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Malawi. Nyika National Park, without locality or collecting details (Mill<br />

1979); Mwanda Mt., 2000, Winter 4159 (MAL).<br />

Lippia javanica (Burm.f.) Spreng.<br />

Lippia whytei Mold.<br />

Chichewa: chanzi, vumba; Yao: chisunganjeu, mchenjema<br />

Shrub to 4 m tall, softly woody; leaves strongly aromatic and slightly unpleasant,<br />

opposite or in whorls of 3, 10–100 x 3–45 mm, scabrid and<br />

rugulose (rough) above, serrulate; flowers in a small compact spike, white<br />

to yellowish green; in woodland, scrubby grassland, rocky hillsides; 450–<br />

2,350 m. Widespread in Africa, from South Africa to Angola and Ethiopia.<br />

Not in Asia, despite the species name, which was applied erroneously.<br />

Source. F.T.E.A. Verbenaceae: 30 (1992).<br />

Malawi. Nyika National Park, without locality or collecting details (Mill<br />

1979).<br />

Lippia plicata Baker <br />

Lippia strobiliformis Mold.<br />

Woody herb or shrub 0.6–3 m, stems square, brown to reddish purple;<br />

leaves aromatic (minty), opposite, rarely in threes, ovate to lanceolate, 15–<br />

100 x 7–55 mm, harshly scabrid above, margins crenate-serrate; flowers in<br />

Rinorea convallarioides (Baker f.) Eyles <br />

Understorey shrub or small tree 1.5–7 m tall; leaves simple, alternate, oblong-elliptic,<br />

40–70 x 15–40 mm, sharply serrate or subentire, hairless except<br />

on main veins, petiole 2–4 mm long, channelled above, stipules 3–<br />

4 mm long; flowers axillary; fruit a 3-sided capsule to 13 mm long; in evergreen<br />

forest; only in submontane forest (below 2,000 m) on the eastern<br />

escarpment, but not recorded by Dowsett-Lemaire (1985). Zimbabwe,<br />

Mozambique, and Malawi.<br />

Source. Chapman & White (1970: 126).<br />

Malawi. Nyika Plateau, “eastern escarpment Forests”, no specimen located.<br />

Viscum combreticola Engl. <br />

Relatively large dioecious leafless parasitic shrubs, more or less pendulous<br />

with age, yellowish green, stems flattened and strongly ribbed, but round<br />

with age; berries stalkless, more or less round, 6–7 mm long, warty when<br />

young, almost smooth at maturity, orange; a parasite of various hosts,<br />

but commonly on Combretum and various legumes, including<br />

Brachystegia. Widespread from D.R.C. and Kenya to Angola and South<br />

Africa.<br />

Source. Mill (1979); Polhill & Wiens (1998: 303).<br />

Malawi. Nyika National Park, Chisanga Falls, 1,800 m, 1982, Dowsett-<br />

Lemaire 307 (K); Chisanga Falls, 1,800 m, 1986, R.M. Polhill & D. Polhill<br />

5254 (K, MAL).<br />

Viscum cylindricum Polhill & Wiens.<br />

Leafless, dioecious, parasitic shrub with pendulous branches up to 1–2 m<br />

long, distal branches flattened, 2–4 mm broad; flowers sessile, single, style<br />

indistinct, stigma on elevated mound above tepal scars, berries sessile, oblong,<br />

7–3.5 mm; in forest and high-rainfall Brachystegia woodland on various<br />

hosts. Only known from Malawi.<br />

Source. Polhill & Wiens (1998: 303); Polhill (2001: 593, fig. 211G).<br />

Malawi. Nyika Plateau, 7 km W. of Chelinda, 1972, Wiens 4614 (K, UT);<br />

Nyika National Park, 6.5 km W. of Livingstonia junction, Nya Mkowa,<br />

2,000 m, 1978, Pawek 13846 (BR, K, MAL, MO, WAG).<br />

Viscum goetzei Engl.<br />

Leafless dioecious parasite with flattened stems narrowing to the nodes;<br />

berries with a 1 mm long stalk, 3 mm long, smooth, orange; a parasite<br />

(epiparasite) of Loranthaceae, in evergreen forests to 2,350 m. Southern<br />

Highlands of Tanzania, Nyika Plateau, and Eastern Highlands of Zimbabwe.<br />

Source. Polhill & Wiens (1998: 302); Polhill (2001: 593, fig. 211J, K).<br />

VIOLACEAE<br />

VISCACEAE<br />

roundish terminal clusters, white/yellow to pink or lilac, scented; in grassland<br />

