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Vol. 127, No. 1 August, 1964
THE
BIOLOGICAL BULLETIN
PUBLISHED BY THE MARINE BIOLOGICAL LABORATORY
THE MARINE BIOLOGICAL LABORATORY
SIXTY-SIXTH REPORT, FOR THE YEAR 1963—SEVENTY-SIXTH YEAR
I. TRUSTEESAND EXECUTIVECOMMITTEE(AS OFAUGUST 16, 1963) 1
STANDING COMMITTEES
II. ACT OF INCORPORATION 4
III. BYLAWSOF THE CORPORATION 4
IV. REPORT OF THE DIRECTOR 6
Addenda:
1. Memorials 8
2. The Staff 9
3. Investigators, Lalor, Lillie and Grass Fellows, and Students 14
4. Fellowships and Scholarships 27
5. Training Programs 28
6. Tabular View of Attendance, 1959—1963 29
7. Institutions Represented 29
8. Evening Lectures 31
9. Evening Seminars 32
10. Members of the Corporation 33
V. REPORT OF THE LIBRARIAN 55
VI. REPORT OF THE TREASURER 56
I. TRUSTEES
GERARD SWOPE, JR., Chairman of the Board of Trustees, 570 Lexington Ave., New
York 22, New York.
ARTHUR K. PARPART, President of the Corporation, Princeton University.
JAMES H. WICKERSHAM, Treasurer, 791 Park Avenue, New York 21, New York.
PHILIP B. ARMSTRONG, Director, State University of New York, College of Medicine
at Syracuse.
C. LLOYD CLAFF, Clerk of the Corporation, Randolph, Mass.
Copyright ©1964, by the Marine Biological Laboratory
This One
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E6H1-F1Z-C8BL

2 MARINE BIOLOGICAL LABORATORY
EMERITI
WILLIAM R. AMBERSON, Marine Biological Laboratory.
C. LALORBURDICK,The Lalor Foundation.
W. C. CURTIS,504 West Mount Avenue, Columbia, Missouri.
PAUL S. GALTSOFF, Woods Hole, Massachusetts.
E. B. HARVEY,Woods Hole, Massachusetts.
CHARLES W. METZ, Woods Hole, Massachusetts.
M. H. JAcoBs, University of Pennsylvania.
F. P. KNOWLTON, Syracuse University.
W. J. V. OSTERHOUT, The Rockefeller Institute.
S
CHARLES PACKARD, Woods Hole, Massachusetts.
A. C. REDFIELD,Woods Hole, Massachusetts.
A. H. STURTEVANT, California Institute of Technology.
LESTER G. BARTH, Columbia University.
TO SERVE UNTIL 1967
JOHN B. BUCK, National Institutes of Health.
AURIN M. CHASE, Princeton University.
SEYMOUR S. COHEN, University of Pennsylvania, School of Medicine.
TERU HAYASHI, Columbia University.
LEWIS KLEINHOLZ, Reed College.
ALBERT I. LANSING, University of Pittsburgh.
S. MERYLROSE,Tulane University.
TO SERVE UNTIL 1966
F. A. BROWN, JR., Northwestern University.
F'. D. CARLSON, The Johns Hopkins University.
SEARS CROWELL, Indiana University.
W. D. MCELROY, The Johns Hopkins University.
C. LADD PROSSER, University of Illinois.
E. A. SCHARRER, Albert Einstein College of Medicine.
ALBERT SZENT-GYöRGYI, Marine Biological Laboratory.
WM. RANDOLPH TAYLOR, University of Michigan.
TO SERVE UNTIL 1965
ERIC G. BALL, Harvard Medical School.
D. W. BRONK,The Rockefeller Institute.
MAC V. EDDS, JR., Brown University.
RUDOLF KEMP'roN, Vassar College.
I. M. KLOTZ,Northwestern University.
ARNOLD LAZAROW, University of Minnesota Medical School.
MORRIS ROCKSTEIN, Medical Research Building, University of Miami.
GEORGEWALD,Harvard University, Biological Laboratories.
TO SERVE UNTIL 1964
ARTHUR L. COLWIN, Queens College.
E. G. BUTLER, Princeton University.
K. S. COLE,The National Institutes of Health.
S. KUFFLER, Harvard Medical School.
C. B. Mx'rz, Institute for Space Biosciences, Florida State University.
ROBERTS RUGH, College of Physicians and Surgeons.
G. T. SCOTT,Oberlin College.
E. ZwILLING,Brandeis University.

TRUSTEES
STANDING COMMITTEES
EXECUTIVE COMMITTEE OF THE BOARD OF TRUSTEES
ARTHUR K. PARPART, ex officio, Chairman
GERARD SWOPE, JR., ex officio
JAMEs H. WICKERSHAM, ex officio
PHILIP B. ARMSTRONG, ex officio
TERU HAYASHI, 1966
M. A. LAUFFER,Chairman
SEYMOUR S. COHEN
KEITH PORTER
IRVING M. KLOTZ
ALBERT I. LANSING, Chairman
CLIFFORD HARDING
DAVID POTTER
RUDOLF KEMPTON, Chairman
W. J. ADELMAN
GEORGE SCOTT
TERU HAYASHI, Chairman
A. C. CLEMENT
BOSTWICK KETCHUM
EDGAR ZWILLING, Chairman
MAC V. EDDS, JR.
J. W.HASTINGS
H. H. SELIGER,Chairman
P. M. FAILLA
WALTER VINCENT
WILLIAM D. MCELROY, Chairman
MAC V. EDDS, JR.
MAC V. EDDS, JR., Chairman
ERIC G. BALL
WILLIAM D. MCELROY
THE LIBRARY COMMITTEE
THE APPARATUS COMMITTEE
WILLIAM D. MCELROY, 1966
ERIC G. BALL, 1965
SEARS CROWELL, 1965
MAC V. EDDS, JR., 1964
I. M. KLOTZ,1964
C. LAnDPROSSER
JAMES LASH
STANLEY WATSON
ARNOLD LAZAROW
WILLIAM D. MCELROY
L. I. REBHIJN
THE SUPPLY DEPARTMENT COMMITTEE
THE INSTRUCTION COMMITTEE
SEARS CROWELL
WALTER HERNDON
JOHN SIMMONS, JR.
ROBERT K. CRANE
ROGER 0. ECKERT
DEWITT STETTEN
THE BUILDINGS AND GROUNDS COMMITTEE
THE RADIATION COMMITTEE
J. W.GREEN
DANIEL GROSCH
MoRRIs ROCKSTEIN
R. L. GREIF
C. C. SPEIDEL
WALTER WILSON
THE RESEARCH SPACE COMMITTEE
TERU HAYASHI
EDGAR ZWILLING
THE COMMITTEE FOR NOMINATION OF OFFICERS
SEARS CROWELL
IRVINGM. Ki.o'rz
TERU HAYASHI
3

4 MARINE BIOLOGICAL LABORATORY
No. 3170
II. ACT OF INCORPORATION
COMMONWEALTH OF MASSACHUSETTS
Be It Known, That whereas Aipheus Hyatt, William Sanford Stevens, William T.
Sedgwick, Edward G. Gardiner, Susan Minns, Charles Sedgwick Minot, Samuel Wells,
William G. Farlow, Anna D. Phillips, and B. H. Van Vieck have associated themselves
with the intention of forming a Corporation under the name of the Marine Biological
Laboratory, for the purpose of establishing and maintaining a laboratory or station for
scientific study and investigation, and a school for instruction in biology and natural his
tory, and have complied with the provisions of the statutes of this Commonwealth in such
case made and provided, as appears from the certificate of the President, Treasurer, and
Trustees of said Corporation, duly approved by the Commissioner of Corporations, and
recorded in this office;
Now, therefore, I, HENRY B. PIERCE, Secretary of the Commonwealth of Massachu
setts, do hereby certify that said A. Hyatt, W. S. Stevens, W. T. Sedgwick, E. G. Gardi
ner, S. Minns, C. S. Minot, S. Wells, W. G. Farlow, A. D. Phillips, and B. H. Van Vieck,
their associates and successors, are legally organized and established as, and are hereby
made, an existing Corporation, under the name of the MARINE BIOLOGICAL LAB
ORATORY, with the powers, rights, and privileges, and subject to the limitations, duties,
and restrictions, which by law appertain thereto.
Witness my official signature hereunto subscribed, and the seal of the Commonwealth
of Massachusetts hereunto affixed, this twentieth day of March, in the year of our Lord
One Thousand Eight Hundred and Eighty-Eight.
[SEAL] HENRY B. PIERCE,
Secretary of the Commonwealth.
III. BYLAWS OF THE CORPORATION OF THE MARINE
BIOLOGICAL LABORATORY
(Revised August 15, 1963)
I. The members of the Corporation shall consist of persons elected by the Board
of trustees.
IT. The officers of the Corporation shall consist of a Chairman of the Board of
Trustees, President, Director, Treasurer and Clerk.
III. The Annual Meeting of the members shall be held on the Friday following the
second Tuesday in August in each year at the Laboratory in Woods Hole, Massachusetts,
at 9:30 A.M., and at such meeting the members shall choose by ballot a Treasurer and a
Clerk to serve one year, and eight Trustees to serve four years, and shall transact such
other business as may properly come before the meeting. Special meetings of the
members may be called by the Trustees to be held at such time and place as may be
designated.
IV. Twenty-five members shall constitute a quorum at any meeting.
V. Any member in good standing may vote at any meeting, either in person or by
proxy duly executed.
VI. Inasmuch as the time and place of the Annual Meeting of members are fixed by
these bylaws, no notice of the Annual Meeting need be given. Notice of any special
meeting of members, however, shall be given by the Clerk by mailing notice of the time

BYLAWS OF THE CORPORATION S
and place and purpose of such meeting, at least fifteen ( 15) days before such meeting, to
each member at his or her address as shown on the records of the Corporation.
VII. The Annual Meeting of the Trustees shall be held promptly after the Annual
Meeting of the Corporation at the Laboratory in Woods Hole, Massachusetts. Special
meetings of the Trustees shall be called by the Chairman, the President, or by any seven
Trustees, to be held at such time and place as may be designated, and the Secretary
shall give notice thereof by written or printed notice, mailed to each Trustee at his
address as shown on the records of the Corporation, at least one ( 1) week before the
meeting. At such special meeting only matters stated in the notice shall be considered.
Seven Trustees of those eligible to vote shall constitute a quorum for the transaction
of business at any meeting.
VIII. There shall be three groups of Trustees:
(A) Thirty-two Trustees chosen by the Corporation, divided into four classes, each
to serve four years. After having served two consecutive terms of four years each
Trustees are ineligible for re-election until a year has elapsed. In addition, there shall
be two groups of Trustees as follows:
(B) Trustees ex officio, who shall be the Chairman, the President, the Director, the
Treasurer, and the Clerk.
(C) Trustees Emeriti, who shall be elected from present or former Trustees by the
Corporation. Any regular Trustee who has attained the age of seventy years shall
continue to serve as Trustee until the next Annual Meeting of the Corporation, where
upon his office as regular Trustee shall become vacant and be filled by election by the
Corporation and he shall become eligible for election as Trustee Emeritus for life. The
Trustees ex officio and Emeriti shall have all the rights of the Trustees, except that
Trustees Emeriti shall not have the right to vote.
The Trustees and officers shall hold their respective offices until their successors are
chosen and have qualified in their stead.
IX. The Trustees shall have the control and management of the affairs of the Cor
poration. They shall elect a Chairman of the Board of Trustees who shall be elected
annually and shall serve until his successor is selected and qualified and who shall also
preside at meetings of the Corporation. They shall elect a President of the Corporation
who shall also be the Vice Chairman of the Board of Trustees and Vice Chairman of
meetings of the Corporation, and who shall be elected for a term of five years and shall
serve until his successor is selected and qualified, except that such term shall not run
beyond the Annual Meeting of the Board following his 65th birthday; candidates over the
age of 65 shall be elected on an annual basis. They shall appoint a Director of the
Laboratory for a term not to exceed five years, provided the term shall not exceed
one year if the candidate has attained the age of 65 years prior to the date of the appoint
ment. They may choose such other officers and agents as they may think best. They
may fix the compensation and define the duties of all the officers and agents; and may
remove them, or any of them except those chosen by the members, at any time. They
may fill vacancies occurring in any manner in their own number or in any of the
offices. The Board of Trustees shall have the power to choose an Executive Committee
from their own number, and to delegate to such Committee such of their own powers
as they may deem expedient. They shall from time to time elect members to the
Corporation upon such terms and conditions as they may think best.
X. The Associates of the Marine Biological Laboratory shall be an unincorporated
group of persons (including associations and corporations) interested in the Laboratory

6 MARINE BIOLOGICAL LABORATORY
and shall be organized and operated under the general supervision and authority of
the Trustees.
XI. The consent of every Trustee shall be necessary to dissolution of the Marine
Biological Laboratory. In case of dissolution, the property shall be disposed of in such
manner and upon such terms as shall be determined by the affirmative vote of two-thirds
of the Board of Trustees.
XII. The account of the Treasurer shall be audited annually by a certified public
accountant.
XIII. These bylaws may be altered at any meeting of the Trustees, provided that the
notice of such meeting shall state that an alteration of the bylaws will be acted upon.
RESOLUTIONS ADOPTED AT TRUSTEE MEETING AUGUST 16,
1963—EXECUTIVE COMMITTEE
I. RESOLVED:
(A) The Executive Committee is hereby designated to consist of ten members as
follows: ez officio members who shall be the Chairman of the Board of Trustees,
President, Director and Treasurer; six additional Trustees, two of whom shall be
elected by the Board of Trustees each year, to serve for a three-year term.
(B) The President shall act as Chairman of the Executive Committee and the
Chairman of the Board of Trustees as Vice Chairman. A majority of the members
of the Executive Committee shall constitute a quorum and a majority of those present
at any properly held meeting shall determine its action. It shall meet at such times
and places and upon such notice and appoint such sub-committees as the Committee
shall determine.
(C) The Executive Committee shall have and may exercise all the powers of the
Board during the intervals between meetings of the Board of Trustees except those
powers specifically withheld from time to time by the Board or by Law.
(D) The Executive Committee shall keep appropriate minutes of its meetings, and
its actions shall be reported to the Board of Trustees.
IT. RESOLVED:
The elected members of the Executive Committee shall be constituted as a standing
“¿Committeefor the Nominations of Officers,†responsible for making nominations at
the annual meeting of the Corporation and of the Board of Trustees, for candidates to
fill each office as the respective terms of office expire. (Chairman of the Board, Presi
dent, Director, Treasurer, and Clerk.)
IV. REPORT OF THE DIRECTOR
To: THE TRUSTEES OF THE MARINE BIOLOGICAL LABORATORY
Gentlemen:
I submit herewith the report of the seventy-sixth session of the Marine Bio
logical Laboratory.

1. Bylaws
REPORT OF THE DIRECTOR 7
At a special meeting of the Board of Trustees (August 15, 1963) changes in
the bylaws were adopted creating a new position of Chairman of the Board to be
responsible principally for overall planning and executive action in the business,
financial, legal and public relations areas. The President is to be responsible for
overall planning and executive administration in the scientific area. The President
will normally be a scientist. The office of Vice-President was abolished as re
dundant. Terms for all of the officers are specified and the responsibilities of the
Trustees are spelled out. At a subsequent meeting the composition and powers
of the Executive Committee were established by resolution. These changes are
incorporated in the bylaws as published in this report.
2. Plant Development
It was noted in the report last year that the Laboratory received a grant from
the National Science Foundation for a boat to be constructed as a replacement of
the Nereis which has been in operation since 1925. This new boat will be com
pleted in September, 1964, is 40 feet long, has a 14-foot beam, and is powered by a
160-horse-power diesel engine.
An additional grant from the National Science Foundation provides funds for
extensive alterations in the library including a relocation of the offices and service
rooms of the librarians, refurbishing the stacks, an extension of the reading room,
improved lighting and new furniture. Part of this work will be completed during
the winter of 1963—64,the remainder the following winter. This same grant pro
vides funds for laboratory modifications which will concentrate the special apparatus
in one area and additional funds for a wharf in front of the main laboratory extend
ing out parallel to the salt water intake dock in the same location as the former
Cayadetta dock.
3.PlanningCommittee
At its Annual Meeting the Board of Trustees authorized the appointment by
the Executive Committee of an ad hoc committee on long-range planning, par
ticularly in reference to modern dormitory, housing and dining hall facilities. This
committee was set up by the Executive Committee and presented a report to the
Executive Committee at its mid-winter meeting. The report was accepted by the
Board of Trustees and the committee was instructed to explore ways of funding
its recommendations.
4. Laboratory Attendance and Turnover
Elsewhere in this report is a “¿Tabular View of Attendance, 1959—1963.†It
will be noted that there has been a gradual increase in the scientific population at
the Laboratory, resulting in considerable measure through an increased number
of research assistants.
The full-time research personnel in the Laboratory this past winter numbered
fifty-three. They were housed in twenty-five laboratories. There were twenty

8 MARINE BIOLOGICAL LABORATORY
five transient workers at the Laboratory, thirteen engaged in research activities,
twelve reading in the library.
It has sometimes been suggested that the Marine Biological Laboratory is a
closed organization with little turnover in research personnel. A survey of the
situation revealed that there will be 76 investigators and 20 library readers in
residence at the Laboratory during the summer of 1964 who were not present in
1963, and 155 investigators and 34 library readers in 1964 who were not present
in 1959.
The three graduate training programs—Gamete Physiology, Comparative
Physiology and Neuromuscular Physiology—Serve as instruments for personnel
change. There were 35 trainees in these three programs in 1963, only three of
them repeaters. The fellowship programs, Lalor and Grass, included ten fellows
in 1963, non of them repeaters.
Obviously there is a continuous flow of new personnel into the Laboratory,
insuring a wide opportunity in the use of its facilities.
5. Personnel Changes
Because of his appointment as a Career Professor, Dr. Clark P. Read had to
resign as head of the training program in Invertebrate Zoology but continues his
interest in this program as a Consultant. We are very fortunate in attracting
another very able invertebrate zoologist, Dr. W. D. Russell Hunter, to head up
this training program.
6 Deaths
We note with sorrow the deaths of two Trustees Emeriti, Dr. F. P. Knowlton
and Dr. W. J. V. Osterhout. Dr. Knowlton was a member of the staff of the
physiology course from 1909—1926, a Trustee from 1923—1945 and a Trustee
Emeritus from 1945 until his death in 1963. Dr. Osterhout was a Trustee from
1918—1940and then a Trustee Emeritus until his death on April 9, 1964.
1. MEMORIALS
LAWRASON RIGGS
by CHARLESPACKARD
Lawrason Riggs, former Treasurer and President of this Corporation, died in Janu
ary, 1963. Few of us realized the extraordinary ability and wide range of interests of
this quiet and unassuming man. He was a successful lawyer in New York City.
He found time to carry on the duties of Treasurer from 1924 to 1942, and to serve in the
same capacity for the Oceanographic Institution from 1930 to 1950. This he did with
the help of one part-time secretary. His ability as Treasurer is shown by the fact that
during the depression years of the 1930s the income from our Endowment Fund
remained fairly constant.
When Dr. Lillie died, Mr. Riggs was unanimously chosen to be President of the
Corporation. Those of us who were privileged to serve with him will always remember
his deep devotion to the Laboratory, his sympathetic understanding of its problems
and his wise advice.

REPORT OF THE DIRECTOR 9
In whatever capacity he served he was always asked to continue year after year.
He was Warden and Senior Warden of his New York Church for 45 years, an officer
of this Laboratory for 20 years and of the Oceanographic Institution for 20 years.
His interests outside the law and the Woods Hole laboratories were exceptional.
They were summed up by his friend, the Rector of the Church of the Ascension, in
these words : “¿His life was one adventure after another—climbing mountains, gazing at
the stars through his new telescope, walking through strange lands, picking up odd bits
of archaeological finds, sailing the waters from Woods Hole to everywhere. He was
at home among the classics.â€
Indeed, during his last stay in the hospital he had beside him a well-worn copy of
the Greek New Testament.
His deep affection for this Laboratory was shown when he announced his retirement
as President. As he spoke of his long association with the Laboratory his voice broke,
and in the middle of a sentence he said “¿I cannot speak further.â€
The Laboratory has lost a most valuable and devoted friend of many years.
REINHARD DOHRN
by ALBERT SZENT-GYöRGYI
It is befitting to remember here a man, Reinhard Dohrn, who was taken from our
rows by death this year, though he did not belong to the Woods Hole family of re
searchers. He belonged to the wider circle of marine biologists, among whom he played
a leading role, less through his own research than by providing for the research of
others, with his devotion to science, his ability of organization, and his very high
human qualities.
He was the Director of the Zoological Station in Naples, founded by his father,
Anton Dohrn. He saved his institution through the turmoil of political events. This he
did, less by political cunning than by his sincerity, honesty, modesty and idealism.
To me he was the embodiment of all the higher and lofty ideals that science and
humanities stand for.
He kept, always, the most friendly relations with Woods Hole in a spirit of brother
hood and not of competition. He parted at an old age, had a grand life full of struggle
and noble endeavor, as a man's life should be, so we can bear no grudge to fate, but,
all the same, his parting is a great loss to us all and to marine biology. We lost a great
and very good friend, and a grand human being.
2. THE STAFF
EMBRYOLOGY
I. INSTRUCTORs
JAMES D. EBERT, Director, Department of Embryology, Carnegie Institution of Wash
ington, in charge of course.
ALLISON L. BURNETT, Associate Professor of Biology, Western Reserve University.
JAMES W. LASH, Assistant Professor of Anatomy, University of Pennsylvania.
A. A. MoscoNA, Professor of Zoology, University of Chicago.
ARTHUR B. PARDEE, Professor of Biology, Princeton University.
JOHN W. SAUNDERS, JR., Professor of Biology, Marquette University.
II. LABORATORY ASSISTANTS
JOHN J. COFFEY,Department of Biology, The Johns Hopkins University.
SIDNEY B. SIMPSON, JR., Department of Zoology, Tulane University.

