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The enigmatic Cooper's and Coxs Sandpipers<br />
summ?r winter<br />
,<br />
V<br />
~<br />
824'17 1 82742<br />
13 June 1933 :C, Novem4er t 966<br />
USA ~ .uSlfahe.<br />
r ac]uIt lemale Iemale<br />
summyr winler Summer<br />
ir<br />
C melanotos<br />
8307'S r"} "~- - .2á&4 :3<br />
swnm- winrer<br />
C paramefanotos<br />
B7997 826013<br />
i. r. ~. 7 April 1902 28 Navember 196 1<br />
AuslraEia AuShaifa<br />
r .,Ie 111' _, r adult male<br />
C ferruginea<br />
FICUfaE 1 Mantle-feathers (smaller) and scapulars of Calydris melanotos (upper) . C paramefanotos<br />
(centre) and C ferruginea (lower) . Diagonal shading indicates brighter-rufous colouring<br />
: stippling is shades of grey or grey-brown SAM registration numbers of specirnens and<br />
their localities are given (trom Cox 1987 )<br />
wing-length is 0.7 mm beyond the upper limit of C acuminata (145 mm . Ma-chani et<br />
al 1986) but the remaining measurements are intermediate between those of that<br />
species and C ferruginea . The measurements of both types of C paramelanotosfall<br />
within the range of variatlon in C melanotos or C ferruginea, or are intermediate<br />
between the two (Cox 1987, contra Buckley 1988)-<br />
identificatiort of trapped birds<br />
Al least two birds seen in Australia and identified as C paramelanotos more<br />
resembled C cooperi (Cox 1989a) . Another trapped at Stockton, New Soutl Wales,<br />
on 21 March 1981 was identified as a probable hybrid of C acuminata and C<br />
ferruginea (Lane et al 1981) but was later regarded to have been an example of C<br />
pararnelanotosthat may proveto be a hybrid (Marchant et al 1986, Cox 1987, Pringle<br />
1987) . More recently, it was noted to have resembled C cooperi (Cox 1989a) and 1<br />
have since examined photographs that show it had the primary-shafts white . shadin g<br />
60