139- 

142. 

182

463 

464 

465 

466 

467 

468 

469 

470 

471 

472 

473 

474 

475 

476 

477 

478 

479 

480 

481 

482 

483 

484 

485 


Petz, W. (2003) Ciliate biodiversity in Antarctic and Arctic freshwater habitats - a bipolar comparison 

European Journal of Protistology, 39, 491-494. 

Pryzbylak, R. (2003) The climate of the Arctic. Kluwer Academic Publishers, Dordrecht, 270pp. 

Remmert, H., & Wisniews, W. (1970) Low resistance to cold of polar animals in summer. Oecologia, 

41, 111-112. 

Røen, U. (1981) The Branchiopoda Crustacea of three high-arctic areas: Ellesmere Island NWT 

Canada Greenland north of 74°N and Svalbard. Verhandlungen Internationale Vereinigung für 

theoretische und angewandte Limnologie, 21, 1541-1544. 

Rose, M.D. & Polis, G.A. (2000) On the insularity of islands. Ecography, 23, 693-701 . 

Sagerup, K., Henriksen, E.O., Skorping, A., Skaares, J.U. & Gabrielsen, G.W. (2000). Intensity of 

parasitic nematodes increases with organochlorine levels in the glaucous gull. Journal of Applied 

Ecology, 37, 532-539. 

Säwström, C., Mumford, P., Marshall, W., Hodson, A., & Laybourn-Parry, J. (2002) The microbial 

communities and primary productivity of cryoconite holes in an Arctic glacier Svalbard 79°N. Polar 

Biology, 25, 591-596. 

Sendstad, E. (1980) Accelerated biodegradation of crude oil on arctic shorelines. Proceedings of the 

Arctic Marine Oil Spill Program Technical Seminar June 3-5. 

Sendstad, E., Sveum, P., Endal, L.J., Brattbakk, I., & Rønning, O.I. (1984). Studies on a seven years 

old seashore crude oil spill on Spitsbergen Proceedings of the Seventh Annual Arctic Marine Oilspill 

Program Technical Seminar June 12-14 1984 Edmonton Alberta p 60-74. 

Serbezov, D. (2002). Thermal responses in life history traits of a cosmopolitan Collembola 

(Hypogastrura viatica) from a high arctic site. Cand. Scient. thesis University of Oslo. 35pp. 

Sinclair, B.J., Vernon, P., Klok, C.J., & Chown, S.L. (2003) Insects at low temperatures: an 

ecological perspective. Trends in Evolution and Ecology, 18, 257-262. 

Skidmore, P. (1997) Zoogeographical notes on the muscid fauna of Greenland and the North 

Atlantic. In Ashworth AC, Buckland PC, Sadler JP, editors. Studies in Quaternary Entomology - An 

Inordinate Fondness for Insects. Quaternary Proceedings 5. John Wiley & Sons, Chichester, UK. p 

247-253. 

Smol, J.P., Wolfe, A.P., Birks, H.J.B., Douglas, M.S.V., Jones, V.J., Korhola, A., Pienitz, R., Ru, K., 

Sorvari, S., Antoniades, D., Brooks, S.J., Fallu, M.A., Hughes, M., Keatley, B.E., Laing, T.E., 

Michelutti, N., Nazarova, L., Nymai, M., Paterson, A.M., Perren, B., Quinlan, R., Rautio, M., Saulnier 

Talbot, E., Siitonen, S., Solovieva, N., & Weckstrom, J. (2005) Climate-driven regime shifts in the 

biological communities of arctic lakes. Proceedings of the National Academy of Sciences USA, 102, 

4397–4402. 

Sømme, L. (1996) Anhydrobiosis and cold tolerance in tardigrades. European Journal of 

Entomology, 93, 349-357. 

Sømme, L. (1999) Cold adaptations in terrestrial invertebrates. In Margesin R. Scinner F, editors. 

Cold-adapted organisms. Ecology physiology enzymology and molecular biology, Springer, Berlin. p 

137-164. 

