122POACEAE5b. Culm sheaths green, without spots, ± glossy but not oily ............................................................................ 2. P. oleosus4b. Culm sheaths not glossy, ± pruinose, often with setae or marginal cilia.6a. Culm sheath ligules usually truncate, 1–2 mm.7a. Culms initially glabrous but pruinose, culm sheaths glabrous or with sparse setae ............................. 3. P. amarus7b. Culms initially pubescent; culm sheaths setose; leaf blades asymmetrical, especially at base.8a. Culm sheath blades deflexed or revolute; leaves 3 or 4(–8) per ultimate branch ................... 4. P. intermedius8b. Culm sheath blades erect; leaves 1 or 2 per ultimate branch ...................................................... 5. P. truncatus6b. Culm sheath ligules ± arcuate, 3–8 mm.9a. Culm sheaths thickly papery or thinly leathery, ligules ca. 3 mm ................................................ 6. P. altiligulatus9b. Culm sheaths leathery, setose, ligules 5–8 mm.10a. Culm sheaths without spots, thickly pruinose and with sparse, brown setae, marginsdistally pink, ligules ca. 5 mm; leaf sheath auricles truncate to vaulted, 1–2.5 mm .............. 7. P. incarnatus10b. Culm sheaths with brown spots and verrucate setae, margins not pink, ligule ca. 8 mm;leaf sheath auricles triangular, 3–4 mm ............................................................................. 8. P. maculosoides3b. Culm sheaths with well-developed auricles and oral setae.11a. Culm sheaths mainly glabrous, basally piliferous ..................................................................................... 9. P. juxianensis11b. Culm sheaths with setae or marginal cilia.12a. Internodes with small cavity or subsolid.13a. Internodes subsolid, initially strigose and finely ridged; leaf sheath auricles absent; oralsetae absent to 3, erect, to 5 mm .................................................................................................. 10. P. solidus13b. Internodes with small cavity, initially glabrous, inconspicuously ridged; leaf sheathauricles ovate to elliptical, oral setae radiating, to 13 mm ............................................... 1<strong>1.</strong> P. hsienchuensis12b. Internodes with large cavity.14a. Culm sheath blades intensely crinkled; internodes ca. 35 cm ................................................... 12. P. rugatus14b. Culm sheath blades smooth; internodes to 33 cm.15a. Internodes to 18.5 cm, culm wall ca. 3 mm thick ........................................................ 13. P. yixingensis15b. Internodes to 33 cm, culm wall 7–8 mm thick.16a. Culm sheath ligule truncate, ca. 1 mm, auricles subcircular or falcate,small, oral setae 3–5 mm ............................................................................... 14. P. wuyishanensis16b. Culm sheath ligule arcuate, ca. 10 mm, auricles oval to elliptical, oralsetae ca. 10 mm ............................................................................................... 15. P. sanmingensis<strong>1.</strong> Pleioblastus maculatus (McClure) C. D. Chu & C. S. Chao,Acta Phytotax. Sin. 18: 3<strong>1.</strong> 1980.斑 苦 竹 ban ku zhuSinobambusa maculata McClure, Lingnan Univ. Sci. Bull.9: 64. 1940; Arundinaria chinensis C. S. Chao & G. Y. Yang;A. kwangsiensis (W. Y. Hsiung & C. S. Chao) C. S. Chao & G.Y. Yang; A. maculata (McClure) C. D. Chu & C. S. Chao ex K.M. Lan (1981), not Hackel (1903); Pleioblastus kwangsiensisW. Y. Hsiung & C. S. Chao; P. longispiculatus B. M. Yang; P.maculatus var. longitubus Li & Wu.Culms 3–8 m, <strong>1.</strong>5–4 cm in diam., initially green, denselyglaucous; internodes subcylindrical; nodes prominent, brown,yellow-green when old, densely hairy, with prominent setosesheath scar. Culm sheaths deciduous, brown-red, oily, glossy,slightly purple, with unevenly scattered brown spots, basallybrown setose, margins without cilia; auricles absent or veryreduced, brown, dotlike or ovate; oral setae few, erect orcurved; ligule often truncate, dark brown-red, entire; bladependulous, linear-lanceolate, slightly hairy, scabrous, remotelyserrulate or nearly entire, revolute. Leaves 3–5 per ultimatebranch, soon deciduous; sheath margin sparsely pubescent;auricles and oral setae absent; ligule truncate, 1–2 mm, hairy,margin ciliolate; pseudopetiole ca. 4 mm; blade lanceolate, 8.8–18.5 × <strong>1.</strong>3–2.9 cm, base cuneate, apex elongate. Inflorescencepaniculate; florets 8–15 per spikelet. Glumes 2; lemma lustrous;palea keels ciliolate; lodicules 3, subequal in size, ciliolate atapex. Ovary bottle-shaped, ca. 8 mm; styles ca. <strong>1.</strong>5 mm; stigmas3, plumose. Caryopsis ellipsoid. New shoots early May–early Jun.● Often growing in dense forests, also planted as an ornamental.Fujian, Guangdong, Guangxi, Guizhou, Jiangsu, Jiangxi, Sichuan, Yunnan;cultivated northward to S Shaanxi.The shoots are edible, but must be treated first. The culms areused for light duties; they are fragile and not suitable for weaving.2. Pleioblastus oleosus T. H. Wen, J. Bamboo Res. 1(1): 24.1982.油 苦 竹 you ku zhuAcidosasa lentiginosa W. T. Lin & Z. J. Feng; Arundinariaoleosa (T. H. Wen) Demoly; A. chinensis C. S. Chao &G. Y. Yang; Pleioblastus longiinternodius B. M. Yang; P. longispiculatusB. M. Yang; Polyanthus longispiculatus (B. M. Yang)C. H. Hu.Culms diffuse, 3–5 m, 1–3 cm in diam.; internodes initiallygrass-green, yellow when old, cylindrical, 18–20(–26)cm, grooved above branches, glossy, glabrous; wall ca. 3 mmthick; nodes prominent, deciduously brown setose. Culmsheaths light green, slightly lustrous, base light brown setose,otherwise glabrous; ligules truncate or slightly concave, 1–2mm, margin ciliolate; blade erect or reflexed, green, lanceolate.
