Obituary : Jane Brotherton Walker (1925-2009)

OBITUARY
Jane Brotherton Walker
Jane Brotherton Walker was born in Nairobi, Kenya on 31 January 1925. As a child
growing up on a farm in Kenya, Jane was home-schooled by her mother during her
primary school years, and then went to England where she matriculated in 1944 at the
County High School for Girls in Retford. While at high school Jane lived with her
aunt and cousins, and because the Second World War was in progress, did not see her
parents for a protracted period. It was during this time that Jane contracted
poliomyelitis and spent several months in a plaster cast in hospital. Over the years the
after-effects of the disease progressively affected her ability to walk, particularly
during her later years. Jane obtained BSc (Honours) and MSc degrees at Liverpool
University in 1948 and 1959 respectively, and was awarded a DSc degree on her
published works by the University of the Witwatersrand, Johannesburg in 1983.
Jane was first employed in 1949 in the Research Branch of Her Majesty’s Overseas
Civil Service as a Research Officer in the East African Veterinary Research
Organization, Muguga, Kenya. She progressed in this organization to Senior
Scientific Officer and then to Principal Scientific Officer, before voluntarily retiring
from Her Majesty’s Service in 1966 to take up a post as Senior Professional Officer at
the Veterinary Research Institute, Onderstepoort, upon the retirement of Dr Gertrud
Theiler. Here Jane spent the rest of her working life. At the Institute she progressed
through the ranks of promotion to Chief Professional Officer, Chief Veterinary
Researcher and Specialist Scientist until her retirement on pension in 1990. After
retirement she continued to work at Onderstepoort in an honorary capacity for 3 days
a week until 1998. During her working life, including her active years after
retirement, Jane was sole, senior or co-author of 53 scientific publications and five
books as well as the descriptions of 18 new tick species. She, herself, illustrated many
of the ticks with meticulously executed line drawings. For the illustrations in her
magnum opus, “The Rhipicephalus species ticks of the World”, Jane used the services
of André Olwage, a young commercial artist imbued with a gentleness and patience
that he sorely needed with Jane’s insistence on the exactness of every detail. Her
legacy in her chosen field of the taxonomy of ticks resides in her published works as
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well as in the training she gave, and the wisdom she imparted to Ivan Horak, Arthur
Spickett, Heloise Heyne, Trevor Petney and Dmitry Apanaskevich.
Jane’s prowess as a scientist was recognized by her peers who conferred on her three
of the most prestigious awards in the field of biological sciences in South Africa,
namely the 1988 Elsdon-Dew Medal of the Parasitological Society of Southern
Africa, for services rendered to Parasitology in Africa, the Agricultural Science and
Technology Woman of the Year Award for 1998, and the Theiler Memorial Trust
Award in 1998 for exceptional service rendered to Veterinary Science in Africa. Her
scientific writing skills were soon recognized at Onderstepoort and she was a member
of the Editorial Committee of the Onderstepoort Journal of Veterinary Research from
1969 until 2000, a function she performed extremely punctiliously to the dismay, but
nevertheless to the ultimate benefit of many a sloppy author.
Jane had her own likes and dislikes amongst the tick genera. She was without doubt
the expert on the genus Rhipicephalus, with which she had an ongoing love affair. She
was greatly distressed when it was proposed that the genus Boophilus become a
subgenus of her beloved Rhipicephalus, a subject which the protagonists of this
change preferred not to discuss with her. Jane also had a deep affection for the
African Amblyommas, but despite being involved in the description of six Ixodes
species and one argasid tick, she never liked ticks of the genus Ixodes or of the family
Argasidae.
While working in Kenya Jane owned a horse, which she rode side-saddle, and also
travelled in her Hillman Mynx automobile to localities that are now nearly
inaccessible in a 4x4 vehicle. She also traveled to South Africa in the Hillman Mynx
when she moved here from Kenya. In South Africa she owned a Mitshubishi Colt,
especially fitted with hand-controls to compensate for the weakness in her legs, and it
served as her daily mode of transport from her home to Onderstepoort.