Sømme, L. (1999) Jordbunnsfaunaens populasjonsdynamikk og tilpasning i arktiske omgivelser. In 

Bengtson SA, Mehlum F, Severinsen T, editors. Terrestrisk økologisk forskningsprogram på 

Svalbard, Svalbardtundraens Økologi. Meddelelser 150 in Tromsø Norsk Polarinstitutt. p 47-54. 

Sømme, S. (1934). Contributions to the biology of Norwegian fish food animals. 1. Lepidurus arcticus 

Pallas 1793. Avhandlinger av det Norske Videnskapelig Akademi, 6, 1-36. 

Søvik, G. (2003). Observations on ovoviviparity and mixed-parity mode in arctic populations of 

Ameronothrus lineatus (Acari: Oribatida). Acarologia, 43, 393-398. 

Søvik, G. (2004) The biology and life history of arctic populations of the littoral mite Ameronothrus 

lineatus (Acari: Oribatida). Experimental and Applied Acarology, 34, 3-20. 

Søvik, G., & Leinaas, H.P. (2002) Variation in extraction efficiency between juvenile and adult 

oribatid mites: Ameronothrus lineatus (Oribatida: Acari) in a Macfadyen high-gradient canister 

extractor. Pedobiologia, 46, 34-41. 

Søvik, G., & Leinaas, H.P. (2003) Adult survival and reproduction in an arctic mite Ameronothrus 

lineatus (Acari: Oribatida): effects of temperature and winter cold. Canadian Journal of Zoology, 81, 

1579-1588. 

Søvik, G., & Leinaas, H.P. (2003) Long life cycle and high adult survival in an arctic population of the 

mite Ameronothrus lineatus (Acari: Oribatida) from Svalbard. Polar Biology, 26, 500-508. 

Søvik, G., Leinaas, H.P., Ims, R.A., & Solhøy, T. (2003) Population dynamics and life history of the 

oribatid mite Ameronothrus lineatus (Acari: Oribatida) on the high arctic archipelago of Svalbard. 

Pedobiologia, 47, 257-271. 

183

486 

487 

488 

489 

490 

491 

492 

493 

494 

495 

496 

497 

498 

499 

500 

501 

502 

503 

504 

505 

506 

507 

508 

509 

510 


Stapleton, L.M., Crout, N.M.J., Sawstrom, C., Marshall, W.A., Poulton, P.R., Tye, A.M., & Laybourn- 

Parry, J. (2005) Microbial carbon dynamics in nitrogen amended Arctic tundra soil: Measurement and 

model testing. Soil Biology and Biochemistry, 37, 2088-2098. 

Stevens, M.I., Greenslade, P., Hogg, I.D. & Sunnucks, P. 2006. Southern hemisphere springtails: 

Could any have survived glaciation of Antarctica? Molecular Biology And Evolution, 23, 874-882. 

Stien, A., Irvine, R.J., Langvatn, R., Albon, S.D., & Halvorsen, O. (2002) The population dynamics of 

Ostertagia gruehneri in reindeer: a model for the seasonal and intensity dependent variation in 

nematode fecundity. International Journal of Parasitology, 32:991-996. 

Stien, A., Irvine, R.J., Ropstad, E., Halvorsen, O., Langvatn, R., & Albon, S.D. (2002) The impact of 

gastrointestinal nematodes on wild reindeer: experimental and cross-sectional studies. Journal of 

Animal Ecology, 71, 937-945. 

Stien, A., Dallimer, M., Irvine, R.J., Halvorsen, O., Langvatn, R., Albon, S.D., & Dallas, J.F. (2005) 

Sex ratio variation in gastrointestinal nematodes of Svalbard reindeer; density dependence and 

implications for estimates of species composition. Parasitology, 130, 99-107. 

Svenning, M.A., Klemetsen, A. & Olsen, T. (2007). Habitat and food choice of Arctic charr in 

Linnévatn on Spitsbergen, Svalbard: the first year-round investigation in a high Arctic lake. Ecology 

of Freshwater Fish,16, 70-77. 

Sveum, P., & Sendstadt, E. (1984) Oljeforurensing på Svalbard Økologiske studier av vanlig 

strandtyper. Rapport STF21 A84073. SINTEF. Trondheim. p 28. 

