Christoph Oberprieler The Systematics of Anthemis L. - Herbmedit.org

6 Oberprieler: Anthemis in N Africa
12. Species delimitation in the Anthemis boveana group .............................................. 76
13. Species delimitation in the Anthemis pedunculata - Anthemis punctata complex ... 85
14. Systematic descriptions and keys .. ......................................................................... 107
Key to the N African species .... .................. ................... ......... ..... .. ................ 109
Clé des espèces de l'Afrique du Nord .......... ...... ............. ....... .................... ... 111
. Anthemis subg. Anthemis ............... .......... ...................... .................. .............. 114
Anthemis sect. Anthemis ..................................... .. .................. ...... ... 114
Anthemis sect. Hiorthia ..................................... ....................... ....... 227
Anthemis sect. Maruta ..... ................................. .......... ............... .... .. 297
Anthemis subg. Cota .. .............. ......................... ............................................. 307
Anthemis sect. Cota ....................................... .................................. 307
15. Acknowledgements .... .. ........................................................ .................................. 313
16. References ........... .................... ........................................................... ................... 314
17. Index to scientific names ........... ........ .. .. ................................................. .. .... ......... 324
1. Introduction
The genus Anthemis belongs to the family Compositae Giseke, alternatively named Asteraceae
Dumort. It is the name-giving genus of the Anthemideae Cass., a tribe which
presently consists of 109 genera (Bremer & Humphries 1993; adding Castrilanthemum
recently described by Vogt & Oberprieler 1996) and shows an extratropical, mainly Old­
World-centred distribution. According to the most recent phylogenetic reconstruction of
the family (Bremer 1996), the Anthemideae are a member of the subfamily Asteroideae
and form the sister group to the Astereae. While Heywood & Humphries (1977) indicate
around 130 species in the genus, more recent estimates give a total of c. 210 annual,
biennial, or perennial species (Bremer & Humphries 1993).
Since Candolle's account of Anthemis in his Prodromus (Candolle 1838), the genus has
never again been subject to a complete revisionary treatment. However, quite comprehensive
regional treatments like Boissier's (1875) Flora orientalis account and Eig's (1938)
studies on the orientai species of Anthemis, along with revisions made in the course of the
large flora projects of the last 40 years, have resulted in our up-to-date understanding of
the genus over a large part of its distributional range, viz. Europe (Fernandes 1975a,
1975b, 1976, 1983), E Europe and W Asia (Fedorov 1961), Turkey (Grierson & Yavin
1975), the Near East (Feinbrun-Dothan 1978), the Flora iranica area (Iranshahr 1986),
and Saudi Arabia (Ghafoor & AI-Turki 1997). Additional information was contributed by
revisionary treatments of defined infrageneric groups (A. ammanthus group, Greuter 1968;
A. sect. Maruta, Yavin 1970; A. sect. Anthemis, Yavin 1972; A. tomentosa group, Georgiou
1990) and by geographically limited biosystematic revisions (Bulgaria: Kuzmanov &
al. 1981 and Thin 1983; Spain: Benedf i Gonzalez 1987).
In N Africa, as compared to other regions within the distributional range of Anthemis,
the taxonomy of the genus is poorly understood, and a criticaI revision is lacking. While
there are flora treatments available for Algeria (Quézel & Santa 1963), Tunisia (Pottier­
Alapetite 1981), Libya (Alavi 1986), and Egypt (Tackholm 1974), for Morocco there is
only a checklist (Jahandiez & Maire 1934). The main aim of the present study was to fill