10 MARINE BIOLOGICAL LABORATORY
CH. DEVILLERS
BEATRICE MINTZ
N. T. SPRATT, JR.
J. P. TRINKAUS
Cii. DEVILLERS
R. B. CRAWFORD
R. Goss
J. OPPENHEIMER
T. D. BROCK
J. F. ALBRIGHT
R. AUERBACH
J. J. KOLLROS
I. R. KONIGSBERG
F. C. STEWARD
A. C. CLEMENT
A. C. CLEMENT
K. R. PORTER
K. R. PORTER
M. GLIMCHER
D. MAZIA
E. BELL
J. Giwss
S. SPIEGELMAN
A. MONROY
S. COHEN
J. W.HASTINGS
H. HALVOR5ON
M. POLLOCK
M. SUS5MAN
W. RUTTER
A. MONROY
C. LEVINTHAL
C. LEVINTHAL
T. B. FITZPATRICK
G. SZABÔ
A. PARDEE
W. VINCENT
F. H. WILT
J. COLLIER
R. S. EDGAR
R. S. EDGAR
D. BODENSTEIN
G. WEISSMANN
III. LECTURES
Structure and appearance of marginal zone and bilateral
symmetry.
Experimental analysis of differentiation in the mammalian
egg.
Morphogenetic movements in the early chick embryo.
Cellular basis of Fundulus epiboly.
Gastrulation in Fund ulus.
Energetics of development in Fundulus.
Regeneration and transplantation of scales in Fundulu.s.
Contributions, actual and potential, of Fundulus to experi
mental embryology.
Properties of cell surface in microorganisms.
Some dynamic aspects of antibody production.
Spleen and thymus development in relation to immunity.
Tissue responses to thyroid hormone in regeneration.
Clonal analysis of myogenesis.
Development of single plant cells.
The mollusk-annelid pattern of development.
Analysis of development in Ilyanassa.
Developmental cytology I.
Developmental cytology II.
Isolation, identification and properties of enamel protein
and the ultra-structural organization of mineral and
organic components.
Mechanisms of cell division.
Keratin biosynthesis and organization in feather develop
ment.
•¿ Collagen formation and destruction in morphogenesis.
The sequential reading of the genome.
Analysis of development in echinoderms.
Sequential protein synthesis during phage infection.
Rhythmic physiological events in Gonyolax.
The timing of enzyme synthesis during synchronous growth
•¿ in yeast.
Stability of extrachromosomal characters and their possible
role in differentiation.
Cellular differentiation in the slime molds.
Differentiation of pancreatic cells.
Analysis of development in echinoderms.
RNA, messenger, and the problem of differentiation.
The folding of a multichain enzyme and genetic comple
mentation.
The anatomy and physiology of pigmentation.
Cell regulatory mechanisms II.
Messengers, nucleoli and the anatomy of a ribosome.
Characteristics of RNA synthesis in cleaving sea urchin
eggs.
Fractionation of nucleic acid in Ilyanassa embryos.
Developmental genetics I.
Developmental genetics II.
Nerve regeneration in insects.
Vitamin A, lysosomes, and metamorphosis.

REPORT OF THE DIRECTOR It
PHYSIOLOGY
I. CONSULTANTS
MERKELH. JACOBS,Professor of Physiology, University of Pennsylvania.
ARTHUR K. PARPART, Professor of Biology, Princeton University.
ALBERT SZENT-GYöRGYI, Director,
Laboratory.
Institute for Muscle Research, Marine Biological
W. D. MCELROY,Director, McCollum-Pratt Institute, The Johns Hopkins University.
II. INSTRUCTORS
J. WOODLAND HASTINGS, AssociateProfessorof Biochemistry,Universityof Illinois,in
charge of course.
E. A. ADELBERG, Professor of Microbiology, Yale University.
PHILIP E. HARTMAN, Associate Professor of Biology, The Johns Hopkins University.
HARLYN HALVORSON, Professor of Bacteriology, University of Wisconsin.
SHINYA IN0UE, Professor of Cytology, Dartmouth College.
K. E. VAN HOLDE, Associate Professor of Physical Chemistry, University of Illinois.
FRED KARUSH, Professor of Microbiology, University of Pennsylvania School of
Medicine.
ALEXANDER KEYNAN, Israel Institute for Biological Research.
JOHN SPUDICH, University of Illinois.
CAROLYN VEEDER, Swarthmore College.
III. LABORATORYASSISTANTS
IV. LECTURES
H. K. SCHACHMAN The architecture of proteins.
ROBERT LOFTFIELD Comments on the current hypothesis of protein synthesis.
EMANUEL MARGOLIASH Structure and evolution of cytochrome c.
ROD CLAYTON Physical aspects of photosynthesis.
GEORGE WALD The problem of visual excitation.
ROD CLAYTON Primary photochemical reactions in photosynthesis.
WILLIAM HAGINS Physical mechanisms in photoreception.
DAN MAZIA Title to be announced.
S0L SPIEGELMAN A comparison of information transfer in DNA and RNA
genomes.
Joint Sessions of the Embryology and Physiology classes
MARTIN POLLOCK Stability of extrachromosomal characters and their possible
role in differentiation.
MAURICE SU5SMAN Cellular differentiation in the slime molds.
A. MONROY Analysis of development in echinoderms.
JOHN LAW Biological transalkylation reactions.
HERMAN KALCKAR Ectobiology in microorganisms and in mammalian cells.
HowAim HOLTZER Problems of cellular differentiation in cartilage and muscle.
WILLIAM ARNOLD A possible role of conduction electrons in biology.
ALBERT SZENT-GYöRGYI A survey of biochemistry.
G. T. REYNOLDS The use of the image intensifier in microscopic observations.
IRVIN ISENBERG Factors in charge-transfer complexing.

12 MARINE BIOLOGICAL LABORATORY
MARINE BOTANY
I. CONSULTANT
WILLIAM RANDOLPH TAYLOR, Professor of Botany, University of Michigan.
II. INSTRUCTORS
RICHARD C. STARR, Professor of Botany, Indiana University, in charge of course.
PHILIP W. CooK, Department of Botany, University of California.
WALTER R. HERNDON, Professor of Botany, University of Tennessee.
PETER S. DIXON, Hartley Botanical Laboratories, University of Liverpool.
III. LABORATORYASSISTANTS
CHARLES F. CLELAND, Department of Biological Sciences, Stanford University.
JOANNE K. SOLEM, Department of Plant Pathology, Cornell University.
IV. COLLECTOR
AUSTIN E. BROOKS,Department of Botany, Indiana University.
V. LECTURES
LUIGI PR0VA5OLI Problems in the development of culture media for algae.
INVERTEBRATE ZOOLOGY
I. CONSULTANTS
F. A. BROWN,JR., Professor of Zoology, Northwestern University.
LIBBIE H. HYMAN, American Museum of Natural History.
ALFRED C. REDFIELD, Woods Hole Oceanographic Institution.
II. INSTRUCTORS
CLARK P. READ, Professor of Biology, Rice University, in charge of course.
JOHN ANDERSON, Professor of Zoology, Cornell University.
‘¿G. F. GWILLIAM,Associate Professor of Biology, Reed College.
‘¿GEORGEHOLZ, Professor of Microbiology, College of Medicine at Syracuse.
W. D. RUSSELL HUNTER, Lecturer in Zoology, University of Glasgow.
ROGER MILKMAN, Associate Professor of Zoology, Purdue University.
RICHARD C. SANBORN, Professor of Zoology, Purdue University.
IRWIN W. SHERMAN, Assistant Professor of Zoology, University of California, Riverside.
STEPHEN BROWN, Ohio State University.
JAMES MCDANIEL, University of Oklahoma.
III. ASSISTANTS
IV. LECTURES
All special lectures were conducted jointly with those of the training program in com
parative physiology.

REPORT OF THE DIRECTOR
MARINE ECOLOGY
I. CONSULTANTS
JOHN H. RYTHER, Woods Hole Oceanographic Institution.
EUGENE P. ODUM, University of Georgia.
EDWIN T. MOUL, Rutgers University.
II. INSTRUCTORS
W. ROWLANDTAYLOR,Department of Oceanography and the Chesapeake Bay Institute,
The Johns Hopkins University, in charge of course.
OTTO H. KINNE, Biologische Anstalt Helgoland, Hamburg, Germany.
JOHN D. PALMER, Department of Biology, University of Illinois,Chicago.
HOWARD L. SANDERS, Woods Hole Oceanographic Institution.
LAWRENCE B. SLOBODKIN, Department of Zoology, University of Michigan.
III. LABORATORYASSISTANT
DONALD C. GORDON, University of Rhode Island.
STANLEY W. WATSON
GORDON C. RILEY
TALBOT H. WATERMAN
F. A. BROWN,JR.
H. H. SELIGER
EDWIN T. MOUL
GORDON A. RILEY
E. R. BAYLOR
M. R. CARRIKER
R. S. SCHELTEMA
ROBERT G. CONOVER
H. A. TURNER
E. L. MILLS
N. B. MARSHALL
C. LADDPROSSER
IV. LECTURES
13
The biology of marine nitrifying organisms.
Organic aggregates in sea water and their ecological sig
nificance.
Orientation and visual physiology.
Biological rhythms.
Measurement of light in bioluminescent organisms.
Zonation on rocky shores.
Zooplankton-phytoplankton interrelationships.
Langmuir cells and plankton distribution.
Systematic Ecology Program at MBL.
Ecology of sessile and benthonic marine invertebrate larvae.
Zooplankton trophic relationships.
Suggested mechanisms for formation of aggregations of
benthonic marine organisms.
Adaptive significance of life history features in an amphi
pod crustacean.
Deep-sea fishes.
Theoretical aspects of physiological adaptations.
THE LABORATORY STAFF
HOMER P. SMITH, General Manager
MRS. DEBORAH LAWRENCE HARLOW, Librarian
CARL 0. SCHWEIDENBACK, Manager, Supply
Department
IRVINE L. BROADBENT, Office Manager
GENERAL OFFICE
ROBERT KARLER, Superintendent,
Buildings and Grounds
ROBERT B. MILLS, Manager, De
partment of Research Service
Mrs. VIVIEN R. BROWN MRS. JUDITH A. KECK
MRS. FLORENCES. BUTZ MRS. VIVIAN I. MANSON
MRS. MARION C. CHASE MRS. VIRGINIA M. MOREHOUSE
MRS. KAREN M. DRISCOLL

14 MARINE BIOLOGICAL LABORATORY
LIBRARY
MRS. V. BRANDENBURG ALBERT K. NEAL
Miss JANE FESSENDEN MRS. DORIS T. RICKER
MRS. L. JOSEPH
MAINTENANCE OF BUILDINGS AND GROUNDS
ROBERT ADAMS WALTER J. JASKUN
ELDON P. ALLEN DONALD B. LEHY
BERNARD F. CAVANAUGH RALPH H. LEWIS
MANUEL P. DUTRA RUSSELL F. LEwIs
STANLEY C. ELDREDGE HENRY F. POTTER
GARDNER F. GAYTON JAMES S. THAYER
ROBERT GUNNING ROBERT H. WALKER, JR.
DEPARTMENT OF RESEARCH SERVICE
GAIL M. CAVANAUGH MISS RUTH PRELLER
SEAVER R. HARLOW FRANK E. SYLVIA
LOWELL V. MARTIN
SUPPLY DEPARTMENT
ARNO J. BOWDEN PAUL SHAVE
DONALD P. BURNHAM BRUNO F. TRAPASSO
DAVID H. GRAHAM JOHN J. VAL0I5
MRS. E. GREEN JARED L. VINCENT
ROBERT 0. LEHY HALLETT S. WAGSTAFF
ROBERT M. PERRY
DINING HALL AND HOUSING
ROBERT T. MARTIN, Manager, Food Service
MRS. ELIZABETH KUIL, Supervisor, Dining Room
MRS. ELLEN T. NICKELSON, Supervisor, Dormitories
ALAN G. LUNN, Supervisor, Cottage Colony
3. INVESTIGATORS; LALOR,LILLIE ANDGRASSFELLOWS;STUDENTS
Independent Investigators, 1963
ADELBERG,EDWARD A., Chairman, Department of Microbiology, Yale University
ADELMAN, WILLIAM J., JR., Associate Professor of Physiology, University of Maryland
ALJURE, EMILI0, Research Fellow (Rockefeller Fellow, University of Cali, Colombia), Colum
bia University, College of Physicians and Surgeons
ALLEN, ROBERTD., Associate Professor of Biology, Princeton University
ANSERSON, JoHN M., Professor of Zoology, Cornell University
ARMSTRONG,CLAY M., Senior Assistant Surgeon, National Institutes of Health
ARMSTRONG, PHILIP B., Professor and Chairman, Department of Anatomy, State University of
New York College of Medicine at Syracuse
ARNOLD,WILLIAM A., Principal Biologist, Oak Ridge National Laboratory
AUCLAIR, WALTER, Assistant Professor of Zoology, University of Cincinnati
AUSTIN, GEORGE M., Independent Investigator, University of Oregon Medical School
AUSTIN, COLIN RUSSELL, Member of External Scientific Staff, Medical Research Council,
Physiological Laboratory, Cambridge, England

REPORT OF THE DIRECTOR 15
BANG, FREDERIK B., Professor and Chairman of Department of Pathobiology, The Johns
Hopkins University School of Hygiene and Public Health
BARTH, L. G., Professor of Zoology, Columbia University
Bszi., EUGENE, Associate Professor of Biology, Massachusetts Institute of Technology
BENNETT, MICHAEL V. L., Associate Professor of Neurology, Columbia University
BIBRING, THOMAS, Visiting Assistant Professor, Vanderbilt University
BINSTOCK, LEONARD, Electronic Engineer (Instrumentation), National Institutes of Health
BODIAN, DAVID, Director, Department of Anatomy, The Johns Hopkins University School of
Medicine
BRETTHAUER, ROGER K., Postdoctoral Fellow, University of Wisconsin
BRINLEY, F. J., JR., Assistant Professor of Physiology, The Johns Hopkins University School
of Medicine
BROWN, FRANK A., JR., Morrison Professor of Biology, Northwestern University
BRYANT,S. H., Associate Professor of Pharmacology, University of Cincinnati
BULLOCK, THEODORE H., Professor of Zoology, University of California, Los Angeles
BURCH,HELENB., Associate Professor of Pharmacology, Washington University
BURKE, JOSEPH A., Assistant Professor of Biology, Acting Chairman of the Department, Loyola
College, Baltimore
BURNETT, ALLISON L., Associate Professor of Biology, Western Reserve University
BUSH, LOUISE F., Assistant Professor of Zoology, Drew University
CARLSON, FRANCIS D., Professor of Biophysics and Chairman of the Department, The Johns
Hopkins University
CHAET, ALnRD B., Professor of Biology, American University
CHENEY, RALPH HOLT, Professor of Biology (General Physiology), Brooklyn College
CHILD, FRANK M., Assistant Professor of Zoology, University of Chicago
CLAFF, C. LLOYD, Treasurer and Investigator, Single Cell Research Foundation, Inc.
CLARK, ARNOLD M., Professor of Biology, University of Delaware
CLARK, ELOISE E., Assistant Professor in Zoology, Columbia University
CLEMENT, ANTHONY C., Professor of Biology, Emory University
COLE, KENNETH S., Chief, Laboratory of Biophysics, NINDB, National Institutes of Health
COLLIER,JACK R., Marine Biological Laboratory
COLWIN, ARTHUR L., Professor of Biology, Queens College
COLWIN, LAURA HUNTER, Lecturer, Queens College
COOK, PHILIP WILLIAM, Assistant Professor of Botany, University of Vermont
COOPERSTEIN,SHERWIN J., Associate Professor of Anatomy, Assistant Dean of Medical School,
Western Reserve University School of Medicine
COPELAND, EUGENE, Professor and Chairman of Department of Zoology, Tulane University
CORMACK, D. H., Research Associate in Molecular Biophysics, Institute of Molecular Bio
physics, Florida State University
Cosrr.u.o, DONALDP., Kenan Professor of Zoology, University of North Carolina
CRANE,ROBERTK., Professor and Chairman of the Department of Biochemistry, The Chicago
Medical School
CROWELL, SRAR5, Professor of Zoology, Indiana University
D'ALESANDRO,PHILIP A., Assistant Professor, The Rockefeller Institute
DAVIDSON, HAROLD,Physical Science Aide, National Institutes of Health
Dw@, WILLIAMC., JR., Assistant Professor, Michigan State University
DETTBARN, WOLF-DIETRICH, Assistant Professor of Neurology, Columbia University, College of
Physicians and Surgeons
DEVILLER5, CHARLES, Professor, Faculté des Sciences, Sorbonne, Paris
DIECKE, F. P. J., Associate Professor of Physiology, George Washington University, School
of Medicine
DIXON, PETER STANLEY, University Lecturer, University of Liverpool
EBERT, JAMES D., Director, Department of Embryology, Carnegie Institution of Washington
ECKERT,ROGER0., Assistant Professor of Zoology, Syracuse University
EDDS, M. V., JR., Professor and Chairman of the Department of Biology, Brown University
EHRENSTEIN, GERALD, Physiologist, National Institutes of Health
FA1u@, PATRICIAMCCLEMENT,Associate Biophysicist, Argonne National Laboratory

16 MARINE.BIOLOGICAL LABORATORY
FAWCETF, DON W., Hersey Professor of Anatomy and Head of the Department, Harvard Uni
versity Medical School
FEDER, NED, Assistant Professor of Biology, Harvard University
Fox, STEPHEN S., Assistant Professor, University of Michigan
FREEMAN, ALAN R., Fellow in Neurology, Columbia University
FUORTES, M. G. F., Head, Neurophysiology Section, NINDB, National Institutes of Health
FURSHPAN, EDWIN J., Assistant Professor of Neurophysiology and Neuropharmacology, Har
yard Medical School
GAINER, HAROLD,USPHS Fellow, Columbia University
GERMAN, JAMES L., Assistant Professor, Rockefeller Institute
GILBERT, DANIEL L., Physiologist, National Institutes of Health
GRANT, PHILIP, Program Director, Developmental Biology, National Science Foundation
GREIF, ROGERL., Associate Professor of Physiology, Cornell University Medical College
GROSCH, DANIEL S., Professor of Genetics, North Carolina State College
GROSS, PAUL R., Associate Professor of Biology, Brown University
GRUNDFEST, HARRY, Professor of Neurology, Columbia University
GWILLIAM, GILBERT F., Associate Professor of Biology, Reed College
HAGINS, WILLIAM A., Research Physiologist, National Institutes of Health
HAGIWARA, S., Professor of Zoology, Department of Zoology, Brain Research Institute, U. C.
L.A.
HALVORSON, HARLYN 0., Professor of Bacteriology, University of Wisconsin
HARDING, CLIFFORD V., Associate Professor of Physiology, Columbia University, College of
Physicians and Surgeons
HARTMAN, PHILIP E., Associate Professor of Biology, The Johns Hopkins University
HASTINGS,J. WOODLAND,Associate Professor of Biochemistry, University of Illinois
HAYASHI, TERU, Professor of Zoology, Columbia University
HEGYELI, ANDREW, Institutefor Muscle Research, Marine Biological Laboratory
HENLEY, CATHERINE, Research Associate, University of North Carolina
HERNDON, WALTER R., Professor and Head of the Department of Botany, University of Ten
nessee
HERVEY, JOHN P., Senior Electronic Engineer, The Rockefeller Institute
HESSLER, ANITA Y., Research Associate, Marine Biological Laboratory
HILL, ROBERT BENJAMIN, Instructor in Physiology, Dartmouth Medical School
HOLZ, GEORGE G., Professor and Chairman of the Department of Microbiology, S. U. N. Y.,
Upstate Medical Center, College of Medicine
HOPKINS, CHARLES ADRIAN, Lecturer in Parasitology, University of Glasgow; and Rice Uni
versity
H0SKIN, FRANCIS C. G., Assistant Professor, Columbia University, College of Physicians and
Surgeons
HUBBARD, RUTH, Research Associate, Harvard University
HUMPHREYS, TOM, NIH Postdoctoral Fellow, Massachusetts Instituteof Technology
HUNTER, W. D. RUSSELL, University Lecturer in Zoology, University of Glasgow
HWANG, JOSEPH C., Research Associate, Columbia University
INoul, SHINYA, Professor and Chairman of the Department of Cytology, Dartmouth Medical
School
ISENBERG, IRvIN, Institutefor Muscle Research, Marine Biological Laboratory
JANOFF, AARON, Assistant Professor of Pathology, New York University School of Medicine
JOHNSSON, RUTH, Institute for Muscle Research, Marine Biological Laboratory
JOSEPHSON, ROBERT K., Assistant Professor, University of Minnesota
KALEY, GABOR, Assistant Professor of Pathology, New York University School of Medicine
KAMINER, BENJAMIN, Institute for Muscle Research, Marine Biological Laboratory
KANE, ROBERT E., Assistant Professor of Cytology, Dartmouth Medical School
KARUSH, FRED, Professor of Microbiology, University of Pennsylvania School of Medicine
KEMPTON, RUDOLF T., Professor of Zoology, Vassar College
KENYAN, ALEXANDER, Foreign Instructor in Physiology, Israel Institutefor Biological Research
KINNE, OTTo, Director and Professor, Biologische Anstalt Helgoland
KLEINHOLz, L. H., Professor of Biology, Reed College

