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Christoph Oberprieler The Systematics of Anthemis L. - Herbmedit.org

Christoph Oberprieler The Systematics of Anthemis L. - Herbmedit.org

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46 <strong>Oberprieler</strong>: <strong>Anthemis</strong> in N Africa<br />

<strong>The</strong> finding <strong>of</strong> n = 9 and 2n = 18 chromosomes for this variety <strong>of</strong> <strong>Anthemis</strong> secundiramea<br />

agrees with a report by Brullo & al. (1977, sub A. secundiramea subsp. intermedia)<br />

. Brullo (in Bartolo & al. 1979) later found two pairs <strong>of</strong> satellited chromosomes in<br />

plants from Lampedusa which he considered to represent a separate subspecies,<br />

A. secundiramea subsp. lopadusana, on account <strong>of</strong> its karyogram (12m + 2sm + 4st-sat),<br />

as he had found that A. secundiramea subsp. secundiramea lacked satellites. <strong>The</strong> Tunisian<br />

plant material studied here shows two pairs <strong>of</strong> satellited subtelocentric chromosomes<br />

(Table 8), exaetly as the morphologieally similar plants from Lampedusa. This supports<br />

the inc1usion <strong>of</strong> A. secundiramea subsp. lopadusana in A. secundiramea varo cossyrensis<br />

deseribed from Pantellaria.<br />

Table 8. Chromosomes 01 <strong>Anthemis</strong> secundiramea var. cossyrensis (OTU 13;<br />

n = 6).<br />

Chromo- Long arm Short arm Totallength<br />

some pair (~m) (~m) (~m)<br />

Relative length<br />

(%)<br />

Arm ratio<br />

Chromosome<br />

type<br />

3.38 2.76 6.14<br />

Il 2.98 2.36 5.34<br />

III 3.58 0.76 5.12<br />

IV 2.67 2.34 5.01<br />

V 3.29 1.67 4.96<br />

VI 3.24 0.59 4.61<br />

VII 2.51 2.04 4.55<br />

VIII 2.73 1.52 4.25<br />

IX 2.29 1.64 3.93<br />

14.0 1.22<br />

12.2 1.26<br />

11.7 4.71<br />

11.4 1.14<br />

11.3 1.97<br />

10.5 5.49<br />

10.4 1.23<br />

9.7 1.79<br />

9.0 1.39<br />

m<br />

m<br />

st-sat<br />

m<br />

sm<br />

st-sat<br />

m<br />

sm<br />

m<br />

<strong>Anthemis</strong> secundiramea varo urvilleana<br />

n=9<br />

-Me, Paradise Bay, 27 Feb 1994, Mevert & Seifert s.n. (Herb. <strong>Oberprieler</strong>).<br />

This taxon, endemie to Malta, had been studied eytologically by Brullo & al. (1997).<br />

<strong>The</strong> ehromosome number reported here is in agreement with their findings.<br />

<strong>Anthemis</strong> ubensis<br />

n=9<br />

- Tn, Beja, Djebel Goraa, 730 m, 3 May 1994, Vogt 12216 & <strong>Oberprieler</strong> 6521 (Fig. 5B).<br />

- Tn, Beja, Djebel Goraa, 960 m, 3 May 1994, Vogt 12291 & <strong>Oberprieler</strong>6596.<br />

- Tn, Beja, Djebel Goraa, 960 m, 3 May 1994, Vogt 12305 & <strong>Oberprieler</strong>6610.<br />

- Tn, El Kef, Djebel Dyr, 1000 m, 4 May 1994, Vogt 12352 & <strong>Oberprieler</strong> 6657.<br />

- Tn, El Kef, Table de Jughurta, 1200-1270 m, 5 May 1994, Vogt 12467 & <strong>Oberprieler</strong> 6772.<br />

2n= 19<br />

*-Tn, FOfl~t de Kesra, 18 May 1994, Vogt 13524 & <strong>Oberprieler</strong> 7829 (Fig. 5C).<br />

- Tn, Zaghouan, Djebel Zaghouan, 23 May 1994, Vogt 13824 & <strong>Oberprieler</strong> 8129.

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