Supporting Information (SI) Appendix - Proceedings of the National ...

Supporting Information (SI) Appendix 

Genetic Structure and Domestication History of the Grape 

Sean Myles a,b,c , Adam R. Boyko b , Christopher L. Owens e,f , Patrick J. Brown a , Fabrizio Grassi g , 

Mallikarjuna K. Aradhya f,h , Bernard Prins f,h , Andy Reynolds b , Jer-Ming Chia i , Doreen Ware f,i , 

Carlos D Bustamante b , Edward S. Buckler a,f 

a Institute for Genomic Diversity, Cornell University, Ithaca, New York, 14853, USA 

b Department of Genetics, Stanford University School of Medicine, Stanford, California, 94305, 

USA 

c Department of Biology, Acadia University, Wolfville, Nova Scotia, B4P 2R6, Canada 

e Grape Genetics Research Unit, Cornell University, Geneva, New York, 14456, USA 

f United States Department of Agriculture – Agricultural Research Service 

g Botanical Garden, Department of Biology, University of Milan, 20139 Milan, Italy 

h National Clonal Germplasm Respository, University of California, Davis, California, 95616, 

USA 

i Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, 11724, USA 

Corresponding author: Sean Myles, Institute for Genomic Diversity, Cornell University, Ithaca, 

New York, 14853, USA, ph: 607-220-4244, fax: 607-254-6379, email: smm367@cornell.edu

Supplementary Figures 

Table of Contents 

Figure S1: Description of thresholds used for defining pedigree relationships among vinifera 

cultivars. 

Figure S2: Evalution of domestication bottleneck in the grape. 

Figure S3: Selective sweep signal on chromosome 17. 

Figure S4: GWA result for grape color and the associated signals of selection for lighter 

pigmentation. 

Figure S5: Evaluation of haplotype diversity in western and eastern vinifera. 

Figure S6: The inter-SNP distance distribution of SNPs assayed by the Vitis9KSNP array. 

Figure S7: Evaluation of authenticity of sylvestris: PCA of vinifera and sylvestris. 

Figure S8: Evaluation of authenticity of sylvestris: PCA of western vinifera and western 

sylvestris. 

Figure S9: Evaluation of authenticity of sylvestris: PCA of eastern vinifera and eastern 

sylvestris. 

Supplementary Tables 

Table S1: Geographic origins of accessions from the USDA grape germplasm collection. 

Table S2: Confirmation of parent-offspring pairs established by other sources. 

Table S3: Inferred trios. 

Table S4: The most highly connected cultivars. 

Table S5: Clonal and first degree relationships for each cultivar used in the present study. 

Table S6: Details of the sylvestris samples used in the present study. 

Supplementary Methods 

Supplementary Text 

Biases in grape germplasm collections: a cautionary note 

Potential misnaming of cultivars 

Assignment of cultivars as “wine” or “table” grapes

Authenticity of Vitis vinifera subsp. sylvestris 

References 

Supplementary Figures 

Figure S1: Description of thresholds used for defining pedigree relationships among 

vinifera cultivars. A) Identity-by-descent (IBD) distribution from pairwise comparisons among 

950 vinifera accessions. Note the break in the Y-axis: the large majority of pairs of accessions 

have IBD near zero as expected. The dashed line represents the lowest IBD value observed 

between a known parent-offspring pair (IBD = 0.466; see Table S1 for known parent-offspring 

pairs). The dotted line represents the threshold above which pairwise comparisons were 

considered clones (i.e. effectively genetically identical; IBD = 0.95). Comparisons between

samples that yielded values above the dashed line and below the dotted line (i.e. 0.466 ≥ IBD < 

0.95) were considered likely first degree relatives, although such values could represent more 

complex familial relationships generated by backcrossing and other complex crossing schemes 

across interleaved generations and/or related cultivars that produce IBD values equivalent to first 

degree relatives. 

B) Thresholds for defining “likely parent-offspring” relationships. Each dot represents a 

pairwise comparison between accessions with 0.466 ≥ IBD < 0.95. IBD values within these 

boundaries could be obtained for siblings, parent-offspring pairs or other complex pedigree 

relationships. The expected Z0 and Z1 scores for parent-offspring pairs are 0 and 1, respectively. 

However, we expect a large variance in these estimates from the current data for various reasons 

(e.g. ascertainment bias, low and uneven SNP density, genotyping error). To define likely parentoffspring 

pairs, we used the highest Z0 and the lowest Z1 scores from known parent-offspring 

pairs (see Table S1) as thresholds for defining parent-offspring pairs in the present data set. The 

vertical dashed line represents the Z1 threshold (0.580) and the horizontal dashed line represents 

the Z0 threshold (0.121). These two thresholds delimit the cluster of dots in the bottom right 

portion of the plot, which we argue is likely to contain mostly parent-offspring pairs. The 

confirmed parent-offspring pairs from Table S1 are shown in red and are found within this 

cluster. We assign the remaining dots in the bottom right quadrant created by the dashed lines as 

“likely parent-offspring pairs”. First degree relationships (0.466 ≥ IBD < 0.95) that do not fall 

within this quadrant are considered “sibling or equivalent”. 

C) Establishing trios by evaluating mendelian inconsistencies. For each accession that has likely 

parent-offspring relationships with at least 2 other cultivars based on the criteria defined in B) 

above, we calculated the proportion of SNPs inconsistent with mendelian inheritance for each 

possible pair of parents. The histogram of the proportion of SNPs inconsistent with mendelian 

inheritance is shown here and is clearly bi-modal. We infer that the small distribution to the left 

represents trios and the large distribution to the right represents comparisons made among 3 first 

degree relatives that do not form a trio. In support of this definition, we find that all of the 

known trios from the literature (Table S3) fall below this threshold. We chose the largest value 

from the smaller distribution to the left, represented by the vertical dashed line at 0.0135, as the 

threshold below which we assign 3 first-degree relatives as a trio. This value corresponds to 

1.35% of SNPs being inconsistent with mendelian inheritance, which is reasonable considering 

genotyping error. A list of trios, and whether they have been previously confirmed in the 

literature, is given in Table S3.

Figure S2: Evalution of domestication bottleneck in the grape. Panels A and B describe 

levels of haplotype diversity in vinifera and sylvestris. A) Median haplotype diversity in vinifera 

and sylvestris across a range of non-overlapping window sizes. At window sizes < 5 SNPs, 

vinifera has higher haplotype diversity than sylvestris which is likely due to ascertainment bias: 

the SNPs assayed by the Vitis9KSNP genotyping array were chosen because they were highly 

polymorphic within a set of vinifera cultivars. At window sizes > 5 SNPs, sylvestris shows 

higher haplotype diversity than vinifera, although the difference between the two species is 

relatively small. B) P value distribution from comparing haplotype diversity between vinifera 

and sylvestris. For each window size, a one-sided Wilcoxon signed rank test was used to 

compare haplotype diversity between the two species. Red dots represent the –log10 P value 

from testing whether vinifera has higher diversity than sylvestris. Blue dots represent the –log10 

P value from testing whether sylvestris has higher diversity than vinifera. At window sizes < 5 

SNPs, vinifera shows significantly higher haplotype diversity than sylvestris, and the opposite is 

true for window sizes > 5 SNPs. C) The decay of linkage disequilibrium (LD) in vinifera and 

sylvestris. Each point represents the median r 2 value and the mean physical distance from bins of 

500 pairwise SNP comparisons. LD decay is rapid in both vinifera and sylvestris. There is no 

apparent difference in LD decay between vinifera and sylvestris.

Figure S3: Selective sweep signal on chromosome 17. A) Haplotype diversity in vinifera 

relative to haplotype diversity in sylvestris is plotted in 20 SNP windows with a 5bp overlap 

across chromosome 17. Values below the dotted line represent windows within the top 1% of 

reductions in haplotype diversity in vinifera compared to windows genome-wide. B) Average Fst 

from 20 SNP windows with a 5bp overlap across chromosome 17. Values above the dotted line 

represent windows with Fst values within the top 1% most extreme Fst values from windows 

genome-wide. Over 20 known or predicted transcripts are found within the 5Mb sweep region, 

which prevents gene-level dissection of this putatively adaptive locus.

Figure S4: GWA result for grape color and the associated signals of selection for lighter 

pigmentation. A) P values for association between SNPs and grape color across the 19 

assembled chromosomes of the grape. The horizontal dashed line corresponds to a 5% 

significance threshold with Bonferroni correction for 5110 tests. B) Magnification of the 

association result on chromosome 2. The location of the known causal alleles for grape color in 

the VvMybA1 and VvMybA2 genes, which were not genotyped in the present study, is indicated 

by the vertical line. C) Average Fst is shown for 10-SNP windows with a 5-SNP overlap across 

chromosome 2. The horizontal dashed line represents the top 1% Fst threshold from windows 

genome-wide. D) Haplotype diversity in white cultivars relative to red cultivars in sliding 

windows as above. Values below the horizontal dashed line represent windows within the 1% 

most extreme reductions in haplotype diversity in white cultivars from windows genome-wide. 

The elevated LD generated by selection for white grapes has generated a diffuse association peak 

covering 5Mb and several hundred genes. The observation that the strongest signals of 

association and selection are not found directly at the causal locus has been seen in other plants, 

for example in Arabadopsis [1] and maize [2].

Figure S5: Evaluation of haplotype diversity in western and eastern vinifera. A) Median 

haplotype diversity in western and eastern vinifera across a range of non-overlapping window 

sizes. At window sizes < 10 SNPs, western vinifera has slightly higher haplotype diversity than 

sylvestris which is likely due to ascertainment bias: the SNPs assayed by the Vitis9KSNP 

genotyping array were chosen primarily because they were highly polymorphic within a set of 

western vinifera cultivars [3]. At window sizes > 10 SNPs, eastern vinifera shows higher 

haplotype diversity than western vinifera, although the difference between the two populations is 

relatively small. B) P value distribution from comparing haplotype diversity between western 

and eastern vinifera. For each window size, a one-sided Wilcoxon signed rank test was used to 

compare haplotype diversity between the two populations. Red dots represent the –log10 P value 

from testing whether western vinifera have higher diversity than eastern vinifera. Blue dots 

represent the –log10 P value from testing whether eastern vinifera have higher diversity than 

western vinifera. At window sizes < 10 SNPs, western vinifera show significantly higher 

haplotype diversity than eastern vinifera, and the opposite is true for window sizes > 10 SNPs. A 

description of the geographic assignment of cultivars to east and west is provided in Table S1.

Figure S6: The inter-SNP distance distribution of SNPs assayed by the Vitis9KSNP array. 

The histogram summarizes the physical distance between neighbouring pairs of 4951 

polymorphic SNPs in our sample that are anchored to the 19 chromosomes. The excess of small 

inter-SNP distances makes the array particularly well-suited for analyses of LD decay in the 

grape since most of the LD decay occurs within 1kb (see Figure S2).

Figure S7: Evaluation of authenticity of sylvestris: PCA of vinifera and sylvestris. PCA was 

performed on 583 unique vinifera cultivars and 59 sylvestris using 2958 LD-pruned SNPs. The 

proportion of the variance explained by each PC is shown in parentheses along each axis. The 

geographic origin of the samples is described in the legend. The black arrow refers to a vinifera 

cultivar, “Beli Potok”, that is likely mislabeled. See the section “Supplementary Text: 

Authenticity of Vitis vinifera subsp. sylvestris” for interpretations of this plot and an explanation 

of Olmo’s “wild vinifera”.

Figure S8: Evaluation of authenticity of sylvestris: PCA of western vinifera and western 

sylvestris. PCA was performed on 177 unique western vinifera cultivars and 38 western 

sylvestris. The proportion of the variance explained by each PC is shown in parentheses along 

each axis. See the section “Supplementary Text: Authenticity of Vitis vinifera subsp. sylvestris” 

for interpretations of this plot.

Figure S9: Evaluation of authenticity of sylvestris: PCA of eastern vinifera and eastern 

sylvestris. PCA was performed on 91 unique eastern vinifera cultivars, 21 eastern sylvestris and 

12 cultivars labeled as Olmo’s “wild vinifera”. The proportion of the variance explained by each 

PC is shown in parentheses along each axis. See the section “Supplementary Text: Authenticity 

of Vitis vinifera subsp. sylvestris” for interpretations of this plot.

Supplementary Tables 

Vitis vinifera subs. vinifera (N = 404) Vitis vinifera subs. sylvestris (N = 59) 

West Central East West East 

Austria (12) Bulgaria (1) Afghanistan (22) Corsica (2) Armenia (5) 

France (60) Croatia (1) Egypt (2) Germany (1) Azerbaijan (1) 

Germany (21) Greece (88) India (3) Italy (14) Dagestan (1) 

Italy (52) Hungary (19) Iran (4) Sardinia (4) Georgia (5) 

Portugal (20) Macedonia (1) Iraq (4) Spain (13) Pakistan (6) 

Spain (8) Romania (2) Israel (4) Tunisia (4) Turkey (2) 

Switzerland (3) Serbia (12) Lebanon (1) Turkmenistan (1) 

Tunisia (1) Pakistan (20) 

Russia (33) 

Turkey (4) 

Uzbekistan (5) 

Yemen (1) 

Table S1: Geographic origins of accessions from the USDA grape germplasm collection. 

The 583 unique vinifera cultivars used in the present study were grouped into their countries and 

regions of origin based mainly on data retrieved from the Germplasm Resource Information 

Network (http://www.ars-grin.gov/). The geographic origins of 404 vinifera cultivars are shown 

here and an additional 179 cultivars are of unknown geographic origin. The 59 sylvestris 

accessions (N = 59) were grouped into countries and regions of origin based on their sampling 

location.

Cultivar A Cultivar B IBD Z0 Z1 Reference 

Barbableu (DVIT 1651) Olivette Noir (DVIT 1306) 0.617 0.048 0.67 GRIN 

Cereza (DVIT 2907) Pais (DVIT 484) 0.515 0.104 0.763 * 

Cereza (DVIT 2907) Muscat of Alexandria (DVIT 841) 0.565 0.054 0.763 * 

CG 1481 (DVIT 2432) Thompson Seedless (DVIT 535) 0.572 0.057 0.742 GRIN 

Chardonnay (DVIT 687) Pinot Noir (DVIT 915) 0.518 0.037 0.89 * 

Clotilde Prosperi (DVIT 568) Italia (IP 65) (DVIT 577) 0.53 0.048 0.843 GRIN 

Durif (DVIT 895) Syrah (DVIT 1053) 0.546 0.081 0.747 * 

Ehrenfelser (DVIT 2317) White Riesling (DVIT 1015) 0.593 0.088 0.639 * 

Flora (DVIT 720) Semillon (DVIT 954) 0.548 0.071 0.763 GRIN 

Flora (DVIT 720) Gewurztraminer (DVIT 738) 0.608 0.055 0.674 GRIN 

French Colombard (DVIT 

726) Chenin Blanc (DVIT 691) 0.478 0.086 0.873 GRIN 

Italia (DVIT 424) Muscat Hamburg (DVIT 860) 0.549 0.062 0.778 * 

Kerner (DVIT 3104) Frankenthal (DVIT 408) 0.495 0.079 0.852 GRIN 

Kerner (DVIT 3104) White Riesling (DVIT 1016) 0.513 0.077 0.819 GRIN 

Kishmish Of Vir (DVIT 

2071) Black Kishmish (DVIT 1326) 0.602 0.054 0.689 GRIN 

Mueller-Thurgau (DVIT 

2337) White Riesling (DVIT 1015) 0.502 0.121 0.754 * 

Mueller-Thurgau (DVIT 

2337) Optima (DVIT 3105) 0.66 0.05 0.58 * 

Muscat Hamburg (DVIT 468) Frankenthal (DVIT 408) 0.517 0.051 0.865 GRIN 

Muscat of Alexandria (DVIT 

841) Muscat Hamburg (DVIT 468) 0.54 0.048 0.825 GRIN 


841) Sultana Moscata (DVIT 527) 0.553 0.044 0.807 * 

NA (PI 588425.02) Sylvaner (DVIT 968) 0.5 0.06 0.879 GRIN 

NA (PI 588668.04) Ezerjo (DVIT 715) 0.532 0.109 0.717 GRIN 

Petite Bouschet (DVIT 894) Alicante Bouschet (DVIT 625) 0.495 0.074 0.862 GRIN 

Pinot (DVIT 733) Gamay Burgoyne (DVIT 734) 0.478 0.081 0.882 * 

Pinot (DVIT 733) Chardonnay (DVIT 687) 0.517 0.03 0.906 * 

Plavac Mali (DVIT 1119) Zinfandel (DVIT 1340) 0.54 0.068 0.784 * 

Primiera (DVIT 491) Delizia Di Vaprio (DVIT 571) 0.554 0.087 0.718 GRIN 

Red Veltliner (DVIT 928) Rothgipfler (DVIT 935) 0.51 0.07 0.84 * 

Reichensteiner (DVIT 3107) Mueller Thurgau (DVIT 2337) 0.544 0.054 0.805 GRIN 

Rothgipfler (DVIT 935) Traminer (DVIT 991) 0.552 0.084 0.728 * 

Ruby Seedless (DVIT 2625) Sultana Moscata (DVIT 527) 0.541 0.077 0.765 * 

Schonburger (DVIT 1124) Pinot Noir (DVIT 1066) 0.556 0.033 0.821 GRIN 

Seedless Emperor (DVIT 

2759) Ruby Seedless (DVIT 2625) 0.551 0.075 0.748 **

Sovrana (DVIT 525) Delizia Di Vaprio (DVIT 571) 0.508 0.087 0.81 GRIN 

Sovrana (DVIT 525) Frankenthal (DVIT 408) 0.559 0.035 0.814 GRIN 

Sultana Moscata (DVIT 527) Thompson Seedless (DVIT 535) 0.513 0.105 0.765 * 

Sylvaner (DVIT 1075) Traminer (DVIT 991) 0.479 0.094 0.855 * 

Syrah (DVIT 1053) Durif (DVIT 895) 0.546 0.081 0.747 * 

Thomuscat (DVIT 536) Muscat of Alexandria (DVIT 841) 0.55 0.082 0.737 GRIN 

Thomuscat (DVIT 536) Thompson Seedless (DVIT 535) 0.552 0.072 0.751 GRIN 

Touriga (DVIT 990) Tarrango II (DVIT 2658) 0.466 0.078 0.911 * 

Trieste-pirovano 299 (DVIT 

2164) Moscato Dadda (DVIT 2109) 0.536 0.103 0.722 GRIN 

Zweigelt-rebe (DVIT 2692) 

Limberger (Blaufraenkisch) 

(DVIT 797) 0.494 0.074 0.864 * 

Table S2: Confirmation of parent-offspring pairs established by other sources. Each row 

represents a previously described parent-offspring relationship, the reference for which is 

provided in the “Reference” column. The term “GRIN” in the Reference column refers to the 

Germplasm Resource Information Network (http://www.ars-grin.gov/). The asterisk in the 

Reference column refers to Sefc et al. [4]. Cultivar names are shown and GRIN IDs are 

provided in parentheses. The IBD and Z scores are shown. The lowest IBD score from this table 

(IBD = 0.466) was used as a lower bound for defining first degree relationships between pairs of 

accessions in the present data set. The lowest Z1 score (0.580) and the highest Z0 score (0.121) 

were used as thresholds for defining “likely parent-offspring” pairs in our sample.

