Geology and Paleontology of Canal Zone and Adjoining ... - USGS

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GASTROPODS: EULIMIDAE, MARGINELLIDAE TO HELMINTHOGLYPTIDAE 377 higher and wider than others, forming a narrow hump. Outer lip broken back. Upper limb of anal sinus heavily bordered. Aperture moderately long. Height (almost complete) 27.6 mm, diameter (in cluding hump) 10.3 mm. An unnamed clavine turrid from the Chagres sand stone is represented by an imperfect specimen. The entire apertural face of the shell and part of the ad- apertural face are corroded. The relatively narrow axial ribs and strongly constricted, weakly sculptured anal fascicle are distinctive. This species evidently is new and has no known close allies. Occurrence: Chagres sandstone (late Miocene), locality 208. Genus Darbya Bartsch? Bartsch, Smithsonian Misc. Colln., v. 91, no. 2, p. 22, 1934. Type (orthotype): Darbya lira Bartsch, living off north coast of Puerto Rico. Submenus Buridrillia Olsson? Olsson, Bull. Am. Paleontology, v. 27, no. 106, p. 51, 1942. Type (orthotype): ClatUrodrillla panarica Olsson, Pliocene, Pacific coast of Central America straddling Costa Rica- Panama boundary. Though the protoconch of Buridrillia is unknown and Darbya is so far represented by only the type species, Powell (1966, p. 63) was justified in pointing out their close alliance and possible identity. As mat ters now stand, the species of Buridrillia are moder ately large and inflated, whereas the single species of Darbya is small and slender. Not all species of Buri drillia have a columellar swelling and only one of the five specimens of Darbya lira shows that feature. This peculiar swelling is hardly comparable to the columel lar plaits of the borsonines and does not belie the clavine features. A worn corroded shell, dredged at a depth of 134 fathoms near Cocos Island (USNM 106888), is indis tinguishable from the type species of Buridrillia, so far as it goes. Cocos Island is off Coiba Island, in the Pacific coastal waters of Panama. Darbya? (Buridrillia?) species Plate 63, figures 6, 7 Small, moderately slender, whorls strongly inflated, pillar of moderate length. Protoconch and early post- protoconch whorls missing. Anal fascicle constricted. Axial ribs wide and widely spaced, attenuated and bent on anal fascicle, 10 on penult whorl. Spiral sculp ture consisting of narrow bands, separated by grooves, in which microscopic axial threads are conspicuous. Spiral bands closely spaced on anal fascicle. A hump on body whorl near aperture. Outer lip broken back to hump. Aperture and siphonal canal of moderate length. As shown by growth lines, anal sinus moder ately deep; border of upper limb thin. Siphonal fasci cle sculptured with closely spaced spiral threads. Height (incomplete) 15 mm, diameter 6 mm. A second unnamed clavine species from the Chagres sandstone, like the preceding species, is represented by an imperfect shell. The outer shell layer and therefore the spiral sculpture are corroded on the apertural face of the shell, except on the lower part of the body whorl. The outline, moderately long siphonal canal and other apertural features, so far as they are preserved, suggest a small species of Buridrillia that lacks a columellar swelling. The fossil species of the subgenus so far described are found in formations of late Miocene or Pliocene age in Ecuador (Olsson, 1964, p. 98-100) and on the Pacific coast of Central America straddling the Costa Rica-Panama boundary (Olsson, 1942, p. 51-53). The spiral sculpture of the Chagres species is stronger than that of the described species. Occurrence: Chagres sandstone (late Miocene), locality 208. Genus Crassispira Swainson Swainson, A treatise on malacology, p. 313, 1840. Type (logotype, Herrmannsen, Indicis generum malacozoorum, v. 1, p. 318, 1847): Pleurotoma Notice Valenciennes in Kiener (cited by Swainson as Pleurotoma Bottce Auct), living, Gulf of California. Subgenus Crassispira s.s. Crassispira henekeni has a narrower aperture and less inflated siphonal fascicle than the type species, as well as fewer and wider axial ribs. In number and width of ribs C. cegis (Woodring, 1928, p. 151, pi. 4, fig. 12; Bow den formation, Jamaica) is more similar to the type species, but, like C. henekeni, has a nar rower aperture. Crassispira (Crassispira) henekeni leptalea Woodring, n. subsp. Plate 58, figure 19; plate 65, figure 5 Of medium size, slender, aperture long and narrow. Protoconch acute, conical, 3-whorled; end marked by faint arcuate axial riblet. Sculpture consisting of slightly protractive, swollen axial ribs (8 on body whorl), overridden by strong, narrow primary spiral threads (7 or 8 on penult whorl). Subsutural thread strong; anal fascicle below subsutural thread bearing faint secondary spiral threads. Outer lip broken back. Anal sinus wide, apex below subsutural thread. Si- phonal fascicle scarcely inflated. Height (practically complete) 30.6 mm, diameter 9.7 mm (type). Height (almost complete) 34 mm, diameter 10.5 mm.