Jane loved classical music, autobiographies, natural science books and the BBC news,
to which she used to listen on overseas channels, and she also took a lively interest in
African politics. Her knowledge of the geography of the world and the exact locations
of cities and towns was astounding. She never owned a TV set or a computer. She was
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a prodigious letter writer, and both her letters and manuscripts were written in pencil,
of which she always had seven at hand. Jane also kept carbon copies of her
manuscripts and most of her correspondence. She also kept nearly every letter,
magazine, journal and book that she received, was given, or purchased, and, if her
home had the space, she may even have been tempted to keep every newspaper!
For the last 19 years of Jane’s life, Florah Kokoele was her faithful house assistant,
seeing to all of her needs. Julia Nkonyane joined Florah in this task during the last 4
years as Jane’s need for assistance increased. These ladies deserve our special thanks.
Upon returning home from work during August 1998, Jane fell from the back stairs of
her home and thereafter ceased to drive her own car, relying on friends for transport.
Walking became increasingly difficult, and unwillingly Jane had to acquire both a
walking ring and a wheel chair, the latter used by her friends as a seat when they came
to visit, or to have tea with her in her bedroom. Jane disliked being house-bound
intensely, and her friends, who took over her regular Friday shopping with Florah,
used to load her in their cars, and after they had parked in the shade at the shopping
centres with her inside, they did the house-shopping with Florah. With the exception
of a few items, only food from Woolworths was purchased. In later years the strain on
Jane’s body to get into a car was too great and her friends then took only Florah and
her shopping list, which was most carefully composed in Jane’s hand-writing, to
purchase the weekly necessities. Friday was shopping day, and no other day would
do, so woe betide the authorities who decreed that certain public holidays must fall on
Fridays!
Before moving to St Kit’s Island in the Caribbean, Eddy and Tammi Krecek had a
home close to Jane’s, and Eddy phoned her in the morning on a nearly daily basis to
enquire of her well-being. He and Tammi were also regular visitors. Eddy and Garth
Holmes also attended to much of the maintenance of Jane’s home. Besides friends and
neighbours who visited her casually, Jane also had a group of friends who visited her
on a regular, almost weekly, basis. Shirley Gilmore visited on Thursday mornings,
Ivan Horak on Friday mornings, and Mike and Mary-Lou Penrith, and Banie and
Naomi Penzhorn on Saturdays and Sundays. During the 2 years that Dmitry
Apanaskevich spent in South Africa Ivan took him to Jane’s for tea on Friday
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mornings and, while Florah and Ivan did the weekly shopping, Jane and Dmitry had
the most lively discussions on tick taxonomy and tick taxonomists.
Jane lived frugally, but her near boundless generosity to those charities that she
considered as worthy will be sorely missed by them, as it will also be by a rural
family whom she supported.
With Jane’s death the scientific world has lost one of its most meticulous and
experienced workers, and the field of tick taxonomy, particularly in Africa, will be
infinitely poorer. Her friends have lost a true Lady and a loyal friend.
Jane died peacefully at home in Pretoria on Friday morning 3 April 2009 while having
tea with Ivan Horak.
IGH
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CURRICULUM VITAE
JANE BROTHERTON WALKER
Personal history
Birth date: 31 January 1925
Place of birth: Nairobi, Kenya
Died: 3 April 2009
Place of death: At home, Pretoria, South Africa
Education
1931-1937: Home-taught by her mother in Kenya.
1938-1944: County High School for Girls, Retford, England.
1944-1948: BSc (Honours), Liverpool University.
1959: MSc, Liverpool University (External student).
Dissertation: “A comparative study of the larvae and nymphae of
ticks belonging to the genus Rhipicephalus in East Africa”.
1960: Attended summer school courses in General Acarology and Medical
and Veterinary Acarology at the Institute of Acarology, University of
Maryland.
1983: DSc, University of the Witwatersrand.
Thesis: “Studies on African ticks (Acarina, Ixodoidea)”.
Professional Career
A. In the Research Branch of Her Majesty’s Overseas Civil Service (HMOCS)
1949: Appointed research Officer (Enytomologist), East African Veterinary
research Organization (EAVRO). Had 3 months post-graduate
training with Dr Gertrud Theiler at the Veterinary Research Institute,
Onderstepoort.
