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Catalogue of the Living Bivalvia of the Eastern Pacific Ocean ...

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No comprehensive inventory <strong>of</strong> <strong>the</strong> bivalve fauna <strong>of</strong> <strong>the</strong><br />

entire eastern <strong>Pacific</strong> has appeared, and it is more than 44<br />

years since A. M. Keen's Check list and bibliography <strong>of</strong> <strong>the</strong><br />

northwestern American marine Mollusca (1937). The<br />

present catalogue was started over fifteen years ago to provide<br />

data for a re-interpretation <strong>of</strong> bivalve geography using<br />

new distributional data, segregated by depth to resolve<br />

effects <strong>of</strong> submergence <strong>of</strong> nor<strong>the</strong>rn species at lower latitude,<br />

and submitted to computer techniques for analysis. This<br />

work is now in final preparation (Bernard & McKinnel MS),<br />

but it appeared useful for taxonomic workers to issue <strong>the</strong><br />

data, and setting down in juxtaposition <strong>the</strong> nor<strong>the</strong>rn and<br />

sou<strong>the</strong>rn representatives <strong>of</strong> various groups may stimulate<br />

fur<strong>the</strong>r study leading to monographie reviews.<br />

I have attempted to include all binomina used regionally,<br />

though <strong>the</strong> early literature is replete with extralimital analogues.<br />

I have built on <strong>the</strong> important faunistic works <strong>of</strong><br />

Carpenter (1872), Dall (1921), Keen (1937), and Burch<br />

(1944-46) for <strong>the</strong> nor<strong>the</strong>rn fauna; Olsson (1961) and Keen<br />

(1971) for tropical America; and Carcelles & Williamson<br />

(1951) and Osorio & Bahamonde (1970) for <strong>the</strong> sou<strong>the</strong>rn<br />

fauna. O<strong>the</strong>r records were obtained by abstracting <strong>the</strong> literature<br />

and more than 5000 titles were searched. I have avoided<br />

mere lucubrations <strong>of</strong> published records, that so <strong>of</strong>ten repeat<br />

and perpetuate errors, and wherever possible type material<br />

was examined and collections consulted contemporaneous to<br />

writings <strong>of</strong> authors where original material is unavailable or<br />

lost. In some cases <strong>the</strong> total nomenclatural confusion makes<br />

determination <strong>of</strong> identity a matter <strong>of</strong> idle but not uninteresting<br />

speculation.<br />

A critical review <strong>of</strong> each group is impossible on so large a<br />

scale and would result in several volumes <strong>of</strong> heroic proportions.<br />

I have let stand a number <strong>of</strong> incorrect generic<br />

usages, as <strong>the</strong>se long-established errors require erection <strong>of</strong><br />

new taxa for <strong>the</strong>ir correction. However, I have corrected<br />

homonyms and have merged a number <strong>of</strong> species. Doubtless,<br />

synonyms and o<strong>the</strong>r oversights have crept in. Besides such<br />

outright errors, two fur<strong>the</strong>r difficulties will try <strong>the</strong> readerpertinent<br />

literature may have been overlooked, and <strong>the</strong> lag in<br />

publication time will result in omission <strong>of</strong> newly described<br />

species. I apologise for <strong>the</strong>se shortcomings and hope <strong>the</strong>y<br />

will be brought to my attention for revision.<br />

FORMAT<br />

Suprageneric taxa are arranged to reflect current concepts<br />

<strong>of</strong> taxonomic relationships, combining stratigraphie, anatomical,<br />

shell ultrastructure and o<strong>the</strong>r criteria with <strong>the</strong> classic<br />

shell morphology. A brief overview <strong>of</strong> <strong>the</strong> present status <strong>of</strong><br />

<strong>the</strong> systematics <strong>of</strong> <strong>the</strong> higher taxa is given below.<br />

Bibliographic references for <strong>the</strong> supraspecific taxa are not<br />

cited, where <strong>the</strong>y are available in <strong>the</strong> Treatise on Invertebrate<br />

Paleontology (Moore 1969), and do not require repetition,<br />

but references to authors <strong>of</strong> names which appeared after<br />

1969, are included. Authorities for corrections <strong>of</strong> names and<br />