378 GEOLOGY AND PALEONTOLOGY OF CANAL ZONE Type: USNM 645777. Type locality: 147h (USGS 6442, i/2 mile (750 me ters) south [southeast] of Gatun, Canal Zone), middle part of Gatun formation. This subspecies, which is smaller and more slender than the nominate subspecies, and has more sharply chiseled spiral sculpture, is rare. Only two specimens, both from the middle part of the Gatun formation, have been found. It is the first record of the species along the west border of the Miocene Caribbean Sea. The nominate subspecies (Sowerby, 1850, p. 50, pi. 10, fig. 6) occurs in the Cercado and Gurabo formations of the Dominican Republic, but is much more abundant in the Gurabo, and also in middle Miocene deposits in Trinidad, Venezuela, and Colombia. The blunt, cylin drical protoconch is not thought to belie the affinities of the Panama form. Crassispira inoequistriata (Li) (Pilsbry, 1931, p. 437, pi. 41, fig. 2), living in Panama Bay, is the most similar living species. No comparable species is known in the western Atlantic Ocean. The deep-water western Atlantic species Pleurotoma (Drillia} horrenda Wat son (1886, p. 308, pi. 26, fig. 4) bears a superficial resemblance. Its aperture, however, is wide and its outer lip strongly flaring. Occurrence: Middle part of Gatun formation, east ern area (middle Miocene), localities 147h, 157. Subgenus Hindsielava Hertlein and Strong Hertlein and Strong, Am. Mus. Nat. History Bull., v. 107, p. 227, 1955. Type (orthotype): Pleurotoma militaris Hinds in Reeve, living, Baja California to Colombia. The more slender outline, more numerous and nar rower axial ribs, stronger spiral sculpture, narrower aperture, and less inflated siphonal fascicle distinguish Hindsiclava from Grassispira s.s. The Oligocene species Pleurotoma abundans Conrad (1848, p. 115, pi. 11, fig. 25; Byram marl, Mississippi) is the earliest Hindsiclava now known. Conrad's illus tration, however, leaves much to be desired. Crassispira (Hindsiclava) censors censors (Sowerby) Plate 58, figures 1, 22; plate 65, figure 6 Pleurotoma censors Sowerby, Geol. Soc. London Quart. Jour., v. 6, p. 50, 1850 (Miocene, Dominican Republic). Guppy, Idem, v. 32, p. 527, pi. 28, fig. 7, 1876 (Miocene, Domini can Republic). Drillia censors (Sowerby), Brown and Pilsbry, Acad. Nat. Sci. Phila. Proc., v. 63, p. 345, 1911 (Miocene, Canal Zone). Maury, Bull. Am Paleontology, v. 5, no. 29, p. 53, pi. 8, figs. 15, 16, 1917 (Miocene, Dominican Republic). Maury, N.Y. Acad. Sci., Scientific Survey of Porto Rico and Vir gin Islands, v. 3, pt. 1, p. 71,1920 (Miocene, Puerto Rico). Pilsbry, Acad. Nat. Sci. Phila. Proc., v. 73, p. 319, pi. 16, fig. 3, 1922 (Miocene, Dominican Republic). Olsson, Bull. Am. Paleontology, v. 9, no. 39, p. 61, pi. 4, figs. 8, 10, 13, 1922 (Miocene, Costa Rica, Canal Zone, Panama). Pal mer, Idem, v. 10, no. 40, p. 11, pi. 2, figs. 7, 8, 1923 (Mio cene, Costa Rica, Dominican Republic). Maury, Idem, v. 10, no. 42, p. 190, pi. 32, fig. 10, 1925 (Miocene, Trini dad). Maury, Brasil Servigo Geol. Mineral. Mon. 4, p. 203, pi. 12, fig. 5, 1925 (Miocene, Dominican Republic). Drillia censors (Guppy), Cossmann, Jour. Conchyliologie, v. 61, p. 20, pi. 2, figs. 8-14, 1913 (Miocene, Martinique, Do minican Republic, Canal Zone). Turris (Crassispira) censors (Sowerby), Rutsch, Schweizer Palseont. Gesell. Abh., v. 55, p. 99, pi. 8, figs. 13-16, 1934 (Miocene, Venezuela). Crassispira censors (Sowerby), Pflug, Acta Humboldtiana, Geol. Palseontologica ser., no. 1, p. 67, pi. 19, figs. 4, 7, 10, 1961 (Miocene, Dominican Republic). Crassispira aff. censors (Sowerby), Jung, Bull. Am. Paleontol ogy, v. 49, no. 223, p. 565, pi. 76, figs. 14, 15, 1965 (Mio cene, Venezuela). 