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Plant Communities of the Midwest - NatureServe

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Ecological groups allow us to think at bigger scales and consider ecological factors that may not<br />

be readily apparent at <strong>the</strong> association level. As Grimm (1983, 1984) demonstrated in his studies <strong>of</strong> <strong>the</strong><br />

prairie-forest border in Minnesota (Figure 8), ecological groups can extend our understanding <strong>of</strong><br />

vegetation pattern beyond site and disturbance factors to historical and climatic factors. By working at<br />

<strong>the</strong> scale <strong>of</strong> “<strong>Midwest</strong>ern Tallgrass Prairies” and “<strong>Midwest</strong>ern Oak Woodlands” we can incorporate<br />

ecological studies that occur over much larger spatial and temporal scales than do many association<br />

studies, and <strong>the</strong>reby we gain a perspective on <strong>the</strong> dynamics <strong>of</strong> individual associations within those groups.<br />

So, this approach allows us to address <strong>the</strong> variety <strong>of</strong> ecological processes and abiotic factors affecting<br />

midwestern plant communities, from <strong>the</strong> moisture/substrate gradient and <strong>the</strong> prairie-forest<br />

physiognomic/fire gradient to <strong>the</strong> interactions <strong>of</strong> both <strong>of</strong> <strong>the</strong>se gradients with climate.<br />

V. PLANT COMMUNITIES OF THE MIDWEST<br />

At this time, <strong>the</strong>re are 588 associations (including 21 <strong>of</strong> uncertain status) and 258 alliances in <strong>the</strong><br />

<strong>Midwest</strong> (Table 7). The total number <strong>of</strong> associations in various states <strong>of</strong> <strong>the</strong> <strong>Midwest</strong> ranged from 74 in<br />

Kansas and Nebraska to 185 in Michigan. By comparison, in 1991, <strong>the</strong> number <strong>of</strong> types catalogued in<br />

natural community classifications from state to state varied from 25 in North Dakota to 191 in Minnesota.<br />

Thus, <strong>the</strong> USNVC has helped to standardize <strong>the</strong> scale <strong>of</strong> community type definition across <strong>the</strong> region.<br />

The midwestern associations can also be aggregated by ecological groups. The number <strong>of</strong> units<br />

by ecological group level is summarized in Table 8. The total number <strong>of</strong> ecological groups in <strong>the</strong><br />

<strong>Midwest</strong> at Level 4 is 152, ranging from 38 in Kansas to 63 in Illinois and Michigan. The complete list <strong>of</strong><br />

associations in order <strong>of</strong> Ecological Group is presented in Table 9. Descriptions <strong>of</strong> each association are<br />

provided in <strong>the</strong> Appendix. A summary <strong>of</strong> <strong>the</strong> fields, as laid out in each association description, is<br />

presented in Box 1, followed by an example description.<br />

Given <strong>the</strong> qualitative but extensive nature <strong>of</strong> <strong>the</strong> work completed to date, it is difficult to estimate<br />

<strong>the</strong> degree to which associations may be added or subtracted in <strong>the</strong> <strong>Midwest</strong>. All twelve states in <strong>the</strong><br />

<strong>Midwest</strong> have helped review <strong>the</strong> associations presented here, and every attempt has been made to include<br />

only vegetation types that meet <strong>the</strong> association concept. Many associations, however, are described only<br />

briefly, and fur<strong>the</strong>r research is needed to understand <strong>the</strong>se types.<br />

There is an urgent need for more extensive, quantitative-based surveys <strong>of</strong> many <strong>of</strong> <strong>the</strong>se types.<br />

Some states, such as Nebraska and Minnesota, are engaged in detailed, plot-based surveys across <strong>the</strong><br />

entire state. In Wisconsin, <strong>the</strong> entire <strong>Plant</strong> Ecology Laboratory vegetation data set that was developed by<br />

Curtis and his students has been computerized (Umbanhower 1990). Analysis <strong>of</strong> <strong>the</strong>se data at <strong>the</strong> local or<br />

state level will, in turn, assist with more rigorous definitions <strong>of</strong> <strong>the</strong> associations. O<strong>the</strong>r states are<br />

collecting data on a project-by-project basis. Ultimately, a national plots database, which allows analyses<br />

to be conducted across jurisdictional lines, will provide <strong>the</strong> quantitative data that will augment and inform<br />

<strong>the</strong> qualitative global descriptions provided here. A collaborative project to develop such a plots database<br />

is now underway at <strong>the</strong> National Center for Ecological Analysis and Syn<strong>the</strong>sis (Peet et al. 2000).<br />

PLANT COMMUNITIES OF THE MIDWEST – 2001<br />

31

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