REPORT OF THE DIRECTOR 17
KRASSNER, STUART M., Guest Investigator, Postdoctoral Research Fellow, The Rockefeller
Institute
KUFFLER, STEPHEN W., Professor of Neurophysiology and Neuropharmacology, Harvard Mcdi
cal School
KUSANO, KIY05HI, Research Associate, Columbia University ; Tokyo Medical and Dental
University
LANSING, ALBERT I., Professor of Anatomy and Chairman of the Department, University of
Pittsburgh
LASH, JAMES W., Assistant Professor of Anatomy, University of Pennsylvania School of
Medicine
LASTER, LEONARD, Chief of Gastroenterology Unit, National Institute of Arthritis and Meta
bolic Diseases
LAUFER, HANS, Assistant Professor of Biology, The Johns Hopkins University
LAZAROW,ARNOLD, Professor and Head of the Department of Anatomy, University of Minnesota
LERMAN, SIDNEY, Associate Professor of Ophthalmology and Assistant Professor of Biochem
istry, University of Rochester School of Medicine and Dentistry
LEVIN, JACK, Fellow, Department of Medicine, The Johns Hopkins Hospital and University
LEVINE, LAWRENCE, Associate Professor of Biochemistry, Brandeis University
LEVINTHAL, CYRUS, Professor of Biophysics, Massachusetts Institute of Technology
LEVY, MILTON, Professor and Chairman of the Department of Biochemistry, New York Uni
versity College of Dentistry
LEWIS, HERMAN W., Program Director, Genetic Biology, National Science Foundation
LIEBMAN, PAUL A., Research Fellow, University of Pennsylvania
L0CHHEAD, JOHN H., Professor of Zoology, University of Vermont
LOEWENSTEIN, WERNER R., Associate Professor of Physiology, Columbia University
LOPEZ, ENRIQUE, Research Associate, Columbia University; National Institute of Cardiology,
Mexico
DE LoRENzo, A. J., Director, Anatomical and Pathological Research Laboratories, The Johns
Hopkins University School of Medicine
LoTE, WARNERB., Associate Professor of Biophysics, The Johns Hopkins University
MACKIE, G. 0., Associate Professor, University of Alberta
MANDRIOTA, FRANK J., Fellow in Neurology, Columbia University
LUTWAK-MANN,CECILIA,Principal Scientific Officer on the Staff of the Agriculture Research
Council of Great Britain, Space Biosciences Institute, Florida State University
MANN, THADDEUS,Director, Agriculture Research Council, Unit of Reproduction, Physiology
and Biochemistry, Cambridge, England, Space Biosciences Institute, Florida State University
MARSLAND, DOUGLAS, Research Professor, Biology Department, New York University
MATEYKO, G. M., Associate Professor of Biology, New York University
MCCARTHY, ELAINE S., Fellow in Research in Neurophysiology, Grass Instrument Company
MCELROY, W. D., Chairman, Department of Biology and Director, McCollum-Pratt Institute,
The Johns Hopkins University
METZ, CHARLES B., Professor of Biology, Florida State University
MILKMAN, ROGER DAWSON, Associate Professor of Zoology, Syracuse University
MOHSEN, T., Assistant Professor, Faculty of Science, Dakar University, Republic of Senegal
MONROY, ALBERTO, Professor of Comparative Anatomy, University of Palermo, Italy
MOORE, JOHN W., Associate Professor of Physiology, Chief, Laboratory of Cellular Neuro
physiology, Duke University
MOORE, RICHARD 0., Professor of Biochemistry, Ohio State University
MORAN, JOSEPH F., JR., Assistant Professor of Biology, Russell Sage College
MOSCONA, A. A., Professor of Zoology, University of Chicago
MULLINS, L. J., Professor of Biophysics, University of Maryland
MUSACCHIA, X. J.,Professor of Biology, Saint Louis University
NACE, PAUL FOLEY, Professor, Molecular Biology Research Unit, McMaster University
NAKAJIMA, SHIGEHIRO, Visiting Fellow and Research Scholar; United Cerebral Palsy Fellow,
Columbia University (University of Tokyo Medical School)
NAKAMURA, YUTAKA, Research Associate, Columbia University (University of Tokyo)
NASATIR, MAIMON, Assistant Professor in Botany, Brown University
NELSON, LEONARD, Associate Professor of Physiology, Emory University

18 MARINE BIOLOGICAL LABORATORY
NoRRIs, W. E., JR., Professor of Biology, Chairman of Department, Southwest Texas State
College
NYBORG,WESLEY L., Professor of Physics, University of Vermont
OVERTON,JANE, Associate Professorial Lecturer, University of Chicago
PALMER, JOHN D., Assistant Professor in Biology, University of Illinois
PARDEE, ARTHUR B., Professor of Biology, Princeton University
PARPART, ARTHUR K., Professor and Chairman of the Department of Biology, Princeton Uni
versity
PASSOW, HERMANN, Professor in Physiology, Head of Department, 11. Physiologisches Institut
d. Universität d. Saarlandes, Homburg, Saar, Germany
PERSON, PHILIP, Chief, Special Dental Research Laboratory, Veterans Administration Hospital,
Brooklyn
POLLOCK, MARTIN RIVERS, Senior Lalor Fellow, National Institute for Medical Research,
London, N.W.7
PORTER, KEITH R., Professor of Biology, Harvard University
POTTER, DAVID D., Associate Professor in Neurophysiology and Neuropharmacology, Harvard
Medical School
PROSSER, CLIFFORD LADD, Professor of Physiology and Head of the Department, University of
Illinois
RABIN, HARVEY, Assistant Professor, The Johns Hopkins University School of Hygiene
READ,C. P., Professor of Biology, Rice University
REBHUN, LIONEL I., Assistant Professor of Biology, Princeton University
REUBEN, JOHN P., Assistant Professor of Neurology, Columbia University
REYNOLDS,GEORGET., Professor of Physics; Director, Elementary Particles and Cosmic Ray
Laboratory, Princeton University
ROCKSTEIN, MORRIS, Professor of Physiology, University of Miami School of Medicine
ROSE, S. MERYL, Professor of Anatomy, Tulane University
ROSENBERG, PHILIP, Research Associate, Columbia University, College of Physicians and
Surgeons
ROSENKRANZ, HERBERT S., Assistant Professor of Microbiology, Columbia University, College
of Physicians and Surgeons
RUGH, ROBERTS,Associate Professor of Radiology, Columbia University, College of Physicians
and Surgeons
RUSTAD, RONALD C., Assistant Professor of Physiology, Florida State University
SAITO, NOZOMU, Visiting Fellow and Research Scholar, United Cerebral Palsy Fellow, Colum
bia University (Tokyo Medical and Dental University)
SANBORN, RICHARD C., Professor of Zoology, Purdue University
SANDERS, HOWARD L., Research Associate, Woods Hole Oceanographic Institution
SATIR, PETER, Instructor in Zoology and College Biology, University of Chicago
SATO, HIDEMI, Assistant Professor, Dartmouth Medical School
SAUNDERS, JOHN W., Professor and Chairman of the Department of Biology, Marquette Uni
versity
SCHMEER, SISTER M. R05ARII, Chairman of Biological Education, Archdiocese, New York City
SCHUEL, HERBERT, USPHS Postdoctoral Fellow, Biology Division, Oak Ridge National Lab
oratory
Scorr, SISTERFLORENCEMARIE, Professor of Biology and Chairman of the Department, Seton
Hill College
Scorr, GEORGET., Professor and Chairman of the Department of Biology, Oberlin College
SEAEL5, ROBERTL., Assistant Professor, University of Virginia
SELIGER, HOWARD H., Associate Professor, McCollum-Pratt Instituteand Department of Biol
ogy, The Johns Hopkins University
SHEMIN, DAVID, Professor of Biochemistry, Columbia University
SHERMAN, IRWIN W., Assistant Professor of Zoology, University of California, Riverside
SHIvERs, CHARLES ALEX, USPHS Postdoctoral Fellow, Florida State University
SIcHEL, F. J., Professor and Chairman of the Physiology and Biophysics Departments, College
of Medicine, University of Vermont
SIDDIQUI,WASIM A., Guest Investigator, The Rockefeller Institute
SIMMONS, JOHN E., JR., Assistant Professor of Biology, Emory University

REPORT OF THE DIRECTOR 19
SJODIN, RAYMOND A., Associate Professor of Biophysics, University of Maryland
SLOBODKIN, L. B., Associate Professor of Zoology, University of Michigan
SPEIDEL, CARL C., Professor of Anatomy, University of Virginia
SPINDEL, WILLIAM, Professor of Chemistry, Rutgers University
STARR, RICHARD C., Professor of Botany, Indiana University
STEINHARDT, JACINTO, Professor of Chemistry, Georgetown University
STEPHENS, GROVER C., Associate Professor of Zoology, University of Minnesota
STEVENSON, J. RoSs, Jacques Loeb Associate in Marine Biology (Kent State University)
STOECKNIUS, WALTHER, Assistant Professor, The Rockefeller Institute
STRITTMATTER, PHILIPP, Associate Professor of Biochemistry, Washington University
STUNKARD, HORACE W., Research Associate, American Museum of Natural History
SUDAK, FREDERICK N., Assistant Professor of Physiology, Albert Einstein College of Medicine
SURGENOR, DOUGLAS M., Dean of the Medical School, Chairman of the Department of Bio
chemistry, State University of New York at Buffalo
SZABÔ, GEORGE, Associate Biologist in Dermatology, Massachusetts General Hospital, Asso
ciate in Anatomy, Harvard Medical School
SZENT-GYöRGYI,ALBERT, Director and Chief Investigator, Institute for Muscle Research, Marine
Biological Laboratory
SZENT-GYöRGYI, ANDREW, Professor of Cytology, Dartmouth Medical School
TASAKI, ICHIJI, Acting Chief, Laboratory of Neurobiology, NIMH, National Institutes of
Health
TAYLOR, ROBERT E., Associate Chief, Biophysics Laboratory, NINDB, National Institutes of
Health
TAYLOR, WILLIAM RANDOLPH, Professor of Botany and Curator of Algae in the Herbarium,
University of Michigan
TAYLOR, WALTER ROWLAND, Assistant Professor of Oceanography, The Johns Hopkins Uni
versity
TOMIZAWA,HENRY H., Senior Research Associate, Associate Professor of Biochemistry, Fels
Research Institute, Antioch College
TORCH, REUBEN, Associate Professor of Zoology, University of Vermont
TRAGER,WILLIAM, Associate Professor, The Rockefeller Institute
TRAVIS, DAvID M., Assistant Professor of Pharmacology, Therapeutics and Medicine, Univer
sity of Florida
TRINKAUS,JOHN PHILIP, Associate Professor of Biology, Yale University
TROLL, WALTER, Associate Professor, New York University, Institute of Industrial Medicine
TWAROG,BETTYM., Assistant Professor, New York University Medical School
TWEEDELL, KENYON S., Associate Professor of Biology, University of Notre Dame
ULBRICHT, WERNER W., Assistant Professor of Physiology, Duke University
USHERWOOD, P. N. R., United Cerebral Palsy Fellow, Columbia University; Department of
Zoology, University of Glasgow
VAN HOLDE, K. E., Associate Professor of Chemistry, University of Illinois
VASINGTON, FRANK D., Postdoctoral Fellow, The Johns Hopkins University
VILLEE, CLAUDE A., Associate Professor of Biological Chemistry, Harvard Medical School
DE VILLAFRANCA, GEORGE W., Associate Professor of Zoology, Smith College
VINCENT, W. S., Associate Professor of Anatomy, University of Pittsburgh
WALD, GEORGE, Professor of Biology, Harvard University
WEDB, H. MARGUERITE, Associate Professor of Biology, Goucher College
WEBEE, MORTON M., Associate Professor, Saint Louis University School of Medicine
WERMAN, ROBERT, Professor of Psychiatry, Indiana University
WICHTERMAN, RALPH, Professor of Biology, Temple University
WIERCINSKI, FLOYD J., Associate Professor of Biology, Drexel Institute of Technology
WILBER,CHARLESG., Dean of the Graduate School and Professor of Biological Sciences, Kent
State University
WILSON, WALTER L., Associate Professor of Physiology and Biophysics, University of Vermont
WITIENBERG, JONATHAN B., Associate Professor of Physiology, Albert Einstein College of
Medicine
YANAGITA, TAME MASA, Professor in Animal Physiology at Ochanomizu University, Japan

20 MARINE BIOLOGICAL LABORATORY
ZIMMERMAN, ARTHUR M., Assistant Professor, State University of New York, Downstate
Medical Center
Lalor Fellows, 1963
POLLOCK, MARTIN RIvERs, National Institute for Medical Research, London
LAUFER, HANS, The Johns Hopkins University
NASATIR, MAIMON, Brown University
MOHSEN, T., University of Strasbourg, Republic of Senegal, Africa
ROSENKRANTZ, HERBERT S., Columbia University, College of Physicians and Surgeons
Liffie Fellow, 1963
DEVILLERS, CHARLES, Facultédes Sciences, Sorbonne, Paris
Grass Fellows, 1963
BULLOCK, THEODORE H., University of California, Los Angeles
BERNSTEIN, THEODORE W., University of Cincinnati,College of Medicine
LIEBMAN, PAUL A., University of Pennsylvania
MCCARTHY, ELAINE S., Biological Laboratories, Harvard University
Beginning Investigators, 1963
ALVAREZ, JAIME, Tulane University
BAUER, GUSTAV ERIC, University of Minnesota
BELAMARICH, FRANK A., State University of New York at Buffalo
BERNSTEIN, THEODORE W., University of Cincinnati College of Medicine
BURCHILL, BROWER R., Florida State University
CHERVIN, PAUL N., University of Vermont
CHURCHICH, JORGE E., University of Illinois
COHEN, LAWRENCE B., Columbia University
COHEN, WILLIAM DAVID, Columbia University
COUTURE, KENNETH G., Tulane University
ESKRIDGE, ROSEMARY W., State University of New York at Buffalo
FISHER, F. M., JR.,Rice University
GANGULY, BANKU B., Serampore and Presidency College, Calcutta
GRANT,ROBERTJ., Columbia University
GREEN, JONATHAN P., The Johns Hopkins University
HOSTETLER,KARL,Western Reserve University School of Medicine
JACKSON, JAMES A., Western Reserve University
KANNO, YOSHINOBU, Tokyo Medical and Dental University
KRIEBEL, MAHLON E., University of Washington
LANGE, MARY T., Marquette University
LENTZ, THOMAS L., Yale University School of Medicine
LESLIE, ROBERT B., Nottingham, England
LINDALL,ARNOLDW., University of Minnesota
MALKIN, LEONARD I., Brown University
MARCHALONIS, JOHN, The Rockefeller Institute
NAKASE, YASUKIYO, Kitasato Institutein Tokyo
PFOHL, RONALD JOHN, Michigan State University
ROVAINEN, CARL M., Harvard University
SANEL,FRANCEST., Columbia University, College of Physicians and Surgeons
SREBRO,RICHARD, National Institutes of Health
TAKATA, MITSURU, Duke University
TAKENAKA, TOSHIFUMI, National Institutes of Health
WATKINS, DUDLEY TAYLOR, Western Reserve Medical School
WHEELER, MAYNARD B., Columbia University, College of Physicians and Surgeons

REPORT OF THE DIRECTOR 21
Research Assistants, 1963
ACQUAVIVA, PATRICIA, Seton Hill College
AKIN, JOHN R., Tulane University
AKIN, GWYNN MARIE, Tulane University
ALFANO, JOSEPHINE, Cornell University Medical College
ANDREWS, PETE.R M., American University
ARNOLD, JOHN M., University of Minnesota
ARTHUR, ELIZABETH JEAN, Purdue University
ASHMAN, ROBERT F., Columbia University, College of Physicians and Surgeons
AVERY, PATRICIA PALMER, Wheaton College
BAIRD, SPENCER L., Woods Hole
BAKER, CYNTHIA, Harvard Medical School
BALLANTINE, THOMAS VAN NESS, Princeton University
BARKER, KENNETH RAy, University of Mississippi
BARNWELL, FRANKLIN H., Northwestern University
BLAIR, GRACE L., University of Minnesota
BORGMAN, MARTHA, Scarsdale, New York
BOSLER, ROBERT, Harvard Medical School
BRADBURY, JACK, Reed College
BRODIN, ARLEN, University of Minnesota
BROOKS, AUSTIN E., Indiana University
BROWN, STEPHEN C., George Washington University
CARLSON, BLENDA, Florida State University
CARLSON, PAUL E., University of Minnesota
CASSIDY, REv. JOSEPH D., North Carolina State College
CHEN, HENRY L., Harvard University
CHURCHICH, MARGARITA, University of Illinois
CLARK, CHARLOTTE, Harvard University
CLELAND, CHARLES F., Stanford University
‘¿COFFEY, JOHN J., The Johns Hopkins University
CORFF,SANDRAC., State University of New York, Downstate Medical Center
‘¿CROSBY, GALE M., Brandeis University
CROUSE,FRANCESW., Biologische Anstalt Helgoland, Germany
DAS, Asrr, University of Illinois
DASHE, CHARLES K., University of Chicago
DAUGHERTY, WAYNE F., JR., The Johns Hopkins University
DAVIS, ERSKINE D., National Institutes of Health
DIEHL, FRED A., Western Reserve University
DIEHL, NORMA A., Western Reserve University
DOANE, MARSHALL, University of Maryland, School of Medicine
DOEBELI, ROBERT, Harvard Medical School
DONOHOE, CHARLIE W., National Institutes of Health
DOUGHERTY, WILLIAM J., Princeton University
EZELL, STILES D., JR., Bryn Mawr College
FALLON, JOHN F., Marquette University
FINDLAY, MILDam M., Syracuse University
FITZJARRELL,AUSTIN T., Tulane University
FOK, Y. B., University of Maryland
FIUR, ELLEN, Brandeis University
FORAN, ELIZABETH H., Smith College
FORD, LINCOLN E., University of Rochester, School of Medicine
FREEMAN, GARY, University of Chicago
FULLER,CAMERONB., Hunter College
FUSARI, MARGARET H., Boston University
‘¿GALlS, ANNA, Columbia University, College of Physicians and Surgeons
GEDMINTAS, DANA, University of Chicago
•¿GERSH, FRANK S., Reed College

22 MARINE BIOLOGICAL LABORATORY
GERSHFELD, NORMAN L., National Institutes of Health
GIIIRADELLA, HELEN, Cornell University
GIBBS, R. G., University of Wisconsin
GOLDMAN,ROBERTD., Princeton University
GORDON,DONALD C., JR., University of Rhode Island
GOUGH,HARRYM., Brown University
GRABNAR, MIKLAvI, The Johns Hopkins University
GRABSKF., ROBERT, University of Kansas
GamE,STEPHENC., ColumbiaUniversity
GUPTA, RAJ KUMARI, University of Alberta
HABAS, LINDA B., University of Illinois
HALL, SUSAN C., Harvard University
HAMMOND, CONSTANCE, University of Pittsburgh
HARDIE,JoN H., University of Illinois
HARRIS, EDWARD M., Duke University
HAUSCHKA, STEPHEN DENISON, The Johns Hopkins University
HAWRYLKA,EUGENIAA., New York University
HICKMAN, JAMES CRAIG, Oberlin College
HINES, ROSEMARY, Saint Louis University
HOLLENBERG, ROBERTD., Harvard Medical School
HOLLIDAY, MARY P., Seton Hill College
HORWITZ, BARBARA ANN, Emory University
HUFNAGEL, LINDA A., University of Vermont
HUGHES, KENT S., The Chicago Medical School
IMLAY, MARC JAMES, American University
JONES, FRANCES E., Columbia University
KAMINSKY, NORMAN M., George Washington University
KAPICA, SUZANNE, Russell Sage College
KATZ, GEORGE,Columbia University
KIEBLICH, JACOBA, University of Amsterdam
KIMBALL, FRANCES, Reed College
KINSER, GLENN W., JR., Indiana University
KL.oETzEL, JOHN A., The Johns Hopkins University
KRAININ, JAMES, Harvard Medical School
KUNISAWA, RIY0, University of California
LAMBSON, ROGER 0., Tulane University
LANDOLT, CASPER DALE, Texas Western College
LARSEN, KATHERINE, University of Chicago
LEITNER, LEO G., JR., National Institutes of Health
LENTZ, JUDITH P., Yale University
LEWIS, DOROTHY, Institute for Muscle Research
LEWIS, HAZEL, Institute for Muscle Research
LIN, JAMESC. H., North Carolina State College
LIVINGSTON, LAURIE, Goucher College
LLOYD, SAMUEL, Princeton University
LONG,CEDRICW., Princeton University
LUBBEN, BARBARA H., St. John's University
MACNAMARA, GAEL R., Columbia University, College of Physicians and Surgeons
MANGAN, JEROME, University of Cincinnati
MARITATO,CYNTHIAA., University of Pennsylvania
MARTIN, STAR, Harvard Medical School
MCDANIEL,JAMES SCorr, University of Oklahoma
MCELROY, KRISTINE, American University
MCKENZIE, DONALD K., National Institutes of Health
MCLAUGHLIN, JANE A., Institute for Muscle Research
MILLER, RICHARD L., University of Chicago
MOHIT, ROBERTA, New York University School of Medicine
MOULE, MARGARET, McMaster University

