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nomenclatural notes on lycidae with a description of a new species

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ACTA UNIVERSITATIS PALACKIANAE OLOMUCENSIS<br />

FACULTAS RERUM NATURALIUM (2000) BIOLOGICA 38<br />

NOMENCLATURAL NOTES ON LYCIDAE WITH A<br />

DESCRIPTION OF A NEW SPECIES (COLEOPTERA)<br />

Ladislav Bocak 1 and Milada Bocakova 2<br />

1 Department <strong>of</strong> Zoology and Anthropology, Faculty <strong>of</strong> Sciences, Palacký University,<br />

Tř. Svobody 26, 771 46 Olomouc, Czech Republic<br />

2 Regi<strong>on</strong>al Museum, nám. Republiky 5, 771 73 Olomouc, Czech Republic<br />

Received July 31 st , 2000; accepted August 22 nd , 2000<br />

Key words: Tax<strong>on</strong>omy, Lycidae, <strong>new</strong> syn<strong>on</strong>yms, <strong>new</strong> combinati<strong>on</strong>s, <strong>new</strong> <strong>species</strong>, Oriental<br />

and Neotropical Regi<strong>on</strong><br />

Abstract<br />

Teroplas ruber sp. n. is described. The following <strong>new</strong> syn<strong>on</strong>yms are proposed: Slipinskia Bocak et<br />

Bocakova, 1992 (= Aferos Kasantsev, 1992), Lyp<strong>on</strong>ia guerryi Pic, 1939, L. robusta Pic, 1922, L. diversicornis<br />

Pic, 1926 (= L. laticornis Fairmaire, 1899), Scarelus maxwelli Kasantsev et Yang, 1999 (= S. anthracinus<br />

Bocakova et Bocak, 1999). Xylobanus angustior Pic, 1922 and Xylobanus angustior var. lateniger Pic, 1922 are<br />

transferred to the genus Wakarumbia Bocak, 1999, Xylobanus angustior var. lateniger is elevated to the<br />

<strong>species</strong> status.<br />

Introducti<strong>on</strong><br />

During our stay in the Nati<strong>on</strong>al Museum <strong>of</strong> Natural history in Paris we found<br />

additi<strong>on</strong>al type material, which clarify the identity <strong>of</strong> some taxa introduced to literature<br />

by Maurice Pic in the first half <strong>of</strong> 20 th century. Additi<strong>on</strong>ally, we brought some<br />

comments <strong>on</strong> the syn<strong>on</strong>ymy <strong>of</strong> several taxa and <strong>on</strong>e <strong>new</strong> <strong>species</strong> found in the material<br />

<strong>of</strong> the Natural History Museum in Paris.<br />

Depositories:<br />

LMBC – Authors’ collecti<strong>on</strong><br />

MHNP – Nati<strong>on</strong>al Museum <strong>of</strong> the Natural History, Paris, France<br />

ZMPA – Muzeum Zoologii PAN, Warszawa, Poland<br />

41


Subfamily Platerodinae<br />

Teroplas Gorham, 1884<br />

Teroplas Gorham, 1884: 243. (Type <strong>species</strong>: Caloclad<strong>on</strong> fusculus Gorham, 1881, by<br />

m<strong>on</strong>otypy).<br />

Remark: This American genus can be easily distinguishable from other Platerodinae<br />

in having l<strong>on</strong>g antennae reaching at least apical quarter <strong>of</strong> elytra in both sexes. Its<br />

male genitalia can be discriminated by l<strong>on</strong>g parameres and two ventrodistal projecti<strong>on</strong>s<br />

<strong>of</strong> phallus. A more detailed descripti<strong>on</strong> was given by Bocakova (in press).<br />

Including the <strong>species</strong> described below Teroplas comprises three <strong>species</strong>.<br />

Teroplas ruber sp. n.<br />

Type material: Holotype, male, “Cubilguitz, Vera Paz, Champi<strong>on</strong>” (MNHN).<br />

Etymology. Named in reference to body colorati<strong>on</strong>.<br />

Diagnosis. Teroplas ruber sp. n. differs from other known Teroplas in having red<br />

body colorati<strong>on</strong>, short phallus, and in paramerae c<strong>on</strong>tiguous dorsally.<br />

