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Pandit Govind Ballabh Pant Memorial Lecture: II

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commonly known as the pitcher plant, is endimic to Khasi Hills. The genus<br />

Nepanthes has 67 species out of which one species each is 64 species are found in<br />

South east Asia, Malaysia, North Queensland and New Calendonia. The foregoing<br />

examples are pointers to the Gondwanaland influence.<br />

The Brahmaputra valley contains extensive areas with natural vegetationswamps<br />

and grassland- with large herbivours fauna typical of alluvial grasslands.<br />

Examples are rhinoceros, buffalo, swamp deer, hog deer, pygmy hog and hespid<br />

hare (Rodgers and Panwar, 1988). Being tropical/subtropical evergreen forest there<br />

is high degree of species richness and high degree evolutionary activity resulting in<br />

pockets of high degree of endemism. Added to it is the fact that with the rise of<br />

Himalaya, there were many micro-climates due to altitudinal and latitudinal factors<br />

leading to large number of small, rather isolated, pockets. Such a situation not only<br />

isolated geographically many widespread species but also presented newer sites for<br />

colonization.<br />

That this has been and still is a region of high evolutionary activity is clear from<br />

the cytogeographic studies on genera like Rhododendron, Camelia, Magnolia,<br />

Buddleia etc. of Janaki Ammal (1950, 1952 a-b, 1953, 1954, Janaki Ammal and<br />

Saunders 1952). The north Eastern Himalaya and North Eastern States together<br />

with South Eastern Provinces of China (like Yunnan and Szechwan) are very active<br />

centres. There are ecologically and phytogeographically highly diversified and<br />

floristically among the richest in the world. Here diploids (2x) geographic ranges of<br />

tetraploids (4x), hexaploids, (6x) octoploids, (8x) and do-decaploids (12x). Thus the<br />

higher the level of ploidy, narrower is the geograpic range. The higher ploids are<br />

essentially endemics confined to narrow region. These are some very meaningful<br />

data from chromosome geography. During colonization of the new habitats new<br />

polypoid races and species of the foregoing genera were produced which seem to<br />

have had a high survival value (Fig. 5 and 6). The highest level of polyploidy in these<br />

genera is seen in the species growing in this region.<br />

Fig. 5- Distribution of polyploides of Rhododenderons in North Eastern India Source:<br />

Janaki Ammal 1954<br />

Fig. 6- Distribution of polyploids of Buddleia in North Eastern India Source: Janaki<br />

Ammal 1954.<br />

There are few temperate species of the Assam Himalaya found in the Nilgiris,<br />

which is another floristrically rich area. The species are: Ternstroemia japonica,<br />

Rhododedron arboreum, Hypericum hookerianum, Thalictrum javantium,<br />

Cotoneaster buxifolia, Parnassia wightiana, Lonicera ligustrina, Gaultheria<br />

fragrantissima, Symplocos laurinana etc. (Nayar, 1977). Some botanists believe that<br />

the presence of such floristic element in Nilgiris is an indication of their being<br />

Pleistocene relicts. According to this view, during Pleistocene glaciation, temperate<br />

flora and fauna moved south. On retreat of the glaciation, temperate relicts were left<br />

at higher altitudes of southern mountains, and continuous distribution between<br />

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