Demographic characteristics of northern spotted owls in the Eastern ...

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Table 3. Reproductive indices of Spotted owls on the Cle Elum Study Area, Okanogan-Wenatchee National Forest, Washington, 1989-2012. 24 Fecundity of female owls 1 Mean brood size of successful nests 2 Year n mean se n mean se 1989 11 0.77 0.12 9 1.89 0.11 1990 31 0.66 0.08 23 1.78 0.09 1991 47 0.45 0.07 25 1.68 0.11 1992 49 0.81 0.06 42 1.88 0.09 1993 46 0.11 0.04 6 1.67 0.21 1994 46 0.75 0.08 32 2.16 0.11 1995 41 0.40 0.07 21 1.57 0.11 1996 35 0.76 0.07 30 1.77 0.09 1997 33 0.06 0.04 3 1.33 0.33 1998 36 0.72 0.09 27 1.93 0.13 1999 27 0.30 0.08 10 1.60 0.16 2000 31 0.52 0.08 20 1.60 0.11 2001 27 0.48 0.09 16 1.63 0.13 2002 19 0.37 0.11 9 1.78 0.15 2003 22 0.66 0.10 16 1.81 0.14 2004 23 0.48 0.10 13 1.69 0.13 2005 20 0.50 0.11 11 1.82 0.12 2006 15 0.57 0.12 10 1.70 0.15 2007 17 0.44 0.11 9 1.67 0.17 2008 10 0.45 0.16 5 1.80 0.20 2009 10 0.55 0.16 6 1.83 0.17 2010 8 0.75 0.16 6 2.00 0.00 2011 10 0.55 0.16 6 1.83 0.17 2012 6 0.58 0.16 5 1.40 0.24 Total 620 0.52 0.02 360 1.79 0.03 1 Sample size (n) includes those females checked for reproductive status by August 31. Fecundity is the number of females fledged per female, assuming a 50:50 sex ratio. 2 Mean brood size of nests that produced at least 1 young, and where the number of fledged young was determined by August 31
Table 4. Reproductive indices of Spotted owls on the Cle Elum Study Area, Okanogan-Wenatchee National Forest, Washington, 1989-2012. proportion (π) of females that nested 1 proportion (π) of nesting females that fledged young 2 proportion (π ) of all females that fledged young 3 year n π 95% CI 4 n π 95% CI 4 n π 95% CI 4 1989 8 0.88 0.47 - 1.00 7 1.00 0.59 - 1.00 11 0.82 0.48 - 0.98 1990 20 0.90 0.68 - 0.99 17 0.94 0.71 - 1.00 31 0.74 0.55 - 0.88 1991 34 0.65 0.46 - 0.80 22 0.82 0.60 - 0.95 47 0.53 0.38 - 0.68 1992 47 1.00 0.92 - 1.00 43 0.88 0.75 - 0.96 49 0.86 0.73 - 0.94 1993 41 0.17 0.07 - 0.32 7 0.86 0.42 - 1.00 46 0.13 0.05 - 0.26 1994 40 0.93 0.80 - 0.98 37 0.78 0.62 - 0.90 46 0.70 0.54 - 0.82 1995 35 0.66 0.48 - 0.81 23 0.87 0.66 - 0.97 41 0.51 0.35 - 0.67 1996 34 0.97 0.85 - 1.00 33 0.91 0.76 - 0.98 35 0.86 0.70 - 0.95 1997 27 0.15 0.04 - 0.34 4 0.75 0.19 - 0.99 33 0.09 0.02 - 0.24 1998 34 0.91 0.76 - 0.98 31 0.84 0.66 - 0.95 36 0.75 0.58 - 0.88 1999 20 0.60 0.36 - 0.81 12 0.75 0.43 - 0.95 27 0.37 0.19 - 0.58 2000 27 0.81 0.62 - 0.94 22 0.91 0.71 - 0.99 31 0.65 0.45 - 0.81 2001 23 0.74 0.52 - 0.90 17 0.82 0.57 - 0.96 27 0.59 0.39 - 0.78 2002 17 0.59 0.33 - 0.82 10 0.80 0.44 - 0.97 19 0.47 0.24 - 0.71 2003 20 0.95 0.75 - 1.00 18 0.78 0.52 - 0.94 22 0.73 0.50 - 0.89 2004 20 0.75 0.51 - 0.91 15 0.80 0.52 - 0.96 23 0.57 0.34 - 0.77 2005 19 0.58 0.34 - 0.80 11 0.91 0.59 - 1.00 20 0.55 0.32 - 0.77 2006 13 0.92 0.64 - 1.00 12 0.67 0.35 - 0.90 15 0.67 0.38 - 0.88 2007 16 0.63 0.35 - 0.84 10 0.90 0.55 - 1.00 17 0.53 0.28 - 0.77 2008 6 0.83 0.36 - 1.00 5 0.80 0.28 - 0.99 10 0.50 0.19 - 0.81 2009 6 0.83 0.36 - 1.00 5 1.00 0.48 - 1.00 10 0.60 0.26 0.88 2010 5 1.00 0.48 - 1.00 5 0.80 0.28 - 0.99 8 0.75 0.35 - 0.97 2011 5 0.80 0.28 - 0.99 4 0.75 0.19 - 0.99 10 0.60 0.26 - 0.88 2012 4 1.00 0.40 - 1.00 4 1.0 0.40 - 1.00 6 0.83 0.36 - 0.99 Total 521 0.73 0.69 - 0.77 374 0.85 0.81 - 0.88 620 0.58 0.54 - 0.62 1 Sample size (n) includes females that were checked for nesting status before June 15 2 Sample size (n) includes nesting females that were checked for reproductive status by August 31 3 Sample size (n) includes all females that were checked for reproductive status by August 31. The sample size for this index is commonly larger than other indices because we often cannot make the required visits to determine nesting status before the June15 cutoff due to limited access or low response rates for non-nesting females. 4 Exact confidence limits for the binomial proportion using the F distribution, Collett (1991) 25
Page 1 and 2: 1 WILDLIFE ECOLOGY TEAM WILDLIFE HA
Page 3 and 4: Spotted Owl calls at each establish
Page 5 and 6: Figure 4. Number of vacant territor
Page 7 and 8: 7 Density Study Area The DSA survey
Page 9 and 10: 9 Figure 7. Percent of Spotted Owl
Page 11 and 12: 11 Reproductive Rates All of the 4
Page 13 and 14: 13 Figure 10. Hypothesized annual f
Page 15 and 16: 15 Figure 12. Looking NE toward the
Page 17 and 18: 17 Figure 12. Northern Spotted Owl
Page 19 and 20: Franklin, A. B., Burnham, K. P., Wh
Page 21 and 22: 21 Publications and presentations i
Page 23: 23 Table 2. Survey effort for the C

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