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UTRECHT MICROPALEONTOLOGICAL BUllETINS

UTRECHT MICROPALEONTOLOGICAL BUllETINS

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drites are trace-fossils consisting of a system of regularly branching tunnels<br />

of uniform diameter, formed by some animal working from a fixed basepoint<br />

at or just below the seafloor. It is a cosmopolitan genus, reported with<br />

certainty from the Ordovicium to the Tertiary. Our Trubi specimens show a<br />

layered, carbonate filling which is more indurated and of lighter colour than<br />

the surrounding sediment. Preferential induration of such tunnel fillings has<br />

been noted before by Kennedy (1970). The entire genus Chondrites undoubtedly<br />

belongs to the Fodichnia and has to be regarded as feeding structures<br />

of sediment-eating animals and not as dwelling burrows of filter-feeding<br />

annelids (Seilacher, 1964). Simpson regards these burrows as the result of<br />

sipunculoid (= unsegmented) worms working from a fixed centre at the<br />

surface of the sediment and producing tunnels by an extensile proboscis; the<br />

branching pattern may be caused by phobotaxis (= behaviour to avoid<br />

penetrating its own tunnel or the tunnel of others in its passage through the<br />

sediment).<br />

A bed with Chondrites is of marine origin and the water was not greatly<br />

agitated by wave-action (Simpson, 1957, p. 494). Warme et al. (1973) stated<br />

that Chondrites has little value as a bathymetric indicator, being cosmopolitan<br />

and crossing both bathymetric and lithologic facies boundaries. But<br />

the presence of this trace fossil requires that the bottom waters be oxygenated<br />

when the traces were formed, and that the sediments contain sufficient<br />

food.<br />

The middle part of the Trubi in section 4 with its well-bedded appearance<br />

resembles the lower part of the Trubi on top of the basal 6 to 7 m of vaguely<br />

bedded, compact limestones. A variable rate of deposition caused by a<br />

fluctuating supply of terrigenous fines seems to be valid for this part of the<br />

section as well. The presence of clay-seams in horizontal partings and the<br />

presence of Chondrites point to sedimentation conditions comparable to<br />

those described for the lower part of the section.<br />

A low rate of sedimentation enabling burrowers to homogenize the<br />

sediment returns in the topmost Trubi part at Punta Piccola. Bottom conditions<br />

apparently remained favourable for bottom-dwelling life. Pyritisation<br />

in boreholes seems to be responsible for the concentric rings around the<br />

observed pipes. The upward decrease in diameter of the burrows may point<br />

to an unknown change in biotope favouring smaller sized burrowers.

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