REPORT OF THE DIRECTOR 23
MULCARE, DONALD J., University of Notre Dame
MUNDAY, JOHN C., JR., University of Illinois
NICHOLLS, JOHN G., Harvard Medical School
PALKA, JOHN M., University of California at Los Angeles
PARR, EARL L., Columbia University
PEARLSTEIN, ROBERT M., Institute for Muscle Research
PENICNAK, A. JOHN, University of Massachusetts
PERRY, BARBARA, Institute for Muscle Research
PETRIE, Ro@ H., Vanderbilt University School of Medicine
PHILPOTT, DELBERT E., Institute for Muscle Research
PRESTIDGE, LOUISE S., Princeton University
QUINN, CONSTANCE, Syracuse University
RAY, PATRIcLA A., Seton Hill College
REED, RAY, Indiana University Medical Center
RAw, ROBERT E., Miami University, Oxford
RICHMOND, ARTHUR PRATT, Boston University
RIES, JAMES J., National Institutes of Health
ROBBINS, NORMAN, The Rockefeller Institute
R0BIS0N, GEORGEALAN, Western Reserve University
ROSEN, BARRY, Brown University
ROSENTHAL, JEAN, Albert Einstein College of Medicine
SANDLIN, RONALD A., National Institutes of Health
SANGER, JOSEPH WILLIAM, Dartmouth Medical School
SCHNITZLER, RONALD M., University of Vermont
SCHOEPF, CLAUDE, Columbia University
SCHULKIND, NANCY M., Smith College
SCH WEND, MARY J., Albert Einstein College of Medicine
SHARPLESS, T. K., Princeton University
SHAW, CHARIS, Tulane University
SHILLING, JANE 0., Mount Holyoke College
SIEGAL, MARIAN R., Brandeis University
SILBERMAN, LESTER, State University of New York, Downstate Medical Center
SILVEIRA, MARINA, University of Sao Paulo, Brasil
SIMMONS, NORWOODN., National Institutes of Health
SIMPSON, SIDNEY B., Tulane University
SKEHAN, PHILIP J., Syracuse University
SMITH, ARLAN E. S., Harvard University
SMITH, ROBERT HARLAN, American University
SMITH, STEPHEN D., Tulane University
SOLEM, JOANNE K., Cornell University
SPEAR, JOSEPH F., Loyola College
SPECHT, PHILIP C., Syracuse University
SPUDICH, JAMES A., University of Illinois
SPUDICH, JOHN L., University of Illinois
STAHL, RUTH C., The Johns Hopkins University
STEADY, HENRY M., Yale University
STERN, DORIS N., Florida State University
STERN, SAMUEL, Florida State University
SZENT-GYORGYI, GYULA, Institute for Muscle Research
SZENT-GYöRGYI, MARTA, Institute for Muscle Research
TAMM, SIDNEY L., The University of Chicago
TAYLOR, ELLEN L., University of Vermont
THORNTON, WILLIAM E., Vanderbilt University
ToLls, HELEN, Wesleyan University
TOOMAJIAN, LEE C., Smith College
TROTTER, ROBERT T., University of Vermont
U@ta, A5T0N, American University
UT5uMI, SAYAKA, University of Pennsylvania School of Medicine

24 MARINE BIOLOGICAL LABORATORY
VAN NORMAN, E., Princeton University
VEEDER, CAROLYN, Swarthmore College
VIRKAR, R. A., University of Minnesota
WASSERMAN, ELEANOR S., Brandeis University
WEINBERG, ROBERT, Massachusetts Institute of Technology
WINDSOR, MARY LINDA, The Johns Hopkins University
WILLIAMS, DANIEL B., Western Reserve University
WOODSIDE, KENNETH H., University of Rochester School of Medicine and Dentistry
ZEIDENBERG, PHILLIP, Columbia University, College of Physicians and Surgeons
ZWICK, MARTIN, Massachusetts Institute of Technology
Library Readers, 1963
BALL, ERIC G., Professor of Biological Chemistry, Harvard Medical School
BAYLOR, MARTHA B., Independent Investigator, Marine Biological Laboratory
BERNE, ROBERT M., Professor of Physiology, Western Reserve University, School of Medicine
Bumnn, ELMERG., Henry Fairfield Osborn Professor of Biology, Princeton University
CHASE, AURIN M., Professor of Biology, Princeton University
CLIFFORD, SISTER ADELE, Professor of Biology, College of Mt. St. Joseph
COHEN, SEYMOUR S., Professor of Biochemistry, University of Pennsylvania
DAVIS, BERNARD D., Head of the Department of Bacteriology and Immunology, Harvard Mcdi
cal School
DUBOIS, ARTHURB., Professor of Physiology, University of Pennsylvania School of Medicine
EDER, HOWARD A., Professor of Medicine, Albert Einstein College of Medicine
EISEN, HERMAN N., Professor and Chairman, Department of Microbiology, Washington Uni
versity School of Medicine
FARMANFARMAIAN, A., Faculty of Medicine, Pahlavi University, Iran
FRIES, E. F. B., Professor of Biology, City College of New York
GABRIEL, MORDECAIL., Associate Professor of Biology, Brooklyn College
GINSBERG, HAROLD S., Professor and Chairman of the Department of Microbiology, University
of Pennsylvania
GLUSMAN, MURRAY, Assistant Professor, Columbia University
GamN, JAMESW., Professor of Physiology, Rutgers University
GREEN, MAURICE, Associate Professor of Microbiology, Saint Louis University
HANDLER, PHILIP, James B. Duke Professor of Biochemistry and Chairman of the Department
of Biochemistry, Duke University
HAUGAARD, NIEL5, Associate Professor of Pharmacology, University of Pennsylvania
HIATT, HOWARD H., Assistant Professor of Medicine, Harvard University
HURWITZ, CHARLES, Assistant Professor of Microbiology, VA Hospital, Albany
ISSELBACHER,KURTJ., Chief, Gastrointestinal Unit and Assistant Professor of Medicine, Massa
chusetts General Hospital and Harvard Medical School
JACOBS, MERKEL H., Professor Emeritus, University of Pennsylvania
KALTENBACH, JANE C., Assistant Professor of Zoology, Mount Holyoke College
KARNOVSKY, MANFRED L., Professor of Biological Chemistry, Harvard University
KEOSIAN, JOHN, Professor of Biology, Rutgers, The State University
KLEIN, MORTON, Professor of Microbiology, Temple University Medical School
KLOTZ, IRVING M., Professor of Chemistry and Biology, Northwestern University
LAUFFER, MAX A., Andrew Mellon Professor of Biophysics, University of Pittsburgh
LEIGHTON, JOSEPH, Professor of Pathology, University of Pittsburgh School of Medicine
LINEAWEAVER, THOMAS H., III, Marine Biological Laboratory
MARKS, PAUL A., Associate Professor of Medicine, Columbia University, College of Physicians
and Surgeons
MCDONALD, SISTER ELIZABETH SETON, Chairman of the Department of Biology, College of
Mt. St. Joseph
NEEDLEMAN, SAUL B., Research Biochemist and Instructor in Biochemistry, Abbott Laboratories
NOVIKOFF, ALEXANDER B., Research Professor, Albert Einstein College of Medicine
RAPPORT, MAURICE M., American Cancer Society Professor of Biochemistry, Albert Einstein
College of Medicine

REPORT OF THE DIRECTOR 25
ROSENBERG, EVELYN K., Associate Professor of Pathology, New York University Medical Center
ROTH, JAY S., Professor of Biochemistry, University of Connecticut
ROWLAND, LEWIS P., Assistant Professor of Neurology, Neurological Institute, Columbia Uni
versity
SCHENCK, J. R., Associate Research Fellow, Abbott Laboratories
SCHNEIDERMAN, HOWARD A., Professor and Chairman of the Department of Biology, Western
Reserve University
SPIEGEL, MELVIN, Associate Professor of Biology, Dartmouth College
SPIRTES,M. A., Associate Professor of Pharmacology, Hahnemann Medical College
STETTEN, DEWITT, JR., Dean of the Medical School, Rutgers, The State University
WARNER, ROBERT C., Professor of Biochemistry, New York University School of Medicine
WAINI0, WALTER W., Chairman of the Department of Physiology and Biochemistry, Rutgers,
The State University
WEISS, LEON, Associate Professor of Anatomy, The Johns Hopkins University School of Medi
cine
WHEELER, GEORGEE., Assistant Professor of Biology, Brooklyn College
WILSoN, THOMAS HASTINGS, Associate Professor of Physiology, Harvard Medical School
YNTEMA, CHESTER L., Professor of Anatomy, State University of New York, Upstate Medical
Center
ZACKS, SUMNER I., Assistant Professor of Pathology, Pennsylvania Hospital
Students, 1963
All students listed completed the formal course program, June 17—July 27. Asterisk indi
cates students completing Post-Course Research Program, July 28—August 31.
** Post-Course only.
ECOLOGY
*ARONSON, WENDY SUE, Vassar College
*BUOGELN, RICHARD G., University of Hawaii
DANIELS, CHARLES I., Duke University
*DEXTER, DEBORAH M., University of North Carolina
FEINGOLD, ALAN 0., Swarthmore College
GEBELEIN, CONRAD, The Johns Hopkins University
*HOLZAPFEL, CHRISTINE, Goucher College
KNOPF, GARRY N., University of Colorado
MYERS, JUDITH, Chatham College
O'BRIEN, JAMES F., Fordham University
*RAED, KENNETH J., Cornell University
*ROBINSON, MARY I., University of Chicago
STEEN, REBECCA H., Yale University
*THAYER, GORDON,Oberlin College
WEST, MARYJANE, University of Michigan
*Zusy, DENNIS R., Northwestern University
EMBRYOLOGY
*BATTIKH, HANI K., Qousair, Horns, Syria
BERGER, HILLARD, Vanderbilt University
*BURDICK,MORTONL., The Johns Hopkins University
CASSENS, GLORIA A., Indiana University
DAVID, JOHN R., New York University, School of Medicine
*DAVIDSON, RICHARD L., Western Reserve University
*EPSTEIN, MICHAEL J., University of Texas
*HARTLINE, DANIEL, Harvard University
*KIMMEL, CHARLES B., The Johns Hopkins University
*KOHNE, DAVID E., Purdue University
*MCWHINNIE, DOLORES, Marquette University

26 MARINE BIOLOGICAL LABORATORY
*PATTON, DOROTHY E., University of Chicago
POLLACK, ROBERT E., Brandeis University
*@r@EDER, RONALD H., Massachusetts Instituteof Technology
*SHOSTAK, STANLEY, Brown University
SPRING,ELINORJ., Harvard Medical School
*STACKHOUSE, HAMILTON LEE, University of Arizona
STEEN, TRYGVE, Yale University
*TERMAN,STANLEYA., Brown University
*YUYAMA, SHUHEI, Florida State University
BOTANY
BF.NADE, LEONARD E., George Mason College of University of Virginia
*CAMPI, JAMES R., Indiana University
*CAREFOOT, JOHN R, Indiana University
CLEMENTS, SUSAN D., Montana State College
@CoSTLow, JUDSON, Indiana University
Cox, EDMOND R., JR.,Middle Tennessee State College
HAMMACK, ROBERT E., University of Alabama
*HARRIS, DENNY 0., Indiana University
*H0DGE,MARTHAD., University of Michigan
JOHNSON, CHARLES W., Wabash College
KEHLENBACK, MRS. EDNA K., Syracuse University
*KNOX, JOHN, Drew University
*LIDDLE,LARRYB., University of Illinois
PERTZ, ANITA R., City College of New York
*RICHARDSON, WILLIAM N., Earlham College
*SAWA, TAKASHI, University of Louisville
*SCHOFIELD, EDMUND A., JR., Clark University
SHEN, EUGENE, University of Texas
*WALNE, PATRICIAL., University of Texas
WEINKAUFF, ANN MARY, Oberlin College
PHYSIOLOGY
*BARLOW, ROBERT B., JR., The Rockefeller Institute
*BOWNDS, MARLIN D., Harvard University
*COWNAN, ANNAMMA, Marquette University
CONNELLY, YVONNE, Northwestern University
DESA, RICHARD JOHN, University of Illinois
*ELZINGER, MARSHALL, University of Illinois
FRATANTONI, JOSEPH, Cornell Medical College
*GARRICK, LINDA S.,The Johns Hopkins University
*GREY, ROBERT D., Washington University
*HAHN, WILLIAM E., Tulane University
*HARRISON, STEPHEN C., Harvard University
*HAYNES, JULIAN F., Western Reserve University
*HILDEN, SHIRLEY, Stanford University
HUMPHREY, RICHARD, The Johns Hopkins Hospital
*JACOBSON, EUGENE D., Walter Reed Army Instituteof Research
*JAcKLETT, JON W., University of Oregon
*LEICHTUNG, BEN H., Northwestern University
MADRID, FELIX, University of Miami
*MITTENTHAL,JAY E., The Johns Hopkins University
NAUMAN, DOROTHY C., Harvard University
@PHILLIP5, ALLEN T., Michigan State University
PoNTIcoRvo, LAURA,Columbia University
SAHA, KUNAL, Calcutta, India
*SEDEROFF, RONALD R., University of California, Los Angeles

REPORT OF THE DIRECTOR 27
*SHAPER, JOEL H., University of California
*STEPHENS, RAYMOND E., University of Pittsburgh
TRIVUS, ROBERT H., Hahnemann Medical College
tTUNIS, MARY JANE, University of California
**MUESING, RICHARD
INVERTEBRATE ZOOLOGY
BARBOUR, STEPHEN D., Temple University
BENSON, WOODRUFF W., Emory University
BROWN, JAMES H., Cornell University
*BUSSER, JOHN H., University of Rhode Island
CARTER, EUGENIE D., Drew University
CAVICCHI, PHYLLIS A., Mt. Holyoke College
CHAMBERLAIN, ELIZABETH, American University
*CHENEY, SA.IIAH D., Earlham College
CONNELLY, SISTER MARY EDWARD, St. Bonaventure University
*Cp.AIG,NESSLYC., Reed College
DABNEY, MICHAEL, Oberlin College
DALSHEIMER, GEORGE H., The Johns Hopkins University
DESANTO, ROBERT S., Columbia University
*DILLARD, WALTER L., Oklahoma University
DOMINIQUE, SIsTER M., University of Notre Dame
DYSON, MARTHA M., Oberlin College
ETZLER, MARILYNN, Washington University
FEDER,NED, Harvard University
GRossI, JOSEPHC., JR., College of Steubenville
IMLAY, MARC J., American University
JENKINS, VIRGINIA, University of Massachusetts
JONES, HELEN E., Cornell University
KARSTADT, M@nIA L., University of California, Berkeley
LEE, SANDRA, Vassar College
*LESH, GEORGIA E., Western Reserve tjniversity
*LEVANDOWSKY, MICHAEL, City College of New York
LEVOWITZ, AMY, Queens College
LLOYD, MARGARET C., Bryn Mawr College
*LONG, NANCY D., University of California
MACNAB, ANN K., Rice University
*MOREAN, EDITH, University of Minnesota
MYREN, RICHARD T., Cornell University
POGANY, Gn.amT C., Tulane University
*SMITH, CHARLEs C., Oklahoma University
*SMITH, CLARE D., University of Arizona
STADLER, JOAN, University of Michigan
STERN, EDWARD L., University of Chicago
*SUMMERS, WILLIAM C., University of Minnesota
TEITELBAUM, DOROTHY J., Columbia University
VIANNEY, SISTER M. JEAN, University of Notre Dame
4. FELLOWSHIPS AND SCHOLARSHIPS, 1963
Bio Club Scholarship:
MICHAEL LEVANDOWSKY, Invertebrate Zoology Course
ANITA PERTZ, Botany Course
Gary N. Calkins Memorial Scholarship:
WILLIAM SUMMERS, Invertebrate Zoology Course
Lucretia Crocker Scholarship:
JOHN R. CAREFOOT, Botany Course
MARY I. ROBINSON, Ecology Course

28 MARINE BIOLOGICAL LABORATORY
Edwin Linton Memorial Endowment of the
Washington and Jefferson College:
JOHN BUSHNELL
B. G. Conklin Fund:
Turtox Scholarship Fund:
I. INSTRUCTORS
C. B. METZ
C. R. AUSTIN
A. MONROY
L. NELSON
II. LECTURES
C. A. SHIVERS
D. GROSCH
R. SAGER
E. ADELBERG
W. 0. NELSON
M. NASATIR
M. OLSEN
T. MANN
I. W. ROWLANDS
I. INSTRUCTORS
S. W. KUFFLER
D. D. POTTER
E. J. FURSHPAN
R. L. DAVIDSON,Embryology Course
D. E. KOHNE, Embryology Course
DOROTHY PATTON, Embryology Course
PATRICIA L. WALNE, Botany Course
5. TRAINING PROGRAMS
FERTILIZATION AND GAMETE PHYSIOLOGY TRAINING PROGRAM
(No lectures given—only seminars)
I. INSTRUCTORS
C. LADDPROSSER
G. KALEY
A. JANOFF
LEWIS KLEINHOLZ
B. ZWEIFACH
II. ASSISTANTS
JOHN MUNDAY
LINDA HABAS
ASIT DAS
Florida State University, in charge of program
Cambridge University
University of Palermo
Emory University
Immunological studies on fertilization in frogs
Reproductive processes in insects
Studies on sexuality in Chiamydomonas
Bacterial conjugation
Inhibition of spermatogenesis
Peculiar propagatory processes in plants
Parthenogenetic development in chicken and turkey eggs
Relation between the androgenic and spermatogenic activity of the
male gonad
Intraperitoneal insemination
NEUROPHYSIOLOGY TRAINING PROGRAM
Harvard Medical School, in charge of program
Harvard Medical School
Harvard Medical School
COMPARATIVE PHYSIOLOGY TRAINING PROGRAM
University of Illinois, in charge of program
New York University
New York University
Reed College
New York University
University of Illinois
University of Illinois
University of Illinois

III. LECTURES
*C L. PROSSER
*LEwIS KLEINHOLZ
GABOR KALEY
AARON JANOFF
*FRANK A. BROWN, JR.
*EUGENE COPELAND
*TALBOT WATERMAN
*W H. SAWYER
GROVER STEPHENS
*CLARK READ
CHESTER HYMAN
*BENJAMIN ZWEIFACH
G. DEVILLAFRANCA
*JOHN KANWISHER
T. H. BULLOCK
REPORT OF THE DIRECTOR
Physiological adaptation
Comparative physiology of muscle
Invertebrate endocrinology
Renin-angiotensin systems in control of peripheral circulation
Biochemistry of leucocytes
Biological rhythms
Reception of magnetic fields
Chloride- and other secreting cells
Arthropod vision
Comparative biochemistry of pituitary polypeptides
Utilization of dissolved organic material by invertebrates
Metabolism of endoparasites
Comparison of open and closed circulations
Contractile endothelial cells and control of microcirculation
Properties of contractile proteins from invertebrate muscles
Temperature tolerance of insects
Properties of cardiac ganglia of crustaceans
* Seminars only. The others were lecture-discussions or these plus seminars.
6. TABULAR VIEW OF ATTENDANCE, 1959-1963
1959
INVESTIGATORS—TOTAL 427
Independent 215
Under instruction 45
Library Readers 51
Research Assistants 116
STUDENTS—TOTAL 134
Invertebrate Zoology 55
Embryology 23
Physiology 27
Botany 20
Ecology 15
TOTAL ATTENDANCE 561
Less persons represented as both
investigators and students
INSTITUTIoNs REPRESENTED—TOTAL 143
By investigators 98
By students 73
SCHOOLS AND ACADEMIES REPRESENTED
By investigators 2
By students 12
FOREIGN INSTITUTIONS REPRESENTED 38
By investigators 29
By students 9
Abbott Laboratories
Alabama, University of
Albert Einstein Medical School
American Museum of Natural History
American University
AntiochCollege
7. INSTITUTIONS REPRESENTED, 1963
29
1960 1961 1962 1963
458 458 494 490
231 224 235 227
42 32 44 34
50 49 56 51
135 151 159 178
122 130 121 124
49 40 38 40
20 21 20 20
28 28 28 28
18 19 20 20
13 22 15 16
580 586 615 614
4 2 1 4 5
557 578 585 611 609
144 132 118 120
83 107 81 83
61 70 57 70
5 3 3 4
2 2
14 28 31 21
11 21 17 15
3 7 14 6
Middle Tennessee State College
Minnesota, University of
Montana State University
Mount Holyoke College
New York State University, College of Medi
cine at Brooklyn

30 MARINE BIOLOGICAL LABORATORY
Argonne National Laboratory
Arizona, University of
Brandeis University
Brooklyn College
Brown University
Bryn Mawr College
California, University of
Carnegie Institution
Chatham College
of Washington
Chicago Medical School
Chicago, University of
Cincinnati, University of
City College of New York
Clark University
College of Mt. St. Joseph on the Ohio
College of Steubenville
Colorado, University of
Columbia University
Columbia University, College of
Physiciansand Surgeons
Connecticut, University of
CornellUniversity
Cornell University Medical College
Dartmouth College
Dartmouth College Medical School
Delaware,University of
Drew University
Drexel Institute of Technology
Duke University
Earlham College
Emory University
Florida State University
Florida, University of
Fordham University
George Washington University,
School of Medicine
Georgetown University
Goucher College
Hahnemann Medical School
Harvard University
Harvard University Medical School
Hawaii, University of
Illinois, University of
Indiana University
Institute for Muscle Research
Johns Hopkins University
Johns Hopkins University, School of Medicine
Kent State University
Louisville, University of
Loyola College
Marquette University
Maryland, University of
Massachusetts General Hospital
Massachusetts Institute of Technology
Massachusetts, University of
Miami, University of
Michigan State University
Michigan, University of
New York State University, College of
Medicine at Syracuse
New York University, Bellevue Medical
Center
New York University, School of Dentistry
New York University, Washington Square
College
North Carolina State College
North Carolina, University of
Northwestern University
Notre Dame, University of
Oak Ridge National Laboratory
Oberlin College
Ohio State University
Oklahoma, University of
Oregon, University of
Pennsylvania, University of
Pennsylvania Medical School, University of
Pittsburgh, University of
Princeton University
Purdue University
Queens College
Reed College
Rhode Island, University of
Rice University
Rochester, University of, School of Medicine
Rockefeller Institute
Russell Sage College
Rutgers University
Saint Bonaventure University
Saint Louis University
Seton Hill College
Single Cell Research Foundation, Inc.
Smith College
Southwest Texas State College
Stanford University
State University of New York at Buffalo
Swarthmore College
Syracuse University
Temple University
Tennessee, University of
Texas, University of
Tulane University
Vanderbilt University
Vermont, University of
Veterans Administration Hospital
Virginia, University of
Wabash College
Walter Reed Army Institute of Research
Washington University
Washington, University of
Washington and Jefferson College
Washington University Medical School
Western Reserve University
Western Reserve University, School of
Meoicine
Wisconsin, University of
Yale University

