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Fruit Fly Expert Identification System and Systematic Information

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Norrbom Status of Knowledge 39<br />

prep.). There are also keys for the faunas of the following more<br />

restricted areas: Britain (White 1988), northern China (Zia &<br />

Chen 1938), central Europe (Merz 1994[3343]), eastern Europe<br />

(Richter 1970), Israel (Freidberg & Kugler 1989), Japan (Ito<br />

1983-1985), <strong>and</strong> Korea (Kwon 1985).<br />

Revisions, monographs <strong>and</strong> keys are listed in the References<br />

section (“REFS”) under each family group <strong>and</strong> genus<br />

group taxon in the database. Because of the unstable higher<br />

classification of the Tephritidae, users should be forewarned<br />

that many keys to genera or higher groups may be less reliable<br />

than regional or local keys that do not strictly follow a classification.<br />

The following publications include keys to genera of<br />

higher taxa of Tephritidae:<br />

Keys to subfamilies <strong>and</strong> tribes have been provided by<br />

Hering (1947), Aczél (1953[24]), Hardy (1973, 1974[1943]),<br />

Foote (1980), Ito (1983), Kwon (1985), Hancock<br />

(1986[1890]), Freidberg & Kugler (1989), White (1988), <strong>and</strong><br />

Merz (1994[3343]). Most of them conflict in classification <strong>and</strong><br />

are difficult to use.<br />

Phytalmiinae — There is no comprehensive generic key.<br />

Acanthonevrini — Most Oriental <strong>and</strong> Australasian genera<br />

were included in one or more of the following works: Hardy<br />

(1973, 1974[1943], 1980[1949], 1986[1962], 1988), Kapoor<br />

(1993), <strong>and</strong> Permkam & Hancock 1995[3795]. The Afrotropical<br />

genera were included under Trypetinae in the key of Hancock<br />

(1986[1890]), <strong>and</strong> the key to the species of four<br />

Afrotropical genera by Munro (1967[3521]) is also useful.<br />

Blepharoneurini — The four genera have never been keyed<br />

as a group.<br />

Epacrocerini — Three of the genera were revised by Hardy<br />

(1982[1954]), <strong>and</strong> all four were included in the key of Hardy<br />

(1988[1964]).<br />

Phascini — The six genera were included under Acanthonevrina<br />

<strong>and</strong> Gastrozonina by Hardy (1986[1962],<br />

1988[1964]).<br />

Phytalmiini — There is no comprehensive generic key.<br />

McAlpine & Schneider (1978) keyed four of the genera. Some<br />

genera have been keyed <strong>and</strong>/or revised under Adramini or<br />

Gastrozonina by Hardy (1986[1961], 1988[1964]).<br />

Trypetinae — The genera of the USA <strong>and</strong> Canada are<br />

included in the key of Foote et al. (1993), <strong>and</strong> those of the<br />

Neotropical Region <strong>and</strong> Nearctic Mexico were keyed by Foote<br />

(1980). The Afrotropical genera, except the Dacini, were keyed<br />

by Hancock (1986[1890], 1991[1895]). Many Palearctic genera<br />

were keyed by Hendel (1927[2109]), <strong>and</strong> others were<br />

included in keys to local areas listed above under that region.<br />

Most Oriental <strong>and</strong> Australasian genera were included in keys<br />

by Hardy (1973, 1974[1943], 1987[1963], 1988[1964]),<br />

Kapoor (1993), Permkam & Hancock (1995[3794],<br />

1995[3795]), <strong>and</strong> Shiraki (1933, 1968).<br />

Adramini — There is no comprehensive generic key.<br />

Many, but not all, of the Oriental <strong>and</strong> Australasian genera were<br />

included in keys by Hardy (1973, 1974,<br />

1983[1958], 1986[1961]), Kapoor (1993), <strong>and</strong> Permkam &<br />

Hancock 1995[3795]), <strong>and</strong> the Afrotropical genera were keyed<br />