POACEAE 123Leaves 3 or 4 per ultimate branch; sheath glabrous; auricles andoral setae often absent, rarely 2, short; ligules rounded ortruncate, ca. 2 mm, slightly hairy, apex not uniform; pseudopetiole2–5 mm; blade linear-lanceolate, 12–20 × <strong>1.</strong>3–2.2 cm,slightly hairy, secondary veins 5–7-paired, base broadly cuneate,margin serrulate, apex caudate. Inflorescence paniculate,lateral; florets 11–13 per spikelet. Glumes 2–4, apex rounded,beaked; lemma subglabrous, apex acute; palea about as long aslemma, keels ciliolate, apex acuminate; lodicules 3, thick, ±rhomboid, margin ciliolate. Ovary cylindrical; stigmas 2 or 3.● Mountain slopes; ca. 800 m. Fujian, Jiangxi, Yunnan, Zhejiang.A plant cultivated in the West under the names Brachystachyumdensiflorum as well as P. oleosus is not in fact this species but an Oligostachyum.Pleioblastus oleosus is possibly a synonym of P. maculatus.The shoots are edible, and the culms are often used for weaving.3. Pleioblastus amarus (Keng) P. C. Keng, Techn. Bull. Natl.Forest. Res. Bur. 8: 14. 1948.苦 竹 ku zhuCulms 3–5 m, <strong>1.</strong>5–2 cm in diam.; internodes green orgreen-purple, spotted, yellow-green when old, cylindrical, 27–29(–38) cm, basally weakly flattened above branches, glabrousor hairy; wall ca. 6 mm thick; nodes prominent; intranode ca. 6mm. Leafy branchlets sometimes pendulous. Culm sheathsgreen, densely glaucous, abaxially glabrous or hairy, densely soat base, margins ciliate and apically orange or withered; auriclesobscure; ligule truncate, 1–2 mm, densely glaucous, ciliolate;blade narrowly lanceolate, margins serrulate. Leaves 3 or 4 perultimate branch; sheath straw-colored, glabrous; auricles andoral setae absent; ligule purple-red, ca. 2 mm; pseudopetiole ca.2 mm; blade elliptic-lanceolate, 4–20 × 2–2.9 cm, secondaryveins 4–8-paired, base cuneate, margin serrulate, apex acuminate.Inflorescence racemose or paniculate. Spikelets 3–6, 4–7cm, glaucous, pedicels hairy; florets 8–13; rachilla internodes4–5 mm. Glumes 3–5, enlarged upward; lemma ovate-lanceolate;palea longer than lemma, rarely equal in length, ciliate,glaucous and slightly hairy between keels, apex acute. Lodicules3, ovate or obovate, posterior narrow, apical marginsciliolate. Anthers light yellow. Ovary narrow, glabrous; stylesshort; stigmas 3. New shoots Apr–Jun, fl. Apr–May.● Plains to low hills, frequently cultivated. Anhui, Fujian, Guizhou,Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang.1a. Leafy branchlets pendulous .............. 3b. var. pendulifolius1b. Leafy branchlets not obviously pendulous.2a. Culm sheaths glabrous or with deciduouswhite hairs at base ............................. 3c. var. tubatus2b. Culm sheaths ± hairy, brown setose atbase.3a. Culms green-purple, purplespotted, densely hairy .... 3d. var. hangzhouensis3b. Culms initially green, yellow-greenwhen old, glaucous, powderyspotted,glabrous ....................... 3a. var. amarus3a. Pleioblastus amarus var. amarus苦 竹 ( 原 变 种 ) ku zhu (yuan bian zhong)Arundinaria amara Keng, Sinensia 6(2): 148. 1935; A.varia Keng; Indocalamus varius (Keng) P. C. Keng; Pleioblastusbrevinodus W. T. Lin & Z. J. Feng; P. longqishanensisN. X. Zhao & Z. Yu Li; P. amarus f. huangshanensis C. L.Huang; P. subrectangularis T. P. Yi & H. Long; P. varius(Keng) P. C. Keng; P. yingdeensis W. T. Lin & Z. M. Wu.Culm sheaths ± hairy, brown setose at base. Culms initiallygreenish, glabrous, glaucous, yellow-green when old, powdery-spotted.Branchlets not obviously pendulous. New shootsJun, fl. Apr–May.● Anhui, Fujian, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan,Yunnan, Zhejiang.The shoots are bitter and inedible. The culms are used for wovenbaskets and containers, umbrella handles, and furniture.3b. Pleioblastus amarus var. pendulifolius S. Y. Chen, ActaPhytotax. Sin. 21: 413. 1983.垂 枝 苦 竹 chui zhi ku zhuLeafy branchlets pendulous. Culm sheaths not glaucous;ligule truncate, slightly concave. New shoots mid–May to earlyJun.● Low hills, slopes. Zhejiang.This variety is cultivated as an ornamental for its pendulous habit.3c. Pleioblastus amarus var. tubatus T. H. Wen, Bull. Bot.Res., Harbin 3(1): 93. 1983.胖 苦 竹 pang ku zhuCulm sheaths green, glossy, hard, glabrous or nearly so atbase, acute at apex, amplexicaul at middle part; auricles oblong,small, transversely extended at both sides of sheath apex, marginuniformly rough ciliate, cilia erect and short; blade roundedat apex.● Low hills, plains. Zhejiang.3d. Pleioblastus amarus var. hangzhouensis S. L. Chen & S.Y. Chen, Acta Phytotax. Sin. 21: 408. 1983.杭 州 苦 竹 hang zhou ku zhuInternodes 28–32(–38) cm; culms green-purple, purplespotted, roughly hairy, hairs dense and inverse. Culm sheathpurple-green, glossy, not glaucous; auricles absent; blade linear-lanceolate.New shoots late Apr–May.● Lowlands, small hills. Zhejiang.The plants are used for fencing and supporting vegetables and aregrown as ornamentals. The culms are not suitable for weaving.4. Pleioblastus intermedius S. Y. Chen, Acta Phytotax. Sin.21: 408. 1983.华 丝 竹 hua si zhuCulms 3–4 m, 1–2 cm in diam.; internodes cylindrical butslightly concave at base above branches, 21–22 cm, initiallygreen and densely glaucous, becoming dark green or yellowgreenwith persistent dark powder, deciduously setose; wall ca.6 mm thick, pith lamellate; nodes slightly prominent. Culmsheaths gradually deciduous, green, about as long as internodes