Descripti<strong>on</strong>. Body dark brown, <strong>on</strong>ly pr<strong>on</strong>otum red, scutellum and elytra redbrown.<br />

Head <strong>with</strong> large eyes, eye diameter 1.8 times l<strong>on</strong>ger than interocular distance.<br />

Antennae filiform, reaching apical <strong>on</strong>e tenth <strong>of</strong> elytra. Antennomere 1 stout, 2 small,<br />

3 by <strong>on</strong>e-quarter shorter than 4. Maxillary palpi short, terminal palpomere securiform.<br />

Pr<strong>on</strong>otum trapezoidal, transverse, widest at basal margin, anterior margin semicircularly<br />

produced forwards, sides elevated. Posterior angles obliquely produced backwards.<br />

Scutellum obl<strong>on</strong>g, weakly emarginate apically. Elytra el<strong>on</strong>gate, slightly widened<br />

apically, 2.4 times as l<strong>on</strong>g as their width medially. Elytra provided <strong>with</strong> dense brown<br />

pubescence. Each elytr<strong>on</strong> <strong>with</strong> 9 weak costae, reticulate cells irregular, but distinct.<br />

Terminal abdominal sternum el<strong>on</strong>gate. Legs slender, compressed, tibiae straight, their<br />

spurs small, trochanters el<strong>on</strong>gate. Male genitalia <strong>with</strong> phallus as l<strong>on</strong>g as paramerae,<br />

ventrodistal projecti<strong>on</strong>s <strong>of</strong> phallus short. Dorsal margins <strong>of</strong> paramerae mutually<br />

c<strong>on</strong>tiguous. Body length: 6.5 mm, humeral width: 1.7 mm. Female unknown.<br />

Distributi<strong>on</strong>: Central America.<br />

Lyp<strong>on</strong>ia (P<strong>on</strong>yalis) laticornis Fairmaire<br />

P<strong>on</strong>yalis laticornis Fairmaire, 1899: 623.<br />

Lyp<strong>on</strong>ia guerryi Pic, 1939: 165 – syn. n.<br />

Lyp<strong>on</strong>ia robusta Pic, 1922: 13 – syn. n.<br />

Lyp<strong>on</strong>ia diversicornis Pic, 1926: 70 – syn. n.<br />

Material examined: Holotype <strong>of</strong> Lyp<strong>on</strong>ia laticornis, male, “Kuatain, Delot<strong>on</strong>, Chine”<br />

(MHNP); Holotype <strong>of</strong> Lyp<strong>on</strong>ia guerryi, female, “Yunnan-Sen 1911”(MHNP);<br />

Holotype <strong>of</strong> Lyp<strong>on</strong>ia robusta, female, “Pe Yen Tsin, Yunnan” (MHNP); Holotype <strong>of</strong><br />

Lyp<strong>on</strong>ia diversicornis, male, “Yunnan” (MHNP).<br />

The revisi<strong>on</strong> <strong>of</strong> Lyp<strong>on</strong>ia was recently published (Bocak, 1999). Although we visited<br />

several times the Natural History Museum in Paris, we found <strong>on</strong>ly recently the types <strong>of</strong><br />

42


Figs 1–5: Teroplas ruber sp. n.: 1 – head and pr<strong>on</strong>otum, 2 – terminal abdominal segments; 3 – basal<br />

antennomeres; 4 – male genitalia.<br />

43


above given <strong>species</strong> described by Maurice Pic. These <strong>species</strong> were left “incertae sedis”<br />

in the revisi<strong>on</strong> and because <strong>of</strong> poor descripti<strong>on</strong>s they were not even classified to<br />

subgenera. The examinati<strong>on</strong> <strong>of</strong> holotypes revealed that they are junior subjective<br />

syn<strong>on</strong>yms <strong>of</strong> Lyp<strong>on</strong>ia (P<strong>on</strong>yalis) laticornis.<br />

Subfamily Ateliinae<br />

Scarelus anthracinus Bocakova et Bocak<br />

Scarelus anthracinus Bocakova et Bocak, 1999: 108 (published April 1999)<br />

Scarelus maxwelli Kasantsev et Yang, 1999: 245 (received December 29, 1998,<br />

accepted May 28, 1999, exact date <strong>of</strong> publicati<strong>on</strong> not available) – syn. n.<br />