Offspring Parent 1 Parent 2 

Mendelian 

inconsistencies 

Reference 

Angelo Pirovano 

(DVIT 564) 

Italia (IP 65) (DVIT 

424) 

Markandi (DVIT 587) 0.00529 

Anna Maria (DVIT 

3156) 

Delizia Di Vaprio 

(DVIT 571) 

Perle de Csaba (DVIT 

2648) 

0.00704 

Arnsburger (DVIT 

2666) 

Mueller Thurgau 

(DVIT 2337) 

Chasselas Tokay 

(DVIT 377) 

0.00809 

Askari (DVIT 343) 

Muscat Hamburg 

(DVIT 468) 


B 15-19 (DVIT 

2171) 

Almeria (DVIT 1079) 

Sultanina Rose (DVIT 

530) 

0.00799 

Badacsony-10 

(DVIT 2667) 

Ezerjo (DVIT 715) NA (DVIT 1978) 0.00692 

Barbableu (DVIT 

348) 


(DVIT 468) 

Olivette Noire (DVIT 

1657) 

0.00591 

Barone Dellaterno 

(DVIT 350) 

Muscat of Alexandria 

(DVIT 841) 


530) 

0.00755 

Beauty Seedless 

(DVIT 351) 

Black Kishmish 

(DVIT 1326) 

Queen Of The 

Vineyard (DVIT 496) 

0.00879 

Bermestia Bianca 

(DVIT 654) 

Trebbiano Bianco 

(DVIT 2067) 

Olivette Vendemien 

(DVIT 482) 

0.00642 

Bien Donnie (DVIT 

2696) 

Queen Of The 


Almeria (DVIT 1079) 0.00768 

Black Prince (DVIT 

361) 


(DVIT 841) 

Pias (DVIT 833) 0.01065 

Bondola (DVIT 

3067) 

Rodites (DVIT 609) Pelaverga (DVIT 892) 0.00623 

Bucklands 

Sweetwater (DVIT 

363) 

Chasselas Dore (DVIT 

373) 


(DVIT 377) 

0.00489 

Cabernet Sauvignon 

(DVIT 677) 

Cabernet Franc (DVIT 

803) 

Sauvignon Blanc 

(DVIT 947) 

0.00842 * 

Calmeria (DVIT 

1084) 

Almeria (DVIT 1079) 


530) 

0.00862 

Cereza (DVIT 2907) 


(DVIT 841) 

Pias (DVIT 833) 0.00898 * 

CG 1481 (DVIT 

2432) 


530) 

Itonychi Lefko (DVIT 

561) 

0.00627 

Chasselas Napoleon 

(DVIT 611) 


(DVIT 377) 

Frankenthal (DVIT 

408) 

0.00608 

Chasselas Tompa 

(DVIT 378) 


(DVIT 611) 


(DVIT 841) 

0.00898 

Conte Rosso (DVIT NA (DVIT 1980) Citronelle (DVIT 381) 0.00762

382) 

Corniola (DVIT 

3162) 


(DVIT 841) 

Pias (DVIT 833) 0.00777 

Durif (DVIT 895) Syrah (DVIT 1053) Durif (DVIT 710) 0.00859 

Early Muscat (DVIT 

394) 


(DVIT 468) 

Queen Of The 


0.00513 

Exotic (DVIT 401) Ribier (DVIT 503) Angelina (DVIT 339) 0.00909 

Ezerfuertu (DVIT 

2672) 

Gewurztraminer 

(DVIT 738) 

Gouveio (DVIT 743) 0.00776 

Flora (DVIT 720) Semillon (DVIT 954) 


(DVIT 738) 

0.01339 

Fort Ross Vine 

(DVIT 1353) 


(DVIT 841) 

Zinfandel (DVIT 

1340) 

0.00995 

Fuji (DVIT 1655) 

Russian Seedless 

(DVIT 2431) 


(DVIT 841) 

0.00958 

Gamay (DVIT 732) Malbec (DVIT 802) 

Gamay Toma (DVIT 

2100) 

0.00601 

Goyura (DVIT 

2637) 


(DVIT 841) 


530) 

0.0066 

Gyongyrizling 

(DVIT 2675) 

White Riesling (DVIT 

1015) 


2648) 

0.00827 GRIN 

Helena (DVIT 756) Refosco (DVIT 1121) 

Zinfandel (DVIT 

1340) 

0.01067 GRIN 

Henab (DVIT 417) Hunisa (DVIT 418) Aswad (DVIT 2087) 0.01038 

Hungarian 

Millenium (DVIT 

761) 

Angelina (DVIT 339) 


(DVIT 377) 

0.00841 

Italia (IP 65) (DVIT 

424) 


373) (DVIT 725) 


(DVIT 468) 

0.00636 

Kerner (DVIT 3104) 


1015) 


408) 

0.00828 GRIN 

Khalili (DVIT 2084) 

Munuka Sufed (DVIT 

2046) 


530) 

0.00526 

Kishmish Hishrau 

(DVIT 1103) 

NA (GVIT 1363.05) 

Black Kishmish 

(DVIT 1326) 

0.00645 

Madam Matijas 

(DVIT 2642) 


(DVIT 841) 


(DVIT 377) 

0.01002 

Maria Pirovano 

(DVIT 588) 


(DVIT 841) 


530) 

0.00799 

Muscat Br39 (DVIT 

1313) 


373) (DVIT 725) 


(DVIT 468) 

0.00692 

Muscat De 

Pantellana (DVIT 

2063) 

Olivette Bauthelet 

(DVIT 590) 


(DVIT 841) 

0.00558 

Muscat Hamburg Muscat of Alexandria Frankenthal (DVIT 0.00719 *

(DVIT 468) (DVIT 841) 408) 

Muscat of 

Alexandria (DVIT 

Black Morocco (DVIT 

360) 

Muscat Galego (DVIT 

859) 

0.00745 

841) 

Muscat Szauter 

(DVIT 863) 

Hunisa (DVIT 418) 


530) 

0.00771 

NA (DVIT 1978) 

Pinot Noir Espinett 

(DVIT 2332) 


859) 

0.00874 



(DVIT 468) 


(DVIT 377) 

0.00564 

NA (GVIT 1614.02) NA (DVIT 1978) 


(DVIT 738) 

0.00711 

NA (PI 340969.03) Carriere (DVIT 953) 


(DVIT 2067) 

0.00943 

NA (PI 588425.02) 

Sylvaner 742 (DVIT 

3180) 


859) 

0.00758 

NA (PI 588668.04) Ezerjo (DVIT 715) NA (DVIT 1978) 0.00829 

Netter (DVIT 474) 


(DVIT 468) 

Dabouki (DVIT 388) 0.01012 

Nicolas Horthy 

number39 (DVIT 

475) 

Madam Matijas 

(DVIT 2642) 

Chasselas Tompa 

(DVIT 378) 

0.00595 

Olmo 25F (DVIT 

2751) 

Yellow Ortlieber 

(DVIT 1020) 


(DVIT 377) 

0.00949 

Orange Muscat 

(DVIT 2167) 


(DVIT 377) 


859) 

0.00818 

Ortega (DVIT 1114) 


(DVIT 2337) 

Siegerrebe (DVIT 

3109) 

0.00673 

Osteiner (DVIT 

2330) 


1015) 

Sylvaner 742 (DVIT 

3180) 

0.00831 GRIN 

Pardinia (DVIT 486) Monukka (DVIT 462) Citronelle (DVIT 381) 0.00788 

Perle (DVIT 2754) 


(DVIT 2337) 


(DVIT 738) 

0.00795 

Perlette (DVIT 488) 

Queen Of The 



530) 

0.00857 GRIN 

Petite Bouschet 

(DVIT 894) 

Aramon (DVIT 633) 

Teoulier Noir (DVIT 

975) 

0.00911 

Queen (DVIT 495) 


(DVIT 468) 


530) 

0.00702 GRIN 

Queen Of The 

Vineyard (DVIT 

496) 


2648) 


Rambola (DVIT 

497) 


360) 

Pias (DVIT 833) 0.00777 

Ranny Vira (DVIT Black Kishmish Chaouch (DVIT 371) 0.00504 GRIN

2684) (DVIT 1326) 

Ribier (DVIT 503) 

Gros Colman (DVIT 

413) 

Robin (DVIT 2651) Ribier (DVIT 503) 

Rotberger (DVIT 

2339) 

Rothgipfler (DVIT 

935) 

Ruby Cabernet 

(DVIT 940) 

Ruby Seedless 

(DVIT 2625) 

Sahibi Sorkh (DVIT 

510) 

Slankamenka (DVIT 

958) 

Sovrana (DVIT 525) 

Sultana Moscata 

(DVIT 527) 

Tarrango II (DVIT 

2658) 

Thomuscat (DVIT 

536) 

Torgoon (DVIT 

2531) 

Unkown Cultivar 

(DVIT 644) 

Vidal 99T (DVIT 

3185) 

Vorosmarthy (DVIT 

548) 

Wittberger (DVIT 

2689) 


1015) 


(DVIT 738) 

Cabernet Sauvignon 

(DVIT 677) 

Sultana Moscata 

(DVIT 527) 


(DVIT 468) 

Queen Of The 



408) 

Red Veltliner (DVIT 

928) 

0.00877 

0.0095 

Carignane (DVIT 848) 0.01011 

Seedless Emperor 

(DVIT 2759) 

0.00787 GRIN 

0.00911 * 

0.0086 

Lal Sorkh (DVIT 442) Husseine (DVIT 420) 0.00837 


(DVIT 841) 

Delizia Di Vaprio 

(DVIT 571) 


(DVIT 841) 

Touriga (DVIT 990) 


(DVIT 841) 

Hosargoon (DVIT 

2503) 

Kontocladi (DVIT 

1036) 


(DVIT 2067) 


(DVIT 611) 


1015) 

Catarratto (DVIT 

2058) 


408) 


530) 


530) 


530) 

0.00831 

0.00821 GRIN 

0.00704 

0.01165 

Nilok (DVIT 2504) 0.00627 

Mavrodaphne (DVIT 

821) 

Jurancon Blanc (DVIT 

1102) 


(DVIT 841) 


408) 

0.00755 GRIN 

0.00595 

0.00777 

0.00871 

0.00643 GRIN 

Table S3: Inferred trios. Each row represents an inferred trio with cultivar names followed by 

GRIN IDs in parentheses for offspring and the two parents. The proportion of SNPs that are 

inconsistent with mendelian inheritance is provided for each trio (see also Figure S1). The term 

“GRIN” in the Reference column refers to the Germplasm Resource Information Network 

(http://www.ars-grin.gov/). The asterisk in the Reference column refers to Sefc et al. [4]. The 

reference column remains empty if the inferred trio was not present in either of the two abovementioned 

sources. See the section “Supplementary Text: Potential misnaming of cultivars” for 

a discussion of sources of error in this table.

Cultivar ID Type 

Number of 1 st 

degree relatives 

Aswad DVIT 2087 table 39 

Muscat of Alexandria DVIT 841 table 30 

Thompson Seedless DVIT 535 table 28 

Kishmish-charni DVIT 2325 table 27 

Asma DVIT 2054 table 23 

Red Roumi DVIT 2083 table 21 

Gewurztraminer DVIT 738 wine 20 

Mathilde DVIT 99 table 19 

Olivette Vendemien DVIT 482 table 18 

Gouveio DVIT 743 wine 17 

Karadzhandal DVIT 2323 table 17 

Appley Towers DVIT 342 table 15 

Muscat Hamburg DVIT 468 table 15 

Lal Sorkh DVIT 442 table 14 

Noir Dautomne DVIT 330 table 14 

Precoce Dastrachan DVIT 321 table 14 

Chasselas Tokay DVIT 377 table 12 

Munuka Sufed DVIT 2046 table 10 

Opale DVIT 483 table 10 

Pinot Noir DVIT 915 wine 10 

Queen Of The Vineyard DVIT 496 table 10 

Rizamat DVIT 2338 table 10 

Cardinal DVIT 2651 table 10 

Table S4: The most highly connected cultivars. Cultivars with 10 or more inferred first degree 

relationships from the present data set are listed.

Cultivar Clones 

030-33 (DVIT 1796) 030-37 (DVIT 1797) 

030-37 (DVIT 1797) 030-33 (DVIT 1796) 

030-44 (DVIT 1798) 030-51 (DVIT 1799) 

030-51 (DVIT 1799) 030-44 (DVIT 1798) 

030-53 (DVIT 1800) 033-50 (DVIT 1801) 

033-50 (DVIT 1801) 030-53 (DVIT 1800) 

034-16 (DVIT 1803) 

034-26 (DVIT 1805) 

034-29 (DVIT 1804) 

034-54 (DVIT 1806) 

034-55 (DVIT 1807) 

035-11 (DVIT 1809) 

035-47 (DVIT 1812) 

035-58 (DVIT 1814) Olmo (035-50) (DVIT 1813) 

Parent 

Offspring 

Ab Jusht (DVIT 333) Ghula Dari (DVIT 2039) Ghula Dari (DVIT 409) 

Adjem Misquet (DVIT 306) 

Affenthaler (DVIT 618) 

Agadia (DVIT 334) 

Aglianico (DVIT 619) 

Agoumastos (DVIT 1024) 

Agria (DVIT 2701) 

Aitah (DVIT 2037) 

Aitah Ghulaman (DVIT 336) 

Ajmi (DVIT 563) 

Ak Schekerek (DVIT 612) 

Al Borla (DVIT 307) 

Mavro Spetzon (DVIT 824), 

Tsoumbrena Mavri (DVIT 

1055), Kara Lakana (DVIT 

773) 

Blue Affenthaler (DVIT 662), 

Kyoho (DVIT 616) 

Rodiares (DVIT 1048), 

Pardala (DVIT 883), 

Kokinorobola (DVIT 777) 

Husseine (DVIT 420), Aitah 

Ghulaman (DVIT 336) 

Husseine (DVIT 420), Aitah 

(DVIT 2037) 

Psarotherico (DVIT 1047), 

Pinger (DVIT 2649) 

Sahibi Sorkh (DVIT 510), 

Fahri (DVIT 402) 



Sibling 

030-44 (DVIT 1798), 030-53 (DVIT 

1800), 034-29 (DVIT 1804) 

030-44 (DVIT 1798), 030-53 (DVIT 

1800), 034-29 (DVIT 1804) 

034-16 (DVIT 1803), 030-33 (DVIT 

1796), 030-53 (DVIT 1800), 034-29 

(DVIT 1804) 

034-16 (DVIT 1803), 030-33 (DVIT 

1796), 030-53 (DVIT 1800), 034-29 

(DVIT 1804) 

034-16 (DVIT 1803), 030-44 (DVIT 

1798), 030-33 (DVIT 1796), 034-29 

(DVIT 1804) 

034-16 (DVIT 1803), 030-44 (DVIT 

1798), 030-33 (DVIT 1796), 034-29 

(DVIT 1804) 

030-44 (DVIT 1798), 030-53 (DVIT 

1800) 

034-55 (DVIT 1807), 034-54 (DVIT 

1806), Olmo (035-07) (DVIT 1808), 035- 

11 (DVIT 1809) 

030-44 (DVIT 1798), 030-33 (DVIT 

1796), 030-53 (DVIT 1800) 

034-55 (DVIT 1807), 034-26 (DVIT 

1805), Olmo (035-07) (DVIT 1808), 035- 

11 (DVIT 1809) 

034-54 (DVIT 1806), 034-26 (DVIT 

1805), Olmo (035-07) (DVIT 1808), 035- 

11 (DVIT 1809) 

034-55 (DVIT 1807), 034-54 (DVIT 

1806), 034-26 (DVIT 1805), Olmo (035- 

07) (DVIT 1808) 

Olmo (035-41) (DVIT 1811) 

Appley Towers (DVIT 342), Noir 

Dautomne (DVIT 330), Precoce 

Dastrachan (DVIT 321), Aswad (DVIT 

2087), Aspruda Santorinih (DVIT 1027) 

Noir Dautomne (DVIT 330), Aswad 

(DVIT 2087) 

Katta Kurgan (DVIT 774) 


Zerk (DVIT 606)

Albana (DVIT 622) Ter Gulmer (DVIT 976) 

Alburla (DVIT 634) 

Aleatico (DVIT 623) 

Alicante Bouschet (DVIT 624) 

Alicante Bouschet (DVIT 625) 

Alicante Branca (DVIT 1025) 

Alicante Ganzin (DVIT 3155) 

Alicante Vermelho (DVIT 626) 

Alloued Zeine (DVIT 337) 

Anatolitico (DVIT 629), 

Alicante Bouschet (DVIT 

625) 



624) 

Almeria (DVIT 1079) Ohanes (DVIT 478) 

Muscat Galego (DVIT 859) 

Alicante Ganzin (DVIT 

3155), Petite Bouschet 

(DVIT 894) 



(DVIT 894) 

Anatolitico (DVIT 629) 

Calmeria (DVIT 1084), B 

15-19 (DVIT 2171), Bien 

Donnie (DVIT 2696) 

Alulu (DVIT 338) Rosa Minna (DVIT 322) Zimsko Belo (DVIT 2664) 

Alvarelhao (DVIT 1058) 

Anacelatta (DVIT 630) 


Angelina (DVIT 339) 


625), Alicante Bouschet 

(DVIT 624) 

Seedless Tokay (DVIT 1308), 

Petinos (DVIT 591), Olho De 

Pargo (DVIT 589), Flame 

Tokay (DVIT 2913), Tokay 

(DVIT 540), Seedless Tokay 

(DVIT 1330) 

Angelo Pirovano (DVIT 564) Clotilde Prosperi (DVIT 568) 

Angevine Oberlin (DVIT 340) 

Angulata (DVIT 341) 

Anna Maria (DVIT 3156) 

Antibo (DVIT 631) 

Appley Towers (DVIT 342) 

Araclinos (DVIT 632) 


Arcathinoh (DVIT 635) 

NA (DVIT 1981), NA (DVIT 

1979), Kepraico (DVIT 787), 

Black Muscat Alexandria 

(DVIT 2111), Karrithato 

(DVIT 770) 

Gaschochi (DVIT 2537), 

Vranac (DVIT 2086) 



(DVIT 894) 

Exotic (DVIT 401), 

Hungarian Millenium 

(DVIT 761) 

Markandi (DVIT 587), 

Italia (IP 65) (DVIT 424) 


2648), Perla Di Csaba 

(DVIT 2042), Delizia Di 

Vaprio (DVIT 571) 

Nebbiolo (DVIT 867) 

Gros Colman (DVIT 413) 

Petite Bouschet (DVIT 

894) 

Korfiatis (DVIT 784) 

Noir Dautomne (DVIT 330) 



Olivette Bauthelet (DVIT 590) 

Royalty (DVIT 937), Saint Macaire 

(DVIT 943) 

Olivette Bauthelet (DVIT 590), Olivette 

Vendemien (DVIT 482), Russian 

Seedless (DVIT 2431) 

Red Roumi (DVIT 2083), Aswad (DVIT 

2087), Karadzhandal (DVIT 2323) 

Aurora (DVIT 345), Aswad (DVIT 2087), 

Olivette Vendemien (DVIT 482), Black 

Ferrara (DVIT 356) 


Nasa Veltliner (DVIT 866) 

Primiera (DVIT 491) 

Gouveio (DVIT 743), Aurora (DVIT 

345), Noir Dautomne (DVIT 330), Mavro 

Spetzon (DVIT 824), Precoce Dastrachan 

(DVIT 321), Aswad (DVIT 2087), 

Mandilaria (DVIT 815), Lady Hutt 

(DVIT 2159), Ribier (DVIT 503), 

Olivette Vendemien (DVIT 482), Robin 

(DVIT 2651), Mathilde (DVIT 99), Asma 

(DVIT 2054), Vustolethi (DVIT 603) 


Mavroliatis (DVIT 1040) Karistino (DVIT 771)

Argant (DVIT 2305) 

Arhaniotico Kokino (DVIT 636) 

Arnsburger (DVIT 2666) 

Aromatico (DVIT 637) 

Hakiki (DVIT 559), Hakiki 

(DVIT 2089) 

Askari (DVIT 343) Laubschers Gem (DVIT 444) 

Askary (DVIT 565) 

Askathari (DVIT 638) 

Asma (DVIT 2054) 

Asma Sohwars (DVIT 309) 

Aspiran Noir (DVIT 639) Queen (DVIT 495) 

Asprocoritho (DVIT 640) 

Asprorombola (DVIT 641) 

Aspruda Ariloghi (DVIT 642) 

Aspruda Halkidos (DVIT 1026) 

Aspruda Mikynon (DVIT 643) 

White Tokay (DVIT 550), 

Stravaganza Vidor (DVIT 

2259), Rhazaki Anatolico 

(DVIT 608) 

Kakourdess Weis (DVIT 

317), Monica (DVIT 461) 

Platirahah (DVIT 920), 

Patrino (DVIT 1356) 

Tavriz (DVIT 2659), 

Longyan (DVIT 2079), 


Barmak Isium (DVIT 310), 

Muscat Caillaba (DVIT 

2389), Schiradzouli Blanc 

(DVIT 514) 

Aspruda Santorinih (DVIT 1027) Royale 4x (DVIT 1666) 

Assirtico (DVIT 645) Atzala (DVIT 646) 

Aswad (DVIT 2087) 

Leanyka (DVIT 794) 

Chasselas Tokay (DVIT 

377), Mueller Thurgau 

(DVIT 2337) 

Semillon (DVIT 954) 


Muscat Hamburg (DVIT 

468) 

Sultanina Rose (DVIT 530) 


317), Cirmisi Sap De 

Sudak (DVIT 313), Tadone 

(DVIT 970) 

Pelaverga (DVIT 892) 


530), Gold (DVIT 410), 


468) 

Dumbrehnah Mavri (DVIT 

708), Asma (DVIT 2054) 

Platani (DVIT 1045), 

Guydouria (DVIT 730) 

Gouveio (DVIT 743), Noir 

Dautomne (DVIT 330), 

Henab (DVIT 417), 

Olivette Vendemien (DVIT 

482), Mathilde (DVIT 99), 

Tadone (DVIT 970), Opale 

(DVIT 483) 

Kishmish-charni (DVIT 2325), Kishmish 

Sorkh (DVIT 437), Muscat Szauter 

(DVIT 863), Lal Sorkh (DVIT 442) 

Migdali (DVIT 830), Gouveio (DVIT 

743), Appley Towers (DVIT 342), Noir 

Dautomne (DVIT 330), Sultanina Rose 

(DVIT 530), Precoce Dastrachan (DVIT 

321), Red Roumi (DVIT 2083), Yaghotti 

No. 1 (DVIT 604), Aswad (DVIT 2087), 

Mandilaria (DVIT 815), Dumbrehnah 

Mavri (DVIT 708), Yaghotti No. 2 (DVIT 

2052), Dschan Im Isium (DVIT 315), 

Olivette Vendemien (DVIT 482), 

Kishmish-charni (DVIT 2325), 

Karadzhandal (DVIT 2323), Mathilde 

(DVIT 99), Vustolethi (DVIT 603), 

Kishmish Of Vir (DVIT 2071), Lal Sorkh 

(DVIT 442) 

Aspruda Santorinih (DVIT 1027) 

Platirahah (DVIT 920), Gouveio (DVIT 

743), Mavro Spetzon (DVIT 824), 

Precoce Dastrachan (DVIT 321), Aswad 

(DVIT 2087) 

Citronelle (DVIT 381), Aurora (DVIT 

345), Agadia (DVIT 334), Appley Towers 

(DVIT 342), Itonychi Lefko (DVIT 561), 

Black Kishmish (DVIT 358), Olivette 

Noire (DVIT 1657), Verdal (DVIT 553), 

Mavro Spetzon (DVIT 824), Alloued 

Zeine (DVIT 337), Precoce Dastrachan 

(DVIT 321), Red Roumi (DVIT 2083),

Atzala (DVIT 646) Assirtico (DVIT 645) 

Augustiatico (DVIT 566) Mavrodaphne (DVIT 821) 

Aurora (DVIT 1341) Hisakasy (DVIT 759) 

Aurora (DVIT 345) Pirovano 47 (DVIT 2047) 

Austrian Seedless (DVIT 1325) 

Austrian Seedless (DVIT 1650) 

B 15-19 (DVIT 2171) 

Badacsony-10 (DVIT 2667) 

Baharat Early (DVIT 2306) 

Sultana (DVIT 526), Sultana 

(DVIT 1321), Austrian 


Sultana (DVIT 526), Sultana 

(DVIT 1321), Austrian 


Orange Muscat (DVIT 2167), 

Moscato Jesus (DVIT 2062), 

Orange Muskat (DVIT 879), 

Tokay (DVIT 2157) 