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POACEAE 717b. Lemma not indurated,
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POACEAE 919a. Palea of uppermost or
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POACEAE 1111a. Culm sheaths glabrou
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POACEAE 134. Bambusa funghomii McCl
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POACEAE 15Culms 8-12 m, 4-6 cm in d
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POACEAE 17nate; florets 4-8; rachil
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POACEAE 19base slightly rounded, ne
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POACEAE 21Branches many, clustered,
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POACEAE 23times laciniate, shortly
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POACEAE 25longer than sheath, abaxi
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POACEAE 27narrowly ovate-triangular
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POACEAE 29Branches many, clustered,
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POACEAE 31Lindley; B. glaucescens (
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POACEAE 33Culms 6-10 m, 3-6 cm in d
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POACEAE 35deciduous, one side smoot
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POACEAE 3776b. Bambusa beecheyana v
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POACEAE 392. Thyrsostachys siamensi
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POACEAE 41late, 12.5-25 × 1.2-2 cm
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POACEAE 4313. Dendrocalamus brandis
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POACEAE 45Yunnan [Myanmar; cultivat
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POACEAE 474b. Leaf blade 3.5-6 cm w
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POACEAE 49● Forested hills; 1000-
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POACEAE 51Arborescent bamboos, some
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POACEAE 535b. Schizostachyum dumeto
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POACEAE 553. Cephalostachyum pergra
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POACEAE 57which name M. bambusoides
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POACEAE 5914b. Culm internodes glab
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POACEAE 6157b. Leaf sheaths glabrou
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POACEAE 631a. Culms 0.5-0.6 m, to 0
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POACEAE 65margins initially brown s
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POACEAE 67The culms are split for w
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POACEAE 69Akad. Wiss. Wien, Math.-N
- Page 65 and 66: POACEAE 71dery, glabrous, margins d
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- Page 75 and 76: POACEAE 81obscure or level; sheath
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- Page 79 and 80: POACEAE 85nent. Branches 6-11 per n
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- Page 95 and 96: POACEAE 101auricles projecting, his
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- Page 127 and 128: POACEAE 133Shrubby bamboos. Rhizome
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- Page 135 and 136: POACEAE 141encircling culm, shorter
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- Page 147 and 148: POACEAE 1539b. Culm internodes 15-3
- Page 149 and 150: POACEAE 155internodes, papery, init
- Page 151 and 152: POACEAE 157Arundinaria armata Gambl
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POACEAE 173linear. Leaves 2-4 per u
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POACEAE 17530. Phyllostachys nigra
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POACEAE 177● Planted. Fujian, Jia
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POACEAE 179veined, apex acuminate w