and various types of woodland, often in disturbed areas; 450–1,950 m.<br />

Mozambique, Malawi, Zambia, Angola, D.R.C., and Tanzania.<br />

Source. Brummitt (1973); F.T.E.A. Verbenaceae: 34 (1992).<br />

Malawi. Nyika National Park, N.W. foot of Nganda, 1972, Brummitt &<br />

Synge 47 (K, MAL, SRGH, FHO, LISC); W. of Vitinthiza Hill, 2000, Salubeni<br />

& Mwanyambo 6763 & 6806 (MAL).<br />

Lippia woodii Mold.<br />

Lippia africana Mold. var. villosa Mold.; Lippia wilmsii Pearson var.<br />

villosa (Mold.) Mold.<br />

Herb or shrublet 150–600 mm tall, with several unbranched stems; leaves<br />

strongly aromatic, opposite, obovate-elliptic to rhomboid, 10–80 x 10–<br />

30 mm, shallowly crenulate, rugose/bullate, silvery hairy below, sparsely<br />

hairy above; flowers in axillary clusters, white, green or yellow, usually<br />

with an orange centre; in open woodland or grassland subjected to frequent<br />

burning; 1,110–2,280 m. Zimbabwe and Mozambique northwards<br />

to Uganda.<br />

Source. F.T.E.A. Verbenaceae: 33 (1992).<br />

Malawi. Nyika National Park, without locality or collecting details (Mill<br />

1979).<br />

Viola abyssinica Steud. ex Oliv. <br />

Chichewa: katongolola<br />

Perennial herb, stems to 600 mm long, trailing or creeping, rooting at nodes,<br />

angular or winged; leaves broadly ovate to roundish, base cordate, margins<br />

crenate-dentate; flowers single, axillary, on pedicels 10–40 mm long,<br />

white to bluish or violet; in moist grassland, moist areas in forest or forest<br />

margins; 1,200–2,520 m. Widespread in the mountains of tropical Africa.<br />

Source. Flora zambesiaca 1(1): 258 (1960).<br />

Malawi. Nyika National Park, without locality, 1946, Brass 17323 (BM, K,<br />

PRE, SRGH); Mpopoti, 2000, Mwanyambo 542 (MAL).<br />

Zambia. Nyika Plateau, near Rest House, 1958, Robson 395 (BM, K, SRGH).<br />

Malawi. Nyika National Park, vicinity of Juniper Forest, 2,190 m, 1972,<br />

Wiens 4612 (K, MO, P).<br />

Viscum loranthicola Polhill & Wiens<br />

Small, leafless, densely branched parasitic shrub, distal internodes rounded<br />

to slightly flattened; flowers sessile, male flowers with anthers fused together;<br />

berries orange, sessile, round, smooth, 3 mm diameter; epiparasite<br />

on several genera of Loranthaceae. Burundi, Tanzania, Zambia, and northern<br />

Malawi.<br />

Source. Polhill & Wiens (1998: 297);<br />

Malawi. Rumphi-Chisenga Road, half a km S. of Nyika Road junction,<br />

1,500 m, 1972, Wiens 4607 (K, MAL, MO, UT); 16 km E. of Barrier, 1,650 m,<br />

1976, Phillips 1338 (K, MO, SRGH, WAG).<br />

Viscum shirense Sprague<br />

Viscum junodii Engl., nom. nud.<br />

Leafless dioecious parasite, pendulous with age, stems flattened; berries<br />

with a stalk of 1–1.5 mm, round to ellipsoid, 3–5 mm long, smooth, orange;<br />

a parasite of various hosts, commonly on Bridelia, in high rainfall<br />

woodland, riverine forest, and on margins of montane forest; up to 1,700 m.<br />

D.R.C., Tanzania, Malawi, Mozambique, Zambia, and Zimbabwe. The<br />

citations given below might be misidentifications for allied species, but V.<br />

shirense is certainly recorded from near Chisenga on the Chambo R. and<br />

around Wenya. Furthermore, the closely similar V. congdonii Polhill &<br />

Wiens, known from Jembya Forest Reserve, is likely to occur. Differences<br />

are given in the sources cited above.<br />

Source. Polhill & Wiens (1998: 306); Polhill (2001: 595).<br />

Malawi. Nyika National Park, Kasyaula & Zovochipolo Forests (Dowsett-<br />

Lemaire 1985).<br />

Zambia. Nyika National Park, Manyenjere, Kasoma & Chowo Forests<br />

(Dowsett-Lemaire 1985).<br />

Plants of the Nyika Plateau<br />

283<br />

VISCACEAE

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