REPORT OF THE DIRECTOR
FOREIGN INSTITUTIONS REPRESENTED
Facultédes Sciences, Paris
University of Liverpool, England
Seramport and Presidency College, Calcutta,
India
Biologische Anstalt Helgoland, Germany
University of Alberta, Edmonton, Canada
Agricultural Research Council of Great Britain
Faculty of Science, Dakar University, Republic
of Senegal, Africa
Kitasato Institute, Tokyo, Japan
Universitãt d'Saarlandes, Homburg/Saar,
Germany
National Institute for Medical Research,
London, England
Tokyo Medical and Dental University,
Japan
Ochanomizu University, Japan
Pahlavi, University of, Shiraz, Iran
Embassy of the Syrian Arab Republic,
Syria
McMaster University, Hamilton, Ontario
Physiological Laboratory, Cambridge,
England
Edinburgh University, Edinburgh, Scotland
University of Glasgow, Scotland
Inst. de Biologia, Mexico
University of Palermo, Italy
University of Sao Paulo, Brasil
SUPPORTING INSTITUTIONS, AGENCIES, AND INDIVIDUALS
Abbott Laboratories
Associates of the Marine Biological Laboratory
Atomic Energy Commission
CIBA Pharmaceutical Products, Inc.
The Commonwealth Fund
Josephine B. Crane Foundation
Duke University
Eli Lilly and Company
Dr. and Mrs. David W. Gaiser
The Grass Foundation
Mr. and Mrs. William H. Greer, Jr.
Hoffman-LaRoche, Inc.
Mr. and Mrs. George F. Jewett, Jr.
The Lalor Foundation
National Institutes of Health
National Science Foundation
Office of Naval Research
The Rockefeller Foundation
Dr. and Mrs. Henry W. Sandeen
Schering Foundation, Inc.
Scientific American, Inc.
E. R. Squibb and Sons
Gerald Swope, Jr.
The Upjohn Company
Wallace Laboratories
James H. Wickersham
8. FRIDAY EVENING LECTURES, 1963
July 5
TERU HAYASHI Contractile proteins and movement
Columbia University
July 12
THEODORE H. BULLOCK Ways and means of integrating IfI nervous
University of California at Los An- systems. I
geles
Alexander Forbes Lecturer at the MBL
July 15
Monday
THEODORE H. BULLOCK Ways and means of integrating in nervous
systems. II
July 19
LAWRENCE B. SLOBODKIN Experimental studies of population dynamics
University of Michigan
July 26
CHARLES DEVILLERS Mechanics of gastrulation
Université de Paris
Faculté de Science
(Sorbonne), France
31

32 MARINE BIOLOGICAL LABORATORY
August 2
ARTHUR B. PARDEE Regulation of biochemical reactions
Princeton University
August 9
ELVIN A. KABAT Immunochemical studies on blood group sub
Columbia Presbyterian Medical Center stances
August 16
MARTIN R. POLLOCK The physiology and evolution of bacterial peni
National Institute for Medical Re- cillinase
search, London, England
Senior Lalor Fellow at the MBL
August 23
PHILIP HANDLER The evolution of proteins
Duke University
July 2 RICHARD DE SA,
J. W. HASTINGSAND
A. E. VATTER
9. TUESDAY EVENING SEMINARS, 1963
JAMES SPUDICH AND
J. W. HASTINGS
LEONARD NELSON
July 9 WARNER E. LOVE AND
NEVENKA M. RUMEN
W. TROLL,
S. BELMAN AND
E. LEVINE
July 16 DOUGLAS MARSLAND
R. H. CHENEY
C. C. SPEIDEL
July 23 R. D. ALLEN
A. J. DELORENZO
July 30 P. PERSON AND
D. PHILPOTT
S. SIMPSON
The Scintillon—a new type of biological
particle: an intracellular crystal which
emits bioluminescent flashes
The inhibition of the bioluminescent oxida
tion of reduced flavin mononucleotide by
2-decenal
Spermatozoan cholinesterase
Heme-heme interaction in lamprey hemo
globin—an explanation
Modification of DNA by a mutagen and
carcinogens
A pressure analysis of the effects of heavy
water (D20) on the form, movement and
plasmagel structure of Amoeba proteus
Variations in resistance to radiation of Ar
bacia
ment
correlated with stages in develop
Motion picture showing comparative resist
ance of Arbacia zygotes to injury by
gamma or ultraviolet irradiation
Polarized light microscopy by means of a
birefringence
rangement
detector and scanning ar
Ultrastructure of a crayfish caudal photo
receptor under conditions of light and
darkness; a tale of hindsight
Limulus
tissue
gill cartilage: a “¿plant-like†animal
Analysis of lizard tail regeneration: the role
of the ependyma
M. M. RAPPORT Ebb-tide antigens in the brain

REPORT OF THE DIRECTOR 33
August 6 W. A. SIDDIQUI A helical structure in ribonucleoprotein
bodies of Entanzoeba invadens
MAX A. LAUFFER 1. A thermodynamic definition of hydration
2. Membrane equilibrium in multicomponent
systems
M. E. KRIEBEL Physiology of the tunicate heart (film)
August 13 RUTH HUBBARDAND Effect of flashes on light and dark adapta
JOHN E. DowI,ING tion
CLARA F. ARMSTRONG Morphological mechanism of intracellular
impulse conduction in striated muscle
A. B. NOVIKOFF Lysosomes in thyroid epithelium of un
treated, TSH-stimulated, and I'31-irradi
ated rats
HANS LAUFER, Nucleo-cytoplasmic interactions in the de
YASUKIYO NAKOSE AND velopment of the salivary gland of Chiro
JEROME VANDERBERG nomus thuinnzi (Diptera)
August 20 F. N. SUDAK Dynamics and energetics of the circulatory
system of dog fish
GEORGE SZABÔ, The pigmentary system of the squid
T. FITZPATRICK AND
G. WILGRAM
F. C. G. HOSKIN Stiniulation by specific quinones of glucose
metabolism by brain
F. M. CHILD Flagellar regeneration: a mechanism ac
counting for its initiation and regulation
10. MEMBERS OF THE CORPORATION, 1963
LIFE MEMBERS
ADOLPH, DR. EDWARD F., University of Rochester, School of Medicine and Den
tistry, Rochester, New York
BRODIE, MR. DONALD, 522 Fifth Avenue, New York 18, New York
COLE, DR. ELBERT C., 2 Chipman Park, Middlebury, Vermont
COWDRY, DR. E. V., 4580 Scott Avenue, St. Louis 10, Missouri
CRANE, MRS. W. Mu1UIAY, 820 Fifth Avenue, New York 21, New York
HF.SS, DR. WALTER, 737 Maple Street, Spartanburg, South Carolina
HISAw, DR. F. L., Biological Laboratories, Harvard University, Cambridge 38,
Massachusetts
JACOBS, DR. M. H., Department of Physiology, University of Pennsylvania, Phila
delphia 4, Pennsylvania
KNOWLTON, DR. F. P., c/o Mr. G. L. Gravelt, Jamesville, Rt. 2, New York
LEWIS, DR. W. H., Johns Hopkins University, Baltimore, Maryland
LOWTHER, DR. FLORENCE, Barnard College, New York, New York
MACDOUGALL, DR. MARY STUART, Mt. Vernon Apartments, 423 Clairniont Avenue,
Decatur, Georgia
MALONE, DR. E. F., 6610 North 11th Street, Philadelphia 26, Pennsylvania
MEANS, DR. J. H., 15 Chestnut Street, Boston, Massachusetts
MEDES, DR. GRACE, 303 Abington Avenue, Philadelphia 11, Pennsylvania

34 MARINE BIOLOGICAL LABORATORY
MOORE, DR. J. PERCY, RD No. 1, Box 347, Chapel Hill, North Carolina
PAYNE, DR. FERNANDUS, Indiana University, Bloomington, Indiana
PLOUGH, DR. H. H., Amherst College, Amherst, Massachusetts
PORTER, DR. H. C., University of Pennsylvania, Philadelphia, Pennsylvania
SCOTT, DR. ERNEST L., Columbia University, New York, New York
SCHRADER, DR. SALLY, Duke University, Durham, North Carolina
TURNER, DR. C. L., Northwestern University, Evanston, Illinois
WAITE, DR. F. G., 144 Locust Street, Dover, New Hampshire
WALLACE, DR. LOUISE B., 359 Lytton Avenue, Palo Alto, California
WARREN, DR. HERBERT 5., 2768 Egypt Road, Audubon, Pennsylvania
WHEDON, DR. A. D., 21 Lawncrest, Danbury, Connecticut
REGULAR MEMBERS
ABELL, DR. RICHARD G.,55 East 2nd Avenue, New York 28,New York
ADELMAN, DR. WILLIAM J., Department of Physiology, University of Maryland
Medical School, Baltimore 1, Maryland
ALBERT, DR. ALEXANDER, Mayo Clinic, Rochester, Minnesota
ALLEN, DR. M. JEAN, Department of Biology, Wilson College, Chambersburg,
Pennsylvania
ALLEN, DR. ROBERT D., Department of Biology, Princeton University, Princeton,
New Jersey
ALSCHER, DR. RUTH, Department of Physiology, Manhattanville College, Purchase,
New York
AMATNIEK, DR. ERNEST, 34 Homer Avenue, Hastings-on-the-Hudson, New York
AMBERSON, DR. WILLIAM R., Woods Hole, Massachusetts
ANDERSON, DR. J. M., Department of Zoology, Cornell University, Ithaca, New
York
ANDERSON, DR. RUBERT S., Medical Laboratories, Army Chemical Center, Mary
land (Send mail to Box 632, Edgewood, Maryland)
ARMSTRONG, DR. PHILIP B., Department of Anatomy, State University of New
York, College of Medicine, Syracuse 10, New York
ARNOLD, DR. WILLIAM A., Division of Biology, Oak Ridge National Laboratory,
Oak Ridge, Tennessee
ATWOOD, DR. KIMBALL C., 702 West Pennsylvania Avenue, Urbana, Illinois.
AUCLAIR, DR. WALTER, Department of Biological Sciences, University of Cincinnati,
Cincinnati 21, Ohio
AUSTIN, DR. C. R., Physiological Laboratory, Downing Street, Cambridge, England
AUSTIN, DR. MARY L., 506k North Indiana Avenue, Bloomington, Indiana
AYERS, DR. JOHN C., Department of Zoology, University of Michigan, Ann Arbor,
Michigan
BAITSELL, DR. GEORGE A., Osborn Zoological Laboratories, Yale University, New
Haven, Connecticut
BALL, DR. ERIC G., Department of Biological Chemistry, Harvard University Med
ical School, Boston 15, Massachusetts
BALLARD, DR. WILLIAM W., Department of Zoology, Dartmouth College, Hanover,
New Hampshire

REPORT OF THE DIRECTOR 35
BALTUS, DR. ELYANE, Laboratoire de Morphologie Animale, 1850 ChausSée de
Wavre, Bruxelles 16, Belgique
BANG, DR. F. B., Department of Pathobiology, Johns Hopkins University School of
Hygiene, Baltimore 5, Maryland
BARD, DR. PHILLIP, Johns Hopkins Medical School, Baltimore, Maryland
BARTH, DR. L. G., Department of Zoology, Columbia University, New York 27,
New York
BARTH, DR. LUCENA, Department of Zoology, Barnard College, New York 27,
New York
BARTLETT, DR. JAMES H., Department of Physics, University of Illinois, Urbana,
Illinois
BAYLOR, DR. E. A., Woods Hole Oceanographic Institution, Woods Hole, Massa
chusetts
BAYLOR, DR. MARTHA B., Marine Biological Laboratory, Woods Hole, Massa
chusetts
BEAMS, DR. HAROLD W., Department of Zoology, State University of Iowa, Iowa
City, Iowa
BECK, DR. L. V., Department of Pharmacology, Indiana University, School of
Experimental Medicine, Bloomington, Indiana
BEHRE, DR. ELINOR M., Black Mountain, North Carolina
BENNETT, DR. MICHAEL V., Department of Neurology, College of Physicians and
Surgeons, New York 32, New York
BENNETT, DR. MIRIAM F., Department of Biology, Sweet Briar College, Sweet
Briar, Virginia
BERG, DR. WILLIAM E., Department of Zoology, University of California, Berkeley
4, California
BERMAN, DR. MONES, Institute for Arthritis and Metabolic Diseases, National In
stitutes of Health, Bethesda 14, Maryland
BERNHEIMER, DR. ALAN W., New York University College of Medicine, New
York 16, New York
BERNSTEIN, DR. MAURICE, Department of Anatomy, Wayne State University Col
lege of Medicine, Detroit 7, Michigan
BERTHOLF, DR. LLOYD M., Illinois Wesleyan University, Bloomington, Illinois
BEVELANDER, DR. GERRIT, Medical Center, University of Texas, Dental Branch,
Houston, Texas 77025
BIGELOW, DR. HENRY B., Museum of Comparative Zoology, Harvard University,
Cambridge 38, Massachusetts
BISHOP, DR. DAVID W., Department of Embryology, Carnegie Institution of Wash
ington, 115 West University Parkway, Baltimore 10, Maryland
BLANCHARD, DR. K. C., Johns Hopkins Medical School, Baltimore 5, Maryland
BLOCH, DR. ROBERT, 518 South 42nd Street, Apt. C7, Philadelphia 4, Pennsylvania
BLUM, DR. HAROLD F., Department of Biology, Princeton University, Princeton,
New Jersey
BODANSKY, DR. OSCAR, Department of Biochemistry, Memorial Cancer Center,
444 East 68th Street, New York 21, New York
BODIAN, DR. DAVID, Department of Anatomy, Johns Hopkins University, 709
North Wolfe Street, Baltimore 5, Maryland

36 MARINE BIOLOGICAL LABORATORY
BOELL, DR. EDGAR J., Osborn Zoological Laboratories, Yale University, New
Haven, Connecticut
BOETTIGER, DR. EDWARD G., Department of Zoology, University of Connecticut,
Storrs, Connecticut
Bow, DR. HAROLDC., Department of Botany, University of Texas, Austin, Texas
BOREI, DR. HANS G., Department of Zoology, University of Pennsylvania, Phila
deiphia 4, Pennsylvania
BOWEN, DR. VAUGHAN T., Woods Hole Oceanographic Institution, Woods Hole,
Massachusetts
BRADLEY, DR. HAROLD C., 2639 Durant Avenue, Berkeley 4, California
BRIDGMAN, DR. ANNA J., Department of Biology, Agnes Scott College, Decatur,
Georgia
BRINLEY, DR. F. J., JR., Department of Physiology, Johns Hopkins Medical School,
Baltimore 5, Maryland
BRONK, DR. DETLEV W., Rockefeller Institute, 66th Street and York Avenue,
New York 21, New York
BROOKS, DR. MATILDA M., Department of Physiology, University of California,
Berkeley 4, California
BROWN, DR. DUGALD E. S., Department of Zoology, University of Michigan, Ann
Arbor, Michigan
BROWN, DR. FRANK A., JR., Department of Biological Sciences, Northwestern Uni
versity, Evanston, Illinois
BROWNELL, DR. KATHERINE A., Department of Physiology, Ohio State University,
Columbus, Ohio
BUCK, DR. JOHN B., Laboratory of Physical Biology, National Institutes of Health,
Bethesda 14, Maryland
BULLOCK, DR. T. H., Department of Zoology, University of California, Los Angeles
24, California
BURBANCK, DR. WILLIAM D., Emory University, Box 15134, Atlanta, Georgia
30333
BURDICK, DR. C. LALOR, The Lalor Foundation, 4400 Lancaster Pike, Wilmington,
Delaware
BURKENROAD, DR. M. D., 3169 Bremerton P1., La Jolla, Calif. 92037
BUTLER, DR. E. G., Department of Biology, P. 0. Box 704, Princeton University,
Princeton, New Jersey
CAMERON, DR. J. A., Baylor College of Dentistry, Dallas, Texas
CANTONI, DR. GIULLIO, National Institutes of Health, Mental Health, Bethesda 14,
Maryland
CARLSON, DR. FRANCIS D., Department of Biophysics, Johns Hopkins University,
Baltimore 18, Maryland
CARPENTER, DR. RUSSELL L., Tufts University, Medford 55, Massachusetts
CARRIKER, DR. MELBOURNE R., Marine Biological Laboratory, Woods Hole,
Massachusetts
CARSON, MISS RACHEL, 11701 Berwick Road, Silver Spring, Maryland
CASE, DR. JAMES, Department of Biology, University of California, Santa Barbara,
Goleta, California
CATTELL, DR. MCKEEN, Cornell University Medical College, 1300 York Avenue,
New York 21, New York

REPORT OF THE DIRECTOR 37
CHAET, DR. ALFRED B., Department of Biology, American University, Washington
16,D.C.
CHAMBERS, DR. EDWARD, Department of Physiology, University of Miami Medical
School, Coral Gables, Florida
CHANG, DR. JOSEPH J., Inst. f. physikal Chemie an der Techn. Hochscule, Aachen,
Germany
CHASE, DR. AURIN M., Department of Biology, Princeton University, Princeton,
New Jersey
CHENEY, DR. RALPH H., Biological Laboratory, Brooklyn College, Brooklyn 10,
New York
CHILD, DR. FRANK M., Department of Zoology, University of Chicago, Chicago 37,
Illinois
CLAFF, DR. C. LLOYD, 5 Van Beal Road, Randolph, Massachusetts
CLARK, DR. A. M., Department of Biological Sciences, University of Delaware,
Newark, Delaware
CLARK, DR. ELOISE E., Department of Zoology, Columbia University, New York
27, New York
CLARK, DR. E. R., 315 5. 41st Street, Philadelphia 4, Pennsylvania
CLARK, DR. LEONARD B., Department of Biology, Union College, Schenectady,
New York
CLARKE, DR. GEORGE L., Biological Laboratories, Harvard University, Cambridge
38, Massachusetts
CLELAND, DR. RALPH E., Department of Botany, Indiana University, Bloomington,
Indiana
CLEMENT, DR. A. C., Department of Biology, Emory University, Atlanta 22,
Georgia
COHEN, DR. SEYMOUR S., Department of Chemistry, University of Pennsylvania
School of Medicine, Philadelphia, Pennsylvania
COLE, DR. KENNETH S., (NINDB), National Institutes of Health, Bethesda 14,
Maryland
COLLETT, DR. MARY E., 34 Weston Road, Wellesley 81, Massachusetts
COLLIER, DR. JACK R., Marine Biological Laboratory, Woods Hole, Massachusetts
COLTON, DR. H. S., Box 699, Flagstaff, Arizona
COLWIN, DR. ARTHUR L., Department of Biology, Queens College, Flushing, New
York
COLWIN, DR. LAURA H., Department of Biology, Queens College, Flushing, New
York
COOPER, DR. KENNETH W., Department of Cytology, Dartmouth Medical School,
Hanover, New Hampshire
COOPERSTEIN, DR. SHERWIN J., Department of Anatomy, Western Reserve Univer
sity Medical School, Cleveland, Ohio
COPELAND, DR. EUGENE, Zoology Department, Tulane University, New Orleans,
Louisiana 70185
COPELAND, DR. MANTON, Bowdoin College, Brunswick, Maine
CORNMAN, DR. IVOR, Hazleton Laboratories, Box 333, Falls Church, Virginia
COSTELLO, DR. DONALD P., Department of Zoology, University of North Carolina,
Chapel Hill, North Carolina