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1953: Specialist study on ticks of the genus Hyalomma with Dr L. Delpy,
Institute Mérieux, Lyons, France.
1956: Appointed Senior Scientific Officer, EAVRO. Assisted Dr H.
Hoogstraal, of the United States Naval Medical Research Unit No. 3
Cairo, with the organization of a 3-month International Scientific
Working Party on Ectoparasites and Arthropod-borne Disease in East
Africa.
1957: Visited veterinary laboratories in Northern and Southern Rhodesia
(Zambia and Zimbabwe), South Africa and Nigeria to see the research
in progress there on arthropod vectors of disease.
1960: Awarded US National Institutes of Health Training Grant to attend
courses on acarology at the University of Maryland, and the
Rockefeller Foundation Travel Grant to visit laboratories and confer
with research workers in the United States and Canada.
1961: Appointed Principal Scientific Officer, EAVRO.
1962: Awarded Welcome Research Travel Grant to attend second meeting
of the FAO/OIE Expert Panel on Tick-borne Disease of Livestock as
an observer and to study tick collections at the United States Naval
Medical Research Unit No. 3, Cairo.
1963: Visited Veterinary Research Institute, Onderstepoort.
1966: Voluntary retirement from HMOCS.
B. Veterinary Research Institute, Onderstepoort, South Africa
1966: Appointed Senior Professional Officer (Entomology/Zoology) at
Veterinary Research Institute, Onderstepoort.
1974: Appointed Chief Professional Officer (Zoology).
1984: Appointed Chief Veterinary Researcher.
1986: Appointed Specialist Scientist.
1990: Retired on pension.
1990-1998: Worked part-time (1-3 days per week) in an honorary capacity at the
Institute.
Experience
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Research: Studies on the taxonomy, hosts and distribution of African ticks,
particularly of the genus Rhipicephalus.
Teaching: Part-time lecturer from 1952-1963 on ticks and other arthropod
parasites of livestock at the Faculty of Veterinary Science, Makerere
College (now Faculty of Veterinary Science, University of Nairobi).
Post graduate Co-promoter of an MSc degree, University of Potchefstroom, South
supervision: Africa.
Committees: Editorial Committee of the East African Wildlife Journal for 4 years.
Edited the Quarterly Bulletin of the Scientific and Technical
Committee of the East African Wildlife Society for the same period.
Editorial Committee of the Onderstepoort Journal of Veterinary
Research from 1969-2000.
Membership of Professional Societies
1953: Elected to Membership of the Institute of Biology, London.
1964: Member, Zoological Society of Southern Africa.
1972: Member, Parasitological Society of Southern Africa.
1982: Elected to Honorary Associate Membership of the South African
Veterinary Association.
Awards
1988: Elsdon-Dew Medal awarded by the Parasitological Society of
southern Africa, for services rendered to Parasitology in Africa.
1998: Agricultural Science and Technology Woman of the Year Award.
1998: Theiler Memorial Trust Award for exceptional service rendered to
Veterinary Science in Africa.
2000-2002: Appointed as an Honorary Professor in the Department of Veterinary
Tropical Diseases, Faculty of Veterinary Science, University of
Pretoria.
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International and National Congresses at which papers were presented by Jane
Walker or by her colleagues on her behalf
1961: Walker, Jane B. “Some observations on the classification and biology
of ticks belonging to the genus Rhipicephalus, with special reference
to the immature stages”. Scientific Conference on the Epidemiology
of Arthropod-borne Diseases, Nairobi, Kenya, 11 January 1961.
1976: Walker, Jane B. 1976. “Ticks and human disease in tropical Africa”.
International Symposium on Medicine in a Tropical Environment,
Pretoria, South Africa, 19-23 July 1976.
1975: Walker, Jane B. “Survey techniques for tick species affecting
domestic animals”. International Workshop on the Ecology and
Control of External Parasites of Cattle of Economic Importance in
Latin America, Cali, Colombia, 25-30 August 1975.
1981: Walker, Jane B., Spickett, A.M., Brett, Susan & Horak, I.G. Poster
“The immature stages of the common ticks of domestic animals in
South Africa”. International Conference on Tick Biology and Control,
Tick Research Unit, Rhodes University, Grahamstown, South Africa,
27-29 January 1981.