Sveum, P., & Sendstadt, E. (1985) Oil polluted seaweeds in the Arctic: short term effects on 

decomposers and fate of the oil. Proceedings of the Eighth Annual Arctic Marine Oilspill Program 

Technical Seminar June 18-20 1985 Edmonton Alberta. p 59-77 

Vernon, V., Vannier, G., & Trehen, P. (1998) A comparative approach to the entomological diversity 

of polar regions. Acta Oecologica, 19, 303-308. 

Wada, N. (1998) Sun tracking flower movement and seed production of mountain avens Dryas 

octopetela L in the high Arctic Ny-Alesund, Svalbard. N I P R Symposium on Polar Biology 

Proceedings, 11, 128-136. 

Weider, L.J., & Hobæk, A. (2000) Phylogeography and arctic biodiversity: a review. Annals 

Zoologica Fennici, 37, 217-231. 

Worland, M.R., Block, W., & Grubor-Lajsic, G. (2000) Survival of Heleomyza borealis (Diptera: 

Heleomyzidae) larvae down to -60°C. Physiological Entomology, 25, 1-5. 

Worland, M.R., & Block, W. (2003). Desiccation stress at -zero temperatures in polar terrestrial 

arthropods. Journal Insect Physiology 49, 193-203. 

Wuthrich, C. (1991) Landschaftsökologische Umweltforschung: Beiträge zu den Wechselwirkungen 

zwischen biotischen und abiotischen Faktoren im hocharktischen Ökosystem Spitzbergen. Die Erde, 

22, 335-352. 

Wuthrich, C. (1992) Stofftransporte Meer-Land: Vogelklifftundren und ornithogene. Böden Stuttgarte 

Geographische Studien, 117, 177-192. 

Wuthrich, C. (1994) Die biologische Aktivität arktischer Böden mit spezieller Berücksichtigung 

ornithogen eutrophierter Gebiete Spitzbergen und Finnmark- Basel Physiogeographica, 17, 1-222. 

Hawes T.C. 2008 Aeolian fallout on recently deglaciated terrain in the high Arctic. Polar Biology 31, 

295-301 

Barhndorff S., Petersen SO., Loeschecke V, Overgaard J and Holmstrup M. 2007 Differences in 

cold and drought tolerance of high arctic and sub-arctic populations of Megaphorura arctica Tullberg 

1876 (Onychiuridae: Collembola). Cryobiology 55,315-323. 

Tumanov DV 2007 Three new species of Macrobiotus (Eutardigrada, Macrobiotidae, tenuis-group= 

from Tien Shan (Kirghizia and Spitsbergen. J. Limnol. 66(suppl.1), 40-48. 

Hågvar S, Heller K and Greve L. 2007. Lycoriella postconspicua Mohrig 1985 (Sciaridae, Diptera) 

new to Svalbard and records of some other Diptera from Svalbard. Norw. J. Entomol. 54,65-68. 

Kukulin VV and Kukulina MM. 2005. Helminthes of the birds of the Barents Sea, fauna, ecology, 

impact on the hosts. Kola Science Centre, Murmansk Marine Biological Institute. 

Gillespie M, Hodkinson ID, Coope EJ, Bird JM and Jonsdottir IS. 2007 Life history and host-plant 

relationships of the rare endemic Arctic aphid Acyrthosiphon calvulus in a changing environment. 

Entomol. Exp. Appl. 

Eastop V.F. & Blackman R.L. 2005. Some new synonyms in Aphididae (Hemiptera: Sternorrhyncha) 

Zootaxa. 1098, 1-36. 

Hullé M., Bonhomme J., Maurice D. & Simon JC. 2008. Is the life cycle of high arctic aphids adapted 

to climate change? Polar Biology 

Evenest A., Carroll J., Christensen G.N., Kallenborn R., Gregor D. & Gabrielsen GW. 2007. Seabird 

guano is an efficient conveyer of persistent organic pollutants (POPs) to Arctic lake ecosystems. 

Environ. Sci. Technol. 41, 1173-1179. 