Baidh Ul Haman (DVIT 346) Black Kishmish (DVIT 358) 

Bakari Mavro (DVIT 653) 

Balta (DVIT 2905) 

Tinta Cao (DVIT 984), Tinta 

Pinheira (DVIT 987), Tinta 

Francisca (DVIT 983) 

Barbableu (DVIT 1651) Barbableu (DVIT 348) 

Barbableu (DVIT 348) Barbableu (DVIT 1651) 

Barbara (DVIT 700) 

Barbera (DVIT 648) 

Barbera (DVIT 649) 

Bargoon (DVIT 2499) 

Barbera (DVIT 648), Barbera 

(DVIT 649) 

Barbara (DVIT 700), Barbera 

(DVIT 649) 

Barbara (DVIT 700), Barbera 

(DVIT 648) 

Barlinka (DVIT 1080) Rangspray (DVIT 1120) 

Platani (DVIT 1045), 


Mathilde (DVIT 99), 

Divromos (DVIT 706), 

Unkown Cultivar (DVIT 

644) 

Precoce Dastrachan (DVIT 

321), NA (DVIT 2044) 


482) 




530), Almeria (DVIT 

1079) 

NA (DVIT 1978), Ezerjo 

(DVIT 715) 


859), Chasselas Tokay 

(DVIT 377), Perle de 

Csaba (DVIT 2648) 


377) 

Saint Macaire (DVIT 943) 


1657), Muscat Hamburg 

(DVIT 468) 



(DVIT 468) 

Barlinka (DVIT 349) Emperor (DVIT 396) Ribier (DVIT 503) 

Klusaine Red (DVIT 2077), Almeria 

(DVIT 1079), Korfiatis (DVIT 784), 

Mandilaria (DVIT 815), Fresno 35-33 

(DVIT 2172), Sochal Kara-kash (DVIT 

2340), Rizamat (DVIT 2338), Hunisa 

(DVIT 418), Kishmish-charni (DVIT 

2325), Robin (DVIT 2651), Karadzhandal 

(DVIT 2323), Asma (DVIT 2054), 

Vustolethi (DVIT 603), Aspruda 

Santorinih (DVIT 1027), Lacturario 

(DVIT 585), Trieste 4x (DVIT 1591), 

Nasa Veltliner (DVIT 866), Berzemina 

(DVIT 656), Danuque (DVIT 387), Netter 

(DVIT 474) 


Citronelle (DVIT 381), Appley Towers 


Almeria (DVIT 1079), Mathilde (DVIT 

99) 

Calmeria (DVIT 1084) 

NA (PI 588668.04) 

Perla Di Csaba (DVIT 2042), Sauvis 

(DVIT 598) 


2087), Karadzhandal (DVIT 2323)

Barmak Isium (DVIT 310) 

Barone Dellaterno (DVIT 350) 

Bastardo (DVIT 2057) 


Beauty Seedless (DVIT 351) 

Beba (DVIT 651) 

Beclan (DVIT 652) 

Beli Potok (DVIT 1840) 


Beli Prokupac (DVIT 1069) 

Bellino (DVIT 352) 

Bequignol (DVIT 2113) 

Bermestia Bianca (DVIT 654) 

Bermestia Violacea (DVIT 655) 

Berurargoon (DVIT 2501) 

Berzamino (DVIT 657) 

Berzemina (DVIT 656) 




Aspruda Mikynon (DVIT 

643), Muscat Caillaba (DVIT 


(DVIT 514) 

Trousseau (DVIT 838), 

Chauche Noir (DVIT 690), 

Trousseau (DVIT 995), Grey 

Riesling (DVIT 745), Beba 

(DVIT 651), Bastardo (DVIT 

650) 


Bastardo (DVIT 2057), 




(DVIT 651) 





Riesling (DVIT 745), 


Bhokari (DVIT 353) Bhokri (DVIT 1081) 

Bhokri (DVIT 1081) Bhokari (DVIT 353) 

Bias Kukuzeti (DVIT 311) 

Bien Donnie (DVIT 2696) 

Black Corinth (DVIT 1052) 

Montils (DVIT 835), 

Morescono (DVIT 461) 


530), Iviar Oliver (DVIT 

578) 

Royalty (DVIT 937), 

Gewurztraminer (DVIT 

738), Trincadeiro (DVIT 

992) 




992) 

Queen Of The Vineyard 

(DVIT 496), Black 

Kishmish (DVIT 1326), 

Kishmish-charni (DVIT 

2325) 




992) 

Cola Tamburo (DVIT 694) 

Ezerjo (DVIT 715) 

Goyura (DVIT 2637), G4-74 (Seedless 

Calmeria) (DVIT 2914), Maria Pirovano 

(DVIT 588), Thomuscat (DVIT 536) 

Sauvignon Gris (DVIT 1123), Chenin 

Blanc (DVIT 691), Flora (DVIT 720) 



Perlette (DVIT 488), NA (GVIT 1504.02) 




Beli Potok (DVIT 1840), Roussaitis 

(DVIT 1049) 

Frankenthal (DVIT 408) Fosters Seedling (DVIT 407) 


738) 


482), Trebbiano Bianco 

(DVIT 2067) 

Almeria (DVIT 1079), 


(DVIT 496), Olivette 

Vendemien (DVIT 482) 

Staphythambelo (DVIT 

962), Liatico (DVIT 796) 


Blanc (DVIT 691), Fer Servadou (DVIT 

2318), Flora (DVIT 720) 

Aswad (DVIT 2087), Mandilaria (DVIT 

815) 

Kokinostaphylo (DVIT 779), Schiraz 

(DVIT 952)

Black Ferrara (DVIT 356) 

Black Hamburg (DVIT 1082) 



Black Kishmish (DVIT 1326) 

Black Kishmish (DVIT 2055) 

Black Hamburg (DVIT 357), 

Marvel Of Vaucluse (DVIT 

456), Black Hamburg (DVIT 

1309) 




456) 



456), Black Hamburg (DVIT 

1309) 

Black Kishmish (DVIT 

2055), Russian Seedless 4x 

(DVIT 1667) 


1326), Russian Seedless 4x 

(DVIT 1667) 

Black Kishmish (DVIT 358) Baidh Ul Haman (DVIT 346) 

Black Malvoisie (DVIT 658) 

Ithani Mavro (DVIT 621), 

Chasselas Rose Royal (DVIT 

2158), Hermitage (DVIT 

1099) 

Black Monukka (DVIT 359) Monukka (DVIT 462) 

Black Morocco (DVIT 360) 

Black Muscat Alexandria (DVIT 

2111) 

Black Prince (DVIT 361) 

NA (DVIT 1981), Angulata 

(DVIT 341), NA (DVIT 


Karrithato (DVIT 770) 


530), Kishmish Hishrau 

(DVIT 1103), Kishmish Of 

Vir (DVIT 2071), Ranny 

Vira (DVIT 2684), Kara 

Dzhidzhigi (DVIT 2322), 

Beauty Seedless (DVIT 

351) 








351) 


2046), Pardinia (DVIT 

486) 

Kritico (DVIT 583), Julius 

Caesar (DVIT 579), Iviar 

Oliver (DVIT 578), 

Rambola (DVIT 497), 

Itonychi Mavro (DVIT 

562) 

Pias (DVIT 833), Iviar 

Oliver (DVIT 578) 

Olivette Noire (DVIT 1657), Almeria 

(DVIT 1079), Olivette Vendemien (DVIT 

482), Zimsko Belo (DVIT 2664) 

Danuque (DVIT 387) 



Kishmish-charni (DVIT 2325) 



2087), Karadzhandal (DVIT 2323) 

Terret Gris (DVIT 980) 

Sultanina Rose (DVIT 530), Khalili 


2325), Kishmish Sorkh (DVIT 437), Lal 

Sorkh (DVIT 442) 

Opale (DVIT 483) 

Olmo 6F V19 (DVIT 2923), Olmo j5-58 

(DVIT 1314), Corniola (DVIT 3162), 

Rambola (DVIT 497) 

Black Rose (DVIT 362) Blackrose 4x (DVIT 2476) Ribier (DVIT 503) Eftakilo (DVIT 575) 

Blackrose 4x (DVIT 2476) Black Rose (DVIT 362) Ribier (DVIT 503) Eftakilo (DVIT 575) 

Blanc Rame (DVIT 659) 

Blaue Elbe (DVIT 660) 

Blauer Fruheburgunder (DVIT 

2307) 

Pinot Noir Espinett (DVIT 

2332), Pinot Noir 2a (DVIT 

2333), Pinot Noir-marianfeld 

(DVIT 2334), Blauer 

Spatburgunder (DVIT 661), 

Pinot Noir (DVIT 915), 

Chenin Blanc (DVIT 691) 

Chardonnay (DVIT 687), 

Muscat Canelli (DVIT 

857), Gamay Burgoyne 

(DVIT 734), NA (GVIT 

1614.02), Gewurztraminer 

(DVIT 738), Precoce De 

Folle Blanche (DVIT 724), Colombard 

(DVIT 695) 

Green Hungarian (DVIT 748) 

Teoulier Noir (DVIT 975)

Blauer Spatburgunder (DVIT 661) 

Blue Affenthaler (DVIT 662) 

Blue Trollinger (DVIT 663) 

Boal Branco (DVIT 664) 

Bonarda (DVIT 671) 

Bondola (DVIT 3067) 

Boyalsing II (DVIT 2505) 

Buaki (DVIT 673) 

Bucklands Sweetwater (DVIT 

363) 

Budrilay (DVIT 2506) 

Vrathiano (DVIT 1012), 

Gamay Beaujolais (DVIT 

733), Pinot Noir (DVIT 


911), Pinot Noir (DVIT 913), 

Pinot Noir (DVIT 914), Late 

Blue Burgundy (DVIT 793), 

Pinot Nero (DVIT 909), 

Serine (DVIT 955), Pinot 

Pernard (DVIT 905), Pinot 

(DVIT 733) 




(DVIT 2334), Pinot Noir 

(DVIT 915), Vrathiano 

(DVIT 1012), Gamay 

Beaujolais (DVIT 733), Pinot 

Noir (DVIT 1066), Pinot Noir 



(DVIT 914), Late Blue 

Burgundy (DVIT 793), Pinot 

Nero (DVIT 909), Blauer 

Fruheburgunder (DVIT 

2307), Serine (DVIT 955), 

Pinot Pernard (DVIT 905), 

Pinot (DVIT 733) 

Kyoho (DVIT 616), 


Frankenthal (DVIT 408), 


1059), Trollinger (DVIT 

2341), Hillbilly (DVIT 2747) 

Psevthosirico (DVIT 592), 

Clairette Blanche (DVIT 

3003), Tsoumbrena Lefki 

(DVIT 997) 

Chasselas Tokay (DVIT 377), 

Chasselas Musque (DVIT 

374), Chassalas Jababer 

(DVIT 2120), Chasselas 

Jaunce Cire (DVIT 2116), 

Chasselas Ciotat (DVIT 372), 

Chasselas Dore (DVIT 689), 

Chasselas Rose (DVIT 376), 

Ciotat (DVIT 2107), Van Der 

Laan (DVIT 543), Chasselas 

Doree Apyrene (DVIT 1643), 

Chasselas Hs 14-33/4 (DVIT 

2310), Chasselas Rouge 

(DVIT 3002) 

Malingre (DVIT 490), 

Yellow Ortlieber (DVIT 

1020), NA (DVIT 1978), 

Schonburger (DVIT 1124) 









1020), NA (DVIT 1978), 


Wittberger (DVIT 2689), 

Kerner (DVIT 3104), 

Fosters Seedling (DVIT 

407), Chasselas Napoleon 

(DVIT 611), Muscat 

Hamburg (DVIT 468), 

Rotberger (DVIT 2339), 

Bellino (DVIT 352), 



693) 

Pelaverga (DVIT 892), 

Rodites (DVIT 609) 

Arnsburger (DVIT 2666), 

Muscat St. Laurent (DVIT 

862), Orange Muscat 

(DVIT 2167), Perla Di 

Csaba (DVIT 487), NA 

(DVIT 1980), Madam 

Matijas (DVIT 2642), 



Napoleon (DVIT 611), 

Bucklands Sweetwater 

(DVIT 363), Olmo 25F 

(DVIT 2751), Bakari 

Mavro (DVIT 653), 


471) 


377), Folle Noire (DVIT 

725) 

Teoulier Noir (DVIT 975) 

Chasselas Tompa (DVIT 378) 

Dalnato I (DVIT 2509)

Burgrave (DVIT 364) 

Cabernet Franc (DVIT 803) 

Cabernet Sauvignon (DVIT 677) 


Calmeria (DVIT 365.9) 


Calzin (DVIT 678) 

Canner (DVIT 367) 

Cannon Hall (DVIT 368) 

Poet Matabon (DVIT 1597), 

Poet Matabon (DVIT 1658) 

Calmeria (DVIT 366), 






Olivette Noire (DVIT 1657), 

Olivette Noir (DVIT 1306) 

Iviar Oliver (DVIT 578), 

Muscat Of Alexandria (DVIT 

466), Red Hanepoot (DVIT 

593), Muscat of Alexandria 

(DVIT 841), Muscat Bowood 

(DVIT 463), Muscat of 

Alexandria (DVIT 465) 

Merlot (DVIT 2682), 

Cabernet Sauvignon (DVIT 

677) 

Sauvignon Gris (DVIT 

1123), Cabernet Franc 

(DVIT 803), Ruby 

Cabernet (DVIT 940) 



1079), G4-74 (Seedless 

Calmeria) (DVIT 2914) 



1079), G4-74 (Seedless 




1079), G4-74 (Seedless 


B 15-19 (DVIT 2171), Olivette 






Zinfandel (DVIT 1340) Helena (DVIT 756) 

Prune De Cazouls (DVIT 

494), Barbableu (DVIT 

348) 


859), Chasselas Tompa 

(DVIT 378), Olmo 6F V19 

(DVIT 2923), Olmo j5-58 

(DVIT 1314), Moscatel 

Ros. De Blas (DVIT 2328), 

Muscat De Ladda (DVIT 

2166), Futuna 699 (DVIT 

2073), Corniola (DVIT 

3162), Muscat Szauter 



Muscat De Pantellana 

(DVIT 2063), Fuji (DVIT 

1655), Fort Ross Vine 


Vaprio (DVIT 571), Malta 

Seedless (DVIT 2327), 


468), Erlihane (DVIT 

2671), Goyura (DVIT 

2637), Maria Pirovano 

(DVIT 588), Opale (DVIT 

483), Mrs Pince Black 

Muscat (DVIT 2160), 

Nonno Giovanni (DVIT 

2257), Neo-Muscat (DVIT 

2645), Barone Dellaterno 

(DVIT 350), Black Prince 

(DVIT 361), Thomuscat 

(DVIT 536), Vorosmarthy 


Morocco (DVIT 360), 

Slankamenka (DVIT 958) 

Aswad (DVIT 2087), Olivette Vendemien 

(DVIT 482), Black Ferrara (DVIT 356) 

July Muscat (DVIT 426)

Cardinal (DVIT 370) 

Carignane (DVIT 1064) 



Carriere (DVIT 953) 

Carvalheira (DVIT 682) 

Casculho (DVIT 2631) 

Robin (DVIT 2651), 

Yugoslavia 360 (DVIT 2344), 

Chaouch 4x (DVIT 1663) 

Carignane (DVIT 848), 

Naboda (DVIT 865), 


Neiretto Verzuolo (DVIT 

875) 





875) 





875) 

Castelao (DVIT 683) Joao Santarem (DVIT 764) 


(DVIT 496), Ribier (DVIT 

503) 

Exotic (DVIT 401), Appley Towers 


Olivette Vendemien (DVIT 482), Fresno 

32-68 (DVIT 2746), Early Muscat (DVIT 

394), Dan Ben Chana (DVIT 2633), 

Flame Seedless (DVIT 1095) 

Ruby Cabernet (DVIT 940) Olmo 34F (DVIT 2752) 



NA (PI 340969.03), 


738), Loin De Loeil (DVIT 

798) 

Catarratto (DVIT 1085) Catarratto (DVIT 2058) Slankamenka (DVIT 958) 

Catarratto (DVIT 2058) Catarratto (DVIT 1085) Slankamenka (DVIT 958) 

Catarratto (DVIT 684) 


Olmo 6F V19 (DVIT 2923), 

Prokupao (DVIT 2926) 

CG 1481 (DVIT 2432) CG 1481 4x (DVIT 2480) 

CG 1481 4x (DVIT 2480) CG 1481 (DVIT 2432) 

Chalian De Drome (DVIT 2309) 

Chamo (DVIT 2730) 

Chan Isium (DVIT 613) 

Chaouch (DVIT 371) 


Clairette De Limoux (DVIT 

2311) 

Chaouchi Politico (DVIT 

599) 



Cardinal (DVIT 370) 

Chaouchi Politico (DVIT 599) Chaouch (DVIT 371) 




561), Sultanina Rose 

(DVIT 530) 


561), Sultanina Rose 

(DVIT 530) 

Schiraz (DVIT 952), 

Ranny Vira (DVIT 2684) 



503) 



Charas (DVIT 685) Tscharas (DVIT 538) Klusaine Red (DVIT 2077) 

Charbono (DVIT 686) 

Jampal (DVIT 763), Rhazaki Arhanon 

(DVIT 554), Tamarez (DVIT 972), 

Unkown Cultivar (DVIT 693) 

Futuna 699 (DVIT 2073), Corniola 

(DVIT 3162), Rambola (DVIT 497), 









Flame Seedless (DVIT 1095)

Chardonay (DVIT 816) 


Chardonnay (DVIT 1332) 

Chardonnay (DVIT 687) 

Chardonnay Musque (DVIT 1652) 

Chassalas Jababer (DVIT 2120) 

Chasselas Ciotat (DVIT 372) 

Chasselas Des Bouches-du-rhone 

(DVIT 2112) 

Chasselas Dore (DVIT 373) 


1020), Chardonnay Musque 

(DVIT 1652), Chardonnay 


Epernay (DVIT 916), Pinot 

Vert Dore (DVIT 918), Pinot 

Musque (DVIT 1117) 




Epernay (DVIT 916), 

Chardonay (DVIT 816), Pinot 

Vert Dore (DVIT 918), Pinot 



1020), Chardonnay (DVIT 

1332), Pinot Noir Epernay 

(DVIT 916), Chardonay 

(DVIT 816), Pinot Vert Dore 

(DVIT 918), Pinot Musque 

(DVIT 1117) 



374), Chasselas Jaunce Cire 

(DVIT 2116), Buaki (DVIT 

673), Chasselas Ciotat (DVIT 

372), Chasselas Dore (DVIT 

689), Chasselas Rose (DVIT 

376), Ciotat (DVIT 2107), 

Van Der Laan (DVIT 543), 

Chasselas Doree Apyrene 

(DVIT 1643), Chasselas Hs 

14-33/4 (DVIT 2310), 

Chasselas Rouge (DVIT 

3002) 






Buaki (DVIT 673), Chasselas 

Dore (DVIT 689), Chasselas 

Rose (DVIT 376), Ciotat 

(DVIT 2107), Van Der Laan 

(DVIT 543), Chasselas Doree 

Apyrene (DVIT 1643), 



(DVIT 3002) 


Folle Noire (DVIT 725), 

Chasselas Napoleon (DVIT 

375) 


2332), Olmo 25F (DVIT 

2751) 



2751) 


2332) 



2751) 
















471) 
















471) 


377), Sauvis (DVIT 598), 

Diamente Nero (DVIT 


(DVIT 468), Lombardia 

(DVIT 447), Conte Rosso 

(DVIT 382) 

Muscat Br39 (DVIT 1313), 

Italia (IP 65) (DVIT 424), 


(DVIT 363) 

Chardonnay (DVIT 687), Muscat Canelli 

(DVIT 857), Gamay Burgoyne (DVIT 

734) 

Gouveio (DVIT 743), Sauvignon Vert 

(DVIT 949) 



734) 

Muscat Canelli (DVIT 857), Gamay 

Burgoyne (DVIT 734), Yellow Ortlieber 

(DVIT 1020) 



734) 

Chasselas Napoleon (DVIT 611)


Chasselas Doree Apyrene (DVIT 

1643) 


2310) 

Chasselas Jaunce Cire (DVIT 

2116) 

Chasselas Michel Tompa (DVIT 

2121) 







Ciotat (DVIT 372), Chasselas 







(DVIT 3002) 












14-33/4 (DVIT 2310), 


3002) 














3002) 








376), Ciotat (DVIT 2107), 




14-33/4 (DVIT 2310), 


3002) 

















471) 
















471) 
















471) 
















471) 



611), Nicolas Horthy 

number39 (DVIT 475) 

Frankenthal (DVIT 408), Muscat 

Hamburg (DVIT 468), Opale (DVIT 

483), Vorosmarthy (DVIT 548)

Chasselas Musque (DVIT 374) 

Chasselas Napoleon (DVIT 375) 


Chasselas Rose (DVIT 376) 


2158) 

Chasselas Rouge (DVIT 3002) 


Chassalas Jababer (DVIT 

2120), Chasselas Jaunce Cire 





376), Ciotat (DVIT 2107), 




14-33/4 (DVIT 2310), 


3002) 

Folle Noire (DVIT 725), 









Dore (DVIT 689), Ciotat 






(DVIT 3002) 


Black Malvoisie (DVIT 658), 

Hermitage (DVIT 1099) 














2310) 
















471) 




(DVIT 363) 


Chasselas Tompa (DVIT 



(DVIT 548) 
















471) 
















471) 


Terret Gris (DVIT 980)

Chasselas Tokay (DVIT 377) 


Chauche Noir (DVIT 690) 

Chenin Blanc (DVIT 691) 

Chenin Noir (DVIT 692) 

Choultu White (DVIT 2909) 

Chrisostaphylo (DVIT 567) 

Churgoon (DVIT 2532) 

Ciminnita (DVIT 1653) 

Ciotat (DVIT 2107) 














(DVIT 3002) 

Chasselas Michel Tompa 

(DVIT 2121) 