1986: Walker, Jane B. & Olwage, A. “The tick vectors of Cowdria
ruminantium (Ixodoidea, Ixodidae, genus Amblyomma) and their
distribution”. International Workshop on Heartwater: past, present
and future, Berg-en-Dal, Kruger National Park, South Africa, 8-11
September 1986.
1994: Penzhorn, B.L., Krecek, R.C., Horak, I.G., Verster, A.J.M., Walker,
J.B., Boomker, J.D.F., Knapp, S.E. & Quandt, S.K.F. “Parasites of
African rhinos: a documentation”. Symposium on Rhinos as Game
Ranch Animals, Onderstepoort, South Africa, 9-10 September 1994.
2000: Horak, I.G. & Walker, Jane B. 1999. “The relationships of some
South African ticks in the genus Rhipicephalus”. Third International
Conference on Ticks and Tick-borne Pathogens: into the 21 st Century.
High Tatera Mountains, Slovakia, 30 August-3 September 1999.
Horak, I.G. & Walker Jane B. 2003. “The taxonomy, distribution and
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iology of Rhipicephalus muhsamae, R. praetextatus, R. simus and R.
gertrudae”. Third National Workshop on Ticks and Tick-borne
Disease in the Sudan. Khartoum, Sudan, 18-20 August 2003.
New tick species described by Jane Walker or by Jane and her co-workers
Rhipicephalus carnivoralis Walker, 1966
Haemaphysalis hyracophila Hoogstraal, Walker & Neitz, 1971
Margaropus wileyi Walker & Laurence, 1973
Argas africolumbae Hoogstraal, Kaiser, Walker, Ledger, Converse & Rice, 1975
Ixodes neitzi Clifford, Walker & Keirans, 1977
Rhipicephalus zambeziensis Walker, Norval & Corwin, 1981
Ixodes brewsterae Keirans, Clifford & Walker, 1982
Ixodes catherinei Keirans, Clifford & Walker, 1982
Ixodes corwini Keirans, Clifford & Walker, 1982
Ixodes macfarlanei Keirans, Clifford & Walker, 1982
Ixodes zairensis Keirans, Clifford & Walker, 1982
Rhipicephalus lounsburyi Walker, 1990
Rhipicephalus neumanni Walker, 1990
Rhipicephalus aquatalis Walker, Keirans & Pegram, 1993
Rhipicephalus exophthalmos Keirans & Walker, 1993
Rhipicephalus interventus Walker, Pegram & Keirans, 1995.
Rhipicephalus oreotragi Walker & Horak, 2000
Rhipicephalus warburtoni Walker & Horak, 2000
PUBLICATIONS
Jane Brotherton Walker
1. Walker, Jane B. 1955. Rhipicephalus pulchellus Gerstäcker, 1873: a description of
the larva and nymph with notes on the adults and on its biology. Parasitology,
45: 95-98.
2. Walker, Jane B. 1956. Rhipicephalus pravus Dönitz, 1910. Parasitology, 46: 243-
246.
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3. Theiler, Gertrud, Walker, Jane B. & Wiley, A.J. 1956. Ticks in the South African
Zoological Survey Collection. Part VIII. Two East African ticks. Onderstepoort
Journal of Veterinary Research. 27: 83-99.
4. Walker, Jane B. 1957. Rhipicephalus humeralis Rondelli, 1926. Parasitology, 47:
145-152.
5. Walker, Jane B. 1959. Notes on the common tick species of East Africa. I.
Introduction. Rhipicephalus appendiculatus and R. pravus. East African Veld, 5:
111-116.
6. Walker, Jane B. 1959. Notes on the common tick species of East Africa. II. More
species belonging to the genus Rhipicephalus. East African Veld, 5: 131-135.
7. Walker, Jane B. 1959. Notes on the common tick species of East Africa. III. The
bont and bont-legged ticks. East African Veld, 5: 157-162.
8. Walker, Jane B. & Wiley, A.J. 1959. Rhipicephalus camelopardalis n. sp.
(Ixodoidea, Ixodidae), a new species of tick from East African giraffes.
Parasitology, 49: 448-453.