184

511 

512 

513 

514 

515 

516 

517 

518 

519 

520 

521 

522 

523 

524 

525 

526 

527 

528 

529 

530 

531 

532 

533 

534 

535 


Petz W., Valbonesi A., Schiftner U., Quesada A. & Ellis-Evans J.C. 2007. Ciliate biogeography in 

Antarctic and Arctic freshwater ecosystems: endemism or global distribution of species? FEMS 

Microbiol. Ecol. 59,396-408 

Geest van G.J., Hessen D.O., Spierenburg P., Dahl-Hansen G.A.P., Christensen G., Faerovig P.J., 

Brehm M., Loonen M.J.J.E. & van Donk. 2007. Goose-mediated nutrient enrichment and planktonic 

grazer control in arctic freshwater ponds. Oecologia 153,653-662. 

Lods-Crozet, B.V., Lencioni, V., Brittain, J.E., Marziali L. & Rossaro B. 2007. Contrasting chironomid 

assemblages in two high Arctic streams on Svalbard. Fund. Appl. Limnol. Archiv für Hydrobiologie 

170,211-222. 

Simon J.-C., Bonhomme J., Blackman R.L., Hullé M., (2008) The winged morph of the high arctic 

aphid, Acyrthosiphon svalbardicum Heikinheimo (Hemiptera: Aphididae): abundance, reproductive 

status and ecological significance. Canadian Entomologist. 140;385-387 

Fuglei E, Stien A, Yoccoz NG 2008 Spatial distribution of Echinococcus multilocularis, Svalbard, 

Norway Emerging Infectious Diseases 14, 73-75 

Petz W., Schiftner U. & Ellis-Evans JC (2006) Biodiversity and quantitative importance of ciliates and 

testate amebae (Protozoa) in the benthos of high Arctic freshwater lakes (Svalbard, Norway). J. 

Eukaryot. Microbiol. 53, 13S (34A). 

Dolnik, OV, Loonen M,J,J,E, 2007 Isospora plectrophenaxia n. sp (Apicomplexa : Eimeriidae), a new 

coccidian parasite found in Snow Bunting (Plectrophenax nivalis) nestlings on Spitsbergen. 

Parasitology Research. 101: 1617-1619. 

Ronowicz M 2005. Species diversity of Arctic gravel beach: case study for species poor habitats. 

Polish Polar Research 26: 287-297 

Vinogradov AV 2006 Zoogeographical peculiarities of Euroasian north continental water-bodies - 

Phylactolaemata and Eurystomata. Linzer biol. Beitr. 38;65-70. 

Bauer R 2002 Comparison of the enchytraeid communities (Oligochaeta) in an Arctic and a subalpine 

area. Natura Jutlandica Occasional Papers. 2002; (2): 25-31 

Wal R van der, Irvine J. Stien, A., Shepherd N, and Albon. 2000. Faecal avoidance and the risk of 

infection by nematodes in a natural population of reindeer. Oecologia-(Berlin). 2000; 124(1): 19-25 

Popiolek M., Szcesna J., Kotusz J., Kuszierz J. and Witkowski A. 2007 The level of infection with 

gastro-intestinal nematodes in Svalbard reindeers from Hornsund area. Polish polar Reserach. 28; 

277-282 

Ávila-Jiménez, M. L., Fjellberg, A. & Coulson, S. J. 2008. First record of Folsomia bisetosella 

Fjellberg, 2005 (Hexapoda, Collembola) from High Arctic islands. Norw. J. Entomol. 55, 129–130. 

Wojtasik B. 2008. Life cycle of Tonnacypris glacialis (Crustacea: Ostracoda). Polish Polar Research 

29;33-34. 

Dariusz J. Gwiazdowicz pers comm. 

Lebedeva N.V. and Lebedev V.D. 2008. Transport of oribatid mites to the polar areas by birds. 

Integrative Acarology. P. 359-367. 

Samchyshyna L., Hansson L.A. and Christoffersen K. 2008 Patterns in the distribution of Arctic 

freshwater zooplankton realated to glaciation history. Polar Biology, 31;1427-1435 

Hammer, M. 1946. The Zoology of East Greenland. Oribatids. Meddelelser om Grønland, 122(1): 1- 

39. 