650) 


482) 









Rose (DVIT 376), Van Der 

Laan (DVIT 543), Chasselas 




(DVIT 3002) 

Cipro Nero (DVIT 380) Kritico (DVIT 583) 

Cirmisi Sap De Sudak (DVIT 313) 
















471) 








992) 


738), Colombard (DVIT 

695), Blanc Rame (DVIT 

659) 

Precoce De Malingre 

(DVIT 490) 

Aurora (DVIT 345), 

Aswad (DVIT 2087), 

Bermestia Bianca (DVIT 

654), Bien Donnie (DVIT 

2696) 
















471) 


360) 


708), Dschan Im Isium 

(DVIT 315), Asma (DVIT 

2054) 

Frankenthal (DVIT 408), Muscat 

Hamburg (DVIT 468), Opale (DVIT 

483), Vorosmarthy (DVIT 548) 



Trousseau (DVIT 838), Sauvignon Gris 

(DVIT 1123), Bequignol (DVIT 2113) 

Gouveio (DVIT 743), Appley Towers 

(DVIT 342), Noir Dautomne (DVIT 330), 

Olivette Noire (DVIT 1657), Red Roumi 

(DVIT 2083), Almeria (DVIT 1079), 

Calmeria (DVIT 1084), Robin (DVIT 

2651), Mathilde (DVIT 99), Asma (DVIT 

2054), Danuque (DVIT 387), Daphne 

(DVIT 570), Black Ferrara (DVIT 356), 


Migdali (DVIT 830)

Citronelle (DVIT 381) 

Clairette Blanche (DVIT 3003) 

Clairette De Limoux (DVIT 2311) 

Bondola (DVIT 3067), 


Tsoumbrena Lefki (DVIT 

997) 

Chalian De Drome (DVIT 

2309) 

Clotilde Prosperi (DVIT 568) Angelo Pirovano (DVIT 564) 


Colombard (DVIT 695) 

Conte Rosso (DVIT 382) 

Corbu Blanc (DVIT 2313) 

Coristano (DVIT 697) 

Cornifesto (DVIT 698) 

Corniola (DVIT 3162) 

Criolla Mesa (DVIT 385) 

Curisti Mici (DVIT 703) 

Dabouki (DVIT 1982) 


Dais-el-anz (DVIT 569) 

Dalnato I (DVIT 2509) 

French Colombard (DVIT 

726) 

Pleven Mychat (DVIT 3171), 

San Juanina Rosada (DVIT 

2927), Criolla Mesa (DVIT 

385) 

Corniola (DVIT 3162), 


San Juanina Rosada (DVIT 

2927) 


708), Ghlicopati (DVIT 741) 

Gamay Burgoyne (DVIT 

734), Somarello Rosso (DVIT 

3188), Gamay Teinturier 

(DVIT 737) 

Damas Rose (DVIT 1092) Damas Rose (DVIT 386) 

Damas Rose (DVIT 386) Damas Rose (DVIT 1092) 

Conte Rosso (DVIT 382), 

Pardinia (DVIT 486) 





Beclan (DVIT 652) 


Grillo (DVIT 752), Mathilde (DVIT 99) 

Gouveio (DVIT 743), Lagorthi (DVIT 

789) 

Chenin Blanc (DVIT 691) Blanc Rame (DVIT 659) 

Citronelle (DVIT 381), NA 

(DVIT 1980) 







Sudak (DVIT 313) 

Dan Ben Chana (DVIT 2633) Dan Ben Hanna (DVIT 2669) Ribier (DVIT 503) 

Dan Ben Hanna (DVIT 2669) Dan Ben Chana (DVIT 2633) Ribier (DVIT 503) 


Daphnata(daphnia?) (DVIT 704) 

Daphne (DVIT 570) 

Dardari (DVIT 2536) 

Dehbina (DVIT 705) 

Delizia Di Vaprio (DVIT 571) 

Demir Kara (DVIT 314) 

Retino Mavro (DVIT 931), Psara (DVIT 

1046), Roussaitis (DVIT 1049) 

Olmo 6F V19 (DVIT 2923), Futuna 699 


Opale (DVIT 483), Black Prince (DVIT 

361) 




361) 

Migdali (DVIT 830), Dschan Im Isium 

(DVIT 315), Asma (DVIT 2054) 


Netter (DVIT 474) Dabouki (DVIT 1982) 

Suputinsky Bietyz (DVIT 

2702) 

Voltzicki (DVIT 1011) 


Anna Maria (DVIT 3156), 

Primiera (DVIT 491), 


Boyalsing II (DVIT 2505) 

Robin (DVIT 2651), Muscat Hamburg 

(DVIT 468), Netter (DVIT 474) 

Robin (DVIT 2651), Muscat Hamburg 

(DVIT 468), Netter (DVIT 474) 

Aswad (DVIT 2087), Black Hamburg 

(DVIT 1082), Olivette Vendemien (DVIT 

482), Mathilde (DVIT 99), Zlahtina 

(DVIT 607) 

Olivette Vendemien (DVIT 482)

Dermatas (DVIT 572) 

Diamente Nero (DVIT 614) 

Diamond Muscat (DVIT 2900) 

Divromos (DVIT 706) 

Dolcetto (DVIT 707) 

Donzelinho De Castelo (DVIT 

1029) 

Donzillinho (DVIT 2634) 

Doradillo (DVIT 2635) 

Dornfelder (DVIT 2705) 

Dschan Im Isium (DVIT 315) 

Duc De Malakoff (DVIT 393) 

Dumbrehnah Mavri (DVIT 708) 

Durif (DVIT 710) 


Early Muscat (DVIT 2911) 



Donzelinho De Castelo 

(DVIT 1029) 

Perla Di Csaba (DVIT 487), 

Mamelon (DVIT 586), 

Muscat Talebot (DVIT 473) 

Curisti Mici (DVIT 703), 

Ghlicopati (DVIT 741) 

Early Muscat (DVIT 394), 

Potamisi (DVIT 922) 



Eftakilo (DVIT 575) Summer Royal (DVIT 2899) 

Ehrenfelser (DVIT 2317) 

Emerald Riesling (DVIT 712) 

Emile Royal (DVIT 395) 

Ribier (DVIT 503), Royal E 


594), Royal (DVIT 596) 


Lombardia (DVIT 447), Opale (DVIT 

483) 

Summer Muscat (DVIT 2902), Flame 


Mavrodaphne (DVIT 821) Mathilde (DVIT 99) 

Gros Manzenc (DVIT 753) 

Cirmisi Sap De Sudak 

(DVIT 313) 


377) 





Durif (DVIT 710), Syrah 

(DVIT 1053) 



Hamburg (DVIT 468) 





1015) 

Muscadelle De Bordelais 

(DVIT 855) 

Black Rose (DVIT 362), 


Emperor (DVIT 396), 



468), Olmo 18-226 D 

(DVIT 2749), Dan Ben 

Chana (DVIT 2633), Gros 

Colman (DVIT 413) 

Emperor (DVIT 396) Barlinka (DVIT 349) Ribier (DVIT 503) 

English Colossal (DVIT 397) 

Erbalus Caluso (DVIT 713) 

Erlihane (DVIT 2671) 

Esparte (DVIT 2912) 

Etraire De Ladhui (DVIT 714) 

Eununi (DVIT 400) 

Exotic (DVIT 401) 


2759) 

Mourvedre Famellestadt 

(DVIT 319) 

Lacturario (DVIT 585), 

Lattuario (DVIT 445) 

Olivette Bauthelet (DVIT 

590), Ruby Seedless 

(DVIT 2625) 

Iviar Oliver (DVIT 578) 

Angelina (DVIT 339), 


Gewurztraminer (DVIT 738) 


Murma Isium (DVIT 320), Korfiatis 

(DVIT 784), Dumbrehnah Mavri (DVIT 

708), Taschli (DVIT 325), Asma (DVIT 

2054) 



Robin (DVIT 2651) 


Black Rose (DVIT 362) 

Mueller Thurgau (DVIT 2337) 


Himrisnky (DVIT 2175), Trincadeiro 

Branco (DVIT 993) 


2087) 

Robin (DVIT 2651)

Ezerfuertu (DVIT 2672) 

Ezerjo (DVIT 715) 

Fahri (DVIT 402) Fahri Kalamak (DVIT 403) 

Fahri Kalamak (DVIT 403) Fahri (DVIT 402) 

Fakdi (DVIT 573) 

Fatai (DVIT 2510) 

Favorita Segaria (DVIT 716) 

Fayoumi (DVIT 2636) 

Feher Som (DVIT 717) 

Feher Szagos (DVIT 718) 

Fer Servadou (DVIT 2318) 

Fernao Pires (DVIT 1063) 


Vital (DVIT 1077), Fernao 

Pires (DVIT 719) 

Vital (DVIT 1077), Fernao 

Pires (DVIT 1063) 

Ferral De Izalva (DVIT 1030) Ferral De Izalva (DVIT 405) 

Ferral De Izalva (DVIT 405) Ferral De Izalva (DVIT 1030) 

Fileri (DVIT 1044) 


Flame Tokay (DVIT 2913) 

Flora (DVIT 720) 

Fogarina (DVIT 721) 

Retino Lefko (DVIT 1067), 

Velana (DVIT 1057) 




Pargo (DVIT 589), Tokay 


(DVIT 1330) 

Gouveio (DVIT 743), 


738) 

Badacsony-10 (DVIT 

2667), NA (PI 588668.04), 

Beli Prokupac (DVIT 

1069) 

Husseine (DVIT 420), 


2046), Katta Kurgan 

(DVIT 774) 

Husseine (DVIT 420), 


2046), Katta Kurgan 

(DVIT 774) 

Gouveio (DVIT 743) 

Millishun (DVIT 2498) Suputinsky Bietyz (DVIT 2702) 



693) 


693) 



(DVIT 761) 


738), Semillon (DVIT 954) 

Raboso Piave (DVIT 927) 

Fokianoh (DVIT 722) Yourutico (DVIT 605) Markandi (DVIT 587) 

Folle Blanche (DVIT 1335) 




Folle Blanche (DVIT 724), 













1102), NA (PI 588326.03) 


1102), NA (PI 588326.03) 


1102), NA (PI 588326.03) 


1102), NA (PI 588326.03) 

Kishmish-charni (DVIT 2325), Lal Sorkh 

(DVIT 442) 

Flora (DVIT 720), Bequignol (DVIT 

2113) 

Mosho Patata (DVIT 1043), Psara (DVIT 


Summer Muscat (DVIT 2902), Fresno 32- 

139 (DVIT 2169), Fresno 32-68 (DVIT 

2746), Perlette (DVIT 488), Robin (DVIT 

2651), Diamond Muscat (DVIT 2900) 


Trousseau (DVIT 838), Sauvignon Gris 

(DVIT 1123), Fer Servadou (DVIT 2318), 


Montils (DVIT 837), Blanc Rame (DVIT 

659), Juracon Noir (DVIT 2321) 






659), Juracon Noir (DVIT 2321)

Folle Noire (DVIT 725) 

Fort Ross Vine (DVIT 1353) 

Fosters Seedling (DVIT 407) 

Fosters White Seedling (DVIT 

2161) 

Frankenthal (DVIT 408) 

Chasselas Dore (DVIT 373), 


375) 

Fosters White Seedling 

(DVIT 2161) 




(DVIT 363) 

Zinfandel (DVIT 1340), 


Frankenthal (DVIT 408) Bellino (DVIT 352) 

Fosters Seedling (DVIT 407) Frankenthal (DVIT 408) Bellino (DVIT 352) 



2341), Hillbilly (DVIT 2747), 












French Colombard (DVIT 726) Colombard (DVIT 695) Chenin Blanc (DVIT 691) Blanc Rame (DVIT 659) 

Fresno 32-139 (DVIT 2169) 

Fresno 32-68 (DVIT 2746) 



Fuji (DVIT 1655) 

Furmint (DVIT 728) 



Russian Seedless (DVIT 

2431) 


Futuna 699 (DVIT 2073) Futuna 700 (DVIT 2076) Iviar Oliver (DVIT 578) 

Futuna 700 (DVIT 2076) Futuna 699 (DVIT 2073) Iviar Oliver (DVIT 578) 

G4-74 (Seedless Calmeria) (DVIT 

2914) 

Gamay (DVIT 732) 

Gamay Beaujolais (DVIT 733) 

Gamay Burgoyne (DVIT 734) 

Gewurztraminer (DVIT 738), 

Traminer (DVIT 991), 








Vrathiano (DVIT 1012), Pinot 








2307), Serine (DVIT 955), 



Somarello Rosso (DVIT 


(DVIT 737), Gamay Rond 


Gamay Toma (DVIT 

2100), Malbec (DVIT 802) 









1020), NA (DVIT 1978), 



2332) 

Sultanina Rose (DVIT 530), Fresno 32-68 

(DVIT 2746), Perlette (DVIT 488), Flame 


Fresno 32-139 (DVIT 2169), Perlette 

(DVIT 488), Robin (DVIT 2651), Flame 


Aswad (DVIT 2087), Fresno 35-75 

(DVIT 2163) 


Moscatel Ros. De Blas (DVIT 2328) 

Olmo 6F V19 (DVIT 2923), Corniola 

(DVIT 3162), Opale (DVIT 483) 


(DVIT 3162), Opale (DVIT 483) 

Sultanina Rose (DVIT 530), Maria 

Pirovano (DVIT 588), Barone Dellaterno 

(DVIT 350), Thomuscat (DVIT 536) 



(DVIT 857), Yellow Ortlieber (DVIT 

1020)

Gamay Rond (DVIT 736) 

Gamay Teinturier (DVIT 737) 

Gamay Toma (DVIT 2100) 

Gaschochi (DVIT 2537) 

Gemeroh (DVIT 1031) 



(DVIT 736) 




(DVIT 737) 



3188), Gamay Rond (DVIT 

736) 

Vranac (DVIT 2086), 







Traminer (DVIT 991) 


2332) 


2332) 

Gamay (DVIT 732) 

Siegerrebe (DVIT 3109), 


2332), Ezerfuertu (DVIT 

2672), Perle (DVIT 2754), 

Sylvaner 742 (DVIT 3180), 

Petite Manseng (DVIT 

2331), Taminga (DVIT 

2656), Trousseau (DVIT 

838), Sauvignon Gris 


1614.02), Chenin Blanc 

(DVIT 691), Verdelho 

(DVIT 1001), Rothgipfler 

(DVIT 935), Flora (DVIT 

720), White Veltliner 

(DVIT 1017), Teoulier 

Noir (DVIT 975), 

Bequignol (DVIT 2113), 

Tinta Madeira (DVIT 985), 





2672), Perle (DVIT 2754), 



















1020) 



1020) 

Vustolethi (DVIT 603) 


Donzillinho (DVIT 2634)


Ghlicopati (DVIT 741) 





708), Curisti Mici (DVIT 

703) 




2672), Perle (DVIT 2754), 




















Ghula Dari (DVIT 2039) Ab Jusht (DVIT 333) Ghula Dari (DVIT 409) 

Ghula Dari (DVIT 409) Khaldar (DVIT 432) Ghula Dari (DVIT 2039) 

Gold (DVIT 410) 

Golden Champion (DVIT 411) 

Verdal (DVIT 553), Rosaki 

(DVIT 502), Verdal (DVIT 

602), Seedless Emperor 

(DVIT 517) 

Gouveio (DVIT 743) Harslevelu (DVIT 755) 

Goyura (DVIT 2637) 

Grassa De Cotnari (DVIT 2080) 

Greco Bianco (DVIT 746) 

Green Hungarian (DVIT 748) 

Green Veltliner (DVIT 749) White Veltliner (DVIT 1017) 

Grey Riesling (DVIT 745) 

Grignolino (DVIT 751) 

Grillo (DVIT 752) 

Gros Colman (DVIT 413) 

Gros Manzenc (DVIT 753) 




Trousseau (DVIT 995), Beba 


650) 

Gungargoon (DVIT 2500) Stophiliatico (DVIT 966) 

Queen (DVIT 495) 

Ezerfuertu (DVIT 2672), 






578) 


738) 




992) 

Appley Towers (DVIT 

342), Ribier (DVIT 503), 

Kadarka (DVIT 768) 

Dornfelder (DVIT 2705) 







Dastrachan (DVIT 321), Red Roumi 

(DVIT 2083), Mandilaria (DVIT 815), 


Chardonay (DVIT 950), Tadone (DVIT 

970), Asma (DVIT 2054), Vustolethi 

(DVIT 603), Aspruda Santorinih (DVIT 

1027), Ezerjo (DVIT 715), Cola Tamburo 

(DVIT 694) 

Malta Seedless (DVIT 2327), Maria 



Blaue Elbe (DVIT 660) 



Citronelle (DVIT 381)


Guydouriha (DVIT 731) 

Gyongyrizling (DVIT 2675) 

Hakiki (DVIT 2089) 

Hakiki (DVIT 559) 

Hakiki (DVIT 559), 

Arhaniotico Kokino (DVIT 

636) 

Hakiki (DVIT 2089), 

Arhaniotico Kokino (DVIT 

636) 

Harslevelu (DVIT 755) Gouveio (DVIT 743) 

Hassaine (DVIT 2638) 

Helena (DVIT 756) 

Henab (DVIT 417) 

Hermitage (DVIT 1099) 

Heroldrebe (DVIT 2676) 

Hibron Blanc (DVIT 758) 

Hillbilly (DVIT 2747) 

Himrisnky (DVIT 2175) 

Kastour Rumi (DVIT 2070), 

Red Roumi Tanta (DVIT 

2260), Red Roumi (DVIT 

2101), Kastour Rumi (DVIT 

2103) 




2158) 




2341), Blue Trollinger (DVIT 

663) 

Xerez (DVIT 1018), Tinta 

Muida (DVIT 986) 

Hisakasy (DVIT 759) Aurora (DVIT 1341) 

Hondromavroudo (DVIT 1028) 

Honigber (DVIT 760) 

Hosargoon (DVIT 2503) 

Humagne (DVIT 3005) 

Hungarian Millenium (DVIT 761) 

Assirtico (DVIT 645) 

Muscat Ottonel (DVIT 

861) 


2648), White Riesling 

(DVIT 1015) 

Ezerfuertu (DVIT 2672), 




Niunai (DVIT 2094) 


Refosco (DVIT 1121) 



Limberger (DVIT 1107) 










Precoce Dastrachan (DVIT 

321), NA (DVIT 2044) 

Kotzifali (DVIT 1034) 

Torgoon (DVIT 2531), 

Shingargoon (DVIT 2603) 


377), Angelina (DVIT 339) 

Perla Di Csaba (DVIT 2042), Madeleine 

Angervine (DVIT 1108) 




(DVIT 2083), Mandilaria (DVIT 815), 


Chardonay (DVIT 950), Tadone (DVIT 


(DVIT 603), Aspruda Santorinih (DVIT 

1027), Ezerjo (DVIT 715), Cola Tamburo 

(DVIT 694) 

Red Roumi (DVIT 2083), Klusaine Red 

(DVIT 2077), Karadzhandal (DVIT 

2323), Munuka Sufed (DVIT 2046) 

Calzin (DVIT 678) 

Red Roumi (DVIT 2083) 



Esparte (DVIT 2912), Trincadeiro Branco 

(DVIT 993) 


Mezes (DVIT 829) 

Parargoon (DVIT 2502)


Husseine (DVIT 420) 

Aitah (DVIT 2037), Aitah 

Ghulaman (DVIT 336) 

Husseine (DVIT 576) Tschilaci (DVIT 539) 

Husseine (rish Baba) (DVIT 421) 

Niunai (DVIT 2094), 

Pirovano 5 (DVIT 2051), Siar 

(DVIT 522), Spin Ugad 

(DVIT 2602), Niunai (DVIT 

2104), Nimrang number58 

(DVIT 476), Nocera (DVIT 

477) 

Hycales (DVIT 422) Perruno (DVIT 893) 

Imperator (p. 264) (DVIT 423) 



Ithani Lefko (DVIT 620) 

Ithani Mavro (DVIT 621) 

Itonychi Lefko (DVIT 561) 

Itonychi Mavro (DVIT 562) 



Victor (DVIT 546) 



Itonychi Lefko (DVIT 561), 

Olivette Blanche (DVIT 479), 

Rish Baba (DVIT 3096) 



2158), Hermitage (DVIT 

1099) 

Ithani Lefko (DVIT 620), 

Olivette Blanche (DVIT 479), 




593), Cannon Hall (DVIT 





Red Roumi (DVIT 2083), 

Henab (DVIT 417), Muscat 

Szauter (DVIT 863) 



Karadzhandal (DVIT 

2323), Hassaine (DVIT 

2638) 


468) 

Angelo Pirovano (DVIT 


(DVIT 468), Folle Noire 

(DVIT 725) 




(DVIT 725) 

CG 1481 (DVIT 2432) 

CG 1481 (DVIT 2432) 


360) 




(DVIT 2923), Olmo j5-58 































(DVIT 2077), Kali Sag (DVIT 2917) 

La Reinha (DVIT 792) 


Muscat Br39 (DVIT 1313), Imperator (p. 