9. Walker, Jane B. 1960. Notes on the common tick species of East Africa. IV. The
blue tick, the yellow dog tick and the fowl tick. East African Veld, 6: 9-13.
10. Walker, Jane B. 1960. Notes on the common tick species of East Africa. Cooper,
McDougall & Robertson (E.A.) Ltd: Nairobi.
11. Walker, Jane B. 1962. Notes on the common tick species of East Africa. 2 nd
edition, revised. Cooper, McDougall & Robertson (E.A.) Ltd: Nairobi.
12. Dinnik, J.A., Walker, Jane B., Barnett, S.F. & Brocklesby, D.W. 1963. Some
parasites obtained from game animals in western Uganda. Bulletin of Epizootic
Diseases of Africa. 11: 37-44.
13. Walker, Jane B. & Parsons, B.T. 1964. The laboratory rearing of Amblyomma
sparsum Neumann, 1899. Parasitology, 54: 173-175.
14. Walker, Jane B. 1966. Rhipicephalus carnivoralis sp. nov. (Ixodoidea, Ixodidae).
A new species of tick from East Africa. Parasitology, 56: 1-12.
15. Walker, Jane B. 1966. Rhipicephalus reichenowi Zumpt, 1943: a re-description of
the male and female and descriptions of the nymph and larva, together with an
account on its known hosts and distribution. Parasitology, 56: 457-469.
16. Clifford, C.M. & Walker, Jane B. 1966. Host and distribution records for several
species of the genus Ixodes (Acarina: Ixodidae) that occur in East Africa.
Review de Zoologie et de Botanique Africaines, 74: 155-168.
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17. Sweatman, G.K., Walker, Jane B. & Bindernagel J.A. 1969. Stages in the
development of Choriopsoroptes kenyensis gen. et sp. n. (Acari: Sarcoptiformes:
Psoroptidae), a body mange mite from captive African buffalo, Syncerus caffer,
in Kenya. Journal of Parasitology, 55: 1298-1310.
18. Walker, Jane B. 1970. Notes on the common tick species of East Africa. 3 rd
edition, revised. Cooper, McDougall & Robertson (E.A.) Ltd: Nairobi.
19. Hoogstraal, H., Walker, Jane B. & Neitz, W.O. 1971. Notes on African
Haemaphysalis ticks. VII. H. (Rhipistoma) hyracophila sp. n. (Ixodoidea:
Ixodidae), a parasite of hyraxes in South Africa. Journal of Parasitology, 57:
417-425.
20. Walker, Jane B. 1973. Acari of the Ethiopian Region. D. Suborder Ixodoidea
Leach, 1815. Department of Agricultural Technical Services, Republic of South
Africa, Entomology Memoir No. 29: 42-45.
21. Walker, Jane B., Bezuidenhout, J.D. 1973. Treatment of tick-infested tortoises.
Journal of the South African Veterinary Association, 44: 381.
22. Walker, Jane B. & Laurence, B.R. 1973. Margaropus wileyi sp. nov. (Ixodoidea,
Ixodidae), a new species of tick from the reticulated giraffe. Onderstepoort
Journal of Veterinary Research, 40: 13-21.
23. Hoogstraal, H., Kaiser, M.N., Walker, Jane B., Ledger, J.A., Converse, J.D. &
Rice, R.C.A. 1975. Observations on the sub-genus Argas (Ixodoidea: Argasidae:
Argas). 10. A. (A.) africolumbae, n. sp., a Pretoria virus-infected parasite of
birds in southern and eastern Africa. Journal of Medical Entomology, 12: 194-
201.
24. Clifford, C.M., Walker, Jane B. & Keirans, J.E. 1976. Ixodes (Afrixodes) bakeri
Arthur & Clifford, 1961 (Acarina: Ixodidae): descriptions of the male and
immature stages from rodents and insectivores and notes on its biology in South
Africa. Onderstepoort Journal of Veterinary Research, 43: 105-112.
25. Clifford, C.M., Walker, Jane B. & Keirans, J.E. 1977. Ixodes (Afrixodes) neitzi, n.
sp. (Acarina: Ixodidae) from the mountain reedbuck in South Africa.
Onderstepoort Journal of Veterinary Research, 44: 143-150.