Rognes, K. (1983b) "Additions to the Norwegian fauna of calyptrate Diptera (Tachinidae, 

Calliphoridae, Muscidae)", Fauna norv. ser. B 88-93, 

Prestrud, K.W.,Dubey, J.P., Asbakk, K., Fuglei, E. and Su, C. (2008) First isolate of Toxoplasma 

gondii from Arctic fox (Vulpes lagopus) from Svalbard. Veterinary Parasitology, 151, 110-114. 

Prestrud, K.W., Asbakk, K., Fuglei, E., Mork, T., Stien, A., Ropstad, E., Tryland, M., Gabrielsen, 

G.W., Lydersen, C., Kovacs, K.M., Loonen, M.J.J.E., Sagerup, K. and Oksanen, A. (2007) 

Serosurvey for Toxoplasma gondii in arctic foxes and possible sources of infection in the high Arctic 

of Svalbard. Veterinary Parasitology. 150:6-12. 

Clark, M.S., Thorne, M.A.S., Purać, J., Grubor- Lajšić, G., Kube M., Reinhardt, R., and Worland, 

M.R. (2007) Surviving extreme polar winters by desiccation: clues from Arctic springtail (Onychiurus 

arcticus) EST libraries BMC Genomics 8:475 

Jonsdottir, I.S. (2005) Terrestrial ecosystems on Svalbard: heterogeneity, complexity and fragility 

from an Arctic Island perspective. Biology and Environment. 105B 155-165 

Hallingback T. (2007) Vilka flugor lurar mossorna pa Svalbard? Fauna och Flora (Stockholm) 102 

Issue: 4 Page(s): 34-35 

Hawes, T.C. (2007) Ballooning in high Arctic linyphiids: a case of behavioural atrophy? Bulletin of the 

British Arachnological Society 14 39-42 

185

536 

537 

538 

539 

540 

541 

542 

543 

544 

545 

546 

547 

548 

549 

550 

551 

552 

553 

554 

555 

556 

557 

558 


Maiolini, B.; Lencioni, V.; Brittain, J.E. (2006) Invertebrate drift in a high Arctic stream under natural 

and experimental light conditions: preliminary results. Internationale Vereinigung fuer Theoretische 

und Angewandte Limnologie Verhandlungen 29, 2201-2204 

Uvarov, A. V. 2004 (2005) Relationship between head capsule length and body length in some 

Svalbard species of Collembola. Russian Entomological Journal 13 107-113 

Gislason, G. M. (2005) Origin of freshwater fauna of the North-Atlantic islands: present distribution in 

relation to climate and possible migration routes. Internationale Vereinigung fuer Theoretische und 

Angewandte Limnologie Verhandlungen 29, 198-203 

Leinaas, H.P. (2002) UV tolerance, pigmentation and life forms in high Arctic Collembola. Ecological 

Studies 153, 123-134 

Van Donk, E.; De Lange, H.J.; Faafeng, B.A. (2001) Impact of UV-radiation on grazing activity of 

Daphnia middendorffiana and use of Chlamydomonas reinhardtii as UV-biodosimeter in an arctic 

freshwater lake (Spitsbergen, Norway). Internationale Vereinigung fuer Theoretische und 

Angewandte Limnologie Verhandlungen 27, 2031-2035 

Hawes T.C., Hines A., Viant M.R., Bale J.S. Worland M.R., Convey P. (2008) Metabolomic 

Fingerprint Of Cryo-Stress In A Freeze Tolerant Insect. CryoLetters 29; 505-515. 

Gwiazdowicz D.J. and Rakowski R. (2009) A redescription of Protolaelaps parvanalis (Thor, 1930) 

(Acari: Ascidae) from Spitsbergen. Entomologica Fennica. 20;281-286. 

Gwiazdowicz D.J. and Gulvik M.E. (2008) Mesostigmatid mites (Acari, Mesostigmata) in Svalbard. 

