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CYPERACEAE<br />

Malawi. Nyika National Park, Nganda, 1972, Synge 368 (BR, EA, K, LISC,<br />

MAL, P, PRE, SRGH, UPS).<br />

Carex aethiopica Schkur.<br />

Perennial, tufted and leafy; leaves often red at base; inflorescence several<br />

erect spikes from upper nodes of culm, glumes reddish brown; along forest<br />

margins near watercourses. Occurring mainly in South Africa.<br />

Malawi. Nyika National Park, Juniper Forest area, 2000, Smook 10917 (PRE,<br />

MAL).<br />

Carex brassii Nelmes<br />

Perennial, tufted and leafy; inflorescence a slender panicle, glumes light<br />

brown; on stream banks in dense shade. Main known locality is Mt.<br />

Mulanje, Malawi. Closely related to C. spicato-paniculata.<br />

Malawi. Nyika National Park. without locality or collecting details (Patel<br />

1999).<br />

Carex chlorosaccus C.B.Clarke<br />

Carex echinochloe Kunze var. chlorosaccus (C.B.Clarke) Kük.<br />

Perennial, tufted and leafy, rhizome woody; culms obtusely triquetrous;<br />

inflorescence a slender, much-branched panicle; spikelet 6–20 mm long; in<br />

moist forests and along paths and roads. Widespread, but mainly E. Africa<br />

to Ethiopia.<br />

Malawi. Nyika National Park, Juniper Forest, 2000, Smook 10904 (MAL, PRE).<br />

Carex cognata Kunth var. congolensis (Turrill) Lye<br />

Carex congolensis Turrill; Carex pseudo-sphaerogyna Nelmes<br />

Perennial, tufted with small groups of culms arising together from short<br />

scaly stolons; inflorescence of 4–6 erect crowded spikes; in medium-altitude<br />

swamps. Zambia, Uganda, Rwanda, and D.R.C. Very close to var.<br />

cognata.<br />

No country given. Nyika Plateau, 1967, Richards 22474 (UZL).<br />

Carex conferta Hochst. ex A.Rich. var. lycurus (K.Schum.)<br />

Lye<br />

Carex lycurus K.Schum.<br />

Perennial, robust, rhizome thick and creeping, roots pale; culm triangular;<br />

leaf 5–12 mm wide; inflorescence dense panicle; spikelet 5–10 mm long;<br />

upland swamps, along streams, and near lakes. N. Malawi, Tanzania, and<br />

Kenya; possibly also in Zimbabwe.<br />

Malawi. Nyika National Park, without locality or collecting details (Patel<br />

1999).<br />

No country given. 1967, Richards 22455 (UZL).<br />

Carex echinochloe Kunze<br />

Perennial, tufted, leafy, rhizomatous; inflorescence a panicle; in open forest<br />

and woodland. Zimbabwe, Malawi, Tanzania, Kenya, and D.R.C.<br />

Malawi. Nyika National Park. without locality or collecting details (Patel 1999).<br />

Carex petitiana A.Rich.<br />

Carex fischeri K.Schum.; Carex cuprea (Kük.) Nelmes; Carex<br />

ninagongensis (Kük.) Robyns & Tournay<br />

Perennial, tufted or shortly creeping; culms bluntly triangular; inflorescence<br />

of 5–8 erect or drooping dense spikes, arising singly from leaf sheaths; in<br />

wet habitats along paths and streams. Ethiopia, E. Africa, Rwanda, and<br />

D.R.C. down to Zimbabwe; also in Nigeria and Cameroon.<br />

No country given. 1959, Robinson 3028 (UZL).<br />

Carex spicato-paniculata C.B.Clarke <br />

Perennial, tufted, rhizome thick; culms triangular; inflorescence of 1–2 pyramidal<br />

panicles from each of the upper leaf sheaths; spikelet 5–10 mm<br />

long; open to dense forest. Ethiopia, S.E. tropical Africa, and South Africa.<br />

Malawi. Nyika National Park, Thazima Gate, 2000, Salubeni & Mwanyambo<br />

6759 (MAL); Sangule Kopje area, 2000, Smook 10855 (PRE, MAL, UZL,<br />

SRGH, LMA); Mbuzinandi, 2000, Salubeni & Mwanyambo 6818 (MAL).<br />

Coleochloa setifera (Ridley) Gilly <br />

Densely tufted perennial grass-like plant; leaves convolute-cylindrical, 0.5–<br />