26. Bulman, G.M. & Walker, Jane B. 1979. A previously unrecorded feeding site on
cattle for the immature stages of the spinose ear tick, Otobius megnini (Duges,
1844). Journal of the South African Veterinary Association, 50: 107-108.
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27. Walker, Jane B., Norval, R.A.I. & Corwin, M.D. 1981. Rhipicephalus
zambeziensis sp. nov., a new tick from eastern and southern Africa, together
with a redescription of Rhipicephalus appendiculatus Neumann, 1901 (Acarina,
Ixodidae). Onderstepoort Journal of Veterinary Research, 48: 87-104.
28. Rechav, Y. & Walker, Jane B. 1981. Ixodid ticks on children. South African
Medical Journal, 59: 889.
29. Keirans, J.E., Clifford, C.M. & Walker, Jane B. 1982. The Ixodes (Afrixodes) oldi
group (Acari: Ixodidae) from subSaharan Africa with descriptions of five new
species. Journal of Medical Entomology, 19: 309-329.
30. Norval, R.A.I., Walker, Jane B. & Colborne, J. 1982. The ecology of
Rhipicephalus zambeziensis and Rhipicephalus appendiculatus (Acarina:
Ixodidae) with particular reference to Zimbabwe. Onderstepoort Journal of
Veterinary Research, 49: 181-190.
31. Clifford, C.M., Walker, Jane B. & Keirans, J.E. 1983. Clarification of the status of
Rhipicephalus kochi Dönitz, 1905 (Ixodoidea, Ixodidae). Onderstepoort Journal
of Veterinary Research, 50: 77-89.
32. Horak, I.G., Potgieter, F.T., Walker, Jane B., De Vos, V. & Boomker, J. 1983.
The ixodid tick burdens of various large ruminant species in South African
nature reserves. Onderstepoort Journal of Veterinary Research, 50: 221-228.
33. Walker, Jane B. & Schultz, K.C.A. 1984. Records of the bont tick, Amblyomma
hebraeum, from the angulate tortoise, Chersina angulata, and the leopard
tortoise, Geochelone pardalis. Onderstepoort Journal of Veterinary Research,
51: 171-173.
34. Pegram, R.G., Clifford, C.M., Walker, Jane B. & Keirans, J.E. 1987. Clarification
of the Rhipicephalus sanguineus group (Acari, Ixodoidea, Ixodidae). I. R.
sulcatus Neumann, 1908 and R. turanicus Pomerantsev, 1936. Systematic
Parasitology, 10: 3-26.
35. Pegram, R.G., Keirans, J.E., Clifford, C.M. & Walker, Jane B. 1987. Systematic
clarification of the Rhipicephalus sanguineus group (Acari, Ixodoidea,
Ixodidae). II. R. sanguineus (Latreille, 1806) and related species. Systematic
Parasitology, 10: 27-44.
36. Walker, Jane B. & Olwage, A. 1987. The tick vectors of Cowdria ruminantium
(Ixodoidea, Ixodidae, genus Amblyomma) and their distribution. Onderstepoort
Journal of Veterinary Research, 54: 353-379.
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37. Pegram, R.G., Walker, Jane B., Clifford, C.M. & Keirans, J.E. 1987. Comparison
of populations of the Rhipicephalus simus group: R. simus, R. praetextatus, and
R. muhsamae (Acari: Ixodidae). Journal of Medical Entomology, 24: 666-682.
38. Walker, Jane B., Keirans, J.E., Pegram, R.G. & Clifford, C.M. 1988. Clarification
of the status of Rhipicephalus tricuspis Dönitz, 1906 and Rhipicephalus
lunulatus Neumann, 1907 (Ixodoidea, Ixodidae). Systematic Parasitology, 12:
159-186.
39. Pegram, R.G. & Walker, Jane B. 1988. Clarification of the biosystematics and
vector status of some African Rhipicephalus species (Acarina: Ixodidae). In:
Service, M.W. (ed.) Biosystematics of haematophagous insects. Systematics
Association Special Volume No. 37: 61-76. Clarendon Press: Oxford.