Oksanen, A, Asbakk, K.; Prestrud, K.W.; Aars, J.; Derocher, A.E.; Tryland, M.; Wiig, O.; Dubey, 

J.P.; Sonne, C.; Dietz, R.; Andersen, M.; Born, E.W. (2009) Prevalence of antibodies against 

Toxoplasma gondii in Polar bears (Ursus maritimus) from Svalbard and East Greenland. J. Parasitol 

95; 89-94 

Stien A, Voutilainen L, Haukisalmi V, Fuglei E, Mork T, Yoccoz NG, Ims RA, Henttonen H. (2010) 

Intestinal parasites of the Arctic fox in relation to the abundance and distribution of intermediate 

hosts. Parasitology. 137; 149-157. 

Prestrud, KW; Asbakk, K; Mork, T; Fuglei, E; Tryland, M; Su, CL (2008) Direct high-resolution 

genotyping of Toxoplasma gondii in arctic foxes (Vulpes lagopus) in the remote arctic Svalbard 

archipelago reveals widespread clonal Type II lineage. Veterinary Parasitology 158;121-128. 

Sorensen, KK; Mork, T; Siguroardottir, OG; Asbakk, K; Akerstedt, J; Bergsjo, B; Fuglei, E (2005) 

Acute toxoplasmosis in three wild arctic foxes (Alopex lagopus) from Svalbard; one with co-infections 

of Salmonella Enteritidis PT1 and Yersinia pseudotuberculosis serotype 2b. Research in Vetinary 

Science. 78;161-167. 

Kaya M., De Smet W.H. and Fontaneto D. (2010) Survey of moss-dwelling bdelloid rotifers from 

middle Arctic Spitsbergen (Svalbard). Polar Biol. 33;833-842. 

Jensen S.M., Aars J., Lydersen C., Kovacs K.M. and Åsbakk K. (2010) The prevalence of 

Toxoplasma gondii in polar bears and their marine mammal prey: evidence for a marine transmission 

pathway? Polar Biol. 33;599-606. 

Åkerstedt J., Lillehaug A., Larsen I.L., Eide N.E., Arnemo J.M. and Kjell Handeland. (2010) 

Serosurvey for canine distemper virus, canine adenovirus Leptospira interrogans and Toxoplasma 

gondii free-ranging canids from Scandinavia and Svalbard. J. Wildlife Disease. 46;474-480. 

Marziali, L. Gozzini M, Lencioni, V (2009) Drift patterns of Chironomidae (Insecta, Diptera) in an 

arctic stream (Svalbard Islands): an experimental approach Studi Trentini di Scienze Naturali. 84; 87 

96. 

Ávila-Jiménez, M. L., Coulson S.J , Solhøy T. & Sjöblom A. (2010) Overwintering of Arctic 

arthropods; the case of the invertebrate fauna of Svalbard now and in the future. Polar Research. 

29;127-137. 

Coulson S.J. (2009) Association of the soil mite Diapterobates notatus (Acari, Oribatidae) with 

Cynomya mortuorum (Linnaeus, 1761) (Calliphoridae, Calliphorinae): implications for the dispersal o 

oribatid mites. International Journal of Acarology. 35;175-177. 

Coulson, S.J., Lorentzen, E., Strøm, H. Gabrielsen G.W. (2009) The parasitic tick Ixodes uriae 

(Acari: Ixodidae) on seabirds from Spitsbergen, Svalbard. Polar Research. 28;399-402. 

Coulson S.J., Monson F., Moe B. and Gabrielsen G.W. (2009) The invertebrate fauna of High Arctic 

seabird nests: the microarthropod community inhabiting nests on Spitsbergen, Svalbard. Polar 

Biology. 32;1041-1046. 

van Geest G.J., Sachse R., Brehm M., van Donk E. & Hessen D.O. 2010 Maximizing growth rate at 

low temperatures: RNA:DNA allocation strategies and life history traits of Arctic and temperate 

Daphnia. Polar Biology 33;1255-1262. 

Stur E. & Ekrem T. 2011 Exploring unknown lifestages of Arctic Tanytarsini (Diptera: Chironomidae) 

with DNA barcoding. Zootaxa 2743;27-39. 