Scarelus pedlenburyi Kasantsev et Yang, 1999: 245 – nomen nudum.<br />

Bocak (1995) misunderstood the identity <strong>of</strong> S. riedeli and S. umbrosus <strong>on</strong> the basis<br />

<strong>of</strong> the study <strong>of</strong> <strong>on</strong>e <strong>of</strong> paratypes <strong>of</strong> S. umbrosus (deposited in ZIW) which is not<br />

c<strong>on</strong>specific <strong>with</strong> holotype. Bocakova and Bocak (1999) clarified the situati<strong>on</strong> and<br />

described S. anthracinus. Later, Kasansev and Yang (1999) made the same correcti<strong>on</strong><br />

and they described <strong>new</strong> <strong>species</strong> S. maxwelli. Their article was published later and the<br />

name S. maxwelli is a junior subjective syn<strong>on</strong>ym <strong>of</strong> S. anthracinus. Although Kasantsev<br />

and Yang (1999) designated the name Scarelus maxwelli as a <strong>new</strong> name, they designated<br />

different specimen as a holotype (specimen collected by Dr. Kasansev in Malaysia<br />

in February 1999). Therefore S. maxwelli can not be either c<strong>on</strong>sidered as a <strong>new</strong><br />

replacement name or the designati<strong>on</strong> <strong>of</strong> the holotype would be invalid (ICZN, 1985,<br />

Art. 72 e; ICZN, 1999, Art. 72.7). We c<strong>on</strong>sider S. maxwelli to be a <strong>new</strong>ly described<br />

<strong>species</strong> and c<strong>on</strong>sequently it is not objective but subjective syn<strong>on</strong>ym. The name Scarelus<br />

pedlenburyi designated as “nomen novum” refers to the same <strong>species</strong> by mistake<br />

(Kasantsev and Yang, 1999). It is c<strong>on</strong>sidered to be an unavailable name here.<br />

Kasantsev and Yang (1999) designated also Scarelus luchti as a nomen novum,<br />

although their intenti<strong>on</strong> to describe a <strong>new</strong> <strong>species</strong> is obvious from the c<strong>on</strong>text. The<br />

remaining nomina nova published by Kasantsev and Yang (1999) c<strong>on</strong>form to the Code<br />

(ICZN, 1985, 1999).<br />

Subfamily Metriorrhynchinae<br />

Wakarumbia angustior (Pic) – comb. n.<br />

Xylobanus angustior Pic, 1922: 16<br />

Material examined: Holotype, male, Nord-Celebes, Toli-Toli, Nov.– Dez. 1895, H.<br />

Fruhstorfer (MHNP).<br />

Wakarumbia lateniger (Pic) – comb. n.<br />

Xylobanus angustior var. lateniger Pic, 1922: 16<br />

Material examined: Holotype, female, S. Celebes, Patutuang Jan. 1896, H. Fruhstorfer<br />

(MHNP).<br />

44


W. lateniger was described as a variety <strong>of</strong> Xylobanus angustior. We found that this<br />

tax<strong>on</strong> is separate <strong>species</strong> and transfer it to Wakarumbia.<br />

Subfamily Erotinae<br />

Aferos Kasantsev<br />

Aferos Kasantsev, 1992: 44 (November 30, 1992)<br />

Type <strong>species</strong>: Stadenus aethiops Kleine, 1992<br />

Slipinskia Bocak and Bocakova, 1992: 257 (December 31, 1992)<br />

Type <strong>species</strong>: Stadenus aethiops Kleine, 1992<br />

The descripti<strong>on</strong> <strong>of</strong> Slipinskia was published later and therefore it is a junior<br />

syn<strong>on</strong>ym <strong>of</strong> Aferos. Both genera are based incidentally <strong>on</strong> the same type <strong>species</strong>.<br />

Acknowledgments<br />

We are very grateful to Dr. J. J. Ménier (Paris), Mr. M. Kerley and M. J. D.<br />