1'Crassispira? cf. Crassispira? consors (Guppy), Marks, Idem, v. 33, no. 139, p. 135, 1951 (Miocene, Ecuador). Wrillia consors portoriccensis Hubbard, N.Y. Acad. Sci., Scien tific Survey of Porto Rico and Virgin Islands, v. 3, pt. 2, p. 158, pi. 24, figs. 8, 9, 1920 (Miocene, Puerto Rico; not Drillia portoricosnsis Hubbard, Idem, p. 159, pi. 24, figs. 11, 12). Wrillia consors fittW&roofci Mansfield, U.S. Natl. Mus. Proc., v. 66, p. 16, pi. 3, fig. 10, 1925 (Miocene, Trinidad). Wrillia consors trinitatensis Mansfield, Idem, p. 17, pi. 3, figs. 12, 13, 1925 (Miocene, Trinidad). Turris (Drillia) militaris (Hinds), Gabb, Am. Philos. Soc. Trans., n. ser., v. 15, p. 207, 1873 (Miocene, Dominican Republic). Gabb, Acad. Nat. Sci. Phila. Jour., 2d ser., v. 8, p. 350, 1881 (Pliocene, Costa Rica). Pleurotoma sp. aff. P. alesidota macilenta Dall, Toula, K. k. Geol. Reichsanstalt Jahrb., v. 61, p. 506, pi. 30, fig. 11, 1911 (Miocene, Canal Zone). Pleurotoma (Drillia) Dalli Toula, Idem, p. 506, pi. 30, fig. 12, 1911 (Miocene, Canal Zone). Not Pleurotoma dalli Verrill and Smith, Connecticut Acad. Trans., v. 5, p. 451, pi. 57, figs. 1, la, 1882. Drillia macilenta rectaxis Pilsbry, Acad. Nat. Sci. Phila. Proc., v. 73, p. 319, pi. 16, figs. 1, 2, 1922 (Miocene, Dominican Republic). Surcula Tw&soni Hanna, California Acad. Sci. Proc., 4th ser., v. 13, no. 10, p. 181, 1924 (substitute name for Pleuro toma dalli Toula). Moderately large, slender, aperture long and nar row. Protoconch acute (apex almost invariably bro ken), 21/2- to 3-whorled, last half whorl slightly or decidedly bulging, end marked by appearance of first axial rib. First 3 to 5 axial ribs narrow, arcuate, ex tending practically from suture to suture. Later ribs wider, slightly protractive, cut off at anal fascicle. Ribs progressively narrower and more closely spaced. Sculpture of late whorls consisting of skewed rectan gles formed by strong, slightly protractive axial ribs (22 to 30 on mature penult whorl) and narrower pri mary spiral threads (4 or 5 on mature penult whorl),
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Geology and Paleontology of Canal Z
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UNITED STATES DEPARTMENT OF THE INT
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GEOLOGY AND PALEONTOLOGY OF CANAL Z
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GASTROPODS: ETJLIMIDAE, MARGINELLID
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GASTROPODS: EULIMIDAE. MARGINELLIDA
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GASTROPODS: EULIMIDAE, MARGINELLIDA
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GASTROPODS: ETJLIMIDAE, MARGINELLID
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Cut area (Acmaeidae to O, common; A
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Cut area (Acmaeidas to Ellobiidae)
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Cavolinidae and a few species of ot
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Cydostremiscus (Ponocyctus) pentago
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GASTROPODS: EULIMIDAE, MARGENTELLID
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Abderospira... ....................
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Cancellaria (Cancellaria) Continued
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Daphnella Continued pagera.........
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Page ischnum, Strioterebrum........
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Page Strombidae_ ..................
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PLATES 48-66
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GEOLOGICAL SURVEY PROFESSIONAL PAPE
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16 17 MIDDLE MIOCENE MOLLUSKS FROM
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GEOLOGICAL SURVEY MIDDLE MIOCENE MO
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MIDDLE MIOCENE MOLLUSKS FROM GATUN
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MIDDLE MIOCENE MOLLUSKS FROM GATUN
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Geology and Paleontology of Canal Zone and Adjoining ... - USGS

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