Sørensen, J.G. and Holmstrup,M. (2011) Cryoprotective dehydration is widespread in Arctic 

springtails. J.Insect Physiol 

186

559 

560 

561 

562 

563 

564 

565 

566 

567 

568 

569 

570 

571 

572 

573 

574 

575 

576 

577 

578 

579 

580 

581 

582 


Vestby S.E. 1983 En regional limnologisk undersøkelse av små innsjøer ved Svea. Svalbard. 

Masters project. Institute of Marine Biology and limnology, Department of limnology, University of 

Oslo. 137p. 

Lavikainen A., Laaksonen S., Beckmen K., Oksanen A., Isomursu M. & Meri S. (2011) Molecular 

identification of Taenia spp. In wolves (Canis lupus), brown bears (Ursus arctos) and cervids from 

North Europe and Alaska. Parasitology International 60;289-295. 

Behan-Pelletier V.M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. 

Zootaxa. 2973;1-56. 

Avila-Jimenez M.L. & Coulson S.J. (2011) A Holarctic Biogeographical Analysis of the Collembola 

(Arthropoda, Hexapoda) Unravels Recent Post-Glacial Colonization Patterns. Insects 11;273-296 

Pilskog H.E. (2011) The invertebrate fauna of Svalbard bird nests; ecology and as a facilitating 

colonisation of an Arctic archipelago. University of Bergen and University centre in Svalbard. June 

2011. 

Gwiazdowicz D.J. & Coulson S.J. (2011) High Arctic gamasid mites (Acari, Mesostigmata): 

community composition on Spitsbergen, Svalbard. Polar Reserach 8311 DOI 

10.3402/polar.v30i0.8311 

Unpublished record 

Bengtsson, B.Å. & Johansson, R. 2011. Nationalnyckeln till Sveriges flora och fauna. Fjärilar: 

Bronsmalar-rullvingemalar. Lepidoptera: Roeslerstammiidae-Lyonetiidae. ArtDatabanken, SLU 

Uppsala. 494 pp. 

Bayartogtokh B., Schatz H. & Ekrem T. (2011) Distribution and diversity of the soil mites of Svalbard 

with redescriptions of three known species (Acari: Oribatida). J. Internat. Acarol. 37;467-484. 

Bengtsson and Johansson 2011 

Ávila-Jiménez M.L & Coulson S.J. (2011) Can snow depth predict the distribution of the high Arctic 

aphid Acyrthosiphon svalbardicum (Hemiptera: Aphididae) on Spitsbergen? BMC Ecology 11:25 

Coulson S.J., Ávila-Jiménez M.L., Fjellberg A., Snazell R., & Gwiazdowicz D.J. (2011) On the 

Collembola, Araneae and Gamasida from the Kinnvika region of Nordaustlandet, Svalbard. 

Geografiska Annaler 93 (4); 253-257 

Gwiazdowicz D.G., Teodorowicz E. & Coulson S.J. (2011) Redescription of Arctoseius haarlovi 

Lindquist, 1963 (Acari, Ascidae) from Spitsbergen, Svalbard. Entomologica Fennica. 22;140-148. 

Gwiazdowicz D.G., Teodorowicz E. & Coulson S.J. (2011) Redescription of Zercon solenites 

Haarløv, 1942 (Acari, Zerconidae) with a key to the Svalbard species of the genus Zercon. 

International Journal of Acarology 37(sup1);135-148. 

Gwiazdowicz & S.J. Coulson. (2010) Phoresis of the duetonymphs of Thinoseius spinosus (Acari, 

Eviphididae) on the calliphorid dipteran Protophormia terraenovae on the High Arctic island of 

Spitsbergen (Svalbard) including a key to deutonymphs of Thinoseius. International Journal of 

Acarology 36;233-236 

Gwiazdowicz D.J., S.J.Coulson & M.L.Ávila-Jiménez. (2009) First records of Zercon andrei Sellnick, 

1958 and Zerconopsis muestairi (Schweizer, 1949) (Acari, Mesostigmata) from Bjørnøya. Norwegian 

Journal of Entomology 56, 117–119 

Lindquist, EE & Makarova, OL (2011) Two new circumpolar mite species of the genus Arctoseius 

Thor, (Parasitiformes, Mesostigmata, Ascidae). Zoologichesky Zhurnal. 90;923-941 

Kaczmarek L., Zawierucha K., Smykla J. and Michalczyk L. (2012) Tardigrada of the Revdalen 

(Spitsbergen) with the descriptions of two new species: Bryodelphax parvuspolaris 

(Heterotardigrada) and Isohypsibius coulsoni (Eutardigrada). Polar Biology 35:1013-1026. 