INTRODUCTION<br />

changes in concepts or status are given, but do not appear in<br />

<strong>the</strong> bibliography.<br />

Genera and subgenera are arranged alphabetically. I have<br />

avoided listing <strong>the</strong> nominate genus directly following <strong>the</strong><br />

family or subfamily entry as done in <strong>the</strong> 'Treatise' , as this<br />

merely reflects chronology and is <strong>of</strong>ten disruptive <strong>of</strong> <strong>the</strong><br />

current view <strong>of</strong> natural relationships, which are constantly<br />

changing as more information accumulates.<br />

Each species comprises a separate entry, with <strong>the</strong> current<br />

binomen in heavy type. The original combination and full<br />

bibliographic reference follows for <strong>the</strong> species and its junior<br />

synonyms, arranged chronologically, <strong>the</strong> status indicated,<br />

where necessary, with abbreviated latin terrns. Distribution<br />

outside <strong>the</strong> area <strong>of</strong> concern is briefly indicated. Occurrence in<br />

<strong>the</strong> Clipperton, Cocos, or Galapagos Islands fauna is noted,<br />

as <strong>the</strong>re appear to be features distinct from <strong>the</strong> continental<br />

shelf biota, notably <strong>the</strong> presence <strong>of</strong> a few endemic taxa and<br />

those species with Indo-<strong>Pacific</strong> affinities. The final line <strong>of</strong><br />

each entry includes <strong>the</strong> geographic range to nearest degree <strong>of</strong><br />

latitude and <strong>the</strong> mid-point <strong>of</strong> geographical distribution. The<br />

bathymetric range is in meters, and may differ from previously<br />

published records as I have not cited depths based on<br />

dead and probably displaced material. Thermal ranges are<br />

followed by <strong>the</strong> oldest paleontological occurrence on <strong>the</strong><br />

American continent. Extralimital or incorrect binomina<br />

which have become embedded in <strong>the</strong> literature are presented<br />

in smaller type. Departures from current systematic arrangement<br />

and proposals <strong>of</strong> replacement names are annotated by a<br />

numbered entry appearing at <strong>the</strong> end <strong>of</strong> <strong>the</strong> catalogue section.<br />

It is an irritant to <strong>the</strong> non-taxonomist to be faced with<br />

changes <strong>of</strong> familiar names; however, it is unavoidable to<br />

eventual stability and universality. All genus taxa were<br />

checked with Neave (1939-75), and <strong>the</strong> specific names with<br />

Sherborn (1902-33), and <strong>the</strong> useful compilation by Ruh<strong>of</strong>f<br />

(1980) bridging <strong>the</strong> period between Sherborn and <strong>the</strong> start <strong>of</strong><br />

<strong>the</strong> Zoological Record. Contemporary taxonomy has largely<br />

disgarded <strong>the</strong> typological concept <strong>of</strong> <strong>the</strong> species in favor <strong>of</strong><br />

emphasis on infraspecific variability. Following this I do not<br />

use <strong>the</strong> subspecific designation, as <strong>the</strong> majority <strong>of</strong> wideranging<br />

species are expected to yield several subspecies.<br />

Formal taxonomie citation requires only <strong>the</strong> binomen<br />

(genus and species), however, <strong>the</strong> original author and<br />

publication date are usually appended. The addition <strong>of</strong><br />

pagination, references to illustrations, addition <strong>of</strong> <strong>the</strong> author<br />

or editor <strong>of</strong> <strong>the</strong> work in which <strong>the</strong> description appeared,<br />

citation <strong>of</strong> <strong>the</strong> authority for change <strong>of</strong> genus, etc, is merely<br />

book-keeping with no formal taxonomie status. I have,<br />

generally, followed <strong>the</strong> Anglo-American Cataloguing Rules<br />

(Gorman & Winkler 1978) when citing authorities' names.<br />

The prefix Du, de, von, etc is not included (Orbigny not<br />

d' Orbigny; Blainville not Ducrotay de Blainville). Technically<br />

this is acceptable only when <strong>the</strong> prefix follows <strong>the</strong> first<br />

name, in o<strong>the</strong>r cases <strong>the</strong> name should be quoted in full<br />

(Megerle von Milhlfeld), but as no doubt arises, and considerable<br />

space is saved, I have followed <strong>the</strong> informal European<br />

usage among peers (Megerle).<br />

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