264) (DVIT 423), Lombardia (DVIT 

447), Opale (DVIT 483) 



447), Opale (DVIT 483) 


2087) 



2087) 


Jampal (DVIT 763) Talia (DVIT 971) 

Joao Santarem (DVIT 764) Castelao (DVIT 683) 

Johanna Mathiasz (DVIT 425) Madam Matijas (DVIT 2642) 

Julius Caesar (DVIT 579) Serphiotico (DVIT 956) 

July Muscat (DVIT 426) 

Juracon Noir (DVIT 2321) 

Jurancon Blanc (DVIT 1102) 

Kabuli (DVIT 2497) 









360) 

Joao Santarem (DVIT 764), Rhazaki 

Arhanon (DVIT 554), Unkown Cultivar 

(DVIT 693) 

Jampal (DVIT 763), Rhazaki Arhanon 



Vorosmarthy (DVIT 548) 


Malbec (DVIT 802) Folle Blanche (DVIT 724) 

Vidal 99T (DVIT 3185), 


Kadarka (DVIT 767) Kadarka (DVIT 768) Gros Colman (DVIT 413) 

Kadarka (DVIT 768) Kadarka (DVIT 767) Gros Colman (DVIT 413) 

Kakourdess Weis (DVIT 317) 

Monica (DVIT 461), Aspruda 

Ariloghi (DVIT 642) 

Kali Sag (DVIT 2917) Kali Sahehi (DVIT 427) 

Kali Sahehi (DVIT 427) Kali Sag (DVIT 2917) 

Kandhar (DVIT 2918) 

Kandhari (DVIT 2081) 

Kara Dzhidzhigi (DVIT 2322) 

Kara Lakana (DVIT 773) 

Karabournov (DVIT 2703) 

Karabryimis (DVIT 1032) 

Nunaka Sia (DVIT 2045), 

Nerello Calabrese (DVIT 

874) 


Adjem Misquet (DVIT 306), 


1055) 

Markandi (DVIT 587), Uva 

Romano (DVIT 542), Terra 

Promessa (DVIT 2114), 

Rhazaki (DVIT 2650), 

Rodites (DVIT 505), 

Waltham Cross (DVIT 2343), 

Rhazaki De Crete (DVIT 

501), Memory Of Queen 

Elizabeth (DVIT 458) 


708), Asma (DVIT 2054) 


2046) 


2046) 

Niunai (DVIT 2094), Karadzhandal 

(DVIT 2323) 

Niunai (DVIT 2094), Karadzhandal 

(DVIT 2323) 

Sultanina Rose (DVIT 530) Malta Seedless (DVIT 2327) 


1326), Kishmish-charni 

(DVIT 2325) 


564), Zabalkanski (DVIT 

552), Queen Of The 

Vineyard (DVIT 496), 

Askari (DVIT 343), 

Yourutico (DVIT 605) 

Kwar II (DVIT 2452) 





Tsambato (DVIT 996)

Karadzhandal (DVIT 2323) 

Karistino (DVIT 771) 


Karvouniariz (DVIT 769) 

Kastour Rumi (DVIT 2070) 



Kecskemet (DVIT 2639) 

Keknyelu (DVIT 775) 

Kepraico (DVIT 787) 

Kerner (DVIT 3104) 

Keshmesh (DVIT 580) 




Black Muscat Alexandria 

(DVIT 2111) 

Henab (DVIT 417), Red 

Roumi Tanta (DVIT 2260), 

Red Roumi (DVIT 2101), 


Henab (DVIT 417), Kastour 

Rumi (DVIT 2070), Red 





1979), Black Muscat 

Alexandria (DVIT 2111), 


Sultanina Rose (DVIT 530), 

Thompson Seedless (DVIT 

535), Kishmishi (DVIT 439), 

Ladyfinger Mutant Of 


2430), Sahebi (DVIT 508), 

Shando Khari (DVIT 520), 

Sultanina Marble (DVIT 

529), NA (PI 588236.02), 

Rozovii Kishmish (DVIT 

507), Ladyfinger Mutant of 

Thompson Seedless4X (DVIT 

2433), Midget Thompson 


Luquaci Thompson (DVIT 

450), Seeded Thompson 

(DVIT 1334) 

Khalchili (DVIT 430) Khalili (DVIT 2040) 






Fahri (DVIT 402), Rizamat 

(DVIT 2338) 

Rougitza Rouge (DVIT 

2546) 



1015) 

CG 1481 (DVIT 2432), 


Yaghotti No. 2 (DVIT 

2052), Calmeria (DVIT 

1084), B 15-19 (DVIT 

2171), Khalili (DVIT 

2084), Perlette (DVIT 

488), Favorita Segaria 



Tarrango II (DVIT 2658), 

Malta Seedless (DVIT 

2327), Queen (DVIT 495), 

Goyura (DVIT 2637), 

Kishmish Sorkh (DVIT 

437), Askary (DVIT 565), 

Muscat Szauter (DVIT 


(DVIT 588), Barone 

Dellaterno (DVIT 350), 

Thomuscat (DVIT 536), 

Sultana (DVIT 526), Lal 


Khaldar (DVIT 432) Ghula Dari (DVIT 409) Ghula Dari (DVIT 2039) 

Black Kishmish (DVIT 358), Alloued 

Zeine (DVIT 337), Red Roumi (DVIT 

2083), Klusaine Red (DVIT 2077), 

Aswad (DVIT 2087), Kali Sag (DVIT 

2917), Sicilien (DVIT 523), Sochal Karakash 

(DVIT 2340), Rizamat (DVIT 

2338), Kishmish-charni (DVIT 2325), 

Munuka Sufed (DVIT 2046), Asma 

(DVIT 2054), Kishmish Of Vir (DVIT 

2071), Hassaine (DVIT 2638), Khorestini 


Petrolianos (DVIT 899), Arcathinoh 

(DVIT 635) 



Husseine (DVIT 420), Munuka Sufed 


Wittberger (DVIT 2689), Rotberger 

(DVIT 2339) 


139 (DVIT 2169), Kishmish Hishrau 

(DVIT 1103), Monukka (DVIT 462), 

Kishmish-charni (DVIT 2325), Asma 

(DVIT 2054), G4-74 (Seedless Calmeria) 

(DVIT 2914)

Khalili (DVIT 2040) Khalchili (DVIT 430) 

Khalili (DVIT 2084) 

Khorestini (DVIT 2605) 


Kishmish Early (DVIT 2324) 

Kishmish Hishrau (DVIT 1103) 

Kishmish Of Vir (DVIT 2071) 

Yaghotti No. 2 (DVIT 2052), 

Tagouti Rouge (DVIT 551), 

Takhani (pink T.s.) (DVIT 

531) 


530), Munuka Sufed 

(DVIT 2046) 








351) 


530), Yaghotti No. 1 

(DVIT 604) 




1363.05) 



(DVIT 2325) 

Kishmish Sorkh (DVIT 437) Kishmish Spien (DVIT 438) Sultanina Rose (DVIT 530) 

Kishmish Spien (DVIT 438) Kishmish Sorkh (DVIT 437) Sultanina Rose (DVIT 530) 

Kishmishi (DVIT 436) 

Kishmishi (DVIT 439) 

Tand (DVIT 533), Tallian 

(DVIT 532) 



535), Ladyfinger Mutant Of 


2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 

Keshmesh (DVIT 580), 








(DVIT 1334) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











Monukka (DVIT 462), Kishmish-charni 

(DVIT 2325), Kishmish Sorkh (DVIT 

437) 

Neeli (DVIT 2514), Karadzhandal (DVIT 

2323), Munuka Sufed (DVIT 2046) 

Sultanina Rose (DVIT 530), Red Roumi 

(DVIT 2083), CG 1481 (DVIT 2432), 

Aswad (DVIT 2087), Yaghotti No. 2 

(DVIT 2052), Summer Muscat (DVIT 

2902), Monukka (DVIT 462), Khalili 

(DVIT 2084), Sochal Kara-kash (DVIT 

2340), Rizamat (DVIT 2338), Perlette 

(DVIT 488), NA (GVIT 1504.02), 

Favorita Segaria (DVIT 716), Black 

Kishmish (DVIT 1326), Karadzhandal 

(DVIT 2323), Asma (DVIT 2054), 

Kishmish Sorkh (DVIT 437), Askary 

(DVIT 565), Muscat Szauter (DVIT 863), 

Maria Pirovano (DVIT 588), Lal Sorkh 

(DVIT 442) 


(DVIT 2054) 


Karadzhandal (DVIT 2323), Asma (DVIT 

2054) 

Monukka (DVIT 462), Khalili (DVIT 


Askary (DVIT 565), Lal Sorkh (DVIT 

442) 

Monukka (DVIT 462), Khalili (DVIT 


Askary (DVIT 565), Lal Sorkh (DVIT 

442) 






(DVIT 2914)

Kleinberger (DVIT 776) 

Klusaine Red (DVIT 2077) 

White Riesling x Sauvignon 

Blanc (DVIT 3187), White 

Elbling (DVIT 1014) 

Kokinelia (DVIT 778) Retino Mavro (DVIT 931) 


Kokinostaphylo (DVIT 779) 

Kominato (DVIT 781) 

Kontehgalo (DVIT 581) 

Kontocladi (DVIT 1036) 

Koptcha (DVIT 783) 



Agoumastos (DVIT 1024) 

Korfiatis (DVIT 784) Ladikinah (DVIT 788) 

Kotzifali (DVIT 1034) 

Kraljevina (DVIT 1037) 

Kreatza (DVIT 582) NA (DVIT 1976) 

Kristalli (DVIT 786) 

Kritico (DVIT 583) Cipro Nero (DVIT 380) 

Kule Dary (DVIT 1070) 

Kurutaktas (DVIT 584) 

Kwar II (DVIT 2452) 

Kyoho (DVIT 616) 


Lacturario (DVIT 585) 

Blue Affenthaler (DVIT 662), 


Eununi (DVIT 400), Lattuario 

(DVIT 445) 

Ladikinah (DVIT 788) Korfiatis (DVIT 784) 

Lady Hutt (DVIT 2159) Syrian (DVIT 2162) 

Charas (DVIT 685) 


644) 

Shtur Angur (DVIT 2168) 

Hondromavroudo (DVIT 

1028) 


360) 


2087), Niunai (DVIT 2094), 

Karadzhandal (DVIT 2323), Hassaine 

(DVIT 2638) 


Black Corinth (DVIT 1052) 

Summer Muscat (DVIT 2902) 

Mathilde (DVIT 99) 

Aswad (DVIT 2087), Dschan Im Isium 

(DVIT 315), Mathilde (DVIT 99), Ter 

Gulmer (DVIT 976) 

Kandhari (DVIT 2081) 

Hycales (DVIT 422) 


2087) 

Aswad (DVIT 2087), Dschan Im Isium 

(DVIT 315), Mathilde (DVIT 99), Ter 

Gulmer (DVIT 976) 

Appley Towers (DVIT 342)

Ladyfinger Mutant Of Thompson 


Ladyfinger Mutant of Thompson 

Seedless4X (DVIT 2433) 

Lagorthi (DVIT 789) 

Lagrain (DVIT 790) 

Lal Sorkh (DVIT 442) 

LArvine (DVIT 2640) 




Sahebi (DVIT 508), Shando 

Khari (DVIT 520), Sultanina 

Marble (DVIT 529), NA (PI 

588236.02), Keshmesh 

(DVIT 580), Rozovii 


Ladyfinger Mutant of 






(DVIT 1334) 






2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 







(DVIT 1334) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 












530), Sahibi Sorkh (DVIT 

510) 






(DVIT 2914) 






(DVIT 2914) 


Monukka (DVIT 462), Rizamat (DVIT 

2338), Favorita Segaria (DVIT 716), 

Kishmish-charni (DVIT 2325), Goyura 


2323), Asma (DVIT 2054), Kishmish 

Sorkh (DVIT 437), Askary (DVIT 565), 

Muscat Szauter (DVIT 863), Maria 

Pirovano (DVIT 588), Katta Kurgan 

(DVIT 774)

Late Blue Burgundy (DVIT 793) 












Pinot Noir (DVIT 914), Pinot 



2307), Serine (DVIT 955), 











1020), NA (DVIT 1978), 



Late Edessa (DVIT 443) Zimsko Belo (DVIT 2664) Alulu (DVIT 338) Black Ferrara (DVIT 356) 

Lattuario (DVIT 445) 

Lacturario (DVIT 585), 

Eununi (DVIT 400) 

Laubschers Gem (DVIT 444) Askari (DVIT 343) 

Leanoy (DVIT 2641) 



468) 

Leanyka (DVIT 794) NA (PI 348888.02) Argant (DVIT 2305) 

Lianoroi (DVIT 795) Mandilaria (DVIT 815) 

Liatico (DVIT 796) 



Loin De Loeil (DVIT 798) 

Lombardia (DVIT 447) 

Longyan (DVIT 2078) 

Longyan (DVIT 2079) 

Limberger (DVIT 797), 

Teltfurtu (DVIT 2660) 









(DVIT 514) 







(DVIT 514) 

Loose Perlette (DVIT 2177) Perlette (DVIT 488) 

Luglienga (DVIT 449) Orleans (DVIT 3170) 

Black Corinth (DVIT 

1052) 

Heroldrebe (DVIT 2676), 

Zweigelt-rebe (DVIT 

2692) 



2692) 







2087) 



Orleans (DVIT 880), Mathilde (DVIT 

99), Asma (DVIT 2054), Berzemina 

(DVIT 656) 

Diamente Nero (DVIT 614), Italia (IP 65) 

(DVIT 424), Muscat Hamburg (DVIT 

468) 

Fresno 32-139 (DVIT 2169), Fresno 32- 

68 (DVIT 2746), NA (GVIT 1504.02), 

Kishmish-charni (DVIT 2325), Beauty 

Seedless (DVIT 351), Flame Seedless 

(DVIT 1095)

Luquaci Thompson (DVIT 450) 

Macaroli (DVIT 799) 

Madam Matijas (DVIT 2642) 

Madeleine Angervine (DVIT 

1108) 

Madeleine Celine (DVIT 452) 

Malaga (DVIT 453) 

Malaga Rosada (DVIT 454) 

Malagusiah (DVIT 801) 

Malbec (DVIT 802) 

Malmsey (DVIT 804) 

Malta Seedless (DVIT 2327) 

Malukato (DVIT 805) 

Malvasia (DVIT 806) 

Malvasia Bianca (DVIT 2117) 

Malvasia Rosario (DVIT 809) 

Malvasia Roxa (DVIT 812) 

Mamelon (DVIT 586) 






2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 







Seeded Thompson (DVIT 

1334) 

Johanna Mathiasz (DVIT 

425) 


590) 

Moscatel Ros. De Blas (DVIT 

2328) 

Malvasia (DVIT 806), 


Sultana Moscata (DVIT 527), 

Royal Thompson (DVIT 

1122) 

Malvasia (DVIT 806), 

Malmsey (DVIT 804) 

Malmsey (DVIT 804), 



Duc De Malakoff (DVIT 

393), Muscat Talebot (DVIT 

473) 

Mandilaria (DVIT 815) Lianoroi (DVIT 795) 

Marditza (DVIT 1039) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 

















1015) 

Precoce De Malingre 

(DVIT 490) 


(DVIT 2063), Seedless 

Emperor (DVIT 2759) 






(DVIT 2914) 


Gyongyrizling (DVIT 2675) 

Alicante Branca (DVIT 1025), Alicante 

Vermelho (DVIT 626) 

Iviar Oliver (DVIT 578) Fuji (DVIT 1655) 

Gamay (DVIT 732), 

Juracon Noir (DVIT 2321) 


(DVIT 855) 




(DVIT 2625) 


(DVIT 855) 


(DVIT 855) 


377) 

Nunaka Sia (DVIT 2045), Goyura (DVIT 

2637), Maria Pirovano (DVIT 588), 

Thomuscat (DVIT 536) 



Orleans (DVIT 880), Mathilde (DVIT 

99), Asma (DVIT 2054), Berzemina 

(DVIT 656)

Maria Pirovano (DVIT 588) 

Markandi (DVIT 587) 

Uva Romano (DVIT 542), 

Karabournov (DVIT 2703), 

Terra Promessa (DVIT 2114), 







Marsanne (DVIT 817) Rousseau (DVIT 936) 

Marvel Of Vaucluse (DVIT 456) 


Matum Margoon (DVIT 2530) 

Mavro Spetzon (DVIT 824) 






1055), Kara Lakana (DVIT 

773) 

Mavrodaphne (DVIT 821) Augustiatico (DVIT 566) 

Mavrokorakas (DVIT 822) 

Mavroliatis (DVIT 1040) 

Mavroudi (DVIT 1041) 

Mavroudi School Of Patras (DVIT 

2061) 

Memory Of Queen Elizabeth 

(DVIT 458) 

Pinot Gris (DVIT 907) 


Romano (DVIT 542), 






Rhazaki De Crete (DVIT 501) 

Merlot (DVIT 2682) Merlot (DVIT 826) 

Merlot (DVIT 826) Merlot (DVIT 2682) 

Mermark (DVIT 1042) 

Mezes (DVIT 829) 



578) 







Mavrodaphne (DVIT 821), 

Gouveio (DVIT 743), 


Tadone (DVIT 970) 


Divromos (DVIT 706), 


644) 

Arcathinoh (DVIT 635) 








803) 


803) 

Kishmish-charni (DVIT 2325), Malta 

Seedless (DVIT 2327), Goyura (DVIT 

2637), G4-74 (Seedless Calmeria) (DVIT 

2914), Barone Dellaterno (DVIT 350), 

Thomuscat (DVIT 536), Lal Sorkh (DVIT 

442) 


Citronelle (DVIT 381), Aurora (DVIT 

345), Appley Towers (DVIT 342), Noir 



(DVIT 2083), Korfiatis (DVIT 784), 

Mandilaria (DVIT 815), Olivette 

Vendemien (DVIT 482), Kontocladi 

(DVIT 1036), Asma (DVIT 2054), 

Vustolethi (DVIT 603), Divromos (DVIT 

706), Danuque (DVIT 387), Unkown 

Cultivar (DVIT 644) 





Honigber (DVIT 760)

Midget Thompson Seedless (DVIT 

1307) 

Migdali (DVIT 830) 

Millishun (DVIT 2498) 

Mission Jalama (DVIT 832) 

Monica (DVIT 461) 






2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 





2433), Luquaci Thompson 

(DVIT 450), Seeded 

Thompson (DVIT 1334) 


Romania (DVIT 595) 


317), Aspruda Ariloghi 

(DVIT 642) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











Fatai (DVIT 2510) 

Asma Sohwars (DVIT 

309), Bondola (DVIT 

3067), Niaropes (DVIT 

876) 


708), Asma (DVIT 2054) 

Montanaccio (DVIT 836) Sciacarello (DVIT 1125) Oki di Boi (DVIT 877) 

Montils (DVIT 835) 

Montils (DVIT 837) 

Bias Kukuzeti (DVIT 311), 


Monukka (DVIT 462) Black Monukka (DVIT 359) 


Moscatel Galego (DVIT 842) 

Moscatel Ros. De Blas (DVIT 

2328) 

Moscato Dadda (DVIT 2109) 

Moscato Di Terracina (DVIT 864) 

Montils (DVIT 835), Bias 

Kukuzeti (DVIT 311) 

Muscat Galego (DVIT 859), 

Muscat Canelli (DVIT 856), 

Muscat Blanc (DVIT 2170), 

Muscat Gros Noir Hatif 

(DVIT 464), Muscat Noir 

(DVIT 469), Muscat Rouge 

Fouce (DVIT 1656) 


2046), Pardinia (DVIT 

486) 


(DVIT 855), Iviar Oliver 

(DVIT 578), Moscato 


Orange Muscat (DVIT 

2167), NA (DVIT 1978), 

Muscato Vidor (DVIT 

2256), Aleatico (DVIT 

623), White Calabrian 

(DVIT 2391), NA (PI 

588425.02) 






(DVIT 2914) 

Cirmisi Sap De Sudak (DVIT 313), 

Dumbrehnah Mavri (DVIT 708), Asma 

(DVIT 2054) 


Sultanina Rose (DVIT 530), Khalili 


2325), Kishmish Sorkh (DVIT 437), Lal 


Malaga Rosada (DVIT 454) Iviar Oliver (DVIT 578) Fuji (DVIT 1655) 


2166), Victorias Choice 

(DVIT 3182), Victorias 

Choice (DVIT 2663), 

Victorias Choice number1 

(DVIT 1598) 


Trieste 4x (DVIT 1591)

Moscato Jesus (DVIT 2062) 

Moscato Mavro (DVIT 845) 

Moschardina (DVIT 847) 

Mosho Patata (DVIT 1043) 

Mourvedre Famellestadt (DVIT 

319) 

Mrs Pince Black Muscat (DVIT 

2160) 

Mueller-thurgau (DVIT 853) 


Baharat Early (DVIT 2306), 



Esparte (DVIT 2912) 

Mueller Thurgau (DVIT 

2337) 

Mueller Thurgau (DVIT 2337) Mueller-thurgau (DVIT 853) 

Munuka Sufed (DVIT 2046) 

Murma Isium (DVIT 320) 

Muscadelle De Bordelais (DVIT 

855) 

Muscat Blanc (DVIT 2170) 



Moscatel Galego (DVIT 842), 










859), Moschardina (DVIT 

847) 

Moscato Mavro (DVIT 

845) 



Perle (DVIT 2754), 

Reichensteiner (DVIT 


(DVIT 1015), Ortega 

(DVIT 1114) 


Perle (DVIT 2754), 

Reichensteiner (DVIT 


(DVIT 1015), Ortega 

(DVIT 1114) 

Fahri (DVIT 402), Kali Sag 

(DVIT 2917), Monukka 

(DVIT 462), Khalili (DVIT 

2084), Sicilien (DVIT 523) 


859), Malvasia (DVIT 

806), Emerald Riesling 

(DVIT 712) 






2167), NA (DVIT 1978), 





588425.02) 


(DVIT 598) 

Fileri (DVIT 1044) 

Himrisnky (DVIT 2175), Trincadeiro 


Optima (DVIT 3105), Ehrenfelser (DVIT 

2317) 

Optima (DVIT 3105), Ehrenfelser (DVIT 

2317) 

Rizamat (DVIT 2338), Karadzhandal 

(DVIT 2323), Hassaine (DVIT 2638), 

Khorestini (DVIT 2605), Katta Kurgan 

(DVIT 774) 

Precoce Dastrachan (DVIT 321), Dschan 

Im Isium (DVIT 315)

Muscat Bowood (DVIT 463) 

Muscat Br39 (DVIT 1313) 

Muscat Caillaba (DVIT 2389) 

Muscat Canelli (DVIT 856) 

Muscat Canelli (DVIT 857) 

Muscat Crime (DVIT 858) 

Muscat De Ladda (DVIT 2166) 

Muscat De Pantellana (DVIT 

2063) 













643) 