2.5 mm wide when flattened, upper surface glabrous, lower surface hairy;<br />

298 Plants of the Nyika Plateau<br />

inflorescence a lax panicle of 1–3 fascicles of spikes, spikelets 2–3 mm long;<br />

forming dense clumps on bare rock surfaces, often very common. Widespread<br />

in eastern tropical Africa from South Africa to Kenya; also in Madagascar.<br />

Malawi. Nyika National Park, Fingira Rock, 2000, Winter 4211 (PRE).<br />

Courtoisina cyperoides (Roxb.) Soják <br />

Courtoisina cyperoides (Roxb.) Nees<br />

Tufted annual, has a ‘fenugreek’ odour; inflorescence with many almost<br />

spherical spikes; spikelets 3.8–4.2 mm long, glumes yellowish grey with a<br />

green, strongly winged midrib; in seasonally wet grassland and temporary<br />

pools. Southern Africa, Mozambique, Angola, Zimbabwe, Zambia, and<br />

Tanzania.<br />

Malawi. Nyika National Park, 2.5 km from Thazima Gate, 2000, Burrows<br />

& Maroyi 6816 (MAL, PRE).<br />

Cyperus alopecuroides Rottb.<br />

Juncellus alopecuroides (Rottb.) C.B.Clarke<br />

Robust perennial; culms triangular; basal leaves crowded; inflorescence<br />

umbel-like with 1–many clusters of spikes; spikelet 2.5–8.0 mm long; in<br />

swamps and other wet areas, usually in standing water. Widespread in southern<br />

and tropical Africa; also in Madagascar, India, and Australia.<br />

Malawi. Nyika National Park, without locality or collecting details (Patel<br />

1999).<br />

Cyperus amabilis Vahl<br />

Annual, slender to robust; culms solitary or crowded; leaf-sheaths short,<br />

reddish brown to purple; inflorescence orange-brown; in seasonally wet<br />

habitats, often in sandy soils near roads, lakes, and swamps. Widespread in<br />

tropical and South Africa; also in Madagascar, India, and S. America.<br />

Malawi. Nyika National Park, Chosi Hill, 2000, Burrows & Burrows 6739<br />

(MAL, PRE).<br />

Cyperus angolensis Boeck. <br />

Tall perennial, stoloniferous with solitary culms from swollen culm-bases;<br />

inflorescence globose, dirty white; dry grassland with scrub and scattered<br />

trees, often in places where burning is frequent. Common in tropical Africa,<br />

less frequent in southern Africa.<br />

Source. Brummitt (1973).<br />

Malawi. Nyika National Park, plateau above upper Mondwe Valley, 1972,<br />

Synge 396 (K, MAL).<br />

Cyperus denudatus L.f. var. lucenti-nigricans (K.Schum.)<br />

Kük.<br />

Perennial; culms sharply triangular to almost winged; inflorescence more<br />

usually tightly congested; spikelets blackish; in upland swamps and wet<br />

grasslands. Widespread in tropical Africa.<br />

Malawi. Nyika National Park, near Juniper Forest, 2000, Smook 10913<br />

(PRE, MAL, UZL, SRGH, LMA).<br />

Cyperus distans L.f.<br />

Tufted perennial, rhizome short, thick; culms in a row or solitary; basal<br />

parts covered with leaf-sheaths; inflorescence compound umbel-like, at the<br />

base enclosed by a greenish to purplish tubular prophyll ending in two<br />

teeth or short blades; spikelet 7–20 mm long; wettish grasslands, streamsides,<br />

edges of pools and ditches, roadside banks, and in cultivation. Tropical<br />

and South Africa, Madagascar; Asia, Philippines, and Australia.<br />

Malawi. Nyika National Park, Dembo bridge, 2000, Thera 3024 (MAL).<br />

Cyperus esculentus L. var. esculentus<br />

Perennial, robust, stoloniferous with root tubers; inflorescence of one sessile<br />

and 3–10 stalked spikes; spikelets 5–20 mm long, obtuse, brown or<br />

rust coloured; in seasonally wet grassland and swamps; also a weed of<br />

cultivation. Cosmopolitan, but in warmer areas.<br />

Malawi. Nyika National Park, Thazima Gate, 2000, Burrows & Maroyi 6819<br />

(MAL, PRE).<br />

Cyperus exaltatus Retz.<br />

?Cyperus immensus C.B.Clarke var. taylorii C.B.Clarke

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