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ecological effects of marine protected areas empafish project ...

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EMPAFISH Booklet no. 1 Ecological <strong>effects</strong> <strong>of</strong> Atlanto-Mediterranean MPAs in the EU<br />

2005) bottom morphology did not exert a detectable effect, probably due to<br />

greater habitat similarities among the localities that were being compared.<br />

Studies describing fish assemblages at multiple scales (e.g. Williams and<br />

Hatcher 1983; Galzin 1987; Holbrook et al. 2000; Gust et al. 2001) have<br />

usually found that habitat type explains small-to-medium patchiness in fish<br />

assemblage structure. Distinguishing the relative contribution <strong>of</strong> habitat<br />

structure can help to elucidate the actual <strong>effects</strong> <strong>of</strong> protection (Jennings et al.<br />

1996; García Charton et al. 2000, 2004).<br />

8

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