Brendell (L<strong>on</strong>d<strong>on</strong>) and Mr. T. Huflejt (Warszawa) who enabled us to study the rich<br />

material deposited in their instituti<strong>on</strong>s. The study was partially supported by a grant<br />

from the Grant Agency <strong>of</strong> the Czech Republic no. 206/98/0539 (L. Bocak) and by<br />

a grant no. RK99P03OMG011 from the Ministry <strong>of</strong> Culture <strong>of</strong> the Czech Republic<br />

(M. Bocakova). The support is gratefully acknowledged.<br />

References<br />

BOCAK, L. (1995): Review <strong>of</strong> the genus Scarelus Waterhouse from Sumatra and Malaysia (Coleoptera:<br />

Lycidae). Stutt. Beitr. Naturk. Ser. A, No 535: 1–10.<br />

BOCAK, L. (1999): A revisi<strong>on</strong> and phylogenetic analysis <strong>of</strong> the genus Lyp<strong>on</strong>ia C. O. Waterhouse, 1878<br />

(Coleoptera, Lycidae). Entomol. Basiliensia 21: 59–103.<br />

BOCAK, L. (2000): Revisi<strong>on</strong> <strong>of</strong> the genus Wakarumbia (Coleoptera: Lycidae). Eur. J. Entomol. 97: 271–278.<br />

BOCAK, L. and BOCAKOVA, M. (1990): Revisi<strong>on</strong> <strong>of</strong> the supergeneric classificati<strong>on</strong> <strong>of</strong> the family Lycidae<br />

(Coleoptera). Pol. Pismo Entomol. 59: 623–676.<br />

BOCAKOVA, M. (in press): Revisi<strong>on</strong> and phylogenetic analysis <strong>of</strong> the subfamily Platerodinae (Coleoptera,<br />

Lycidae). Eur. J. Entomol.<br />

BOCAKOVA, M. and BOCAK, L. (1999): New Lycidae from peninsular Malaysia (Coleoptera). Entomol.<br />

Basiliensia 21: 105–109.<br />

FAIRMAIRE, L. (1899): Descripti<strong>on</strong>s de Coléopteres nouveaux recueillis en Chine par M de Latouche.<br />

Ann. Soc. Ent. Fr. 68: 616–642.<br />

GORHAM, H. S. 1881: Malacodermata. Fam. Lycidae. Biol. Centr. Amer. Col. 3 (2): 1–29.<br />

GORHAM, H. S. 1884: Malacodermata. Suplement. Fam. Lycidae. Biol. Centr. Amer. Col. 3(2): 225–272.<br />

ICZN, 1985: Internati<strong>on</strong>al Code <strong>of</strong> Zoological nomenclature. Third Editi<strong>on</strong>. The Intenati<strong>on</strong>al Trust for<br />

Zoological Nomenclature xx+338 pp. L<strong>on</strong>d<strong>on</strong>.<br />

ICZN, 1999: Internati<strong>on</strong>al Code <strong>of</strong> Zoological Nomenclature. Fourth Editi<strong>on</strong>. The Intenati<strong>on</strong>al Trust for<br />

Zoological Nomenclature xxix+305 pp. L<strong>on</strong>d<strong>on</strong>.<br />

KASANTSEV, S. (1992): Revisi<strong>on</strong> <strong>of</strong> the genus Stadenus (Coleoptera Lycidae) <strong>of</strong> Africa. Russ. Entomol. J.<br />

1(1): 37–50.<br />

45


KASANTSEV, S. and YANG, P. S. (1999): Tax<strong>on</strong>omic and Syn<strong>on</strong>ymic Notes <strong>on</strong> Lycidae (Coleoptera) <strong>with</strong><br />

Descripti<strong>on</strong>s <strong>of</strong> New Species from Taiwan. Chinese J. Entomol. 19: 239–247.<br />

PIC, M. (1922): C<strong>on</strong>tributi<strong>on</strong> a l’étude des Lycides. L’Échange Nr. 407: 16 hors texte.<br />

PIC, M. (1926): Sur le genre Lyp<strong>on</strong>ia Wat. (Col. Lycidae). Bull. bimens. Soc. Linn. Ly<strong>on</strong> 5: 70.<br />

PIC, M. (1939): Malacodermes exotiques. L’Echange 475: 165<br />

46<br />

Acta Univ. Palacki. Olomuc.<br />

Fac. rer. nat. (2000)<br />

Biol. 38, 41–46

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