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humus group - Territorio e Sistemi agro-forestali - Università degli ...

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OF<br />

Figure 3<br />

zo<br />

% volume<br />

roots excluded<br />

90<br />

10<br />

noz<br />

OFzo : zoogenic transformed material ><br />

10 % of the volume of the horizon after<br />

removing living roots.<br />

OFnoz : non zoogenic transformed<br />

material >= 90 % of the volume of the<br />

horizon after removing living roots.<br />

ZOOGENIC TRANSFORMED MATERIAL = RECOGNIZABLE<br />

REMAINS AND HUMIC COMPONENTS processed by animals<br />

= leaves, needles and other plant residues more or less degraded<br />

by soil animals, mixed with animal droppings. A fine powdered<br />

and/or grained structure (less than 1 mm) is typical in a terminal<br />

stage of faunal attack in an organic horizon. At this last level of<br />

biotransformation, the substrate (OH horizon) is essentially<br />

comprised of organic animal droppings of varying size (droppings<br />

of epigeic earthworms, macroarthropods such as millipedes,<br />

woodlice and insect larvae, microarthropods such as mites and<br />

springtails and enchytraeids dominate). Within hemorganic<br />

horizons, animal activity leads to different types of A horizons,<br />

depending on the animals’ ability to dig into the mineral soil and<br />

thoroughly mix organic and mineral matter.<br />

NON ZOOGENIC TRANSFORMED MATERIAL = REMAINS AND<br />

HUMIC COMPONENT processed by fungi or other non-faunal<br />

processes = leaves, needles and other plant residues more or<br />

less fragmented and transformed into fibrous matter by fungi.<br />

Recognizable and recent animal droppings are absent or not<br />

detectable by the naked eye in the organic horizons; fungal<br />

hyphae can be recognized as white, brown or yellow strands<br />

permeating the organic or hemorganic substrates; traces of animal<br />

activity (old bite marks, mucus) may sometimes be detectable but<br />

are always marginal. In the last stage of biodegradation of an<br />

organic horizon, non zoogenic substance may essentially be<br />

composed of brown, dry plant residues more or less in powder<br />

form or tiny fragments (OF and OH horizons), or be massive like a<br />

dark wet plastic clay (OH or very organic A horizons).

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