40. Walker, Jane B. 1990. Two new species of ticks from southern Africa whose
adults parasitize the feet of ungulates: Rhipicephalus lounsburyi n. sp. and
Rhipicephalus neumanni n. sp. (Ixodoidea, Ixodidae). Onderstepoort Journal of
Veterinary Research, 57: 57-75.
41. Lessard, P., L'Eplattenier, R., Norval, R.A.I., Kundert, K., Dolan, T.T., Croze, H.,
Walker, Jane B., Irvin, A.D. & Perry, B.D. 1990. Geographical information
systems for studying the epidemiology of cattle diseases caused by Theileria
parva. Veterinary Record, 126: 255-262.
42. Walker, Jane B. 1991. A review of the ixodid ticks (Acari, Ixodidae) occurring in
southern Africa. Onderstepoort Journal of Veterinary Research, 58: 81-105.
43. Walker, Jane B., Keirans, J.E. & Pegram, R.G. 1993. Rhipicephalus aquatilis sp.
nov. (Acari: Ixodidae), a new tick species parasitic mainly on the sitatunga,
Tragelaphus spekei, in East and Central Africa. Onderstepoort Journal of
Veterinary Research, 60: 205-210.
44. Keirans, J.E., Walker, Jane B., Horak, I.G. & Heyne, H. 1993. Rhipicephalus
exophthalmos sp. nov., a new tick species from southern Africa, and
redescription of Rhipicephalus oculatus Neumann, 1901, with which it has
hitherto been confused (Acari: Ixodida: Ixodidae). Onderstepoort Journal of
Veterinary Research, 60: 229-246.
45. Walker, Jane B., Pegram, R.G. & Keirans, J.E. 1995. Rhipicephalus interventus
sp. nov. (Acari: Ixodidae), a new tick species closely related to Rhipicephalus
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tricuspis Dönitz, 1906 and Rhipicephalus lunulatus Neuman, 1907, from East
and Central Africa. Onderstepoort Journal of Veterinary Research, 62: 89-95.
46. Horak, I.G., Braack, L.E.O., Fourie, L.J. & Walker, Jane B. 2000. Parasites of
domestic and wild animals in South Africa. XXXVIII. Ixodid ticks collected
from 23 wild carnivore species. Onderstepoort Journal of Veterinary Research,
67: 239-250.
47. Horak, I.G., Fourie, L.J., Heyne, H., Walker, Jane B. & Needham, G.R. 2002.
Books
Ixodid ticks feeding on humans in South Africa: with notes on preferred hosts,
geographic distribution, seasonal occurrence and transmission of pathogens.
Experimental and Applied Acarology, 27: 113-136.
1. Yeoman, G.H. & Walker, Jane B., assisted by Ross, J.P.J. & Docker, T.M. 1967.
The ixodid ticks of Tanzania. A study of the zoogeographyof the Ixodidae of an
East African country. Commonwealth Institute of Entomology: London.
2. Walker, Jane B. 1974. The ixodid ticks of Kenya. A review of present knowledge
of their hosts and distribution. Commonwealth Institute of Entomology:
London.
3. Walker, Jane B., Mehlitz, D. & Jones, G.E. 1978. Notes on the ticks of Botswana.
German Agency for Technical Coopperation, Ltd (GTZ): Eschborn.
4. Howell, C.J., Walker, J.B. & Nevill, E.M. 1978. Ticks, mites and insects infesting
domestic animals in South Africa. Part 1. Descriptions and biology. Department
of Agricultural Technical Services, Republic of South Africa, Science Bulletin
no. 393.
5. Walker, Jane B., Keirans, J.E. & Horak, I.G. 2000. The genus Rhipicephalus
(Acari, Ixodidae): a guide to the brown ticks of the World. Cambridge
Academic Press: Cambridge.
Conference Proceedings (Published in full)
1. Walker, Jane B. 1961. Some observations on the classification and biology of ticks
belonging to the genus Rhipicephalus, with special reference to the immature
stages. East African Medical Journal, 38: 232-238.
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2. Walker, Jane B. 1977. Ticks and human disease in tropical Africa. Medicine in a
tropical environment. Proceedings of the International Symposium, South Africa
1976. 276-290. A.A. Balkema: Rotterdam, The Netherlands.
3. Walker, Jane B. 1978. Survey techniques for tick species affecting domestic
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