Victorias Choice (DVIT 


(DVIT 2663), Moscato Dadda 


Choice number1 (DVIT 

1598) 




(DVIT 2923), Olmo j5-58 
































468), Folle Noire (DVIT 

725) 






2167), NA (DVIT 1978), 





588425.02) 



2751) 



Trieste 4x (DVIT 1591) 



590) 



Chardonnay (DVIT 687), Gamay 

Burgoyne (DVIT 734), Yellow Ortlieber 

(DVIT 1020)


Muscat Gros Noir Hatif (DVIT 

464) 

Muscat Hamburg (DVIT 1059) 




Muscat Noir (DVIT 469), 

Muscat Rouge Fouce (DVIT 

1656) 




Muscat Noir (DVIT 469), 

Muscat Rouge Fouce (DVIT 

1656) 


Trollinger (DVIT 2341), 

Hillbilly (DVIT 2747), Blue 

Trollinger (DVIT 663) 

Muscat Hamburg (DVIT 468) Muscat Hamburg (DVIT 860) 

Muscat Hamburg (DVIT 860) Muscat Hamburg (DVIT 468) 

Muscat Noir (DVIT 469) 













2167), NA (DVIT 1978), 





588425.02) 













NA (DVIT 1980), 

Imperator (p. 264) (DVIT 




Queen (DVIT 495), Askari 

(DVIT 343), Barbableu 

(DVIT 348), Netter (DVIT 

474) 




NA (DVIT 1980), 

Imperator (p. 264) (DVIT 




Queen (DVIT 495), Askari 

(DVIT 343), Barbableu 

(DVIT 348), Netter (DVIT 

474) 






2167), NA (DVIT 1978), 





588425.02) 


Chasselas Tompa (DVIT 378), Dan Ben 

Chana (DVIT 2633), Lombardia (DVIT 

447) 

Chasselas Tompa (DVIT 378), Dan Ben 

Chana (DVIT 2633), Lombardia (DVIT 

447)

Muscat of Alexandria (DVIT 465) 

Muscat Of Alexandria (DVIT 466) 







(DVIT 463) 

Iviar Oliver (DVIT 578), Red 

Hanepoot (DVIT 593), 

Cannon Hall (DVIT 368), 


841), Muscat Bowood (DVIT 


(DVIT 465) 




(DVIT 2923), Olmo j5-58 


































(DVIT 2923), Olmo j5-58 

































Muscat of Alexandria (DVIT 841) 





368), Muscat Bowood (DVIT 


(DVIT 465) 




(DVIT 2923), Olmo j5-58 































Muscat Of Hungary (DVIT 2390) White Calabrian (DVIT 2391) Muscat Galego (DVIT 859) 

Muscat Oliver (DVIT 2643) 

Muscat Ottonel (DVIT 861) 

Muscat Rouge Fouce (DVIT 1656) 

Muscat St. Laurent (DVIT 471) 


Muscat Szauter (DVIT 472) 



Muscato Vidor (DVIT 2256) 







(DVIT 469) 



393), Mamelon (DVIT 586) 

NA (DVIT 1976) Kreatza (DVIT 582) 


2648) 

Guydouriha (DVIT 731) 






2167), NA (DVIT 1978), 





588425.02) 


377), NA (PI 258975.02) 


377), NA (PI 258975.02) 


Perla Di Csaba (DVIT 2042) 



Iviar Oliver (DVIT 578) Opale (DVIT 483) 


530), Hunisa (DVIT 418) 


377) 


Red Roumi (DVIT 2083), Kishmishcharni 

(DVIT 2325), Askary (DVIT 565), 

Lal Sorkh (DVIT 442)

NA (DVIT 1977) Negru Virtos (DVIT 2644) Prokupac (DVIT 1071) 






NA (GVIT 1363.05) 

NA (GVIT 1504.02) 

NA (GVIT 1508.02) 

NA (GVIT 1512.02) 


(DVIT 341), Kepraico (DVIT 

787), Black Muscat 



Chasselas Des Bouches-durhone 

(DVIT 2112) 

Angulata (DVIT 341), NA 

(DVIT 1979), Kepraico 

(DVIT 787), Black Muscat 



Taifi (DVIT 2174), Tabyrn 

(DVIT 969) 

NA (GVIT 1614.02) Zifir (DVIT 2690) 

NA (PI 258975.02) 

NA (PI 340969.03) 


859), Badacsony-10 (DVIT 

2667), Pinot Noir Espinett 


1614.02), NA (PI 

588668.04) 


377), Sauvis (DVIT 598), 

Diamente Nero (DVIT 


(DVIT 468), Lombardia 

(DVIT 447), Conte Rosso 

(DVIT 382) 

Hisakasy (DVIT 759) 

Kishmish Hishrau (DVIT 

1103) 


(DVIT 496) 


2332), Gewurztraminer 


1978) 


862), Muscat St. Laurent 

(DVIT 471) 

Trebbiano Bianco (DVIT 

2067), Carriere (DVIT 

953) 

NA (PI 348888.02) Leanyka (DVIT 794) Argant (DVIT 2305) 

NA (PI 588236.02) 






2430), Sahebi (DVIT 508), 



529), Keshmesh (DVIT 580), 








(DVIT 1334) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











Taifi (DVIT 2174), Rizamat (DVIT 

2338), Tana-Kuzi (DVIT 2928) 

Perlette (DVIT 488), Kishmish-charni 

(DVIT 2325), Beauty Seedless (DVIT 

351) 

Rizamat (DVIT 2338), Tana-Kuzi (DVIT 

2928), NA (GVIT 1363.05) 






(DVIT 2914)

NA (PI 588326.03) 

NA (PI 588425.02) 

NA (PI 588668.04) Zengo (DVIT 2691) 

NA (PI 588673.06) 

NA (PI 588709.02) 

Naboda (DVIT 865) 

Namonia (DVIT 2511) 



Nebbiolo Bourgu (DVIT 868) 

Nebbiolo Fino (DVIT 869) 

Nebbiolo Michet (DVIT 2921) 

White Riesling (DVIT 1015), 


White Riesling (PI 

588673.02) 


Sylvaner (DVIT 968), 

Sylvaner (DVIT 1075) 





875) 

Nebbiolo Michet (DVIT 

2921), Nebbiolo Bourgu 

(DVIT 868) 

Nebbiolo (DVIT 867), 

Nebbiolo Michet (DVIT 

2921) 

Nebbiolo (DVIT 867), 

Nebbiolo Bourgu (DVIT 868) 




859), Sylvaner 742 (DVIT 

3180) 


(DVIT 715) 


Gyongyrizling (DVIT 

2675), Kerner (DVIT 


(DVIT 2337), Rotberger 

(DVIT 2339), Ehrenfelser 

(DVIT 2317), Osteiner 

(DVIT 2330), Madeleine 



738), Osteiner (DVIT 

2330), NA (PI 588425.02) 


Optima (DVIT 3105) 



Antibo (DVIT 631), 

Ughetta (DVIT 2662) 





Angelina (DVIT 339), Aswad (DVIT 

2087) 

Nebbiolo Tronero (DVIT 871) Negro Amaro (DVIT 872) Sangiovese (DVIT 945) Unknown Cultivar (DVIT 881) 

Neeli (DVIT 2514) 

Neelilay (DVIT 2507) 

Negrette (DVIT 2329) 

Negro Amaro (DVIT 872) 

Spanna (DVIT 961), Pinot 

St.george (DVIT 917) 

Nebbiolo Tronero (DVIT 

871) 

Negru Virtos (DVIT 2644) NA (DVIT 1977) Prokupac (DVIT 1071) 

Neiretta (DVIT 873) 

Neiretto Verzuolo (DVIT 875) 

Neo-Muscat (DVIT 2645) 

Nerello Calabrese (DVIT 874) 

Netter (DVIT 474) 

Niaropes (DVIT 876) 






Kandhar (DVIT 2918) 

Khorestini (DVIT 2605) 

Sangiovese (DVIT 945) Unknown Cultivar (DVIT 881) 


Iviar Oliver (DVIT 578) Opale (DVIT 483) 


Dabouki (DVIT 388), 


468) 


Aswad (DVIT 2087), Dan Ben Chana 

(DVIT 2633)

Nicolas Horthy number39 (DVIT 

475) 

Chasselas Tompa (DVIT 

378), Madam Matijas 

(DVIT 2642) 

Nilok (DVIT 2504) Nosargoon (DVIT 2533) Torgoon (DVIT 2531) Parargoon (DVIT 2502) 

Nimrang number58 (DVIT 476) 



Nocera (DVIT 477) 


Nonno Giovanni (DVIT 2257) 

Norakert PRG 2224 (DVIT 2683) 





2104), Husseine (rish Baba) 


477) 





(DVIT 421), Nimrang 

number58 (DVIT 476), 





(DVIT 2602), Husseine (rish 

Baba) (DVIT 421), Nimrang 












2638) 



2638) 



2638) 



2638) 











Gouveio (DVIT 743), Agadia (DVIT 

334), Appley Towers (DVIT 342), 

Setzlinge Dodreliabi (DVIT 331), Mavro 

Spetzon (DVIT 824), Precoce Dastrachan 

(DVIT 321), Red Roumi (DVIT 2083), 


Hisakasy (DVIT 759), Mathilde (DVIT 


(DVIT 603), Alburla (DVIT 634) 

Nosargoon (DVIT 2533) Nilok (DVIT 2504) Torgoon (DVIT 2531) Parargoon (DVIT 2502) 

Nunaka Sia (DVIT 2045) 

Nerello Calabrese (DVIT 

874), Kandhar (DVIT 2918) 

Ohanes (DVIT 478) Almeria (DVIT 1079) 

Oki di Boi (DVIT 877) 

Olho De Pargo (DVIT 589) 



Petinos (DVIT 591), Flame 



(DVIT 1330) 

Olivette Bauthelet (DVIT 590) Malaga (DVIT 453) 

Olivette Blanche (DVIT 479) 





Calmeria (DVIT 1084), B 

15-19 (DVIT 2171), Bien 

Donnie (DVIT 2696) 

Montanaccio (DVIT 836) 



(DVIT 761) 


(DVIT 2063), Seedless 


CG 1481 (DVIT 2432) 


Olivette Vendemien (DVIT 482), Black 

Ferrara (DVIT 356) 


Alicante Branca (DVIT 1025), Alicante 

Vermelho (DVIT 626) 


2087)

Olivette Noir (DVIT 1306) 

Olivette Noire (DVIT 1657) 

Olivette Rose (DVIT 481) 

Olivette Noire (DVIT 1657), 

Canner (DVIT 367) 

Canner (DVIT 367), Olivette 

Noir (DVIT 1306) 

Olivette Vendemien (DVIT 482) Ciminnita (DVIT 1653) 

Olmo (035-07) (DVIT 1808) 

Olmo (035-41) (DVIT 1811) 

Olmo (035-50) (DVIT 1813) 035-58 (DVIT 1814) 

Olmo 18-226 D (DVIT 2749) 

Olmo 25F (DVIT 2751) 

Olmo 34F (DVIT 2752) 

Olmo 6F V19 (DVIT 2923) 

Olmo j5-58 (DVIT 1314) 

Opale (DVIT 483) 


Orange Muscat (DVIT 2167) 

Orange Muskat (DVIT 879) 

Prokupao (DVIT 2926), 










Orleans (DVIT 3170) Luglienga (DVIT 449) 

Orleans (DVIT 880) 

Ortega (DVIT 1114) 

Osteiner (DVIT 2330) 

Otscha Bala (DVIT 617) 



348) 



348) 

Aurora (DVIT 345), 



654), Bien Donnie (DVIT 

2696) 



377), Muscat Canelli 

(DVIT 857), Yellow 

Ortlieber (DVIT 1020) 









Olivette Noire (DVIT 1657), Red Roumi 

(DVIT 2083), Almeria (DVIT 1079), 

Calmeria (DVIT 1084), Robin (DVIT 

2651), Mathilde (DVIT 99), Asma (DVIT 

2054), Danuque (DVIT 387), Daphne 

(DVIT 570), Black Ferrara (DVIT 356), 


034-55 (DVIT 1807), 034-54 (DVIT 

1806), 034-26 (DVIT 1805), 035-11 

(DVIT 1809) 

Olmo (035-50) (DVIT 1813) 

Olmo (035-41) (DVIT 1811) 





Iviar Oliver (DVIT 578) Black Prince (DVIT 361) 













2337) 



1015) 

Chasselas Tompa (DVIT 378), Diamente 

Nero (DVIT 614), Futuna 699 (DVIT 

2073), Corniola (DVIT 3162), Muscat 

Szauter (DVIT 472), Italia (IP 65) (DVIT 

424), Neo-Muscat (DVIT 2645), Black 

Morocco (DVIT 360) 

Perle (DVIT 2754), Mueller Thurgau 

(DVIT 2337), White Riesling (DVIT 

1015) 


(DVIT 598) 


(DVIT 598) 

Precoce Dastrachan (DVIT 321), 

Mandilaria (DVIT 815), Taschli (DVIT 

325)

Oubeidy (DVIT 2173) 

Pais (DVIT 484) Pias (DVIT 833) 

Parargoon (DVIT 2502) 

Pardala (DVIT 883) 

Pardinia (DVIT 486) 

Patrino (DVIT 1356) 

Peagudo (DVIT 887) 

Pearl of Zola (DVIT 2646) 


Perbos 139 (DVIT 2145) 





Perle de Csaba (DVIT 2648) 


Agoumastos (DVIT 1024), 


Platirahah (DVIT 920), 

Aspruda Halkidos (DVIT 

1026) 

Romania (DVIT 595), 



393), Mamelon (DVIT 586), 


Perle (DVIT 2754), Riesling 

Muscat (DVIT 2336) 

Perle (DVIT 2647), Riesling 

Muscat (DVIT 2336) 

Perlette (DVIT 488) Loose Perlette (DVIT 2177) 

Perruno (DVIT 893) Hycales (DVIT 422) 

Petinos (DVIT 591) 

Petite Bouschet (DVIT 894) 

Petite Manseng (DVIT 2331) 



Olho De Pargo (DVIT 589), 

Flame Tokay (DVIT 2913), 

Tokay (DVIT 540), Seedless 


Olmo 6F V19 (DVIT 


3162), Rambola (DVIT 

497), Black Prince (DVIT 

361) 

Citronelle (DVIT 381), 

Monukka (DVIT 462) 

Tinta Madeira (DVIT 985) 


2648), Perla Di Csaba 

(DVIT 2042) 




876) 

Pearl of Zola (DVIT 2646), 



377) 



(DVIT 2337) 



(DVIT 2337) 


2675), Pearl of Zola (DVIT 

2646), Anna Maria (DVIT 

3156), Muscat Oliver 

(DVIT 2643), Orange 



(DVIT 496) 






(DVIT 761) 


Teoulier Noir (DVIT 975), 



738) 

Zlahtina (DVIT 607) 

Nilok (DVIT 2504), Hosargoon (DVIT 

2503) 


Perle de Csaba (DVIT 2648), 

Gyongyrizling (DVIT 2675), Muscat 

Oliver (DVIT 2643), Orange Muscat 

(DVIT 2167), Queen Of The Vineyard 

(DVIT 496) 




Fresno 32-139 (DVIT 2169), Fresno 32- 

68 (DVIT 2746), NA (GVIT 1504.02), 

Kishmish-charni (DVIT 2325), Beauty 

Seedless (DVIT 351), Flame Seedless 

(DVIT 1095) 


Nasa Veltliner (DVIT 866)

Petrolianos (DVIT 899) 

Peverella (DVIT 900) 

Pfeffer (DVIT 1116) Robin Noir (DVIT 933) 

Pias (DVIT 833) Pais (DVIT 484) 

Picpoule Noir (DVIT 902) 

Pinger (DVIT 2649) 


Pinot Blanc (DVIT 1065) 

Pinot Gris (DVIT 907) 

Pinot Musque (DVIT 1117) 

Pinot Nero (DVIT 909) 

Psarotherico (DVIT 1047), Al 

Borla (DVIT 307) 















Blauer Fruheburgunder 

(DVIT 2307), Serine (DVIT 

955), Pinot Pernard (DVIT 

905) 

Mavroudi School Of Patras 

(DVIT 2061) 







Vert Dore (DVIT 918) 

















905), Pinot (DVIT 733) 

Olmo 6F V19 (DVIT 


3162), Rambola (DVIT 

497), Black Prince (DVIT 

361) 









1020), NA (DVIT 1978), 




2751) 









1020), NA (DVIT 1978), 


Karistino (DVIT 771) 





734) 


Pinot Noir-marianfeld (DVIT 

2334) 

Pinot Noir (DVIT 1066) 




2333), Blauer Spatburgunder 












2307), Serine (DVIT 955), 












Pinot Noir (DVIT 913), Pinot 

Noir (DVIT 914), Late Blue 




2307), Serine (DVIT 955), 














Late Blue Burgundy (DVIT 

793), Pinot Nero (DVIT 909), 




905), Pinot (DVIT 733) 

Pinot Noir (DVIT 912) Preto Manteudo (DVIT 923) 


















905), Pinot (DVIT 733) 









1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 





Tinta Amarella (DVIT 982) 




Pinot Noir 2a (DVIT 2333) 

Pinot Noir Epernay (DVIT 916) 

















905), Pinot (DVIT 733) 

















905), Pinot (DVIT 733) 

















905), Pinot (DVIT 733) 







(DVIT 1117) 









1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 




2751) 






734)

Pinot Noir Espinett (DVIT 2332) 

Pinot Pernard (DVIT 905) 

Pinot St.george (DVIT 917) 

Pinot Vert Dore (DVIT 918) 

Pirobelle (DVIT 2700) 

Pinot Noir 2a (DVIT 2333), 

Pinot Noir-marianfeld (DVIT 

2334), Blauer Spatburgunder 












2307), Serine (DVIT 955), 



















955), Pinot (DVIT 733) 

Negrette (DVIT 2329), 

Spanna (DVIT 961) 








Pirovano 47 (DVIT 2047) Aurora (DVIT 345) 

Pirovano 5 (DVIT 2051) 

Pizzutello (DVIT 489) 

Platani (DVIT 1045) 

Platirahah (DVIT 920) 

Plavac Mali (DVIT 1119) 

Pleven Mychat (DVIT 3171) 

Niunai (DVIT 2094), Siar 







Aspruda Halkidos (DVIT 

1026), Patrino (DVIT 1356) 

Corniola (DVIT 3162), San 

Juanina Rosada (DVIT 2927), 










1020), NA (DVIT 1978), 










1020), NA (DVIT 1978), 




2751) 


(DVIT 496) 


482) 



2638) 

Assirtico (DVIT 645) 


Troja (DVIT 994) 





734), NA (GVIT 1614.02), 

Gewurztraminer (DVIT 738), Precoce De 

Malingre (DVIT 490), Yellow Ortlieber 

(DVIT 1020), NA (DVIT 1978) 




734) 

Citronelle (DVIT 381), Appley Towers 


Almeria (DVIT 1079), Mathilde (DVIT 

99) 







361)




Precoce Dastrachan (DVIT 321) 

Precoce De Malingre (DVIT 490) 

Burgrave (DVIT 364), Poet 

Matabon (DVIT 1658) 

Burgrave (DVIT 364), Poet 

Matabon (DVIT 1597) 



Preto Manteudo (DVIT 923) Pinot Noir (DVIT 912) 

Prieska (DVIT 924) 

Primiera (DVIT 491) 




Hisakasy (DVIT 759) 

Choultu White (DVIT 

2909), Pinot Noir Espinett 


Celine (DVIT 452) 

Delizia Di Vaprio (DVIT 

571) 

Prokupac (DVIT 1071) Smederevka (DVIT 1074) Negru Virtos (DVIT 2644) 

Prokupao (DVIT 2926) 

Prosecco Tondo (DVIT 926) 

Prune De Cazouls (DVIT 494) 

Psara (DVIT 1046) 

Psarotherico (DVIT 1047) 

Psevthosirico (DVIT 592) 

Olmo 6F V19 (DVIT 2923), 


Al Borla (DVIT 307), Pinger 

(DVIT 2649) 



3003), Tsoumbrena Lefki 

(DVIT 997) 

Queen (DVIT 495) Aspiran Noir (DVIT 639) 

Queen Of The Vineyard (DVIT 

496) 

Raboso Piave (DVIT 927) 

Raisin De Palestine (DVIT 1072) 

Rambola (DVIT 497) 

Scolokertek Kiralynoje 

(DVIT 516) 

Rangspray (DVIT 1120) Barlinka (DVIT 1080) 





1657) 




530), Gold (DVIT 410), 


468) 



2648), Pirobelle (DVIT 


488), NA (GVIT 1504.02), 

Robin (DVIT 2651), Early 

Muscat (DVIT 394), Bien 

Donnie (DVIT 2696), 


351) 

Fogarina (DVIT 721) 

Pias (DVIT 833), Black 

Morocco (DVIT 360) 

Chaouch (DVIT 371), 



(DVIT 2325) 



(DVIT 342), Setzlinge Dodreliabi (DVIT 

331), Noir Dautomne (DVIT 330), 

Murma Isium (DVIT 320), Mavro 

Spetzon (DVIT 824), Red Roumi (DVIT 

2083), Aswad (DVIT 2087), Orleans 


(DVIT 2054), Vustolethi (DVIT 603), 







Fileri (DVIT 1044), Roussaitis (DVIT 

1049), Coristano (DVIT 697) 



(DVIT 3162), Black Prince (DVIT 361)

Red-white Veltliner (DVIT 929) Red Veltliner (DVIT 928) 

Red Fujii (DVIT 3173) 

Red Hanepoot (DVIT 593) 

Red Ohanez (DVIT 2757) 




Red Roumi Tanta (DVIT 2260) 








Zabalkanski (DVIT 552), 

Sabalkanskoi (DVIT 3077), 


Zabalkanski (DVIT 552), 









Roumi (DVIT 2101), Kastour 

Rumi (DVIT 2103) 

Rothgipfler (DVIT 935), 


Red Veltliner (DVIT 928) 