38 MARINE BIOLOGICAL LABORATORY
COSTELLO, DR. HELEN MILLER, Department of Zoology, University of North Caro
lina, Chapel Hill, North Carolina
CRANE, MR. JOHN 0., Woods Hole, Massachusetts
CRANE, DR. ROBERT K., Department of Biochemistry, The Chicago Medical School,
Chicago 12, Illinois
CROASDALE, DR. HANNAH T., Dartmouth College, Hanover, New Hampshire
CROUSE, DR. HELEN V., Department of Botany, Columbia University, New York
27, New York
CROWELL, DR. SEARS, Department of Zoology, Indiana University, Bloomington,
Indiana
CSAPO, DR. ARPAD I., Rockefeller Institute, 66th Street and York Avenue, New
York 21, New York
CURTIS, DR. MAYNIE R., Box 8215, University Branch, Coral Gables 46, Florida
CURTIS, DR. W. C., 504 Westmount Avenue, Columbia, Missouri
DAN, DR. JEAN CLARK, Misaki Biological Station, Misaki, Japan
DAN, DR. KATSUMA, Misaki Biological Station, Misaki, Japan
DANIELLI, DR. JAMES F., Department of Medicinal Chemistry, University of Buf
falo School of Pharmacy, Buffalo 14, New York
DAVIS, DR. BERNARD D., Harvard Medical School, 25 Shattuck Street, Boston 15,
Massachusetts
DAWSON, DR. A. B., Biological Laboratories, Harvard University, Cambridge 38,
Massachusetts
DAWSON, DR. J. A., 129 Violet Avenue, Floral Park, Long Island, New York
DEANE, DR. HELEN W., Albert Einstein College of Medicine, New York 61, New
York
DETTBARN, DR. WOLF-DIETRICH, Department of Neurology, College of Physicians
and Surgeons, New York 32, New York
DILLER, DR. IRENE C., Institute for Cancer Research, Fox Chase, Philadelphia 11,
Pennsylvania
DILLER, DR. WILLIAM F., 2417 Fairhill Avenue, Glenside, Pennsylvania
DODDS, DR. G. S., West Virginia University School of Medicine, Morgantown,
West Virginia
DOLLEY, DR. WILLIAM L., Trevillans, Virginia
DOOLITTLE, DR. R. F., Chemistry II, Karolinska Institute, Stockholm 60, Sweden
DOWBEN, DR. ROBERT, Biology Department, Massachusetts Institute of Technology,
Cambridge, Massachusetts
DURYEE, DR. WILLIAM R., George Washington University, 2300 K Street N.W.,
Washington, D. C.
EBERT, DR. JAMES DAVID, Director, Carnegie Institute of Washington, 115 W.
University Parkway, Baltimore 10, Maryland
ECKERT, DR. ROGER 0., Department of Zoology, Syracuse University, Syracuse 10,
New York
EDDS, DR. MAC V., JR., Department of Biology, Brown University, Providence 12,
Rhode Island
EDWARDS, DR. CHARLES, Department of Physiology, University of Minnesota,
Minneapolis 14, Minnesota
EICHEL, DR. HERBERT J., Department of Biological Chemistry, Hahnemann Medical
College, Philadelphia, Pennsylvania

REPORT OF THE DIRECTOR 39
EISEN, DR. HERMAN, Department of Medicine, Washington University, St. Louis,
Missouri
ELLIOTT, DR. ALFRED M., Department of Zoology, University of Michigan, Ann
Arbor, Michigan
ESSNER, DR. EDWARD S., Department of Pathology, Albert Einstein College of
Medicine, New York 61, New York
EVANS, DR. TITUS C., State University of Iowa College of Medicine, Iowa City,
Iowa
FAILLA, DR. P. M., Radiological Physics Division, Argonne National Laboratory,
Argonne, Illinois
FARMANFARMAIAN, DR. ALLAHvERDI, Professor of General Physiology, Faculty of
Medicine, Pahiavi University, Shiraz, Iran
FAURE-FREMIET, DR. EMMANUEL, College de France, Paris, France
FAWCETT, DR. D. W., Department of Anatomy, Harvard Medical School, Boston
15, Massachusetts
FERGUSON, DR. F. P., Division of General Medical Sciences, National Institutes
of Health, Bethesda 14, Maryland
FERGUSON, DR. JAMEs K. W., Connought Laboratories, University of Toronto,
Toronto, Ontario, Canada
FIGGE, DR. F. H. J., University of Maryland Medical School, Lombard and Green
Streets,Baltimore1,Maryland
FINGERMAN, DR. MILTON, Department of Zoology, Newcomb College, Tulane Uni
versity, New Orleans 18, Louisiana
FISCHER, DR. ERNST, Department of Physiology, Medical College of Virginia,
Richmond, Virginia
FISHER, DR. FRANK M., JR., Department of Biology, Rice University, Houston 1,
Texas
FISHER, DR. JEANNE M., Department of Biochemistry, University of Toronto,
Toronto, Ontario, Canada
FISHER, DR. KENNETH C., Department of Biology, University of Toronto, Toronto,
Ontario, Canada
FORBES, DR. ALEXANDER, 16 Divinity Avenue, Cambridge, Massachusetts
FRAENKEL, DR. GOTTFRIED S., Department of Entomology, University of Illinois,
Urbana, Illinois
FREYGANG, DR. WALTER H., JR., 6247 29th St. N.W., Washington 15, D. C.
FRIES, DR. ERIK F. B., Box 605, Woods Hole, Masachusetts
FUORTES, DR. MICHAEL C. F., NINDB, National Institutes of Health, Bethesda 14,
Maryland
FURSHPAN, DR. EDWIN J., Department of Neurophysiology, Harvard Medical
School, Boston 15, Massachusetts
FURTH, DR. JACOB, 99 Fort Washington Avenue, New York 32, New York
FYE, DR. PAUL M., Woods Hole Oceanographic Institution, Woods Hole, Massa
chusetts
GABRIEL, DR. MORDECAI,Departnient of Biology, Brooklyn College, Brooklyn 10,
New York
GAFFRON, DR. HANS, Department of Biology, Florida State University, Institute of
Molecular Biophysics, Tallahassee, Florida

40 MARINE BIOLOGICAL LABORATORY
GALL, DR. JOSEPH G., Department of Zoology, University of Minnesota, Minne
apolis 14, Minnesota
GALTSOFF, DR. PAUL S., Woods Hole, Massachusetts
GILBERT, DR. DANIEL L., National Institutes of Health, Laboratory of Biophysics,
NINDB, Bethesda 14, Maryland
GILMAN, DR. LAUREN C., Department of Zoology, University of Miami, Coral
Gables, Florida
GINSBERG, DR. HAROLD S., Department of Microbiology, University of Pennsyl
vania School of Medicine, Philadelphia 4, Pennsylvania
GOLDSMITH, DR. TIMOTHY H., Department of Zoology, Yale University, New
Haven, Connecticut
GOLDSTEIN, DR. LESTER, Department of Zoology, University of Pennsylvania, Phila
delphia 4, Pennsylvania
GOODCHILD, DR. CHAUNCEY G., Department of Biology, Emory University, Atlanta
22, Georgia
GOODRICH, DR. H. B., Department of Biology, Wesleyan University, Middletown,
Connecticut
GOTSCHALL, DR. GERTRUDE Y., 315 East 68th Street, Apt. 9M, New York, New
York 10021
GRAHAM, DR. HERBERT, U. S. Fish and Wildlife Service, Woods Hole, Massa
chusetts
GRAND, MR. C. G., Cancer Institute of Miami, 1155 N. W. 15th Street, Miami,
Florida
GRANT, DR. PHILIP, National Science Foundation, 1951 Constitution Avenue,
Washington25,D. C.
GRAY, DR. IRVING E., Department of Zoology, Duke University, Durham, North
Carolina
GREEN, DR. JAMES W., Department of Physiology, Rutgers University, New Bruns
wick, New Jersey
GREEN, DR. MAURICE, Department of Microbiology, St. Louis University Medical
School, St. Louis, Missouri
GREGG, DR. JAMES H., Department of Biological Sciences, University of Florida,
Gainesville, Florida
GREGG, DR. JOHN
Carolina
R., Department of Zoology, Duke University, Durham, North
GREIF, DR. ROGER L., Department of Physiology, Cornell University Medical Col
lege, New York 21, New York
GRIFFIN, DR. DONALD F., Biological Laboratories, Harvard University, Cambridge
38, Massachusetts
GROSCH, DR. DANIEL S., Department of Genetics, Gardner Hall, North Carolina
State College, Raleigh, North Carolina
GROSS, DR.. PAUL, Department
Island
of Biology, Brown University, Providence 12, Rhode
GRUNDFEST, DR. HARRY, College of Physicians and Surgeons, Columbia University,
New York 32, New York
GUTTMAN, DR. RITA, Department of Physiology, Brooklyn College, Brooklyn 10,
New York
GWILLIAM, DR. G. F., Department of Biology, Reed College, Portland 2, Oregon

REPORT OF THE DIRECTOR 41
HAJDU, DR. STEPHEN, National Institutes of Health, Bethesda 14, Maryland
HALL, DR. FRANK G., Department of Physiology, Duke University Medical School,
Durham, North Carolina
HAMBURGER, DR. VIKTOR, Department of Zoology, Washington University, St.
Louis, Missouri
HAMILTON, DR. HOWARD L., Department of Biology, University of Virginia, Char
lottesville, Va.
HANCE, DR. ROBERT T., RR #3, 6609 Smith Road, Loveland, Ohio
HARDING, DR. CLIFFORD V., JR., Columbia University, College of Physicians &
Surgeons,630 W. 168thSt.,New York, New York 10032
HARNLEY, DR. MORRIS H., Washington Square College, New York University,
New York 3, New York
HARTLINE, DR. H. KEFFER, Rockefeller Institute, 66th Street and York Avenue,
New York 21, New York
HARTMAN, DR. FRANK A., Ohio State University, Hamilton Hall, Columbus, Ohio
HARTMAN, DR. P. E., Department of Biology, Johns Hopkins University, Baltimore
18, Maryland
HARVEY, DR. ETHEL BROWNE, Marine Biological Laboratory, Woods Hole, Massa
chusetts
HASTINGS, DR. J. WOODLAND, Division of Biochemistry, University of Illinois,
Urbana, Illinois
HAUSCHKA, DR. T. S., Roswell Park Memorial Institute, 666 Elm Street, Buffalo 3,
New York
HAXO, DR. FRANCIS T., Division of Marine Botany, Scripps Institution of Ocean
ography, University of California, La Jolla, California
HAYASHI, DR. TERU, Department of Zoology, Columbia University, New York 27,
New York
HAYDEN, DR. MARGARET A., 34 Weston Road, Wellesley 81, Massachusetts
HAYWOOD, DR. CHARLOTTE, P. 0. Box 14, South Hadley, Massachusetts
HENDLEY, DR. CHARLES D., 615 South Avenue, Highland Park, New Jersey
HENLEY, DR. CATHERINE, Department of Zoology, University of North Carolina,
Chapel Hill, North Carolina
HERNDON, DR. WALTER R., Botany Department, University of Tennessee, Knox
ville, Tennessee
HERVEY, MR. JOHN P., Box 735, Woods Hole, Massachusetts
HESSLER, DR. ANITA Y., Marine Biological Laboratory, Woods Hole, Massa
chusetts
HIATT, DR. HOWARD H., Department of Medicine, Harvard Medical School, Bos
ton 15, Massachusetts
HIBBARD, DR. HOPE, Department of Zoology, Oberlin College, Oberlin, Ohio
HILL, DR. SAMUEL E., 135 Brunswick Road, Troy, New York
HIRSHFIELD, DR. HENRY I., Department of Biology, New York University, Wash
ington Square College, New York 3, New York
HOADLEY, DR. LEIGH, Biological Laboratories, Harvard University, Cambridge 38,
Massachusetts
HODES, DR. ROBERT, Department of Pediatrics, Mount Sinai Hospital, New York
39, New York

42 MARINE BIOLOGICAL LABORATORY
HODGE, DR. CHARLES, IV, Department of Biology, Temple University, Philadelphia
22, Pennsylvania
HOFFMAN, DR. JOSEPH, National Heart Institute, National Institutes of Health,
Bethesda 14, Maryland
HOLLAENDER, DR. ALEXANDER, Biology Division, Oak Ridge National Laboratory,
Oak Ridge, Tennessee
HOLZ, DR. GEORGE G., JR., Dept. of Microbiology, S. U. N. Y., Upstate Medical
Center, Syracuse, New York
HOPKINS, DR. HOYT 5., 59 Heatherdell Road, Ardsley, New York
HOSKIN, DR. FRANCIS C. G., Department of Neurology, College of Physicians &
Surgeons, New York 32, New York
HUNTER, DR., FRANCIS R., University of the Andes, Calle 18-a Carrera I-E,
Bogota, Colombia, South America
HUNTER, DR. W. D. R., Department of Zoology, Lyman Hall, Syracuse University,
Syracuse 10, New York
HURWITZ, DR. J., Department of Molecular Biology, Albert Einstein College of
Medicine, Eastchester Road and Morris Park Ave., Bronx, New York 10461
HUTCHENS, DR. JOHN E., Department of Physiology, University of Chicago, Chi
cago 37, Illinois
HYDE, DR. BEAL B., Department of Plant Sciences, University of Oklahoma, Nor
man, Oklahoma
HYMAN, DR. LIBBIE H., American Museum of Natural History, Central Park West
at 79th Street, New York 24, New York
IN0UE, DR. S., Department of Cytology, Dartmouth Medical School, Hanover,
New Hampshire
IRvING, DR. LAURENCE, Laboratory of Zoophysiology, University of Alaska, Col
lege, Alaska
ISENBERG, DR. IRVIN, Institute for Muscle Research, Marine Biological Labora
tory, Woods Hole, Massachusetts
ISELIN, MR. COLUMBUS O'D., Woods Hole, Massachusetts
ISSELBACKER,
chusetts
DR. KURT J., Massachusetts General Hospital, Boston 14, Massa
JENNER, DR.CHARLES E., Departmentof Zoology,Universityof NorthCarolina,
Chapel Hill, North Carolina
JOHNSON, DR. FRANK H., Department of Biology, Princeton University, Princeton,
New Jersey
JONES, DR. E. RUFFIN, JR., Department of Biological Sciences, University of
Florida, Gainesville, Florida
JONES, DR. RAYMOND F., Department of Biology, Princeton University, Princeton,
New Jersey
JOSEPHSON, DR. R. K., Department of Zoology, University of Minnesota, Minne
apolis 14, Minnesota
KAAN, DR. HELEN W., Marine Biological Laboratory, Woods Hole, Massachusetts
KABAT, DR. E. A., Neurological Institute, College of Physicians and Surgeons,
New York 32, New York
KAMINER, DR. B., Institute for Muscle Research, Marine Biological Laboratory,
Woods Hole, Massachusetts

REPORT OF THE DIRECTOR 43
KANE, DR. ROBERT E., Department of Cytology, Dartmouth Medical School,
Hanover, New Hampshire
KARUSH, DR. FRED, Department of Microbiology, University of Pennsylvania
School of Medicine, Philadelphia 4, Pennsylvania
KAUFMAN, DR. B. P., Department of Zoology, University of Michigan, Ann Arbor,
Michigan
KEMP, DR. NORMAN E., Department of Zoology, University of Michigan, Ann
Arbor, Michigan
KEMPTON, DR. RUDOLF T., Department of Zoology, Vassar College, Poughkeepsie,
New York
KEOSIAN, DR. JOHN, Department of Biology, Rutgers University, Newark 2, New
Jersey
KETCHUM, DR. BosTwIcK H., Woods Hole Oceanographic Institution, Woods
Hole, Massachusetts
KEYNAN, DR. ALEXANDER, Institute for Biological Research, Ness-Ziona, Israel
KILLE, DR. FRANK R., State Department of Education, Albany 1, New York
KIND, DR. C. ALBERT, Department of Zoology, University of Connecticut, Storrs.
Connecticut
KINDRED, DR. J. E., University of Virginia, Charlottesville, Virginia
KING, DR. ROBERT L., State University of Iowa, Iowa City, Iowa
KINGSBURY, DR. JOHN M., Department of Botany, Cornell University, Ithaca,
New York
KISCH, DR. BRUNO, 71 Maple Street, Brooklyn 25, New York
KLEIN, DR. MORTON, Department of Microbiology, Temple University, Philadel
phia, Pennsylvania
KLEINHOLZ, DR. LEWIS H., Department of Biology, Reed College, Portland 2,
Oregon
KLOTZ, DR. I. M., Department of Chemistry, Northwestern University, Evanston,
Illinois
KOLIN, DR. ALEXANDER, Department of Biophysics, California Medical School,,
Los Angeles 24, California
KORR, DR. I. M., Department of Physiology, Kirksville College of Osteopathy,.
Kirksville, Missouri
KRAHL, DR. M. E., Department of Physiology, University of Chicago, Chicago 37,.
Illinois
KRANE, DR. STEPHEN M., Massachusetts General Hospital, Boston 14, Massa
chusetts
KRAUSS, DR. ROBERT, Department of Botany, University of Maryland, Baltimore,
Maryland
KREIG, DR. WENDELL J. S., 303 East Chicago Avenue, Chicago, Illinois
KUFFLER, DR. STEPHEN W., Department of Pharmacology, Harvard Medical
School, Boston 15, Massachusetts
KUNITZ, DR. MOSES, Rockefeller Institute, 66th Street and York Avenue, New
York 21, New York
LAMY, DR. FRANCOIS, Department of Anatomy, University of Pittsburgh School
of Medicine, Pittsburgh 13, Pennsylvania
LANCEFIELD, DR. D. E., Queens College, Flushing, New York

44 MARINE BIOLOGICAL LABORATORY
LANCEFIELD, DR. REBECCA C., Rockefeller Institute, 66th Street and York Aye
nue, New York 21, New York
LANDIS, DR. E. M., Harvard Medical School, Boston 15, Massachusetts
LANSING, DR. ALBERT I., Department of Anatomy, University of Pittsburgh Medi
cal School, Pittsburgh 13, Pennsylvania
LASH, DR. JAMES W., Department of Anatomy, University of Pennsylvania School
of Medicine, Philadelphia 4, Pennsylvania
LAUFER, DR. H., Department of Biology, Johns Hopkins University, Baltimore 18,
Maryland
LAUFFER, DR. MAX A., Department of Biophysics, University of Pittsburgh, Pitts
burgh 13, Pennsylvania
LAWLER, DR. H. CLAIRE, Department of Biochemistry and Neurology, College of
Physicians and Surgeons, New York 32, New York
LAVIN, DR. GEORGE I., 6200 Norvo Road, Baltimore 7, Maryland
LAZAROW, DR. ARNOLD, Department of Anatomy, University of Minnesota Medical
School, Minneapolis 14, Minnesota
LEDERBERG, DR. JOSHUA, Department of Genetics, Stanford Medical School, Palo
Alto, California
LEE, DR. RICHARD E., Cornell University College of Medicine, New York 21, New
York
LEFEVRE, DR. PAUL G., University of Louisville School of Medicine, Louisville,
Kentucky
LEHMANN, DR. FRITZ, Zoologische Institut, University of Berne, Berne, Switzer
land
LEVINE, DR. RACHMIEL, New York Medical College, Department of Medicine,
5th Avenue at 106th Street, New York 29, New York
LEVY, DR. MILTON, Department of Biochemistry, New York University School of
Dentistry, New York 10, New York
LEWIN, DR. RALPH A., Scripps Institution of Oceanography, La Jolla, California
LEWIS, DR. HERMAN W., Genetic Biology Program, National Science Foundation,
Washington 25, D. C.
LEWIS, DR. IVEY F., 800 Rugby Road, Charlottesville, Virginia
LING, DR. GILBERT, 307 Berkeley Road, Merion, Pennsylvania
LITTLE, DR. E. P., 216 Highland Street, West Newton, Massachusetts
LLOYD, DR. DAVID P. C., Rockefeller Institute, 66th Street and York Avenue, New
York 21, New York
LOCHHEAD, DR. JOHN H., Department of Zoology, University of Vermont, Burling
ton, Vermont
LOEB, DR. R. F., 950 Park Avenue, New York 28, New York
LOEWENSTEIN, DR. WERNER R., Department of Physiology, College of Physicians
and Surgeons, New York 32, New York
LOFTFIELD, DR. ROBERT B., Massachusetts General Hospital, Boston, Massachu
setts
LORAND, DR. LASZLO, Department of Chemistry, Northwestern University, Evans
ton, Illinois
DE LORENZO, DR. ANTHONY, Anatomical and Pathological Research Laboratories,
Johns Hopkins Hospital, Baltimore 5, Maryland

REPORT OF THE DIRECTOR 45
LOVE, DR. WARNER E., 1043 Marlau Drive, Baltimore 12, Maryland
LUBIN, DR. MARTIN, Department of Pharmacology, Harvard Medical School,
Bostorr 15, Massachusetts
LYNCH, DR. CLARA J., Rockefeller Institute, 66th Street and York Avenue, New
York 21, New York
LYNN, DR. W. GARDNER, Department of Biology, Catholic University of America,
Washington 17, D. C.
MCCANN, DR. FRANCES, Department of Physiology, Dartmouth Medical School,
Hanover, New Hampshire
MCCOUCH, DR. MARGARET SUMWALT, University of Pennsylvania Medical School,
Philadelphia 4, Pennsylvania
MCDONALD, SISTER ELIZABETH SETON, Department of Biology, College of Mt. St.
Joseph, Mt. St. Joseph, Ohio
MCDONALD, DR. MARGARET R., Waldermar Medical Research Foundation, Sunny
side Blvd. and Waldermar Road, Woodbury R.D., New York
MCELROY, DR. WILLIAM D., Department of Biology, Johns Hopkins University,
Baltimore 18, Maryland
MAAS, DR. WERNER K., New York University College of Medicine, New York
City, New York
MAGRUDER, DR. SAMUEL R., Department of Anatomy, Tufts Medical School, 135
Harrison Avenue, Boston, Massachusetts
MANWELL, DR. REGINALD D., Department of Zoology, Syracuse University, Syra
cuse 10, New York
MARKS, DR. PAUL A., Columbia University, College of Physicians and Surgeons,
New York 32, New York
MARSHAK, DR. ALFRED, Departnient of Radiology, Jefferson Medical College,
Philadelphia 7, Pennsylvania
MARSLAND, DR. DOUGLAS A., 48 Church Street, Woods Hole, Massachusetts
MARTIN, DR. EARL A., 682 Rudder Road, Naples, Florida 33940
MATHEWS, DR. SAMUEL A., Thompson Biological Laboratory, Williams College,
Williamstown, Massachusetts
MAZIA, DR. DANIEL, Department of Zoology, University of California, Berkeley 4,
California
MEINKOTH, DR. NORMAN, Department of Biology, Swarthmore College, Swarth
more, Pennsylvania
METZ, DR. C. B., Institute for Space Biosciences, Florida State University, Talla
hassee, Florida
METZ, DR. CHARLES W., Box 714, Woods Hole, Massachusetts
MIDDLEBROOK, DR. ROBERT, Dartmouth Medical Center, Hanover, New Hampshire
MILKMAN, DR. ROGER D., Department of Zoology, Syracuse University, Syracuse
10, New York
MILLER, DR. J. A., JR., Department of Anatomy, Tulane University Medical
School, New Orleans 18, Louisiana
MILNE, DR. LORUS J., Department of Zoology, University of New Hampshire,
Durham, New Hampshire
MOE, MR. HENRY A., Guggenheim Memorial Foundation, 551 Fifth Avenue, New
York 17, New York