by Hancock (1986[1890]).<br />

Carpomyini — There is no key strictly to the genera of this<br />

group, although they are included in various regional or local<br />

keys. Norrbom (1994[3663]) provided a key to four American<br />

genera of Carpomyina.<br />

Dacini — All three genera of Dacini were included in the<br />

key to economically important genera of White & Elson-Harris<br />

(1992). Munro (1984) revised the Afrotropical species, although<br />

most of his proposed generic <strong>and</strong> suprageneric names<br />

have not been accepted. Drew (1989[232]) keyed the genera,<br />

subgenera <strong>and</strong> species of the Australasian Region. For the<br />

Oriental Region, most species are included in one or more of<br />

the following works: Hardy (1973, 1974[1943], 1982[1952],<br />

1983[1956]), Wang & Zhao (1989), Tseng, Chen & Chu<br />

(1992), Kapoor (1993), <strong>and</strong> Drew & Hancock (1994[1238]). A<br />

key to the Afrotropical genera of Ceratitidina was provided by<br />

Hancock (1987), <strong>and</strong> Hancock & Drew (1994[1901]) <strong>and</strong> Permkam<br />

& Hancock (1995[3794]) keyed most of the Oriental <strong>and</strong><br />

Australasian genera. The Afrotropical genera of Gastrozonina<br />

have been keyed by Hancock (1985[1889]), <strong>and</strong> most of the<br />

Oriental <strong>and</strong> Australasian genera have been included in keys<br />

by Hardy (1973, 1974[1943], 1988[1964]) <strong>and</strong> Kapoor (1993).<br />

Ortalotrypetini — Norrbom (1994[3662]) provided a key<br />

to four of the five included genera.<br />

Rivelliomimini — There is no key including all three of<br />

the genera, but the two Afrotropical genera were included in<br />

the key of Hancock (1986[1890]).<br />

Toxotrypanini — The three genera were included in the<br />

keys of Foote (1980) <strong>and</strong> Foote et al. (1993).<br />

Trypetini — Han (1992) provided a key to most of the<br />

world genera of Trypetina <strong>and</strong> the Chetostoma group. The three<br />

genera of Acidoxanthina are included in keys by Hancock<br />

(1987) <strong>and</strong> Hardy (1987).<br />

Tephritinae — The genera of the USA <strong>and</strong> Canada were<br />

included in the key of Foote et al. (1993), <strong>and</strong> those of the<br />

Neotropical Region <strong>and</strong> Nearctic Mexico were keyed by Foote<br />

(1980). Most Afrotropical genera have been keyed by Bezzi<br />

(1924[470], 1924[469], 1924[472]), Munro (1957[3511]), <strong>and</strong><br />

Freidberg (1987). Many Palearctic genera were keyed by Hendel<br />

(1927[2109]), <strong>and</strong> others were included in keys to local<br />

areas listed above under that region. Most Oriental <strong>and</strong> Australasian<br />

genera were included in keys by Hardy (1973,<br />

1974[1943], 1988[1965]), Hardy & Adachi (1956), Hardy &<br />

Delfinado (1980), Hardy & Drew (1996), Kapoor (1993), Shiraki<br />

(1933, 1968), <strong>and</strong> Zia (1937). The Australian genera were<br />

revised by Hardy & Drew (1996).<br />

Acrotaeniini — There is no key strictly to the genera of<br />

this group. Foote (1980) keyed some of the genera under<br />

Platensinini, but he included others under Trypetinae, Tephritini<br />

<strong>and</strong> Aciurini.<br />

Dithrycini — There is no comprehensive generic key for<br />

this group. Freidberg & Kaplan (1992) revised the Afrotropical<br />

species of Oedaspidina, <strong>and</strong> they <strong>and</strong> Freidberg & Mansell<br />

(1995) provided keys to the Old World genera. The genera of<br />

Cecidocharina have not been keyed as a group, but were included<br />

in keys by Aczél (1953[24]), Foote (1980) <strong>and</strong> Foote et<br />

al. (1993).<br />

Eurostini — The three genera were included in the key of<br />

Foote et al. (1993).<br />

Eutretini — There is no comprehensive generic key for this<br />

group. That by Munro (1952[3503]) is obsolete. Foote (1980)<br />

keyed most of the American genera under Ditrichini. Freidberg

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