(DVIT 2923), Olmo j5-58 







































Gouveio (DVIT 743), Noir Dautomne 

(DVIT 330), Itonychi Lefko (DVIT 561), 

Black Kishmish (DVIT 358), Alloued 

Zeine (DVIT 337), Precoce Dastrachan 

(DVIT 321), Klusaine Red (DVIT 2077), 

Aswad (DVIT 2087), Niunai (DVIT 

2094), Henab (DVIT 417), Olivette 

Vendemien (DVIT 482), Sochal Karakash 

(DVIT 2340), Rizamat (DVIT 


Karadzhandal (DVIT 2323), Mathilde 

(DVIT 99), Asma (DVIT 2054), Hassaine 

(DVIT 2638), Lacturario (DVIT 585), 



Red Roumi (DVIT 2083)

Red Veltliner (DVIT 928) 

Ree (DVIT 131) 

Red-white Veltliner (DVIT 

929) 

Rothgipfler (DVIT 935), 


Refosco (DVIT 1121) Refosco (DVIT 930) Helena (DVIT 756) Syrah (DVIT 1053) 

Refosco (DVIT 930) Refosco (DVIT 1121) Helena (DVIT 756) Syrah (DVIT 1053) 

Reichensteiner (DVIT 3107) 

Retino Lefko (DVIT 1067) 

Fileri (DVIT 1044), Velana 

(DVIT 1057) 

Retino Mavro (DVIT 931) Kokinelia (DVIT 778) 

Rhazaki (DVIT 2650) 

Rhazaki Anatolico (DVIT 608) 

Rhazaki Arhanon (DVIT 554) 

Rhazaki De Crete (DVIT 501) 

Rhazaki Mavro (DVIT 555) 



Riesling Italico (DVIT 932) 

Riesling Muscat (DVIT 2336) 













2259), Asprocoritho (DVIT 

640) 









(DVIT 458) 

Royal E (DVIT 506), Emile 

Royal (DVIT 395), Ribier 

(DVIT 594), Royal (DVIT 

596) 


(DVIT 506), Emile Royal 


596) 

Perle (DVIT 2754), Perle 

(DVIT 2647) 



Olivette Blanche (DVIT 479) 


2337) 


















468), Olmo 18-226 D 









468), Olmo 18-226 D 






(DVIT 2337) 

CG 1481 (DVIT 2432) 




Joao Santarem (DVIT 764), Jampal 

(DVIT 763) 





2087)

Rizamat (DVIT 2338) 



Cardinal (DVIT 370), 


Robin Noir (DVIT 933) Pfeffer (DVIT 1116) 

Rodiares (DVIT 1048) 



Romania (DVIT 595) 

Rondinella (DVIT 2652) 


Agoumastos (DVIT 1024), 
















503) 







Satino (DVIT 946), 

Bondola (DVIT 3067) 




876) 

Rosa Minna (DVIT 322) Alulu (DVIT 338) Zimsko Belo (DVIT 2664) 

Rosaki (DVIT 502) 

Rotberger (DVIT 2339) 

Rothgipfler (DVIT 935) 

Verdal (DVIT 553), Golden 

Champion (DVIT 411), 

Verdal (DVIT 602), Seedless 




1015) 

Red Veltliner (DVIT 928), 


738) 

Rougitza Rouge (DVIT 2546) Steinschiller (DVIT 964) Kecskemet (DVIT 2639) 

Roussaitis (DVIT 1049) 

Roussas (DVIT 1050) Roussico (DVIT 1051) 

Rousseau (DVIT 936) Marsanne (DVIT 817) 

Roussico (DVIT 1051) Roussas (DVIT 1050) 

Royal (DVIT 596) 


(DVIT 506), Emile Royal 


594) 






468), Olmo 18-226 D 





2087), Taifi (DVIT 2174), Sochal Karakash 

(DVIT 2340), Kishmish-charni 


2323), Munuka Sufed (DVIT 2046), NA 

(GVIT 1363.05), Lal Sorkh (DVIT 442) 








Wittberger (DVIT 2689), Kerner (DVIT 

3104) 

Volitza Lefki (DVIT 1010), Voltzicki 

(DVIT 1011), Psara (DVIT 1046), Fileri 

(DVIT 1044), Beli Potok (DVIT 1841), 



Royal E (DVIT 506) 

Royal Thompson (DVIT 1122) 

Royale 4x (DVIT 1666) 

Royalty (DVIT 937) 

Rozovii Kishmish (DVIT 507) 

Ribier (DVIT 503), Emile 

Royal (DVIT 395), Ribier 


596) 

Malta Seedless (DVIT 2327), 

Sultana Moscata (DVIT 527) 

Aspruda Santorinih (DVIT 

1027) 






2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 








(DVIT 1334) 






468), Olmo 18-226 D 







(DVIT 2625) 





Platirahah (DVIT 920), Gouveio (DVIT 

743), Mavro Spetzon (DVIT 824), 


(DVIT 2087) 

Trousseau (DVIT 838) Alicante Ganzin (DVIT 3155) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 
















(DVIT 2914) 

Rubired (DVIT 938) Saint Macaire (DVIT 943) Balta (DVIT 2905) Alicante Ganzin (DVIT 3155) 

Ruby Cabernet (DVIT 940) 

Ruby Seedless (DVIT 2625) 

Russian Seedless (DVIT 2431) 

Russian Seedless 4x (DVIT 1667) 

Sabalkanskoi (DVIT 3077) 


1326), Black Kishmish 

(DVIT 2055) 

Zabalkanski (DVIT 552), Red 

Ohanez (DVIT 2757), Red 

Ohanez (DVIT 499) 


Cabernet Sauvignon (DVIT 

677) 


2759), Malta Seedless 

(DVIT 2327) 

Fuji (DVIT 1655) Alicante Vermelho (DVIT 626) 








351) 


Kishmish-charni (DVIT 2325)

Sahebi (DVIT 508) 






2430), Shando Khari (DVIT 

520), Sultanina Marble 


588236.02), Keshmesh 

(DVIT 580), Rozovii 








(DVIT 1334) 

Sahibi (DVIT 509) Sahibi Sorkh (DVIT 510) 

Sahibi Sorkh (DVIT 510) Sahibi (DVIT 509) 

Saint Emilion (DVIT 942) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











Husseine (DVIT 420), Lal 


Husseine (DVIT 420), Lal 







(DVIT 2914) 

Trebbiano Bianco (DVIT 2067) 

Saint Macaire (DVIT 943) Rubired (DVIT 938) Balta (DVIT 2905) Alicante Ganzin (DVIT 3155) 

Salomani (DVIT 597) 

San Juanina Rosada (DVIT 2927) 

San Martino (P. 215) (DVIT 513) 

Sangiovese (DVIT 945) 

Sangioveto (DVIT 2065) 

Satino (DVIT 946) 

Corniola (DVIT 3162), 



Schiradzouli Violet (DVIT 

515) 

Sauvignon Blanc (DVIT 947) Sauvignon Gris (DVIT 1123) 

Sauvignon Gris (DVIT 1123) Sauvignon Blanc (DVIT 947) 

Sauvignon Vert (DVIT 948) Sauvignon Vert (DVIT 949) 

Sauvignon Vert (DVIT 949) Sauvignon Vert (DVIT 948) 

Sauvis (DVIT 598) 

Schiradzouli Blanc (DVIT 514) 

Schiradzouli Violet (DVIT 515) 

Schiraz (DVIT 952) 


Schwin (DVIT 2604) 






643) 

San Martino (P. 215) (DVIT 

513) 






361) 

Negro Amaro (DVIT 872) Unknown Cultivar (DVIT 881) 



738), Cabernet Sauvignon 

(DVIT 677) 


738), Cabernet Sauvignon 

(DVIT 677) 

Trousseau (DVIT 838), Chenin Blanc 

(DVIT 691), Flora (DVIT 720), 


Trousseau (DVIT 838), Chenin Blanc 

(DVIT 691), Flora (DVIT 720), 




NA (DVIT 1980) Orange Muscat (DVIT 2167) 

Chaouch (DVIT 371), 



2332) 

Black Corinth (DVIT 1052)

Sciacarello (DVIT 1125) Montanaccio (DVIT 836) Oki di Boi (DVIT 877) 

Scolokertek Kiralynoje (DVIT 

516) 

Seeded Thompson (DVIT 1334) 


(DVIT 496) 






2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 








450) 

Seedless Emperor (DVIT 2759) English Colossal (DVIT 397) 

Seedless Emperor (DVIT 517) 

Seedless Tokay (DVIT 1308) 

Seedless Tokay (DVIT 1330) 

Semillon (DVIT 954) 



Rosaki (DVIT 502), Verdal 

(DVIT 602) 

Angelina (DVIT 339), Petinos 

(DVIT 591), Olho De Pargo 

(DVIT 589), Flame Tokay 

(DVIT 2913), Tokay (DVIT 

540), Seedless Tokay (DVIT 

1330) 






(DVIT 540) 



2648), Pirobelle (DVIT 


488), NA (GVIT 1504.02), 

Robin (DVIT 2651), Early 

Muscat (DVIT 394), Bien 

Donnie (DVIT 2696), 


351) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 













(DVIT 2625) 



(DVIT 761) 



(DVIT 761) 

Flora (DVIT 720), 

Aromatico (DVIT 637) 

Sereksiya Rosavi (DVIT 2653) Shtur Angur (DVIT 2168) Koptcha (DVIT 783) 







(DVIT 2914) 



Nasa Veltliner (DVIT 866)

Serine (DVIT 955) 
















(DVIT 2307), Pinot Pernard 

(DVIT 905), Pinot (DVIT 

733) 

Serphiotico (DVIT 956) Julius Caesar (DVIT 579) 

Setzlinge Dodreliabi (DVIT 331) 

Shando Khari (DVIT 520) 

Shingargoon (DVIT 2603) 

Shtur Angur (DVIT 2168) 

Siar (DVIT 522) 

Sicilien (DVIT 523) 

Sidezitis (DVIT 557) 

Sidezitis Proimo (DVIT 558) 






2430), Sahebi (DVIT 508), 


529), NA (PI 588236.02), 









(DVIT 1334) 

Sereksiya Rosavi (DVIT 

2653) 


Pirovano 5 (DVIT 2051), 

Spin Ugad (DVIT 2602), 


Husseine (rish Baba) (DVIT 

421), Nimrang number58 


477) 

Siegerrebe (DVIT 3109) Triplett 30-47 (DVIT 3111) 










1020), NA (DVIT 1978), 



360) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 












Koptcha (DVIT 783) 



2638) 


2046) 


738), Ortega (DVIT 1114) 

Catarratto (DVIT 2058), 



Noir Dautomne (DVIT 330), Precoce 

Dastrachan (DVIT 321) 






(DVIT 2914) 



Karadzhandal (DVIT 2323)

Smederevka (DVIT 1074) Prokupac (DVIT 1071) Negru Virtos (DVIT 2644) 

Sochal Kara-kash (DVIT 2340) 

Somarello Rosso (DVIT 3188) 



(DVIT 737), Gamay Rond 

(DVIT 736) 

Souzao (DVIT 959) Souzao No. 809 (DVIT 960) 

Souzao No. 809 (DVIT 960) Souzao (DVIT 959) 


Spanna (DVIT 961) 

Spin Ugad (DVIT 2602) 

Staphythambelo (DVIT 962) 

Negrette (DVIT 2329), Pinot 

St.george (DVIT 917) 








Staphythostaphylo (DVIT 

963) 

Staphythostaphylo (DVIT 963) Staphythambelo (DVIT 962) 

Stein (DVIT 1127) 


2332) 


Delizia Di Vaprio (DVIT 

571) 



2638) 


1052) 


1052) 

Steinschiller (DVIT 964) Rougitza Rouge (DVIT 2546) Kecskemet (DVIT 2639) 

Stophiliatico (DVIT 966) Gungargoon (DVIT 2500) 

Stravaganza Vidor (DVIT 2259) 

Sultana (DVIT 1321) 

Sultana (DVIT 526) 

Sultana Moscata (DVIT 527) 


Rhazaki Anatolico (DVIT 

608), Asprocoritho (DVIT 

640) 

Sultana (DVIT 526), Austrian 


Austrian Seedless (DVIT 

1650) 

Sultana (DVIT 1321), 

Austrian Seedless (DVIT 

1325), Austrian Seedless 

(DVIT 1650) 

Malta Seedless (DVIT 2327), 

Royal Thompson (DVIT 

1122) 






(DVIT 2625) 


2087), Rizamat (DVIT 2338), Kishmishcharni 

(DVIT 2325), Karadzhandal 

(DVIT 2323) 



1020) 





Thomuscat (DVIT 536)

Sultanina Marble (DVIT 529) 


Summer Muscat (DVIT 2902) 






2430), Sahebi (DVIT 508), 


NA (PI 588236.02), 









(DVIT 1334) 





2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 









(DVIT 1334) 

Summer Royal (DVIT 2899) Eftakilo (DVIT 575) 

Suputinsky Bietyz (DVIT 2702) 




Sylvaner 742 (DVIT 3180) 

Syrah (DVIT 1053) 

Yugoslavia 2798 (DVIT 

2731) 


NA (PI 588709.02), Sylvaner 

(DVIT 968) 



(DVIT 1075) 


(DVIT 968), Sylvaner (DVIT 

1075) 

Syrian (DVIT 2162) Lady Hutt (DVIT 2159) 

CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 











CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 
















(DVIT 2914) 






(DVIT 2914) 

Sultanina Rose (DVIT 530), Kontehgalo 


2325), Diamond Muscat (DVIT 2900), 


Black Rose (DVIT 362) 

Dardari (DVIT 2536) Fatai (DVIT 2510) 



2330), NA (PI 588425.02) 



2330), NA (PI 588425.02) 



2330), NA (PI 588425.02) 



Sylvaner 742 (DVIT 3180) 



Refosco (DVIT 1121), Viognier (DVIT 

2342) 


Tabyrn (DVIT 969) 


Tagouti Rouge (DVIT 551) 

Taifi (DVIT 2174) 

Takhani (pink T.s.) (DVIT 531) 

Taifi (DVIT 2174), NA 

(GVIT 1508.02) 


Kishmish Early (DVIT 2324), 


531) 

NA (GVIT 1508.02), Tabyrn 

(DVIT 969) 



Tagouti Rouge (DVIT 551) 

Talia (DVIT 971) Jampal (DVIT 763) 

Tallian (DVIT 532) 

Tamarez (DVIT 972) 

Taminga (DVIT 2656) 

Tana-Kuzi (DVIT 2928) 

Tand (DVIT 533) 

Tannat (DVIT 973) 

Tand (DVIT 533), Kishmishi 

(DVIT 436) 

Kishmishi (DVIT 436), 

Tallian (DVIT 532) 

Tarrango I (DVIT 2657) Tarrango II (DVIT 2658) 

Tarrango II (DVIT 2658) Tarrango I (DVIT 2657) 

Taschli (DVIT 325) 

Tavriz (DVIT 2659) 







(DVIT 514) 

Teinturier Male (DVIT 974) Teoulier Noir (DVIT 975) 




Teoulier Noir (DVIT 975) Teinturier Male (DVIT 974) 

Ter Gulmer (DVIT 976) Albana (DVIT 622) 

Teroldego (DVIT 977) 

Terra Promessa (DVIT 2114) 












Mathilde (DVIT 99), Asma 

(DVIT 2054) 



(DVIT 604) 



(DVIT 604) 


738) 


530), Touriga (DVIT 990) 




738), Muscat Crime (DVIT 


(DVIT 894) 



2692) 


738), Muscat Crime (DVIT 


(DVIT 894) 








2928), NA (GVIT 1363.05) 



(DVIT 2054) 


2928), NA (GVIT 1363.05) 


(DVIT 2054) 

Joao Santarem (DVIT 764), Rhazaki 

Arhanon (DVIT 554), Unkown Cultivar 

(DVIT 693) 

Joao Santarem (DVIT 764), Unkown 

Cultivar (DVIT 693) 

Taifi (DVIT 2174), NA (GVIT 1363.05) 

Orleans (DVIT 880), Dschan Im Isium 

(DVIT 315) 



Korfiatis (DVIT 784)

Terret Blanc (DVIT 978) 

Terret Bourret (DVIT 979) 


Terret Gris (DVIT 980), 

Terret Bourret (DVIT 979), 

Terret Schwarz (DVIT 1318) 


Terret Blanc (DVIT 978), 



Terret Bourret (DVIT 979), 


Terret Noir (DVIT 1054) Valdepenas (DVIT 1068) 




Terret Bourret (DVIT 979) 

Thiako (DVIT 934) Thiakon (DVIT 981) 

Thiakon (DVIT 981) Thiako (DVIT 934) 

Thompson Seedless (DVIT 535) 



Tinta Cao (DVIT 984) 

Tinta Francisca (DVIT 983) 

Tinta Madeira (DVIT 985) 

Tinta Mole (DVIT 988) 

Tinta Muida (DVIT 986) 

Tinta Pinheira (DVIT 987) 


Kishmishi (DVIT 439), 



2430), Sahebi (DVIT 508), 



529), NA (PI 588236.02), 









(DVIT 1334) 

Balta (DVIT 2905), Tinta 

Pinheira (DVIT 987), Tinta 

Francisca (DVIT 983) 

Balta (DVIT 2905), Tinta Cao 

(DVIT 984), Tinta Pinheira 

(DVIT 987) 

Trincadeiro Branco (DVIT 

993) 

Himrisnky (DVIT 2175), 

Xerez (DVIT 1018) 

Balta (DVIT 2905), Tinta Cao 

(DVIT 984), Tinta Francisca 

(DVIT 983) 

Macaroli (DVIT 799) Ithani Mavro (DVIT 621) 




CG 1481 (DVIT 2432), 




1084), B 15-19 (DVIT 








2327), Queen (DVIT 495), 













578) 




738), Peagudo (DVIT 887) 







(DVIT 2914) 

Malta Seedless (DVIT 2327), G4-74 

(Seedless Calmeria) (DVIT 2914), Maria 


(DVIT 350) 


Esparte (DVIT 2912), Himrisnky (DVIT 

2175) 


(DVIT 993) 

Tintureiro (DVIT 989) Trincadeiro (DVIT 992) Trousseau (DVIT 838) Unkown Cultivar (DVIT 693)



Torgoon (DVIT 2531) 



Tourigan National (DVIT 628) 

Traminer (DVIT 991) 

Trebbiano Bianco (DVIT 2067) 

Trentham Black (DVIT 537) 

Trieste-pirovano 299 (DVIT 2164) 



Trifera (DVIT 600) 




Orange Muskat (DVIT 879) 





Tokay (DVIT 2913), Seedless 


Touriga (DVIT 990), 

Tourigan National (DVIT 

628) 

Tourigan National (DVIT 


Touriga (DVIT 990), Touriga 

(DVIT 1061) 




Trieste 4x (DVIT 1591), 



2164), Trieste 4x (DVIT 

1669) 

Trieste 4x (DVIT 1591), 


2164) 







(DVIT 761) 

Nilok (DVIT 2504), 


Tarrango II (DVIT 2658) 






2672), Perle (DVIT 2754), 

















Vidal 99T (DVIT 3185), 

NA (PI 340969.03), 


654), Vidal 804 (DVIT 

3183) 


2166) 


2166) 


2166) 


(DVIT 598) 



Saint Emilion (DVIT 942) 




Trincadeiro (DVIT 992) Tintureiro (DVIT 989) Trousseau (DVIT 838) Unkown Cultivar (DVIT 693) 

Trincadeiro Branco (DVIT 993) Tinta Mole (DVIT 988) 

Triplett 30-47 (DVIT 3111) Siegerrebe (DVIT 3109) 

Troja (DVIT 994) 


738), Ortega (DVIT 1114) 

Plavac Mali (DVIT 1119) 

Esparte (DVIT 2912), Himrisnky (DVIT 

2175)

Trollinger (DVIT 2341) 

Trousseau (DVIT 838) 

Trousseau (DVIT 995) 

Tsambato (DVIT 996) 



1059), Hillbilly (DVIT 2747), 







650) 




Grey Riesling (DVIT 745), 

Beba (DVIT 651), Bastardo 

(DVIT 650) 













992) 




992) 

Tscharas (DVIT 538) Charas (DVIT 685) Klusaine Red (DVIT 2077) 

Tschilaci (DVIT 539) Husseine (DVIT 576) 

Tsoumbrena Lefki (DVIT 997) 

Tsoumbrena Mavri (DVIT 1055) 

Turkish Grape (DVIT 541) 


Unknown Cultivar (DVIT 1060) 

Unknown Cultivar (DVIT 881) 



Uva Romano (DVIT 542) 

Vaftra (vaspa) (DVIT 998) 




3003) 



Kara Lakana (DVIT 773) 










Valdepenas (DVIT 1068) Terret Noir (DVIT 1054) 

Valdepenas (DVIT 999) 




Mavrodaphne (DVIT 821), 

Kontocladi (DVIT 1036) 

Vital (DVIT 1077), Boal 








Vuthomato (DVIT 610) 






Karabryimis (DVIT 1032) 





Negro Amaro (DVIT 872), Sangiovese 

(DVIT 945) 


Joao Santarem (DVIT 764), Jampal 


Trincadeiro (DVIT 992)

Van Der Laan (DVIT 543) 

Velana (DVIT 1057) 

Venus (seedless) (DVIT 1130) 

Verdal (DVIT 553) 

Verdal (DVIT 602) 

Verdea (DVIT 1000) 

Verdelho (DVIT 1001) 

Vermentino Favorita (DVIT 1002) 

Vermentino Favorita (DVIT 545) 

Vernaccia Bianca (DVIT 1004) 

Vernaccia Di San Gimignano 

(DVIT 1003) 

Vernaccia Sarda (DVIT 1005) 

Vernatschia Alternatures (DVIT 

1076) 

Vertzami (DVIT 1007) 


Victorias Choice (DVIT 2663) 

Victorias Choice (DVIT 3182) 

Victorias Choice number1 (DVIT 

1598) 

Vidal 804 (DVIT 3183) 