46 MARINE BIOLOGICAL LABORATORY
MONROY, DR. ALBERTO, Institute of Comparative Anatomy, University of Palermo,
Italy
MOORE, DR. GEORGE M., Department of Zoology, University of New Hampshire,
Durham, New Hampshire
MOORE, DR. JOHN A., Department of Zoology, Columbia University, New York
27, New York
MOORE, DR. JOHN W., Department of Physiology, Duke University Medical Cen
ter, Durham, North Carolina
MOORE, DR. R. 0., Department of Biochemistry, Ohio State University, Columbus
10, Ohio
MORAN, DR. JOSEPH, Department of Biology, Russell Sage College, Troy, New
York
MORRILL, DR. JOHN B., JR., Department of Biology, Wesleyan University, Middle
town, Connecticut
MOSCONA, DR. A. A., Department of Zoology, University of Chicago, Chicago 37,
Illinois
MOUL, DR. E. T., Department of Botany, Rutgers Ufliversity, New Brunswick,
New Jersey
MOUNTAIN, MRs. J. D., Charles Road, Mt. Kisco, New York
MULLINS, DR. LORIN J., Department of Biophysics, University of Maryland School
of Medicine, Baltimore 1, Maryland
MUSACCHIA, DR. XAVIER, Department of Biology, St. Louis University, St. Louis
4, Missouri
NABRIT, DR. S. M., President, Texas Southern University, 3201 Wheeler Avenue,
Houston 4, Texas
NACE, DR. PAUL FOLEY, Department of Biology, Hamilton College, McMaster
University, Hamilton, Ontario, Canada
NACHMANSOHN, DR. DAVID, Columbia University, College of Physicians and
Surgeons, New York 32, New York
NASATIR, DR. MAIMON, Department of Botany, Brown University, Providence,
Rhode Island
NAVEZ, DR. ALBERT E., 206 Churchill's Lane, Milton 86, Massachusetts
NELSON, DR. LEONARD, Department of Physiology, Emory University, Atlanta 22,
Georgia
NASON, DR. ALVIN, McCollum-Pratt Institute, The Johns Hopkins University,
Baltimore 18, Maryland
NEURATH, DR. H., Department of Biochemistry, University of Washington, Seattle
5, Washington
NICOLL, DR. PAUL A., Block Oak Lodge, RR #2, Bloomington, Indiana
NIU, DR. MAN-CHIANG, Temple University, Philadelphia 22, Pennsylvania
NOVIKOFF, DR. ALEX B., Department of Pathology, Albert Einstein College of
Medicine, New York 61, New York
OCHOA, DR. SEVERO, New York University College of Medicine, New York 16,
New York
ODUM, DR. EUGENE, Department of Zoology, University of Georgia, Athens,
Georgia

REPORT OF THE DIRECTOR 47
OPPENHEIMER, DR. JANE M., Department of Biology, Bryn Mawr College, Bryn
Mawr, Pennsylvania
OSTERHOUT, DR. W. J. V., 450 East 63rd Street, New York 21, New York
OSTERHOUT, DR. MARION IRWIN, 450 East 63rd St., New York 21, New York
PACKARD, DR. CHARLES, Woods Hole, Massachusetts
PAGE, DR. IRVINE H., Cleveland Clinic, Cleveland, Ohio
PALMER, DR. JOHN D., Department of Biology, University of Illinois, Navy Pier,
Chicago 11, Illinois
PARPART, DR. ARTHUR K., Department of Biology, Princeton University, Prince
ton, New Jersey
PASSANO, DR. LEONARD M., Osborn Zoological Laboratories, Yale University,
New Haven, Connecticut
PATTEN, DR. BRADLEY M., University of Michigan School of Medicine, Ann Arbor,
Michigan
PERKINS, DR. JOHN F., JR., Department of Physiology, University of Chicago,
Chicago 37, Illinois
PERSON, DR. PHILIP, Chief, Special Dental Research Program, VA Hospital,
Brooklyn 9, New York
PETTIBONE, DR. MARIAN H., U. S. National Museum, Division of Marine Interbe
brates, Washington 25, D. C.
PHILPOTT, DR. DELBERT E., Department of Biochemistry, University of Colorado
MedicalCenter,4200 East Ninth Ave.,Denver 20,Colorado
PICK, DR. JOSEPH, Department of Anatomy, New York University, Bellevue Medi
cal Center, New York 16, New York
PIERCE, DR. MADELENE E., Department of Zoology, Vassar College, Poughkeepsie,
New York
POLLISTER, DR. A. W., Department of Zoology, Columbia University, New York
27, New York
POND, DR. SAMUEL E., 53 Alexander Street, Manchester, Connecticut
PORTER, DR. KEITH R., Biological Laboratories, Harvard University, Cambridge
38, Massachusetts
POTTER, DR. DAVID, Department of Neurophysiology, Harvard Medical School,
Boston 15,Massachusetts
PROCTOR, DR. NATHANIEL, Department of Biology, Morgan State College, Balti
more 12, Maryland
PROSSER, DR. C. LADD, Department of Physiology, Burrill Hall, University of Illi
nois, Urbana, Illinois
PROVASOLI, DR. LUIGI, Haskins Laboratories, 305 E. 43rd Street, New York 17,
New York
RAMSEY, DR. ROBERT W., Medical College of Virginia, Richmond, Virginia
RANKIN, DR. JOHN S., Department of Zoology, University of Connecticut, Storrs,
Connecticut
RANZI, DR. SILvI0, Department of Zoology, University of Milan, Milan, Italy
RAPPORT, DR. M., Department of Biochemistry, Albert Einstein College of Medi
cine, New York 61, New York
RATNER, DR. SARAH, Public Health Research Institute of the City of New York,
Foot of East 15th Street, New York 9, New York

48 MARINE BIOLOGICAL LABORATORY
RAY, DR. CHARLES, JR., Department of Biology, Emory University, Atlanta 22,
Georgia
READ, DR. CLARK P., Department of Biology, Rice University, Houston, Texas
REBHUN, DR. LIONEL I., Department of Biology, Box 704, Princeton University,
Princeton, New Jersey
RECKNAGEL, DR. R. 0., Department of Physiology, Western Reserve University,
Cleveland, Ohio
REDFIELD, DR. ALFRED C. , Woods Hole, Massachusetts
RENN, DR. CHARLES E., 509 Ames Hall, Johns Hopkins University, Baltimore 18,
Maryland
REUBEN, DR. JOHN P., Department of Neuro'ogy, College of Physicians and Sur
geons, New York 32, New York
REZNIKOFF, DR. PAUL, Cornell University Medical College, 1300 York Avenue,
New York 16, New York
RICH, DR. ALEXANDER, Department of Biology, Massachusetts Institute of Tech..
nology, Cambridge, Massachusetts
RICHARDS, DR. A., 2950 E. Mabel Street, Tucson, Arizona
RICHARDS, DR. A. GLENN, Department of Entomology, University of Minnesota,
St. Paul 1, Minnesota
RICHARDS, DR. OSCAR W., American Optical Company Research Center, South
bridge, Massachusetts
ROCKSTEIN, DR. MoRRIS, Medical Research Building, 1600 N. W. 10th Avenue,
Miami. Florida
ROGICK, DR. MARY D., College of New Rochelle, New Rochelle, New York
ROMER, DR. ALFRED S., Harvard University, Museum of Comparative Zoology,
Cambridge 38, Massachusetts
RON KIN, DR. RAPHAEL R., National Science Foundation Course Content Improve
nient Section, Washington 25, D. C.
Roor, DR. R. \‘V.,Department of Biology, College of the City of New York, New
York City, New York
Roor, DR. \V. S., Department of Physiology, Columbia University College of Phy
sicians and Surgeons, New York 32, New York
ROSE, DR. S. MERYL, Department of Anatomy, Tulane University, New Orleans
18. Louisiana
ROSENBERG, DR. EVELYN K., Department of Pathology, New York University
Bellevue Medical Center, New York 16, New York
ROSENBERG, DR. PHILIP, Department of Neurology, Columbia University, New
York City, New York
ROSENBLUTH, MISS RAJA, Department of Zoology, Columbia University, New
York 27, New York
ROSENTHAL, DR. THEODORE B., Department of Anatomy, University of Pittsburgh
Medical School, Pittsburgh 13, Pennsylvania
ROSLANSKY, DR. JOHN, Department of Zoology, University of Illinois, Urbana,
Illinois
ROTH, DR. JAY S., Department of Zoology and Entomology, University of Con
necticut, Storrs, Connecticut
ROTHENBERG, DR. M. A., Scientific Director, Dugway Proving Ground, Dugway,
Utah

REPORT OF THE DIRECTOR 4@
RUGH, DR. ROBERTS, Radiological Research Laboratory, College of Physicians and
Surgeons, New York 32, New York
RUNNSTRöM, DR. JOHN, Wenner-Gren Institute, Stockholm, Sweden
RUSTAD, DR. RONALD C., Department of Radiology, Western Reserve University,
Cleveland, Ohio 44106
RUTMAN, DR. ROBERT j., General Laboratory Building, 215 5. 34th Street, Phila
delphia, Pennsylvania
RYTHER, DR. JOHN H., Woods Hole Oceanographic Institution, Woods Hole,
Massachusetts
SAGER, DR. R., Department of Zoology, Columbia University, New York 27, New
York
SANBORN, DR. RICHARD C., Department of Biological Sciences, Purdue University,
Lafayette, Indiana
SANDERS, DR. HOWARD L., Woods Hole Oceanographic Institution, Woods Hole,
Massachusetts
SAUNDERS, DR. JOHN, Department of Biology, Marquette University, Milwaukee
3, Wisconsin
SAUNDERS, MR. LAWRENCE, West Washington Square, Philadelphia 5, Pennsyl
vania
SAZ, DR. ARTHUR KENNETH, National Institutes of Health, Bethesda 14, Maryland
SCHACHMAN, DR. HOWARD K., Department of Biochemistry, University of Cali
fornia, Berkeley 4, California
SCHARRER, DR. ERNST, Department of Anatomy, Albert Einstein College of Medi
cine, New York 61, New York
SCHLESINGER, DR. R. WALTER, Department of Microbiology, Rutgers Medical
School, New Brunswick, New Jersey
SCHMIDT, DR. L. H., Christ Hospital, Cincinnati, Ohio
SCHMITT, DR. FRANCIS 0., Department of Biology, Massachusetts Institute of
Technology, Cambridge, Massachusetts
SCHMITT, DR. 0. H., Department of Physics, University of Minnesota, Minne
apolis 14, Minnesota
SCHNEIDERMAN, DR. HOWARD A., Department of Biology, Western Reserve Uni
versity, Cleveland, Ohio
SCHOLANDER, DR. P. F., Scripps Institution of Oceanography, La Jolla, California
SCHOTT@, DR. OSCAR E., Department of Biology, Amherst College, Amherst,
Massachusetts
SCHRAMM, DR. J. R., Department of Botany, Indiana University, Bloomington,
Indiana
SCOTT, DR. ALLAN C., Colby College, Waterville, Maine
SCoTT, DR. D. B. MCNAIR, 15 General Laboratory Bldg., University of Pennsyl
vania, Philadelphia 4, Pennsylvania
SCoTT, SISTER FLORENCE MARIE, Seton Hill College, Greensburg, Pennsylvania
SCOTT, DR. GEORGE I., Department of Zoology, Oberlin College, Oberlin, Ohio
SEARs, DR. MARY, Glendon Road, Woods Hole, Massachusetts
SELIGER, DR. HOWARD H., McCollum-Pratt Institute, Johns Hopkins University,.
Baltimore 18, Maryland
SENFT, DR. ALFRED W., Woods Hole, Massachusetts
SEVERINGHAUS, DR. AURA E., 375 W. 250 Street, New York 71, New York

50 MARINE BIOLOGICAL LABORATORY
SHANES, DR. ABRAHAM, Department of Pharmacology, University of Pennsyl
vania School of Medicine, Philadelphia 4, Pennsylvania
SHAPIRO, DR. HERBERT, 6025 N. 13th Street, Philadelphia 41 , Pennsylvania
SHAVER, DR. JOHN R., Department of Zoology, Michigan State University, East
Lansing, Michigan
SHEDLOVSKY, DR. THEODORE, Rockefeller Institute, 66th Street and York Avenue,
New York 21, New York
SHEMIN, DR. DAVID, Department of Biochemistry, Columbia University, New
York City, New York
SHERMAN, DR. I. W., Division of Life Sciences, University of California, River
side, California
SICHEL, DR. FERDINAND J. M., University of Vermont, Burlington, Vermont
SICHEL, MRS. F. J. M., Department of Biology, Trinity College, Burlington, Ver
mont
SILVA, DR. PAUL, Department of Botany, University of California, Berkeley 4,
California
SIMMONS, DR. JOHN E., JR., Department of Biology, Rice University, Houston 1,
Texas
SLIFER, DR. ELEANOR H., 308 Linsmore Avenue, Glenside, Pennsylvania
SMELSER, DR. GEORGE K., Department of Anatomy, Columbia University, New
York 32, New York
SMITH, DR. DIETRICH C., Department of Physiology, University of Maryland
School of Medicine, Baltimore 1, Maryland
SMITH, MR. HOMER P., General Manager, Marine Biological Laboratory, Woods
Hole, Massachusetts
SMITH, MR. PAUL FERRIS, Woods Hole, Massachusetts
SMITH, DR. RALPH I., Department of Zoology, University of California, Berkeley
4, California
SONNEBORN, DR. T. M., Department of Zoology, Indiana University, Bloomington,
Indiana
SONNENBLICK, DR. B. P., Rutgers University, 40 Rector Street, Newark 2, New
Jersey
SPECTOR, DR. A., Howe Laboratories, Harvard Medical School, Boston 15, Massa
chusetts
SPEIDEL, DR. CARL C., Department of Anatomy, University of Virginia Medical
School, Charlottesville, Virginia
SPIEGEL, DR. MELVIN, Department of Biological Sciences, Dartmouth College,
Hanover, New Hampshire
SPRATT, DR. NELSON T., Department of Zoology, University of Minnesota, Minne
apolis 14, Minnesota
SPYROPOULOS, DR. C. S., Bldg. 9-Rm. 140, National Institutes of Health, Bethesda
14, Maryland
STARR, DR. RICHARD C., Department of Botany, Indiana University, Bloomington,
Indiana
STEINBACH, DR. H. BURR, Department of Zoology, University of Chicago, Chicago
37, Illinois
STEINBERG, DR. MALCOLM S., Department of Biology, Johns Hopkins University,
Baltimore18,Maryland

REPORT OF THE DIRECTOR 51
STEINHARDT, DR. JACINTO, Georgetown University, Washington 7, D. C.
STEPHENS, DR. GROVER C., Department of Zoology, University of Minnesota,
Minneapolis 14, Minnesota
STETTEN, DR. DEWITT, Dean, Rutgers University Medical School, New Bruns
wick, New Jersey
STETTEN, DR. MARJORIE R., Rutgers University Medical School, New Brunswick,
New Jersey
STEWART, DR. DOROTHY, Rockford College, Rockford, Illinois
STOKEY, DR. ALMA G., Department of Botany, Mount Holyoke College, South
Hadley, Massachusetts
STONE, DR. WILLIAM, JR., Ophthalmic Plastics Laboratory, Massachusetts Eye
and Ear Infirmary, Boston, Massachusetts
STRAUS, DR. W. L., JR., Department of Anatomy, Johns Hopkins University Medi
cal School, Baltimore 5, Maryland
STREHLER, DR. BERNARD L., 4115 Westview Road, Baltimore 18, Maryland
STRITTMATTER, DR. PHILIPP, Department of Biological Chemistry, Washington
University Medical School, St. Louis, Missouri
STUNKARD, DR. HORACE W., American Museum of Natural History, Central Park
West at 79th Street, New York 24, New York
STURTEVANT, DR. ALFRED H., California Institute of Technology, Pasadena, Cali
fornia
SUDAK, DR. FREDERICK N., Department of Physiology, Albert Einstein College of
Medicine, New York 61, New York
SULKIN, DR. S. EDWARD, Department of Bacteriology, University of Texas, South
western Medical School, Dallas, Texas
SWOPE, MR. GERARD, JR., 570 Lexington Avenue, New York 22, New York
SZABÔ, DR. GEORGE, Department of Dermatology, Massachusetts General Hospital,
Boston 14, Massachusetts
SZENT GYöRGYI, DR. ALBERT, Institute for Muscle Research, Marine Biological
Laboratory, Woods Hole, Massachusetts
SZENT GYÔRGYI, DR. ANDREW G., Department of Cytology, Dartmouth Medical
School, Hanover, New Hampshire
TASAKI, DR. ICHIJI, Laboratory of Neurobiology, NIMH, National Institutes of
Health, Bethesda 14, Maryland
TAYLOR, DR. ROBERT E., Laboratory of Neurophysiology, NINDB, Bethesda 14,.
Maryland
TAYLOR, DR. WILLIAM RANDOLPH, Department of Botany, University of Michigan,.
Ann Arbor, Michigan
TAYLOR, DR. W. ROWLAND, Department of Oceanography, Johns Hopkins Uni
versity, Baltimore 18, Maryland
TEWINKEL, DR. Lois E., Department of Zoology, Smith College, Northampton,.
Massachusetts
TOBIAS, DR. JULIAN, Department of Physiology, University of Chicago, Chicage
37, Illinois
TRACY, DR. HENRY C., 3595 Mynders #3, Memphis 11, Tennessee
TRAGER, DR. WILLIAM, Rockefeller Institute, 66th Street and York Avenue, New
York 21, New York

52 MARINE BIOLOGICAL LABORATORY
TRAVIS, DR. D. M., Department of Pharmacology, University of Florida, Gaines
ville, Florida
TRINKAUS, DR. J. PHILIP, Department of Zoology, Osborn Zoological Labora
tories, Yale University, New Haven, Connecticut
TROLL, DR. WALTER, Department of Industrial Medicine, New York University
College of Medicine, New York 16, New York
TWEEDELL, DR. KENYON S., Department of Biology, University of Notre Dame,
Notre Dame, Indiana
TYLER, DR. ALBERT, Division of Biology, California Institute of Technology, Pasa
dena, California
URETZ, DR. ROBERT B., Department of Biophysics, University of Chicago, Chi
cago 37, Illinois
DE VILLAFRANCA, DR. GEORGE W., Department of Zoology, Smith College, Nor
thampton, Massachusetts
VILLEE, DR. CLAUDE A., Department of Biological Chemistry, Harvard Medical
School, Boston 15, Massachusetts
VINCENT, DR. WALTER S., Department of Anatomy, University of Pittsburgh,
Pittsburgh 13, Pennsylvania
WAINIO, DR. W. W., Bureau of Biological Research, Rutgers University, New
Brunswick, New Jersey
WALD, DR. GEORGE, Biological Laboratories, Harvard University, Cambridge 38,
Massachusetts
WARNER, DR. ROBERT C., Department of Chemistry, New York University College
of Medicine, New York 16, New York
WARREN, DR. L., c/o Dr. A. Bussard, Institut Pasteur, 25 Rue de Docteur Roux,
Paris, France
WATERMAN, DR. T. H., Department of Zoology, 272 Gibbs Research Laboratory,
Yale University, New Haven, Connecticut
WEBB, DR. MARGUERITE, Department of Physiology, and Bacteriology, Goucher
College, Towson, Baltimore, Maryland
WEISS, DR. LEON P., Department of Anatomy, The Johns Hopkins University
School of Medicine, Baltimore 5, Maryland
WEISS, DR. PAUL A., Laboratory of Developmental Biology, Rockefeller Institute,
66th Street and York Avenue, New York 21, New York
WENRICH, DR. D. H., Department of Zoology, University of Pennsylvania, Phila
delphia 4, Pennsylvania
WERMAN, DR. ROBERT, Institute of Psychiatric Research, Indiana University Medi
cal Center, 1100 W. Michigan Street, Indianapolis 7, Indiana
WHITAKER, DR. DOUGI.ASM., Rockefeller Institute, 66th Street and York Avenue,
New York 21, New York
WHITE, DR. E. GRACE, 1312 Edgar Avenue, Chambersburg, Pennsylvania
WHITING, DR. ANNA R., 535 West Vanderbilt Drive, Oak Ridge, Tennessee
WHITING, DR. PHINEAS, 535 West Vanderbilt Drive, Oak Ridge, Tennessee
WICKERSHAM, MR. JAMES H., 791 Park Avenue, New York 21, New York
WICHTERMAN, DR. RALPH, Department of Biology, Temple University, Philadel
phia 22, Pennsylvania
WIEMAN, DR. H. L., Box 485, Falmouth, Massachusetts

