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Christoph Oberprieler The Systematics of Anthemis L. - Herbmedit.org

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40 <strong>Oberprieler</strong>: <strong>Anthemis</strong> in N Africa<br />

chromosomes. A further karyotype analysis was performed by Benedf i Gonzalez (1987)<br />

who indicates the chromosome formula 12m + 2M + 2st-sat + 2t-sat for plants from near<br />

Barcelona. Karyotype analyses performed for the present study yielded a chromosome<br />

formula <strong>of</strong> 10m + 4sm + 4st-sat for two Moroccan populations and a more symrnetric<br />

karyotype with 14m + 4st-sat for a S Spanish population (Table 3).<br />

<strong>Anthemis</strong> arvensis subsp. sphacelata<br />

n=9<br />

- It, Reggio Calabria, Montalto, 1600 m, 27 May 1994, Vogt 13983 & <strong>Oberprieler</strong> 8288.<br />

This interesting biennial to perennial subspecies <strong>of</strong> <strong>Anthemis</strong> arvensis endemie to the<br />

mountains <strong>of</strong> Sicily and S Italy, had been studied cytologically only once, by Brullo & al.<br />

(1978a) who found a diploid chromosome set <strong>of</strong> 2n = 18 chromosomes (12m + 4sm + 2tsat)<br />

in a Sicilian population. <strong>The</strong> present count from the Calabrian mountains corroborates<br />

this data.<br />

<strong>Anthemis</strong> (subg. <strong>Anthemis</strong>, sect. <strong>Anthemis</strong>) ser. Chrysanthae<br />

<strong>Anthemis</strong> gharbensis<br />

n=9<br />

-Ma, Arbaoua - Moulay Bousselham, lO m, 24 Apr 1993, Vogt 10161 & <strong>Oberprieler</strong> 4609.<br />

-Ma, Forèt de la Mamora, 120 m, 27 Apr 1993, Vogt 10259 & <strong>Oberprieler</strong> 4707.<br />

2n = 18<br />

* - Ma, Forèt de la Mamora, 120 m, 2 May 1987, Podlech 43489.<br />

<strong>The</strong> haploid and diploid chromosome numbers mentioned above were already reported<br />

earlier (<strong>Oberprieler</strong> 1994). <strong>The</strong> karyotype was found to be very symrnetrical, consisting <strong>of</strong><br />

5 pairs <strong>of</strong> metacentic, 2 pairs <strong>of</strong> submetacentric, and 2 pairs <strong>of</strong> satel\ited subtelocentric<br />

chromosomes (10m + 4sm + 4st-sat, Table 4).<br />

Table 4. Chromosomes <strong>of</strong> <strong>Anthemis</strong> gharbensis (OTU 11; n = 8).<br />

Chromo- Long arm Short arm Totallength Relative length Arm ralio Chromosome<br />

some pair (IJm) (IJm) (IJm) (%) type<br />

4.54 3.60 8.14 14.4 1.26 m<br />

Il 3.95 3.18 7.13 12.6 1.24 m<br />

III 4.63 2.28 6.91 12.2 2.03 sm<br />

IV 3.44 2.93 6.37 11.2 1.17 m<br />

V 4.33 1.00 6.11 10.8 4.33 sI-saI<br />

VI 3.32 2.65 5.97 10.5 1.25 m<br />

VII 3.69 1.94 5.63 9.9 1.90 sm<br />

VIII 3.02 2.31 5.33 9.4 1.30 m<br />

IX 3.55 0.84 5.17 9.1 4.22 sI-saI

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