(DVIT 3002) 

Fileri (DVIT 1044), Retino 

Lefko (DVIT 1067) 

Golden Champion (DVIT 

411), Rosaki (DVIT 502), 

Verdal (DVIT 602), Seedless 




Rosaki (DVIT 502), Seedless 


Vermentino Favorita (DVIT 

545) 

Vermentino Favorita (DVIT 

1002) 








1598) 






1598) 




Choice (DVIT 2663), 

Moscato Dadda (DVIT 2109) 
















471) 


738) 




(DVIT 725) 







Trebbiano Bianco (DVIT 

2067) 







447), Opale (DVIT 483)

Vidal 99T (DVIT 3185) 

Viognier (DVIT 2342) 

Violento (DVIT 1008) 

Vital (DVIT 1077) 

Volarova (DVIT 601) 

Volitza Lefki (DVIT 1010) 

Fernao Pires (DVIT 1063), 


Volta (DVIT 2043) Volta (DVIT 547) 

Volta (DVIT 547) Volta (DVIT 2043) 

Voltzicki (DVIT 1011) 


Vranac (DVIT 2086) 

Vrathiano (DVIT 1012) 

Vustolethi (DVIT 603) 

Vuthomato (DVIT 610) 

Waltham Cross (DVIT 2343) 

Wengerka (DVIT 326) 

White Calabrian (DVIT 2391) 

White Corinth (DVIT 549) 

White Elbling (DVIT 1014) 

Gaschochi (DVIT 2537), 


















905), Pinot (DVIT 733) 






Rodites (DVIT 505), Rhazaki 

De Crete (DVIT 501), 


(DVIT 458) 

Muscat Of Hungary (DVIT 

2390) 

White Riesling x Sauvignon 

Blanc (DVIT 3187), 



1102), Trebbiano Bianco 

(DVIT 2067) 


693) 

Syrah (DVIT 1053) 

Roussaitis (DVIT 1049) 

Dehbina (DVIT 705) Roussaitis (DVIT 1049) 



611) 









1020), NA (DVIT 1978), 


Vaftra (vaspa) (DVIT 998) 








Chasselas Tompa (DVIT 378), Madam 

Matijas (DVIT 2642) 






(DVIT 2054), Gemeroh (DVIT 1031)

White Riesling (DVIT 1015) 

White Riesling (DVIT 1016) 

White Riesling (PI 588673.02) 

White Riesling x Sauvignon Blanc 

(DVIT 3187) 

White Tokay (DVIT 550) 

NA (PI 588673.06), White 

Riesling (DVIT 1016), White 

Riesling (PI 588673.02) 


NA (PI 588673.06), White 

Riesling (PI 588673.02) 


NA (PI 588673.06), White 

Riesling (DVIT 1016) 

White Elbling (DVIT 1014), 



2259), Rhazaki Anatolico 

(DVIT 608), Asprocoritho 

(DVIT 640) 

White Veltliner (DVIT 1017) Green Veltliner (DVIT 749) 

Wittberger (DVIT 2689) 

Xarello (DVIT 1150) 

Xerez (DVIT 1018) 

Xylomaherudah (DVIT 1019) 

Yaghotti No. 1 (DVIT 604) 

Yaghotti No. 2 (DVIT 2052) 

Yellow Ortlieber (DVIT 1020) 

Himrisnky (DVIT 2175), 

Tinta Muida (DVIT 986) 


Tagouti Rouge (DVIT 551), 


531) 

Chardonnay Musque (DVIT 

1652), Chardonnay (DVIT 

1332), Pinot Noir Epernay 




(DVIT 1117) 





























738) 



1015) 



(DVIT 2325) 



(DVIT 604) 



2751) 

Yourutico (DVIT 605) Fokianoh (DVIT 722) Markandi (DVIT 587) 

Yugoslavia 2798 (DVIT 2731) 

Suputinsky Bietyz (DVIT 

2702) 




Kerner (DVIT 3104), Rotberger (DVIT 

2339) 


(DVIT 993) 

Asma (DVIT 2054) 


(DVIT 2054) 



734) 

Dardari (DVIT 2536) Fatai (DVIT 2510)

Yugoslavia 360 (DVIT 2344) 

Zabalkanski (DVIT 552) 

Zalovitico (DVIT 1021) 

Robin (DVIT 2651), Cardinal 

(DVIT 370), Chaouch 4x 

(DVIT 1663) 


Red Ohanez (DVIT 2757), 


Zengo (DVIT 2691) NA (PI 588668.04) 

Zerk (DVIT 606) 

Zifir (DVIT 2690) NA (GVIT 1614.02) 



503) 



(DVIT 715) 


2332), Gewurztraminer 


1978) 








Ajmi (DVIT 563) 

Zimsko Belo (DVIT 2664) Late Edessa (DVIT 443) Alulu (DVIT 338) Black Ferrara (DVIT 356) 

Zinfandel (DVIT 1023) 




Zweigelt-rebe (DVIT 2692) 







Fort Ross Vine (DVIT 

1353), Helena (DVIT 756), 

Calzin (DVIT 678), Plavac 

Mali (DVIT 1119) 


1353), Helena (DVIT 756), 




1353), Helena (DVIT 756), 




Pias (DVIT 833), Danuque (DVIT 387) 

Table S5: Clonal and first degree relationships for all 950 vinifera accessions included in 

the present study.

egion country USDA ID latitude longitude Collector 

elevation 

(meters) 

east Georgia DVIT 3357 41°48' 43°46' Ed Stover 823 


east Armenia DVIT 3355 39°5' 46°29' Ed Stover 969 


west Sardinia 

39°18' 08°90' Fabrizio Grassi 

east Georgia 

Fabrizio Grassi 

west Sardinia 




west Italy 


west Italy 


west Italy 


west Germany 


west Tunisia 

36°60' 08°46 Fabrizio Grassi 

west Italy 


west Italy 


west Corsica 


west Italy 



west Italy 


west Italy 


west Italy 


west Italy 


west Italy 


west Corsica 


west Italy 


west Sardinia 


west Italy 



west Tunisia 


west Sardinia 


east Turkey 


west Spain 

42°30' 00°70' Raffael Ocete 

west Italy 


west Tunisia DVIT 2426.6 

Harold Olmo 

east Dagestan 


east Pakistan DVIT 2269 

Austin Goheen 

west Spain 



Austin Goheen 


Austin Goheen 


Austin Goheen

east Azerbaijan 



Austin Goheen 


east Turkmenistan 


east Pakistan DVIT 2262.1 

Austin Goheen 

west Tunisia DVIT 2426 

Harold Olmo 

east Turkey 


west Spain 


west Spain 


west Spain 


west Spain 


west Spain 

38°01' 06 47' Raffael Ocete 

west Spain 


west Spain 


west Spain 

Raffael Ocete 

west Spain 


west Spain 


west Spain 


Table S6: Details of the sylvestris samples used in the present study. See the section 

“Supplementary Text: Authenticity of Vitis vinifera subsp. sylvestris” for a discussion of 

authenticity of these sylvestris samples.

Supplementary Methods 

Genotype data quality filters 

To verify genotypes, we compared genotype calls from the BeadStudio software to Illumina GA 

sequence data from 17 accessions of diverse origin for which a strict set of rules were used to 

call genotypes [3]. We only considered genotypes above the following SNP quality thresholds: 

GenTrain Score ≥ 0.3 and GenCall ≥ 0.2. At these thresholds, we observe 93.18% concordance 

for 60411 genotypes called on both platforms, which is an underestimate of genotyping accuracy 

as reduced representation libraries (RRLs) were sequenced with the Illumina GA, which results 

in heterozygotes being called homozygotes at an unknown rate. In addition, lower concordance 

rates are expected in highly diverse species, where numerous unknown flanking polymorphisms 

cause hybridization issues on the genotyping arrays. Based on 145 pairwise comparisons 

between replicate samples genotyped with the Vitis9KSNP array, we discarded SNPs with 

replication rates < 97%. The mean replication rate for the remaining 6507 SNPs was 0.9981. We 

discarded 307 excessively heterozygous SNPs with HWE p-values < 1e-4 within a group of 

vinifera that was pruned so that no two accessions had an Identity-by-State (IBS) > 0.95. SNPs 

with significant excess homozygosity were left in after visual inspection of cluster plots. An 

additional 727 SNPs were removed because they were monomorphic in the sample analysed here 

and 86 poor quality SNPs were removed after visual inspection of cluster plots. The total number 

of SNPs remaining for analysis was 5387. 

The species, cultivar name and cultivar type (wine or table grape) of each sample was obtained 

from the Germplasm Resources Information Network (GRIN) database of the USDA 

(http://www.ars-grin.gov/). Twenty-three samples labeled as vinifera in GRIN were excluded 

from analysis because they were identified as wild Vitis species or wild/vinifera hybrids based on 

multi-dimensional scaling (MDS) plots of an IBS matrix that included hundreds of samples from 

numerous wild Vitis species and wild/vinifera hybrids. A geographic region of origin was 

assigned to 811 vinifera accessions based on mostly on information from GRIN and the 

geographic origin of each sylvestris accession was assigned based on its collection location 

(Table S1). Samples with genotype call rates < 0.7 were excluded. In total, 950 vinifera and 59 

sylvestris remained for analysis.

Supplementary Text 

Biases in grape germplasm collections: a cautionary note 

The conclusions from the present study are based on samples from the USDA grape germplasm 

collection. The USDA grape germplasm collection harbors one of the most (if not the most) 

extensive collection of grape genetic diversity in the world. Nevertheless, it is unclear to what 

extent this collection represents an unbiased sample of the worldwide genetic diversity of 

vinifera. It is possible, for example, that the proportion of the morphological diversity in the 

collection that is represented by clonal mutants is a poor representation of this proportion in 

vinifera worldwide. It is also possible that the extent to which vinifera cultivars are related by 

first degree relationships is less or more than expected from an exhaustive survey of vinifera 

worldwide. All germplasm collections are biased in complex and often unknown ways. This 

study is therefore an evaluation of genetic diversity in the USDA grape germplasm collection 

and is not an unbiased survey of worldwide genetic variation within vinifera. The reader should 

take this cautionary note into consideration when interpreting the results of the present study. 

Potential misnaming of cultivars 

The cultivar names associated with each DNA sample are, in some cases, clearly incorrect. 

There are numerous examples of such mismatches between names and their inferred genetic 

relatedness (see ftp://ftp.gramene.org/pub/vitis_2010_files/): Kyoho (DVIT 616) is actually 

Affenthaler; Mamelon (DVIT 586) is in fact Perle de Csaba; Gamay (DVIT 732) is likely not a 

true Gamay because of its inferred pedigree relationships. We have therefore not only provided 

the name associated with each DNA sample, but also the DVIT number, which is a unique 

identifier carried by each grapevine in the USDA grape germplasm collection. The naming 

inaccuracies may be due to curation error, sample mix-up or other synonym and homonym 

problems, which are common in germplasm collections. However, DNA sample mix-up is an 

unlikely explanation for most inaccuracies because sample processing was done primarily with 

robotics and we observed no discordance for 145 pairwise comparisons between replicate 

samples placed randomly across sample plates. Thus, we conclude that the cases of cultivar 

misidentification in the USDA grape germplasm collection are likely often due to curation error. 

We are confident that our main conclusions will not be strongly influenced by these inaccuracies

in cultivar naming. The present data set is being used to improve the naming accuracy of the 

USDA grape germplasm collection. 

Assignment of cultivars as “wine” or “table” grapes 

In most cases, table and wine grapes can be distinguished unambiguously. In some instances, 

however, cultivars are used for both purposes or their use is unknown. Therefore, we used our 

best judgment in classifying grapes into these categories when ambiguity arose and left cultivars 

as “unknown” when we could not be confident in a classification. Low degrees of 

misclassification are unlikely to affect our results. 

Authenticity of Vitis vinifera subsp. sylvestris 

The identification and collection of true sylvestris from the wild is notoriously difficult because 

of the existence of feral vinifera that can be misidentified as sylvestris and the ease with which 

wild and cultivated grapevines can cross [5,6]. We made use of samples reported as sylvestris 

from several independent collection expeditions. Table S6 provides details of the sylvestris 

samples used in the present study. Latitude, longitude and elevation are provided where data 

were available. Additional information on the samples from the USDA can be accessed via the 

USDA Germplasm Resources Information Network (GRIN: http://www.ars-grin.gov/). 

Five grapevine biologists collected samples we report here as “sylvestris” (Table S6). We have 

no information regarding the criteria employed by Austin Goheen to identify the six vines he 

collected in Pakistan and donated to the USDA in 1987. However, Dr Goheen was an expert in 

the field and we can safely assume that he used the standard set of distinctive morphological 

traits of sylvestris [7]. Similarly, little to no information is available on the strategy used by Dr 

Olmo in collecting the two samples from Tunisia, but he was also a well-known expert in 

grapevine identification. Furthermore, in PC space (Figure 4) the two Tunisian samples 

collected by Dr Olmo are found close to the two Tunisian samples provided by Dr Grassi, whose 

criteria for identification follow. The largest proportion (42 of 59) of our sylvestris samples come 

from Dr Grassi and Dr Ocete who used well-established guidelines for the identification of 

sylvestris [7]. The most relevant differentiating trait used to distinguish between wild and 

cultivated vines was the mating system: sylvestris is dioecious while vinifera is hermaphroditic

with very few exceptions. Sampling was carried out during bloom, so it was possible to verify 

the flower sex. Moreover, they ensured that the sylvestris samples did not come from 

populations growing adjacent to present-day vineyards or land known to be previously planted 

with vinifera. We note, however, that despite the use of these criteria, it is impossible to rule out 

the possibility of influence from cultivated vines. All sampling locations were at an altitude of 

0-500m with environmental conditions typical for sylvestris: wetlands and forests with high 

humidity, as well as the frequent presence of tree species such as elm, poplar, hawthorn, 

hornbeam and oak on which sylvestris grow as lianas. Ed Stover also used similar standard 

criteria and ensured that the 9 samples used here from Armenia and Georgia did not come from 

populations growing adjacent to present-day vineyards or land known to be previously planted 

with vinifera. Dr Stover collected samples in October and therefore could not confirm flower 

sex. However, the vast majority of the vines was not bearing fruit and was therefore presumably 

male. It is worth noting that results are at least consistent across sampling expeditions: samples 

collected from similar geographic locations on different expeditions generally cluster together in 

PC space (Figure 4, Figures S7-S9). 

It is impossible to claim with absolute certainty that the samples we identify as sylvestris are 

truly wild samples with no genetic input from cultivated grapevines. This is the case for any 

study that makes use of sylvestris samples [e.g. 8,9]. While it is most common to make use of 

morphological criteria exclusively, here we enhance the morphological criteria we have used to 

identify sylvestris with genetic analyses. 

To provide a visualization of the relationships among all of the samples used in the present 

study, a PCA plot was generated using the 2958 LD-pruned SNPs on the set of unique vinifera 

and sylvestris samples (Figure S7). Figure S7 shows that there is one exceptional sample, 

indicated with an arrow, that is labeled as vinifera but clearly clusters among the western 

sylvestris. This sample was donated to the USDA collection in 1983 by Dr Harold Olmo and is 

labeled in GRIN as “Beli Potok” (DVIT 1841), a female vine likely of Serbian origin with 

morphological characteristics that are consistent with it being sylvestris. Olmo’s “wild vinifera” 

samples, indicated by black dots in Figure S7, are discussed below. From Figure S7, it is 

apparent that the western sylvestris are relatively distinct and pull out along both PC1 and PC2.

Figure S8, which includes only western sylvestris and western vinifera, also demonstrates that 

western sylvestris samples do not overlap with vinifera in genetic space. These observations lend 

credibility to the identity of the western sylvestris and suggest that our criteria for authenticating 

sylvestris based strictly on morphology meaningfully capture the genetic relationship between 

sylvestris and vinifera. 

Figure S7 suggests that the eastern sylvestris, however, are largely indistinguishable from 

vinifera. Figure S9, which includes only the eastern sylvestris and the eastern vinifera, clearly 

demonstrates that this is the case. Particularly noteworthy, however, is the tight cluster of black 

dots in the top right of the plot. These samples, along with the other black dots, are labeled as 

eastern vinifera from Afghanistan and were donated in 1983 by Dr Harold Olmo. Additional 

notes associated with these samples indicate that they are “wild vinifera”, which suggests that 

they may be sylvestris. We refer to these samples as Olmo’s “wild vinifera”. Additional 

information on these samples, including images, is available from GRIN (http://www.ars- 

grin.gov). Many of these samples are inter-related: the five samples in the top right corner of the 

plot are all sibs, for example (see ftp://ftp.gramene.org/pub/vitis_2010_files/). 

From Figure S9, it is evident that 11 of the 21 eastern sylvestris samples, within the dotted 

ellipse, cluster discretely to the right side of the plot. One may reason that, because of their 

apparent unique genetic identity compared to vinifera, these 11 sylvestris samples represent true 

sylvestris. This reasoning, however, is not completely sound. Our observation that the 

remaining 10 eastern sylvestris samples overlap with vinifera in genetic space is consistent with 

standard models of grape domestication. Because the grape was most likely domesticated in the 

east and the number of generations that have passed since the domestication event(s) may have 

been severely reduced by vegetative propagation, we do not necessarily expect eastern sylvestris 

to be distinguishable from vinifera using the Vitis9KSNP array data. In addition, we make no 

claims in the present study about a single or multiple domestication events: it is possible, and 

arguably likely, that genetic input into present day vinifera came from sylvestris from several 

geographical areas in the east. The inability to reliably identify true sylvestris samples makes 

testing such hypotheses (i.e. single versus multiple domestication events) intractable and we 

therefore do not attempt to do so. In addition, the clustering of the eastern sylvestris in genetic

space is unlikely to reflect biases in sampling: samples within the group of 11 sylvestris on the 

right side of the plot and within the group of 10 sylvestris samples on the left side of the plot both 

contain individuals collected on independent sampling expeditions by independent samplers. 

Nevertheless, one could argue that the 11 eastern sylvestris samples which cluster to the right in 

Figure S9 and do not overlap with vinifera in genetic space are the most likely true sylvestris 

samples. Therefore, we reanalyzed our data, restricting our “eastern sylvestris” samples to 

include only the 11 samples within the dotted ellipse in Figure S9. We also excluded several 

other samples: Olmo’s 12 “wild vinifera” samples from Afghanistan, because of their unknown 

identity, and the “Beli Potok” vinifera sample which clusters with western sylvestris. With this 

refined sample set, our conclusions based on Fst, haplotype diversity, LD decay and the 3 

population test do not change. Thus, after using genetic data to evaluate the identity of our 

sylvestris samples and restricting our analyses to subsets of the sample in which we are most 

confident, our conclusions remain unchanged. 

Despite our extensive evaluations of the identity of the sylvestris samples above, it is 

nevertheless important to consider how the misidentification of wild material would affect the 

conclusions of the present study. Here we consider the effect of misidentity on two of the main 

conclusions drawn from the analysis of sylvestris data: that vinifera experienced a weak 

population bottleneck and that western vinifera experienced introgression from western 

sylvestris. 

Our conclusions about the strength of the population bottleneck are based on differences in 

haplotype diversity and the decay of LD between vinifera and sylvestris. If the sylvestris 

samples from the present study experienced introgression from cultivated vines, this would 

indeed bias our estimates of diversity in the sylvestris we used to represent the “progenitor” or 

“ancestral sample”. In addition, it is well known that habitat loss has led to a decline in 

population sizes of sylvestris [5,10]. Thus, even an exhaustive sample of contemporary sylvestris 

populations may not fully represent ancestral diversity because of recent population bottlenecks. 

This sampling issue is an inevitable source of potential bias that is applicable to any study of this 

nature. Despite these caveats, we provide the most comprehensive examination of genome-wide 

patterns of LD decay and haplotype diversity in vinifera and sylvestris to date. The identity of

the eastern sylvestris is more questionable than the identity of the western sylvestris because 

eastern sylvestris clusters more closely with vinifera in genetic space (Figures S7) and because it 

is safe to assume that there was a much longer period of potential contact between sylvestris and 

vinifera in the east than in the west. We therefore performed the LD decay and haplotype 

diversity analyses with only the western sylvestris samples instead of the full sylvestris sample. 

Our results remain unchanged and this demonstrates that our conclusions are robust to this 

potential source of misidentification. Moreover, it is clear that levels of haplotype diversity are 

very high in vinifera regardless of the diversity observed in our sample of sylvestris (median 

haplotype diversity of 5 SNP haplotypes = 0.80, Figure 3), which is consistent with a weak 

domestication bottleneck. Finally, it is worth noting that our model of a weak domestication 

bottleneck is consistent with the widespread use of vegetative propagation during the history of 

grapevine breeding [11] and the relatively minor morphological changes between sylvestris and 

vinifera [6]. 

Our analysis using the 3-population test [12] supports a scenario in which vinifera from western 

Europe are the result of an admixture event between vinifera from the east and sylvestris from 

western Europe. Using this same test, we find no support for a scenario in which our “western 

sylvestris” sample is influenced by gene flow from vinifera. To our knowledge, this is the most 

robust statistical framework we can employ to test models of the admixture history of vinifera 

and sylvestris. This test, in conjunction with the PCA results (Figure 4), provides strong support 

for the unique, unadmixed identity of the western sylvestris sample. Because the eastern 

sylvestris sample is almost indistinguishable from the vinifera (Fst < 0.06 to all vinifera 

populations, which is the pattern we expect if domestication occurred in the east and there was a 

weak population bottleneck), the identity of the eastern sylvestris will necessarily rely on the 

morphological criteria used for its identification. However, the identity of the eastern sylvestris 

is extraneous to our main conclusion that western vinifera experienced introgression from 

western sylvestris: the 3-population test used to test this hypothesis does not include eastern 

sylvestris.

In conclusion, we acknowledge that the identification of true sylvestris presents significant 

challenges to robustly testing hypotheses about the domestication of the grape and that any 

conclusion drawn based on data from “sylvestris” samples should be interpreted with caution. 

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