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Explanation Of Gene Action As Related To Physiological

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_lOsomewhere<br />

nearly equally balanced by the humerous other processes which,<br />

themselves, would cause an opposite kind of curvilinearity.<br />

Instead of being surprised at dominance or eplstasls and at the many<br />

cases where herltabillty is rather low, I tend to be astonished that<br />

dominance and eplstasls are not more extreme and that herltabilltles as<br />

.._.-.----.---___.<br />

high as i0_o are often found. I suppose natural selection has required<br />

that to be true to a considerable extent, else the specieswould not have<br />

survived. That is, the Mendelian segregation and recombination would have<br />

produced an intolerably large proportion of individuals so defective in<br />

one way or another that they could not survive, if these various growth<br />

processes had combined with each other in extremely irregular ways.<br />

If a species is dependent for survival on having a certain highly _<br />

complex genetic combination, then it can survive only by becoming almost--_<br />

completely homozygous for most of the genes needed in that combination.<br />

This, I am sure, has happenedas concerns differences between major<br />

groupings of the plant or animal world, such as the differences between<br />

<strong>Gene</strong>ra, Families and Orders. However, breeders are concerned with<br />

differences such as occur within species and even within breeds. If<br />

these differences are many and if each of them is frequent, then I think<br />

the combinations of their various effects must come somewhere near to<br />

being additive, else the species or breed would have difficulty surviving.<br />

Speculations like these seem to serve no useful purpose, except<br />

that they may help explain how the complicated processes which the genes<br />

initiate or guide can still yield viable individuals most of the time<br />

andhow it happens that the end product can be nearly enough in proportion<br />

to the differences in genes transmitted that we often find<br />

heritabilities of i0 to 20_ or even higher.

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