Popovic D., Purac J., Kojic D., Pamer E.L., Worland M.R., Blagojevic D.P. and Grubor-Lajsic G. 

(2011) LEA protein expression during cold indiced dehydration in the Arctic Collembola Megaphorura 

arctica. Arch. Biol. Sci. Belgrade, 63;681-683. 

Chaubert, B., Derocles, S.A.P., Hulle, M., Le Relec, A., Outreman, Y., Simon, J-C. & Tomanovic, Z. 

Two new species of aphid parasitoids (Hyemnoptera, Braconidae, Aphidiinae) from the high arctic 

(Spitsbergen, Svalbard). Zoologischer Anzeiger. 

Gwiazdowicz DJ., Solhøy T., Coulson SJ., Lebedeva NV. And Melekhina EN. First record of 

Vulgarogamasus immanis (Acari, Mesostigmata) in Svalbard. Polish Pol Res 33;35-39 

Coulson et al. Introduction of invertebrates into the High Arctic via imported soils: the case of 

Barentsburg in the Svalbard. Biological Invasions 

Knapp J., Staebler S., Bart J.M., Stien A., Yoccoz N.G., Drogermuller C., Gottstein B., Deplazes P. 

2012. Echinococcus multilocularis in Svalbard, Norway: Microsatellite genotyping to investigate the 

origin of a highly focal contamination. Infection Genetics and Evolution. 12; 1270-1274. 

Davidson R.K., Romig T., Jenkins E., Tryland M. & Robertson L.J. 2012. The impact of globalisation 

on the distribution of Echinococcus multilocularis. Trends in Parasitology 28;239-247. 

187

583 

584 

585 

586 

587 

588 

589 


Davidson R.K., Romig T., Jenkins E., Tryland M. & Robertson L.J. 2012. The impact of globalisation 

on the distribution of Echinococcus multilocularis. Trends in Parasitology 28;239-247. 

Thorne M.A.S., Worland M.R., Feret M.J., Lilley K.S. & Clark M.S. 2012. Proteomics of 

cryoprotective dehydration in Megaphorura arctic Tullberg 1878 (Onychiuridae: Collembola). Insect 

Molecular Biology 2011. 20;303-310. 

Sørensen J.G., Heckmann L-H. & Holmstrup M. 2010. Temporal gene expression profiles in a 

palaearctic springtail as induced by desiccation, cold exposure and during recovery. Func. Ecol. 

24;838-846. 

Carlsson A.M., Irvine R.J., Wilson K., Piertney S.B., Halvorsen O., Coulson S.J., Stein A. & Albon 

S.D. 2012. Disease transmission in an extreme environment: Nematode parasites infect reindeer 

during the Arctic winter. Int. J. Parasitol. 42;789.795 

Zmudczynska K., Olejniczak I., Zwolicki A., Iliszko L., Convey P. & Stempniewicz L. 2012. Influence 

of allochtonous nutrients delivered by colonial seabirds on soil collembolan communities on 

Spitsbergen. Pol. Biol. 35;1233-1245. 

Eide N., Stien A., Prestrud P, Yoccuz N.G. & Fuglei E. 2012. Reproductive responses to spatial and 

temporal prey availability in a coastal Arctic fox population. J. Anim. Ecol. 81;640-648. 

Purac J., Pond D. W., Gruboe-Lajsic G., Kojic D., Blagojevic D.P., Worland M.R. & Clark M.S. 2012. 

Cold hardening induces transfer of fatty acids between polar and nonpolar lipid pools in the Arctic 

collembolan Megaphorura arctica. Physiol. Entomol. 2011. 36;135-140. 

188

Checklist of the Terrestrial and freshwater invertebrate fauna - Unis

Create successful ePaper yourself

Delete template?

Save as template?