REPORT OF THE DIRECTOR 53
WIERCINSKI, DR. FLOYD J., 600 W. Franklin, Apt. 208, Minneapolis, Minnesota
55405
WIGLEY, DR. ROLAND L., U. S. Fish and Wildlife Service, Woods Hole, Massa
chusetts
WILBER, DR. C. G., Marine Laboratories, University of Delaware, Newark, Dela
ware
WILLIER, DR. B. H., Department of Biology, Johns Hopkins University, Baltimore
18, Maryland
WILSON, DR. J. WALTER, Department of Biology, Brown University, Providence
12, Rhode Island
WILSON, DR. T. HASTINGS, Department of Physiology, Harvard Medical School,
Boston 15, Massachusetts
WILSON, DR. WALTER L., Department of Physiology, University of Vermont Col
lege of Medicine, Burlington, Vermont
WIT5CHI, DR. EMIL, Universitãt Basel, Anatomisches Institut, Pestalozzistrasse
20, Basel, Switzerland
WITTENBERG, DR. JONATHAN B., Department of Physiology and Biochemistry,
Albert Einstein College of Medicine, New York 61, New York
WRIGHT, DR. PAUL A., Spaulding Bldg., Department of Zoology, University of
New Hampshire, Durham, New Hampshire
WRINCH, DR. DOROTHY, Department of Physics, Smith College, Northampton,
Massachusetts
YNTEMA, DR. C. L., Department of Anatoniy, State University of New York Col
lege of Medicine, Syracuse 10, New York
YOUNG, DR. D. B., Main Street, North Hanover, Massachusetts
ZIMMERMAN, DR. A. M., Department of Pharmacology, State University of New
York, Downstate Medical Center, Brooklyn 3, New York
ZINN, DR. DONALD J., Department of Zoology, University of Rhode Island, Kings
ton, Rhode Island
ZIRKLE, DR. RAYMOND E., Department of Radiobiology, University of Chicago,
Chicago 37, Illinois
ZORZOLI, DR. ANITA, Department of Physiology, Vassar College, Poughkeepsie,
New York
ZULLO, DR. VICTOR A., Marine Biological Laboratory, Woods Hole, Massachusetts
ZWEIFACH, DR. BENJAMIN, New York University Bellevue Medical Center, New
York 16, New York
Z\VILLING, DR. EDGAR, Department of Biology, Brandeis University, Waltham 54,
Massachusetts
ASSOCIATE MEMBERS
ALDRICH, MISS AMEY OWEN BARBOUR, MR. AND MRS. LUCIUS H.
ALTON, MRS. BENJAMIN BARROWS, MRS. ALBERT
ANDRES, MRS. WILLIAM BARTOW, MR. AND MRS. CLARENCE W.
ARMSTRONG, MRS. PHILIP B. BARTOW, MRS. FRANCIS D.
BACON, MR. AND MRS. ROBERT BARTOW, MRS. PHILIP K.
BALL, MRS. ERIC G. BARTOW, MR. S. R.
BAKALAR, MR. AND MRS. DAVID BEALE, MR. AND MRS. EDWARD

54 MARINE BIOLOGICAL LABORATORY
BELL, MRS. ARTHUR W.
BIGELOW, MRS. ROBERT PAYNE
BRADLEY, MRS. ALBERT L.
BRADLEY, DR. AND MRS. CHARLES
BROWN, MRS. THORNTON
BURDICK, DR. C. LALOR
BURLINGAME, MRS. F. A.
BUTLER, DR. AND MRS. E. G.
CAHOON, MRS. SAMUEL T.
CALKINS, MRS. GARY N.
CALKINS,
JR.
MR. AND MRS. G. NATHAN,
CARLTON, MR. WINSLOW G.
CAREY, MISS CORNELIA
CLARK, MRS. JAMES B.
CLAFF, MRS. C. LLOYD
CLAFF, MR. MARK M.
CLARK, MRS. LEROY
CLARK, EDNA A.
CLARK, MR. AND MRS. W. VAN ALAN
CLOWES, MR. ALLEN W.
CLOWES, MRS. G. H. A.
CLOWES, DR. AND MRS. GEORGE H. A.,
JR.
COLTON, MRS. H. SEYMOUR
CONKLIN, MISS ISABEL
CRAMER, MR. AND MRS. I. D. W.
CRANE, MR. JOHN
CRANE, MISS LOUISE
CRANE, MRS. W. CAi@FY
CRANE, MRS. W. Mu@@
CROWELL, MR. AND MRS. PRINCE S.
CROSSLEY, MISS DOROTHY
CURTIS, DR. AND MRS. WILLIAM D.
DANIELS, MR. AND MRS. BRUCE
DANIELS, MR. AND MRS. F. H.
DAY, MR. AND MRS. POMEROY
DRAPER, MRS. MARY C.
DREYER, MR. AND MRS. F. A.
DUB0IS, DR. ARTHUR B.
ELSMITH, MRS. DOROTHY
EWING, MR. WILLIAM
FERGUSON, MRS. JAMES J.
FISHER, MRS. B. C.
FORBES, DR. ALEXANDER
FRANCIS, MR. LEWIS H., JR.
FROST, MRS. FRANK J.
GAISER, DR. AND MRS. DAVID W.
GALTSOFF, MRS. EUGENIA
GARFIELD, MISS ELEANOR
GARLOCK, MR. ROBERT
GIFFORD, MR. AND MRS. JOHN A.
GIFFORD, MR. AND MRS. PROSSER
GILCHRIST, MR. AND MRS. JOHN M.
GILDEA, DR. AND MRS. E. F.
GILLETTE, MR. AND MRS. ROBERT S.
GLAZEBROOK, MRS. JAMES
GREEN, MISS GLADYS M.
GREER, MR. AND MRS. WILLIAM HER
SHEY, JR.
GULESIAN, MRS. PAUL J.
HAMLEN, MR. AND MRS. J. MONROE
HANNA, MR. AND MRS. THOMAS C.
HARRINGTON, MR. AND MRS. ROBERT D.
HARVEY, DR. AND MRS. RICHARD
HERVEY, MRS. JOHN P.
HIRSCHFELD, MRS. NATHAN B.
HOPKINS, MRS. RALPH H.
HOUSTON, MR. AND MRS. HOWARD E.
JANNEY, MRS. MARY D.
JEWETT, MR.ANDMRS.G. F., JR.
JONES, MR. AND MRS. DEWITT C., JR.
KEITH, MRS. HAROLD C.
KOLLER, DR. AND MRS. LEWIS R.
LEMANN, MRS. BENJAMIN
LOBB, MR. AND MRS. JOHN
LOEB, DR. AND MRS. ROBERT F.
LOVELL, MR. AND MRS. HOLLIS R.
MAKRAUER, MR. S. LANG
MARSLAND, DR. AND MRS. DOUGLAS A.
MARVIN, MRS. WALTER TAYLOR
MAST, MRS. S. 0.
MATHER, MR. FRANK J., III
MCCUSKER, MRS. PAUL T.
MCELROY, DR. AND MRS. W. D.
MCGILLICUDDY, DR. AND MRS. JOHN J.
MCKELVY, MR. JOHN E.
MCLANE, MRS. HUNTINGTON
MCVITTY, MRS. A. E.
MEIGS, MR. AND MRS. ARTHUR
MEIGS, DR. AND MRS. J. WISTER
MELLON, MR. AND MRS. RICHARD P.

MINIs, MR. AND MRS. ABRAM J., JR.
MITCHELL, MRS. JAMES McC.
MITCHELL, MRS. PHILIP
MIXTER, MRS. WILLIAM JASON
MOSSER, MRS. BENJAMIN D.
MOTLEY, MRS. THOMAS
NEWTON, MISS HELEN K.
NICHOLS, MRS. GEORGE
NIMS, MRS. E. D.
THE AARON E. NORMAN FUND, INC.
PACKARD, MRS. CHARLES
PARK, MR. AND MRS. MALCOLM S.
PARPART, DR. AND MRS. ARTHUR K.
PENNINGTON, Miss ANNE H.
PHILIPPE, MR. PIERRE
PUTNAM, MR. WILLIAM A., III
REDFIELD, DR. AND MRS. ALFRED C.
REZNIKOFF, DR. AND MRS. PAUL
RIVINu5, MRS. F. M.
RUDD, MR. AND MRS. H. W. DWIGHT
RUGH, MRS. ROBERTS
SANDS, Miss ADELAIDE G.
SAUNDERS, MR. AND MRS. LAWRENCE
REPORT OF THE LIBRARIAN
SHIVERICK, MRS. ARTHUR
SINCLAIR, MR. AND MRS. W. RICHARD
SON
SMITH, MRS. HOMER P.
SPEIDEL, MRS. CARL C.
STONE, MR. AND MRS. LEO
STONE, DR. WILLIAM, JR.
STONE, MRS. SAMUEL M.
STRAUSS, MR. AND MRS. DONALD B.
STUN KARD, MRS. HORACE
SWOPE, MR. DAVID
SWOPE, MR. AND MRS. GERARD, JR.
SWOPE, Miss HENRIETTA H.
SZENT-GYöRGYI, DR. ALBERT
TOMPKINS, MR. AND MRS. B. A.
WEBSTER, MRS. EDWIN S.
WHITELEY, Miss MABEL W.
WICKERSHAM, MRS. JAMES H.
WILHELM, DR. AND MRS. HILMER J.
WILSON, MRS. EDMUND B.
WOLFINSOHN, MRS. W.
YNTEMA, MRS. CHESTER L.
V. REPORT OF THE LIBRARIAN
The 1963 acquisitions added 68 new titles to our current journal collection,
bringing the total nuniber to 1826. Of this total, the Laboratory subscribed to 568,
sent The Biological Bulletin in exchange for 644 and gratefully received 187 as
gifts. The Woods Hole Oceanographic Institution subscribed to 144, sent its
Collected Reprints in exchange for 215 and accepted 68 as gifts.
To our book shelves we added a total of 353 titles, the Laboratory having pur
chased 108 and the Institution 67. Generous authors contributed 11 books and
122 were presented by the publishers who held summer exhibits in the main lobby.
From various other sources, 45 books were received.
Through purchase, exchange and gift the Laboratory completed 17 journal sets
and partially completed 26 sets, whereas the Institution completed two sets and
partially completed 9 sets. We added 4266 reprints to our collection, 2862 of
which were of current issue.
There were 1089 volumes sent to the bindery. Outgoing inter-library loans
‘¿totalled306 and 76 titles were borrowed for our investigators. The Xerox machine
installed by the Laboratory enabled us to give more efficient photoprint service to
.other libraries and to our own readers.
The Library now contains 82,436 bound volumes and 228,000 reprints.
Several gifts of books and reprints were received, for which the Librarian
‘¿wishes to extend acknowledgment to Drs. Douglas A. Marsland, Roberts Rugh
55

56 MARINE BIOLOGICAL LABORATORY
and Albert Szent-Györgyi. From the estate of Dr. Herbert W. Rand we acquired
60 books. We were extremely happy to get these, as many of them have been used
to replace worn copies, many of which have been long out-of-print.
In November, work was started on Library alterations. These renovations cover
extensive improvements in the stacks, new offices for the Staff, a rare-books room,
a catalogue room and a photoprint room. It is anticipated that these changes
will be advantageous to our readers and will increase the efficiency of the services
rendered by the Staff.
Respectfully submitted,
DEBORAH L. HARLOW,
Librarian
VI. REPORT OF THE TREASURER
The market value of the General Endowment Fund and the Library Fund at
December 31, 1963, amounted to $2,155,489 as against book value of $1,242,896.
This compares with values of $1,994,709 and $1,242,771, respectively, at the end
of the preceding year. The average yield on the securities was 3.64% of the
market value and 6.31% of book value. The total uninvested principal cash in
the above accounts as of December 31, 1963, was $431.33. Classification of the
Securities held in the Endowment Funds appears in the Auditor's report.
The market value of the pooled securities as of December 31, 1963, was $390,026
with uninvested principal cash of $168.26, the market value at December 31, 1962,
being $353,243. The book value of the securities in this account was $315,196 on
December 31, 1963, compared with $302,453 a year earlier. The average yield
on market value was 3.36% and 4.16% of book value.
The proportionate interest in the Pool Fund Account of the various Funds as
of December 31, 1963, is as follows:
Pension Funds 31.117%
General Laboratory Investment 48.642
Other:
Bio Club Scholarship Fund 1.393
Rev. Arsenius Boyer Scholarship Fund 1.705
Gary N. Calkins Fund 1.596
Allen R. Memhard Fund 310
F. R. Lillie Memorial Fund 5.376
Lucretia Crocker Fund 5.819
E. G. Conklin Fund 983
Jewett Memorial Fund 517
M. H. Jacobs Scholarship Fund 702
Anonymous Gift 1.840
Donations from the MBL Associates for 1963 were $4,295.00, as compared with
$5,305 for 1962. Unrestricted gifts from foundations, societies and companies
amounted to $24,990.

REPORT OF THE TREASURER 57
During the year we administered the following grants:
Investigators Training MBL, Institutional
11 NIH 5 NIH 4 NIH
4NSF 2NSF 3NSF
1 Ford 3 ONR
1 Commonwealth 1 AEC
17 7 11
The rateof overheadon grantsto investigators has been increasedto 20%,
based on the amount expended. The overhead on these grants for this year
amounted to $56,494as compared with $43,405fortheprecedingyear.
The LillieFellowshipFund, witha market valueof$104,171and a book value
of $93,018, as well as the investment in the General Biological Supply House, with
a book valueof$12,700,iscarriedintheBalanceSheetitem“¿Other Investments.â€
The General Biological Supply House fiscal year ended June 30, 1963, and had
a profit after taxes of $241,616 as compared to $302,657 in 1962 and $302,851 in
1961 and $314,034in 1960 and $303,300in 1959. In the fiscalyear 1963,the
Marine BiologicalLaboratoryreceiveddividendsfrom the General Biological
Supply House of $42,164 as against $38,100 in 1962 and $33,020 in 1961 and
$30,480 in 1960.
Following is a statement of the auditors:
To the Trustees of the Marine Biological Laboratory, Woods Hole, Massachusetts:
We have examined the balance sheets of the Marine Biological Laboratory as
at December 31, 1963, the related statement of operating expenditures, income and
currentfundand statementoffundsfortheyearthenended. Our examinationwas
made in accordancewith generallyacceptedauditingstandards,and accordingly
includedsuch testsof the accountingrecordsand such otherauditingprocedures
as we considerednecessaryinthecircumstances.We examined and have reported
on financial statements of the Laboratory for the year ended December 31, 1962.
In our opinion,theaccompanyingfinancial statementspresentfairlytheassets,
liabilities and funds of the Marine BiologicalLaboratoryat December 31, 1963
and
basis.
1962, and the results of its operations for the years then ended on a consistent
Boston, Massachusetts
March30, 1964
LYBRAND, Ross BRos. & MONTGOMERY
JAMES H. WICKERSHAM,
Treasurer

58 MARINE BIOLOGICAL LABORATORY
Investments held by Trustee:
MARINE BIOLOGICAL LABORATORY
BALANCE SHEETS
December 31, 1963 and 1962
Investments 1963 1962'
Securities, at cost (approximate market quotation, 1963—
$2,155,000) $1,242,896 $1,242,771
Cash 431 641
Investments of other endowment and unrestricted funds:
Pooled investments, at cost (approximate market quotation, 1963
$390,000) less $5,728 temporary investment of current fund
1,243,327 1,243,412
cash 309,468 296,725
Other investments 120,424 139,677
Cash 14,083 15,135
Accounts receivable 21,131 31
Plant Assets
$1,708,433 $1,694,980
Land, buildings, Library and equipment (note) 4,931,472 4,900,749
Less allowance for depreciation (note) 1,313,162 1,247,149
Current Assets
$3,618,310 $3,653,600
Cash 100,991 118,732
Temporary investment in pooled securities 5,728 5,728
U. S. Treasury bills, due prior to 2/13/64 at cost 74,324 248,253
Accounts receivable (U. S. Government, 1963—$95,142; 1962—$41,579) 140,825 78,099
Inventories of specimens and Bulletins 45,288 41,903
Prepaid insurance and other 7,062 19,006
$374,218 $511,721

REPORT OF THE TREASURER . 59
MARINE BIOLOGICAL LABORATORY
BALANCE SHEETS
December 31,1963 and 1962
Endowment Funds 1963 1962
Endowment funds given in trust for benefit of the Marine Biological
Laboratory $1,243,327 $1,243,412
Endowment funds for awards and scholarships:
Principal 126,980 126,302
Unexpended income 12,077 11,353
139,057 137,655
Unrestricted funds functioning as endowment 206,378 206,378
Retirement fund 108,481 94,048
Pooled investments—accumulated gain 11,190 13,487
Plant Funds
$1,708,433 $1,694,980
Funds expended for plant, less retirements 4,931,472 4,900,749
Less allowance for depreciation charged thereto 1,313,162 1,247,149
Current Liabilities and Funds
$3,618,310 $3,653,600
Accounts payable and accrued expenses 62,763 35,729
Advance subscriptions 10,362 7,524
Unexpended grants—research 110,433 344,437
Unexpended balances of gifts for designated purposes 13,531 9,461
Current fund 177,129 114,570
$ 374,218 $ 511,721
Note.—The Laboratory has since January 1, 1916 provided for reduction of book amounts
of plant assets and funds invested in plant at annual rates ranging from 1% to 5% of the original
cost of the assets.

@ 60
MARINE BIOLOGICAL LABORATORY
MARINE BIOLOGICAL LABORATORY
STATEMENTS OF OPERATING EXPENDITURES, INCOME AND CURRENT FUND
Years Ended December 31, 1963 and 1962
Operating Expenditures
1963 1962
Research and accessory services $ 273,333 $ 237,580
Instruction 147,163 142,394
Library and publications (including book purchases—1963, $25,628;
1962, $24,287) 77,922 72,514
Direct costs on research grants 484,640 378,163
Direct costs on institution support grants 129,642 98,043
1,112,700 928,694
Administration and general 97,339 92,124
Plant operation and maintenance 111,441 102,632
Dormitories and dining 174,030 155,234
Additions to plant from current income 7,793 22,088
Less depreciation included in plant operation and dormitories and
1,503,303 1,300,772
dining above but charged to plant funds 67,472 65,061
Income
1,435,831 1,235,711
Research fees 113,024 99,314
Accessory services (including sales of biological specimens—1963,
$38,019; 1962, $29,898) 103,398 90,247
Instruction fees 28,461 27,439
Library fees, Bulletins, subscriptions and other 43,952 41,314
Dormitories and dining income 129,486 125,176
Grants for support of institutional activities:
Instruction and training 131,564 127,988
Support services 129,642 98,043
General 113,853 79,500
Reimbursements and allowances for direct and indirect costs on specific
research grants 541,134 421,568
Gifts used for current expenses 29,285 39,824
Investment income used for current expenses 134,591 120,697
1,498,390 1,271,110
Excess current income 62,559 35,399
Current fund balance January 1 114,570 79,171
Current fund balance December 31 $ 177,129 $ 114,570

REPORT OF THE TREASURER 61
MARINE BIOLOGICAL LABORATORY
Balance
January
1, 1963
Invested funds $1,694,980
STATEMENT OF FUNDS
Year Ended December 31,1963
Unexpended research
grants $ 344,437 682,189 916,193
$ 110,433
Unexpended gifts for
designated purposes$ 9,461 37,265
29,285 3,910 ________ $ 13,531
Current fund
Total gifts and
$ 114,570 _______
62,559(1)
_______ ________ ______
$ 177,129
________
other receipts... _______ $801,910 _______ $136,547 $1,073,058
________ $19,321
Gifts
Grants for research,
training and
support
Net loss on sales of
securities
Appropriated from cur
rent income and
other
(1) Excess current in
Total
come over expendi
tures
Scholarship awards.
Payments to pensioners
Other
Gifts and Invest- Used for
Other ment Current
Receipts Income Expenses
$ 19,897 $136,547 $ 127,580
37,265
682,189
(1,704)
21,601
62,559
$801,910
Other
Expendi
tures
Balance
December
31, 1963
$15,411 $1,708,433
4,534
10,877
3,910
$19,321

62 MARINE BIOLOGICAL LABORATORY
MARINE BIOLOGICAL LABORATORY
SUMMARY OF INVESTMENTS
December 31, 1963
%of Market %of
Investment
Income
Cost Total Quotations Total 1963
Securities held by Trustee:
General endowment fund:
U. S. Government securities $ 120,358 11.5 $ 119,533 6.6 $ 2,614
Corporate bonds 505,629 49.0 490,820 27.4 20,925
Preferred stocks 54,573 5.2 54,200 3.1 2,829
Common stocks 358,414 34.3 1,124,595 62.9 40,034
1,038,974 100.0 1,789,148 100.0 66,402
General Educational Board
endowment fund:
U. S. Government securities 44,099 21.6 43,885 12.0 1,756
Other bonds 83,181 40.8 83,770 22.8 3,222
Preferred stocks 15,641 7.6 15,750 4.3 1,074
Common stocks 61,001 30.0 222,936 60.9 6,028
203,922 100.0 366,341 100.0 12,080
Total securities held by
Trustee $1,242,896 $2,155,489 78,482
Investments of other endowment
and unrestricted funds:
Pooled investments:
U. S. Government securities 56,292 17.9 55,882 14.3 1,022
Corporate bonds 106,477 33.8 106,769 27.4 5,657
Common stocks 152,427 48.3 227,375 58.3 6,436
315,196 100.0 $ 390,026 100.0 13,115
Less temporary investment
of current fund cash— (5,728) (236)
309,468 12,879
Other investments:
U. S. Government securities 28,072 641
Other bonds 15,032 1,545
Preferred stocks 3,728 130
Common stocks 58,886 43,556
Real estate 14,706 ________
120,424 45,872
Total investment of
other endowment and
unrestricted funds. . $ 429,892 58,751
Total investment income 137,233
Custodian's fees charged thereto.... (686)
Investment income distributed to
funds 136,547
Current investments:
U. S.Treasurybills, due priorto
2/13/64 $ 74,324 $ 75,000 6,775
Temporary investment in pooled
securities $ 5,728 236
7,011
$143,558